282 lines
31 KiB
XML
282 lines
31 KiB
XML
<document id="9F528B5AB8C4D6683AB8DDF97CBC594A" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre & Murienne, Jérôme" docDate="2020" docId="03B7726A135B5320FDA9FC54FEEBFC31" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus donosoi Moret 2020, sp. nov." docType="treatment" docUuid="F31CFECF-7D04-4800-9559-EA234039B5DD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="22" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="20" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="A40C9125CE8DBE193A147DCFBBD1D134" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EED624B9DE6B833BE0F0F4CC9E577ED7">
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<mods:title id="A35BFE4524152B4D5D39DD9553850188">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
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</mods:titleInfo>
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<mods:name id="73C58130C232D37CAADBA7D1224BD43E" type="personal">
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<mods:role id="7BA486A4658D5262E8800AF396F171E7">
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<mods:roleTerm id="E00E1F9457FC0FD792CC6572247AE3EE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2E55EEB22657FC23AA00EF1CF8B6A764">Moret, Pierre</mods:namePart>
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<mods:nameIdentifier id="91C5892E4E17A83399381758E7483187" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
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<mods:affiliation id="C04FA32CA07A6087B196E281DD0C008C">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
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<mods:nameIdentifier id="ED0FAB3B2A44D5082F1E9BBE42AA11E8" type="email">pierre.moret@univ-tlse2.fr&pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name id="270FDE5078774805081FBAF1123ACB8D" type="personal">
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<mods:roleTerm id="0C02B92D64786FB7EC29E41C8D9A8771">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C9B34CC1B6454D995FD3D1E3F09D4392">Murienne, Jérôme</mods:namePart>
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<mods:nameIdentifier id="5B1C7A1D33CC692ED917E380BA341951" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
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<mods:affiliation id="ACCDA8D1ADF7DEC263FACB2C49594E7A">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
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<mods:nameIdentifier id="D194B15FF3B86F1EDDB9E8B5E5F4FE94" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
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<mods:typeOfResource id="F7C063F8A45BA6BFE8B8FEE514A2D61B">text</mods:typeOfResource>
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<mods:titleInfo id="8277D7F9F03C6B15CBB422EB838C355E">
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<mods:title id="81CA4C0182868DF7E62115F2EE1652D6">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="C33912F2428DC867DE66743B2D9C448B">
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<mods:date id="37FC9CD60BCF7A4D704803D41B71C46C">2020</mods:date>
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<mods:detail id="EB853754EE3F54A6D627A27E0B5B63F6" type="pubDate">
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<mods:number id="FD62AB1E616E8499C83F8A8FEA51EE29">2020-05-15</mods:number>
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</mods:detail>
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<mods:detail id="2B9E0E8517EFEE899A2DF4738200C42E" type="volume">
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<mods:number id="080A7AAA1F57B779ABA7D866B907C5F7">646</mods:number>
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</mods:detail>
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<mods:start id="D41925CF3C4ECE8E8F9DE3BC320940C8">1</mods:start>
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<mods:end id="3A41D7CD2CF8537EEAFBC2D253C7020E">55</mods:end>
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<mods:classification id="7FFA78FDA7FEF87BAF7FEE14FB2A6BE9">journal article</mods:classification>
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<mods:identifier id="44E21FEED89340A1C4F6F89D34D32CC5" type="CLB-Dataset">22092</mods:identifier>
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<mods:identifier id="D0BDE47BEE7355A5931F1B85EB6ECB63" type="DOI">10.5852/ejt.2020.646</mods:identifier>
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<treatment id="03B7726A135B5320FDA9FC54FEEBFC31" ID-DOI="http://doi.org/10.5281/zenodo.3848405" ID-GBIF-Taxon="164160404" ID-Zenodo-Dep="3848405" LSID="urn:lsid:zoobank.org:act:F31CFECF-7D04-4800-9559-EA234039B5DD" httpUri="http://treatment.plazi.org/id/03B7726A135B5320FDA9FC54FEEBFC31" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
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<subSubSection id="C30490F7135B5326FDA9FC54FB43FC6F" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8BA1C37C135B5326FDA9FC54FC5AFC4A" blockId="19.[367,1221,942,1039]" box="[600,988,942,969]" pageId="19" pageNumber="20">
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<heading id="D0E97410135B5326FDA9FC54FC5AFC4A" box="[600,988,942,969]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
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<taxonomicName id="4C1EB8FF135B5326FDA9FC54FCFCFC4A" ID-CoL="8MLNS" authority="Moret" authorityName="Moret" authorityYear="2020" box="[600,890,942,969]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="donosoi" status="sp. nov.">
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<emphasis id="B96A1F6E135B5326FDA9FC54FCADFC4A" bold="true" box="[600,811,942,969]" italics="true" pageId="19" pageNumber="20">Dyscolus donosoi</emphasis>
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Moret
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</taxonomicName>
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<taxonomicNameLabel id="A259A215135B5326FC70FC55FC5AFC4A" box="[897,988,943,969]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BA1C37C135B5326FE9EFC2BFB43FC6F" blockId="19.[367,1221,942,1039]" box="[367,1221,977,1004]" pageId="19" pageNumber="20">
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<uri id="FF8FCF7E135B5326FE9EFC2BFB43FC6F" box="[367,1221,977,1004]" pageId="19" pageNumber="20">
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urn:lsid:zoobank.org:act:
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<uuid id="FFB8F9A9135B5326FD7AFC2BFB43FC6F" box="[651,1221,977,1004]" pageId="19" pageNumber="20">F31CFECF-7D04-4800-9559-EA234039B5DD</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30490F7135B5326FD29FC0FFCDAFB8C" box="[728,860,1012,1039]" pageId="19" pageNumber="20" type="description">
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<paragraph id="8BA1C37C135B5326FD29FC0FFCDAFB8C" blockId="19.[367,1221,942,1039]" box="[728,860,1012,1039]" pageId="19" pageNumber="20">
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<figureCitation id="1325DFF9135B5326FD29FC0FFCDAFB8C" box="[728,860,1012,1039]" captionStart="Figs 15–18. 15–16" captionStartId="20.[189,241,1885,1911]" captionTargetBox="[224,1368,330,1828]" captionTargetId="figure@20.[198,1401,304,1841]" captionTargetPageId="20" captionText="Figs 15–18. 15–16. Dyscolus donosoi Moret sp. nov. 15. Habitus of the female holotype (QCAZ, COI voucher PM402-02). 16. Aedeagus, median lobe in lateral and ventral view. 17–18. D. danglesi Moret sp. nov. 17. Habitus of the male holotype (MNHN, COI voucher PM170-01). 18. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829692" httpUri="https://zenodo.org/record/3829692/files/figure.png" pageId="19" pageNumber="20">Figs 15–16</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30490F7135B5326FF4CFBCCFD97FB46" pageId="19" pageNumber="20" type="etymology">
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<paragraph id="8BA1C37C135B5326FF4CFBCCFEC2FBD3" blockId="19.[189,324,1078,1104]" box="[189,324,1078,1104]" pageId="19" pageNumber="20">
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<heading id="D0E97410135B5326FF4CFBCCFEC2FBD3" bold="true" box="[189,324,1078,1104]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
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<emphasis id="B96A1F6E135B5326FF4CFBCCFEC2FBD3" bold="true" box="[189,324,1078,1104]" pageId="19" pageNumber="20">Etymology</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BA1C37C135B5326FF4CFB9FFD97FB46" blockId="19.[189,1399,1125,1222]" pageId="19" pageNumber="20">
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Noun in the genitive case, dedicated to David A. Donoso, assistant professor at the Escuela Politécnica Nacional (Quito), specialist of the ecology of neotropical ants, who took part in the discovery of the
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<typeStatus id="54A57DDE135B5326FF4CFB51FEA2FB46" box="[189,292,1195,1221]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
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of this new species.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30490F7135B5320FF4CFB17FEEBFC31" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="materials_examined">
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<paragraph id="8BA1C37C135B5326FF4CFB17FEEAFA85" blockId="19.[189,364,1260,1287]" box="[189,364,1260,1287]" pageId="19" pageNumber="20">
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<heading id="D0E97410135B5326FF4CFB17FEEAFA85" bold="true" box="[189,364,1260,1287]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
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<emphasis id="B96A1F6E135B5326FF4CFB17FEEAFA85" bold="true" box="[189,364,1260,1287]" pageId="19" pageNumber="20">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA1C37C135B5326FF1BFAE1FEDAFAB6" blockId="19.[189,1399,1307,1439]" box="[234,348,1307,1333]" pageId="19" pageNumber="20">
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<emphasis id="B96A1F6E135B5326FF1BFAE1FEDAFAB6" bold="true" box="[234,348,1307,1333]" pageId="19" pageNumber="20">
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<typeStatus id="54A57DDE135B5326FF1BFAE1FEDAFAB6" box="[234,348,1307,1333]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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||
<paragraph id="8BA1C37C135B5326FF4CFAC4FAF5FA1C" blockId="19.[189,1399,1307,1439]" pageId="19" pageNumber="20">
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<materialsCitation id="3B76C921135B5326FF4CFAC4FAF5FA1C" ID-GBIF-Occurrence="2620303358" collectingDate="2015-03-18" collectionCode="QCAZ" collectorName="P. Moret & C. Ruiz & D. Donoso" country="Ecuador" elevation="2110" latitude="-3.9878056" location="Estacion El Colibri" longLatPrecision="1" longitude="-79.09417" municipality="Parque Nacional Podocarpus" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Zamora-Chinchipe Province" typeStatus="holotype">
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||
<collectingCountry id="F30983EC135B5326FF4CFAC4FECDFADB" box="[189,331,1342,1368]" name="Ecuador" pageId="19" pageNumber="20">ECUADOR</collectingCountry>
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||
•
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||
<specimenCount id="9D1808F5135B5326FE95FAC5FEFCFADA" box="[356,378,1343,1369]" pageId="19" pageNumber="20" type="female">♀</specimenCount>
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||
(lacks the right foreleg and the right middle leg);
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<collectingRegion id="49DA0D9E135B5326FC39FAC4FA9FFADB" box="[968,1305,1342,1368]" country="Ecuador" name="Zamora-Chinchipe" pageId="19" pageNumber="20">Zamora-Chinchipe Province</collectingRegion>
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,
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<collectingMunicipality id="6BC55906135B5326FAD4FAC5FE3BFAF8" pageId="19" pageNumber="20">Parque Nacional Podocarpus</collectingMunicipality>
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,
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<location id="8EC195A7135B5326FE3AFA9BFD33FAF8" LSID="urn:lsid:plazi:treatment:03B7726A135B5320FDA9FC54FEEBFC31:8EC195A7135B5326FE3AFA9BFD33FAF8" box="[459,693,1377,1403]" country="Ecuador" latitude="-3.9878056" longLatPrecision="1" longitude="-79.09417" municipality="Parque Nacional Podocarpus" name="Estacion El Colibri" pageId="19" pageNumber="20" stateProvince="Zamora-Chinchipe Province">Estación El Colibrí</location>
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;
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<geoCoordinate id="EE2AA5BB135B5326FD32FA9AFCDAFAFF" box="[707,860,1376,1404]" degrees="3" direction="south" minutes="59" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="16.1" value="-3.9878056">3°59′16.1″ S</geoCoordinate>
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,
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<geoCoordinate id="EE2AA5BB135B5326FC9DFA9AFB97FAFF" box="[876,1041,1376,1404]" degrees="79" direction="west" minutes="5" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="39.0" value="-79.09417">79°5′39.0″ W</geoCoordinate>
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;
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<quantity id="4CE66E99135B5326FBEEFA98FBFCFAFF" box="[1055,1146,1378,1404]" metricMagnitude="3" metricUnit="m" metricValue="2.11" pageId="19" pageNumber="20" unit="m" value="2110.0">2110 m</quantity>
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a.s.l.
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</elevation>
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;
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<date id="FFA0E5BC135B5326FB3DFA98FAF4FAFF" box="[1228,1394,1377,1404]" pageId="19" pageNumber="20" value="2015-03-18">
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<collectingDate id="EFE41C54135B5326FB3DFA98FAF4FAFF" box="[1228,1394,1377,1404]" pageId="19" pageNumber="20" value="2015-03-18">18 Mar. 2015</collectingDate>
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</date>
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;
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||
<collectorName id="26EBA6AA135B5326FF4CFA7FFEA2FA1C" box="[189,292,1413,1439]" pageId="19" pageNumber="20">P. Moret</collectorName>
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,
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||
<collectorName id="26EBA6AA135B5326FEDEFA7FFE0EFA1C" box="[303,392,1413,1439]" pageId="19" pageNumber="20">C. Ruiz</collectorName>
|
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and
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||
<collectorName id="26EBA6AA135B5326FE33FA7FFDC7FA1C" box="[450,577,1413,1439]" pageId="19" pageNumber="20">D. Donoso</collectorName>
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||
leg.; COI voucher PM402-02, BOLD sequence SUM187-18;
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||
<collectionCode id="ED0F5BB9135B5326FAEAFA7FFAE9FA1C" box="[1307,1391,1413,1439]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="19" pageNumber="20" type="Museum">QCAZ</collectionCode>
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||
.
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||
</materialsCitation>
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</paragraph>
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<paragraph id="8BA1C37C135B5326FF1BFA3CFE43FA63" blockId="19.[189,1399,1476,1646]" box="[234,453,1476,1504]" pageId="19" pageNumber="20">
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||
<emphasis id="B96A1F6E135B5326FF1BFA3CFEE1FA63" bold="true" box="[234,359,1478,1504]" pageId="19" pageNumber="20">
|
||
<typeStatus id="54A57DDE135B5326FF1BFA3CFEE1FA63" box="[234,359,1478,1504]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(2 ♂♂)
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135B5326FF4CFA13FA92F9EE" blockId="19.[189,1399,1476,1646]" pageId="19" pageNumber="20">
|
||
<materialsCitation id="3B76C921135B5326FF4CFA13FCEFF9C9" ID-GBIF-Occurrence="2620303384" collectionCode="GBNP, CISEC" collectorName="C. Ruiz & D. Marin" country="Ecuador" elevation="2140" latitude="-3.988" location="Fundacion Arcoiris" longLatPrecision="78" longitude="-79.093" municipality="Parque Nacional Podocarpus" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Zamora-Chinchipe Province" typeStatus="paratype">
|
||
<collectingCountry id="F30983EC135B5326FF4CFA13FECDF980" box="[189,331,1513,1539]" name="Ecuador" pageId="19" pageNumber="20">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="9D1808F5135B5326FE84FA13FE2FF987" box="[373,425,1512,1540]" pageId="19" pageNumber="20" type="male">1 ♂</specimenCount>
|
||
(lacks middle and hind legs);
|
||
<collectingRegion id="49DA0D9E135B5326FCCEFA13FB11F980" box="[831,1175,1513,1539]" country="Ecuador" name="Zamora-Chinchipe" pageId="19" pageNumber="20">Zamora-Chinchipe Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BC55906135B5326FB5AFA10FECFF9A5" pageId="19" pageNumber="20">Parque Nacional Podocarpus</collectingMunicipality>
|
||
,
|
||
<location id="8EC195A7135B5326FEA7F9F6FDB8F9A5" LSID="urn:lsid:plazi:treatment:03B7726A135B5320FDA9FC54FEEBFC31:8EC195A7135B5326FEA7F9F6FDB8F9A5" box="[342,574,1548,1574]" country="Ecuador" latitude="-3.988" longLatPrecision="78" longitude="-79.093" municipality="Parque Nacional Podocarpus" name="Fundacion Arcoiris" pageId="19" pageNumber="20" stateProvince="Zamora-Chinchipe Province">Fundación Arcoiris</location>
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||
;
|
||
<geoCoordinate id="EE2AA5BB135B5326FDBAF9F7FD37F9A4" box="[587,689,1549,1575]" degrees="3.988" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="55" value="-3.988">3.988º S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2AA5BB135B5326FD31F9F7FCC6F9A4" box="[704,832,1549,1575]" degrees="79.093" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="55" value="-79.093">79.093º W</geoCoordinate>
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||
;
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|
||
<quantity id="4CE66E99135B5326FCBFF9F7FC2FF9A4" box="[846,937,1549,1575]" metricMagnitude="3" metricUnit="m" metricValue="2.1399999999999997" pageId="19" pageNumber="20" unit="m" value="2140.0">2140 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<collectorName id="26EBA6AA135B5326FC08F9F7FBD2F9A5" box="[1017,1108,1548,1575]" pageId="19" pageNumber="20">C. Ruiz</collectorName>
|
||
and
|
||
<collectorName id="26EBA6AA135B5326FB60F9F7FB79F9A4" box="[1169,1279,1549,1575]" pageId="19" pageNumber="20">D. Marín</collectorName>
|
||
leg.; COI voucher CR017, BOLD sequence
|
||
<collectionCode id="ED0F5BB9135B5326FD90F9CAFD32F9C9" box="[609,692,1584,1610]" collectionName="GBNP" pageId="19" pageNumber="20">GBNP</collectionCode>
|
||
014-17;
|
||
<collectionCode id="ED0F5BB9135B5326FCE7F9CAFCEFF9C9" box="[790,873,1584,1610]" collectionName="CISEC" pageId="19" pageNumber="20">CISEC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B76C921135B5326FC7AF9CAFA96F9EE" ID-GBIF-Occurrence="2620303404" collectionCode="GBNP, CISEC" collectorName="C. Ruiz & D. Marin" country="Ecuador" elevation="2140" latitude="-3.988" location="Fundacion Arcoiris" longLatPrecision="78" longitude="-79.093" municipality="Parque Nacional Podocarpus" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
|
||
<specimenCount id="9D1808F5135B5326FC7AF9CAFC3AF9C9" box="[907,956,1582,1610]" pageId="19" pageNumber="20" type="male">1 ♂</specimenCount>
|
||
(lacks middle and hind legs); same collection data as for preceding; COI voucher CR018, BOLD sequence
|
||
<collectionCode id="ED0F5BB9135B5326FBF9F9A9FBDAF9ED" box="[1032,1116,1619,1646]" collectionName="GBNP" pageId="19" pageNumber="20">GBNP</collectionCode>
|
||
015-17;
|
||
<collectionCode id="ED0F5BB9135B5326FB48F9A9FA96F9EE" box="[1209,1296,1619,1645]" collectionName="CISEC" pageId="19" pageNumber="20">CISEC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135B5326FF4CF96EFE52F92D" blockId="19.[189,468,1684,1710]" box="[189,468,1684,1710]" pageId="19" pageNumber="20">
|
||
<heading id="D0E97410135B5326FF4CF96EFE52F92D" bold="true" box="[189,468,1684,1710]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B96A1F6E135B5326FF4CF96EFE52F92D" bold="true" box="[189,468,1684,1710]" pageId="19" pageNumber="20">Diagnostic description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135B5320FF4CF939FB5AFEEF" blockId="19.[189,1399,1731,2004]" lastBlockId="21.[189,1398,267,364]" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
|
||
Habitus:
|
||
<figureCitation id="1325DFF9135B5326FED4F939FEFEF95D" box="[293,376,1731,1758]" captionStart="Figs 15–18. 15–16" captionStartId="20.[189,241,1885,1911]" captionTargetBox="[224,1368,330,1828]" captionTargetId="figure@20.[198,1401,304,1841]" captionTargetPageId="20" captionText="Figs 15–18. 15–16. Dyscolus donosoi Moret sp. nov. 15. Habitus of the female holotype (QCAZ, COI voucher PM402-02). 16. Aedeagus, median lobe in lateral and ventral view. 17–18. D. danglesi Moret sp. nov. 17. Habitus of the male holotype (MNHN, COI voucher PM170-01). 18. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829692" httpUri="https://zenodo.org/record/3829692/files/figure.png" pageId="19" pageNumber="20">Fig. 15</figureCitation>
|
||
. Wingless. Body length:
|
||
<quantity id="4CE66E99135B5326FD64F939FCB8F95E" box="[661,830,1731,1757]" metricMagnitude="-2" metricUnit="m" metricValue="1.2850000000000001" metricValueMax="1.3900000000000001" metricValueMin="1.1800000000000002" pageId="19" pageNumber="20" unit="mm" value="12.85" valueMax="13.9" valueMin="11.8">11.8–13.9 mm</quantity>
|
||
. Body shiny black with a bluish, faintly iridescent lustre on the elytra; femora and tibiae nigropiceous, tarsi and palpi reddish brown; antennomeres 1–3 brunneopiceous, 4–11 reddish brown. Elytral microsculpture transverse. Head elongate, moderately convex, eyes relatively small, not protruding, genae slightly convex; mandibles very long, acute and arcuate at apex. Pronotum large, oval, completely rounded basally, with only one pair of lateral setae, lacking the basal pair; laterobasal depressions shallow. Elytra elongate-oval, convex; base arcuate; striae entire, broad and deeply impressed; intervals flat on disc, subconvex near apex; preapical sinuation obsolete. Third elytral interval without discal setae. Last visible abdominal ventrite. With one pair (♂) or two pairs (
|
||
<specimenCount id="9D1808F5135B5326FACBF843FACBF850" box="[1338,1357,1977,2003]" pageId="19" pageNumber="20" type="female">♀</specimenCount>
|
||
) of setae along its apical margin. Legs slender, fourth metatarsomere with one pair of subapical dorsolateral setae, apical lobes asymmetrical with a big outer lobe and no inner lobe.
|
||
<emphasis id="B96A1F6E135D5320FBF7FED5FB29FECA" box="[1030,1199,303,329]" italics="true" pageId="21" pageNumber="22">Male genitalia</emphasis>
|
||
:
|
||
<figureCitation id="1325DFF9135D5320FB4FFED4FA95FECB" box="[1214,1299,302,328]" captionStart="Figs 15–18. 15–16" captionStartId="20.[189,241,1885,1911]" captionTargetBox="[224,1368,330,1828]" captionTargetId="figure@20.[198,1401,304,1841]" captionTargetPageId="20" captionText="Figs 15–18. 15–16. Dyscolus donosoi Moret sp. nov. 15. Habitus of the female holotype (QCAZ, COI voucher PM402-02). 16. Aedeagus, median lobe in lateral and ventral view. 17–18. D. danglesi Moret sp. nov. 17. Habitus of the male holotype (MNHN, COI voucher PM170-01). 18. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829692" httpUri="https://zenodo.org/record/3829692/files/figure.png" pageId="21" pageNumber="22">Fig. 16</figureCitation>
|
||
. Median lobe arcuate with a long, spatulate apex, endophallus unarmed.
|
||
<emphasis id="B96A1F6E135D5320FC62FEA8FBD0FEEF" box="[915,1110,338,364]" italics="true" pageId="21" pageNumber="22">Female genitalia</emphasis>
|
||
: unstudied.
|
||
</paragraph>
|
||
<caption id="DF6193F4135C5321FF4CF8A7FEEDF863" ID-DOI="http://doi.org/10.5281/zenodo.3829692" ID-Zenodo-Dep="3829692" httpUri="https://zenodo.org/record/3829692/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,241,1885,1911]" targetBox="[224,1368,330,1828]" targetPageId="20">
|
||
<paragraph id="8BA1C37C135C5321FF4CF8A7FEEDF863" blockId="20.[189,1399,1885,2016]" pageId="20" pageNumber="21">
|
||
<emphasis id="B96A1F6E135C5321FF4CF8A7FE1DF8F4" bold="true" box="[189,411,1885,1911]" pageId="20" pageNumber="21">Figs 15–18. 15–16</emphasis>
|
||
.
|
||
<taxonomicName id="4C1EB8FF135C5321FE58F8A7FD4FF8FB" authority="Moret" authorityName="Moret" authorityYear="2020" box="[425,713,1885,1912]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="donosoi" status="sp. nov.">
|
||
<emphasis id="B96A1F6E135C5321FE58F8A7FDFCF8F4" box="[425,634,1885,1911]" italics="true" pageId="20" pageNumber="21">Dyscolus donosoi</emphasis>
|
||
Moret
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A259A215135C5321FD20F8A4FCAAF8FB" box="[721,812,1886,1912]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96A1F6E135C5321FCCAF8A7FCDEF8F4" bold="true" box="[827,856,1885,1911]" pageId="20" pageNumber="21">15</emphasis>
|
||
. Habitus of the female holotype (QCAZ, COI voucher PM402-02).
|
||
<emphasis id="B96A1F6E135C5321FE4BF87AFE51F819" bold="true" box="[442,471,1920,1946]" pageId="20" pageNumber="21">16</emphasis>
|
||
. Aedeagus, median lobe in lateral and ventral view.
|
||
<emphasis id="B96A1F6E135C5321FBB7F87AFB09F819" bold="true" box="[1094,1167,1920,1946]" pageId="20" pageNumber="21">17–18</emphasis>
|
||
.
|
||
<taxonomicName id="4C1EB8FF135C5321FB6FF87BFAF1F818" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1182,1399,1920,1947]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="danglesi" status="sp. nov.">
|
||
<emphasis id="B96A1F6E135C5321FB6FF87BFAA0F819" box="[1182,1318,1920,1946]" italics="true" pageId="20" pageNumber="21">D. danglesi</emphasis>
|
||
Moret
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A259A215135C5321FF4CF85EFE90F83D" box="[189,278,1956,1982]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96A1F6E135C5321FEEDF859FEBFF83E" bold="true" box="[284,313,1955,1981]" pageId="20" pageNumber="21">17</emphasis>
|
||
. Habitus of the male holotype (MNHN, COI voucher PM170-01).
|
||
<emphasis id="B96A1F6E135C5321FBCFF859FBDDF83E" bold="true" box="[1086,1115,1955,1981]" pageId="20" pageNumber="21">18</emphasis>
|
||
. Aedeagus, median lobe in lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA1C37C135D5320FF4CFE69FEE5FE2E" blockId="21.[189,355,403,429]" box="[189,355,403,429]" pageId="21" pageNumber="22">
|
||
<heading id="D0E97410135D5320FF4CFE69FEE5FE2E" bold="true" box="[189,355,403,429]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B96A1F6E135D5320FF4CFE69FEE5FE2E" bold="true" box="[189,355,403,429]" pageId="21" pageNumber="22">Comparisons</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135D5320FF4CFE38FC8BFD0F" blockId="21.[189,1399,450,652]" pageId="21" pageNumber="22">
|
||
Owing to the unique combination of a rounded basal half of pronotum, absence of pronotal hind setae, and asetose third interval of the elytra,
|
||
<taxonomicName id="4C1EB8FF135D5320FD60FE1FFC35FE7C" authority="Moret" authorityName="Moret" authorityYear="2020" box="[657,947,485,511]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="donosoi" status="sp. nov.">
|
||
<emphasis id="B96A1F6E135D5320FD60FE1FFCE4FE7C" box="[657,866,485,511]" italics="true" pageId="21" pageNumber="22">Dyscolus donosoi</emphasis>
|
||
Moret
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A259A215135D5320FC4AFE1FFB91FE7C" box="[955,1047,485,511]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has no close relatives among the described Ecuadorian
|
||
<emphasis id="B96A1F6E135D5320FE1CFDF2FDE7FDA1" box="[493,609,520,546]" italics="true" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C1EB8FF135D5320FE1CFDF2FDDAFDA1" authorityName="Dejean" authorityYear="1831" box="[493,604,520,546]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Dyscolus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In our molecular analysis (Fig. 2)
|
||
<taxonomicName id="4C1EB8FF135D5320FC08FDF3FB4FFDA0" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1017,1225,520,547]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="donosoi" status="sp. nov.">
|
||
<emphasis id="B96A1F6E135D5320FC08FDF3FBFCFDA1" box="[1017,1146,520,546]" italics="true" pageId="21" pageNumber="22">D. donosoi</emphasis>
|
||
Moret
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A259A215135D5320FB21FDF3FAADFDA0" box="[1232,1323,521,547]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
forms a clade with
|
||
<taxonomicName id="4C1EB8FF135D5320FEAEFDD6FD0AFDC6" authority="Moret, 1993" authorityName="Moret" authorityYear="1993" box="[351,652,555,582]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="saxatilis">
|
||
<emphasis id="B96A1F6E135D5320FEAEFDD6FE6FFDC6" box="[351,489,555,581]" italics="true" pageId="21" pageNumber="22">D. saxatilis</emphasis>
|
||
Moret, 1993
|
||
</taxonomicName>
|
||
, but the bootstrap support for this relationship is very weak.
|
||
<taxonomicName id="4C1EB8FF135D5320FF4CFDB5FE15FDEA" authorityName="Moret" authorityYear="1993" box="[189,403,591,617]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="saxatilis">
|
||
<emphasis id="B96A1F6E135D5320FF4CFDB5FE15FDEA" box="[189,403,591,617]" italics="true" pageId="21" pageNumber="22">Dyscolus saxatilis</emphasis>
|
||
</taxonomicName>
|
||
is a páramo high-altitude species with black non-metallic integuments, broad head, short mandibles, and short and robust appendages.
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135D5320FF4CFD49FE9BFD4E" blockId="21.[189,285,691,717]" box="[189,285,691,717]" pageId="21" pageNumber="22">
|
||
<heading id="D0E97410135D5320FF4CFD49FE9BFD4E" bold="true" box="[189,285,691,717]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B96A1F6E135D5320FF4CFD49FE9BFD4E" bold="true" box="[189,285,691,717]" pageId="21" pageNumber="22">Habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135D5320FF4CFD18FB47FC9C" blockId="21.[189,1398,738,799]" pageId="21" pageNumber="22">
|
||
Montane forest on the Eastern slope of the
|
||
<taxonomicName id="4C1EB8FF135D5320FD43FD18FC86FD7F" authorityName="Stal" authorityYear="1866" box="[690,768,738,764]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="54" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
|
||
, at around
|
||
<quantity id="4CE66E99135D5320FC71FD18FBA4FD7F" box="[896,1058,738,764]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="21" pageNumber="22" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100–2200 m</quantity>
|
||
a.s.l. Active at the beginning of the night (7–8.30 pm), on the trunk of a medium-sized tree, one meter above ground.
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135D5320FF4CFCBCFE6DFCE3" blockId="21.[189,491,838,864]" box="[189,491,838,864]" pageId="21" pageNumber="22">
|
||
<heading id="D0E97410135D5320FF4CFCBCFE6DFCE3" bold="true" box="[189,491,838,864]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B96A1F6E135D5320FF4CFCBCFE6DFCE3" bold="true" box="[189,491,838,864]" pageId="21" pageNumber="22">Geographic distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA1C37C135D5320FF4CFC8FFEEBFC31" blockId="21.[189,1399,885,946]" pageId="21" pageNumber="22">
|
||
Only known from the
|
||
<typeStatus id="54A57DDE135D5320FE35FC8FFE71FC0C" box="[452,503,885,911]" pageId="21" pageNumber="22">type</typeStatus>
|
||
locality in Southern
|
||
<collectingCountry id="F30983EC135D5320FD1EFC8FFCD5FC0C" box="[751,851,885,911]" name="Ecuador" pageId="21" pageNumber="22">Ecuador</collectingCountry>
|
||
, in the Parque Nacional Podocarpus. Probably microendemic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |