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<document id="C22E3FD51941296C2A89066E97D412E7" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A1351532FFDBDFB5AFEEBFB06" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus sulcipedis Moret 2020, sp. nov." docType="treatment" docUuid="611661EF-D8A2-4E28-8622-349EC9F6C992" docUuidSource="ZooBank" docVersion="8" lastPageNumber="27" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="26" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2D4380E774019655C1862C575FD7B809" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1A3B4F3DD2FE3017E9F7A1587F48DCF2">
<mods:title id="2E2D2F4E7057A975FBE37313457FE676">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="C8F8BF17BC1F8C36B26BF7C97D656FA5" type="personal">
<mods:role id="61909356DE3CA23BBAA27D5F586F261C">
<mods:roleTerm id="2E4DB922E9AE12E4D06C0364FDC4B692">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A015BBD3EE0D299C52D39E4D9DFAF981">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="86674D88155DC28D3C780442C137DC22" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="A98D76C1E5942AE4980FAE1BC29D1635">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="478B7FDD8E65DA23DC0CE9791152993C" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="27839D6FE52ECA6313E7330FDC80CA21" type="personal">
<mods:role id="E6A7C6BB522B5D8D60AD328C45220277">
<mods:roleTerm id="B90003CF7B39ADE325BCBBBB0D8A8AD0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="178EEF05E9C9E06429D84AC203DF3BC8">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="BC8C1BEA42E78FE6100C8BA1120837CA" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="DDEF93CBE95CAFEECEEC9F55F2C6BBB8">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="CD559E760B5856A3383AFB3908ECD8E6" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="03FF9BD82E31B55F3608F194BAFC0720">text</mods:typeOfResource>
<mods:relatedItem id="2896C997441227C71DC32C640298E1A8" type="host">
<mods:titleInfo id="CBCF6DDA38C0A95C5CDCA5E094870E63">
<mods:title id="702A06BBABBF31FDCDE8F438234542A1">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="F063CC3B3F7D71C53870108EC35C9534">
<mods:date id="7F59089F7E52253AD8D01B3665FBD1D1">2020</mods:date>
<mods:detail id="4B738992D77FC2C250DA01AEB3844DD7" type="pubDate">
<mods:number id="46D5B2779F6F92C90D1A54E64D19D30F">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="B4E586BF4FF45C12FDEF10BAA14C90EE" type="volume">
<mods:number id="E4EE59397E30AE1E7EE38FD00D7CA975">646</mods:number>
</mods:detail>
<mods:extent id="F0F6ABA91D727BA069E1691FFA95DAC2" unit="page">
<mods:start id="6EF63013AF3B276E9F437EE0012BFAD1">1</mods:start>
<mods:end id="C21ECB9FF2185E5EBED618B4E62C771C">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="20DAF0F57A2E1908693E7D2B6EA2CE14">journal article</mods:classification>
<mods:identifier id="73A4A6625F80E85D247221109E410494" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="E7F71252D75D4076E9232D15F1B26DD0" type="DOI">10.5852/ejt.2020.646</mods:identifier>
<mods:identifier id="71E376C91F88CD317252CCC58AE9B86C" type="GBIF-Dataset">c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b</mods:identifier>
<mods:identifier id="7E141AF69AE7FC5CA36773E112E817AD" type="Zenodo-Dep">3829682</mods:identifier>
<mods:identifier id="C54AC01260F1D195BC81098149BA8BF1" type="ZooBank">4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B</mods:identifier>
</mods:mods>
<treatment id="03B7726A1351532FFDBDFB5AFEEBFB06" ID-DOI="http://doi.org/10.5281/zenodo.3848369" ID-GBIF-Taxon="164160411" ID-Zenodo-Dep="3848369" LSID="urn:lsid:zoobank.org:act:611661EF-D8A2-4E28-8622-349EC9F6C992" httpUri="http://treatment.plazi.org/id/03B7726A1351532FFDBDFB5AFEEBFB06" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<subSubSection id="C30490F71351532CFDBDFB5AFB3AFB5E" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BA1C37C1351532CFDBDFB5AFC6EFB38" blockId="25.[375,1212,1184,1281]" box="[588,1000,1184,1211]" pageId="25" pageNumber="26">
<heading id="D0E974101351532CFDBDFB5AFC6EFB38" box="[588,1000,1184,1211]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="4C1EB8FF1351532CFDBDFB5AFC01FB38" ID-CoL="8M7VQ" authority="Moret" authorityName="Moret" authorityYear="2020" box="[588,903,1184,1211]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sulcipedis" status="sp. nov.">
<emphasis id="B96A1F6E1351532CFDBDFB5AFCB1FB38" bold="true" box="[588,823,1184,1211]" italics="true" pageId="25" pageNumber="26">Dyscolus sulcipedis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151351532CFC7FFB5BFC6EFB38" box="[910,1000,1185,1211]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C1351532CFE86FB39FB3AFB5E" blockId="25.[375,1212,1184,1281]" box="[375,1212,1219,1245]" pageId="25" pageNumber="26">
<uri id="FF8FCF7E1351532CFE86FB39FB3AFB5E" box="[375,1212,1219,1245]" pageId="25" pageNumber="26">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A91351532CFD64FB39FB3AFB5E" box="[661,1212,1219,1245]" pageId="25" pageNumber="26">611661EF-D8A2-4E28-8622-349EC9F6C992</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71351532CFD29FB1DFCDDFA82" box="[728,859,1255,1281]" pageId="25" pageNumber="26" type="description">
<paragraph id="8BA1C37C1351532CFD29FB1DFCDDFA82" blockId="25.[375,1212,1184,1281]" box="[728,859,1255,1281]" pageId="25" pageNumber="26">
<figureCitation id="1325DFF91351532CFD29FB1DFCDDFA82" box="[728,859,1255,1281]" captionStart="Figs 19–24" captionStartId="24.[189,241,1879,1905]" captionTargetBox="[306,1275,267,1823]" captionTargetId="figure@24.[269,1319,262,1835]" captionTargetPageId="24" captionText="Figs 19–24. Dyscolus spp., habitus (19–21) and aedeagus, median lobe in lateral view (22–24). 19. D. rugitarsis Moret sp. nov., male holotype (QACZ). 20. D. sulcipedis Moret sp. nov., male holotype (QACZ). 21. D. marini Moret sp. nov., female paratype. 22. D. rugitarsis Moret sp. nov. 23. D. sulcipedis Moret sp. nov. 24. D. marini Moret sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829694" httpUri="https://zenodo.org/record/3829694/files/figure.png" pageId="25" pageNumber="26">Figs 20, 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71351532CFF4CFAD2FC45FAF1" box="[189,963,1320,1394]" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BA1C37C1351532CFF4CFAD2FEC2FAC1" blockId="25.[189,324,1320,1346]" box="[189,324,1320,1346]" pageId="25" pageNumber="26">
<heading id="D0E974101351532CFF4CFAD2FEC2FAC1" bold="true" box="[189,324,1320,1346]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B96A1F6E1351532CFF4CFAD2FEC2FAC1" bold="true" box="[189,324,1320,1346]" pageId="25" pageNumber="26">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFAADFC45FAF1" blockId="25.[189,963,1367,1394]" box="[189,963,1367,1394]" pageId="25" pageNumber="26">Compound Latin adjective meaning ‘with sulcate feet’ (i.e., tarsi).</paragraph>
</subSubSection>
<subSubSection id="C30490F71351532FFF4CFA62FEEBFB06" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BA1C37C1351532CFF4CFA62FEEAFA31" blockId="25.[189,364,1432,1458]" box="[189,364,1432,1458]" pageId="25" pageNumber="26">
<heading id="D0E974101351532CFF4CFA62FEEAFA31" bold="true" box="[189,364,1432,1458]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B96A1F6E1351532CFF4CFA62FEEAFA31" bold="true" box="[189,364,1432,1458]" pageId="25" pageNumber="26">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1351532CFF1BFA3DFEDAFA62" blockId="25.[189,1399,1479,1576]" box="[234,348,1479,1505]" pageId="25" pageNumber="26">
<emphasis id="B96A1F6E1351532CFF1BFA3DFEDAFA62" bold="true" box="[234,348,1479,1505]" pageId="25" pageNumber="26">
<typeStatus id="54A57DDE1351532CFF1BFA3DFEDAFA62" box="[234,348,1479,1505]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFA10FBB8F9A4" blockId="25.[189,1399,1479,1576]" pageId="25" pageNumber="26">
<materialsCitation id="3B76C9211351532CFF4CFA10FBB8F9A4" ID-GBIF-Occurrence="2620303422" collectingDate="2016-08-04" collectionCode="QCAZ" collectorName="P. Moret &amp; S. Aguirre &amp; E. Moreno" country="Ecuador" elevation="3240" latitude="-4.711861" location="Parque Nacional Yacuri" longLatPrecision="1" longitude="-79.44035" municipality="Parque Nacional Yacuri" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Loja Province" typeStatus="holotype">
<collectingCountry id="F30983EC1351532CFF4CFA10FECDF987" box="[189,331,1514,1540]" name="Ecuador" pageId="25" pageNumber="26">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F51351532CFE96FA12FEF8F987" box="[359,382,1512,1540]" pageId="25" pageNumber="26" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E1351532CFE7FFA10FDBEF987" box="[398,568,1514,1540]" country="Ecuador" name="Loja" pageId="25" pageNumber="26">Loja Province</collectingRegion>
,
<collectingMunicipality id="6BC559061351532CFDB4FA10FCE4F987" box="[581,866,1514,1540]" pageId="25" pageNumber="26">Parque Nacional Yacuri</collectingMunicipality>
, Waypoint 167;
<geoCoordinate id="EE2AA5BB1351532CFBD7FA10FB31F987" box="[1062,1207,1514,1540]" degrees="4.711861" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="-4.711861">4.711861º S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB1351532CFB34FA10FAF4F987" box="[1221,1394,1514,1541]" degrees="79.440355" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="-79.44035">79.440355º W</geoCoordinate>
;
<elevation id="0033244F1351532CFF4CF9F7FED0F9A4" box="[189,342,1549,1576]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="25" pageNumber="26" unit="m" value="3240.0">
<quantity id="4CE66E991351532CFF4CF9F7FE90F9AB" box="[189,278,1549,1576]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="25" pageNumber="26" unit="m" value="3240.0">3240 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1351532CFE92F9F7FE71F9A4" box="[355,503,1549,1575]" pageId="25" pageNumber="26" value="2016-08-04">
<collectingDate id="EFE41C541351532CFE92F9F7FE71F9A4" box="[355,503,1549,1575]" pageId="25" pageNumber="26" value="2016-08-04">4 Aug. 2016</collectingDate>
</date>
;
<collectorName id="26EBA6AA1351532CFDF2F9F4FDECF9AB" box="[515,618,1550,1576]" pageId="25" pageNumber="26">P. Moret</collectorName>
,
<collectorName id="26EBA6AA1351532CFD84F9F7FD6BF9A4" box="[629,749,1549,1575]" pageId="25" pageNumber="26">S. Aguirre</collectorName>
and
<collectorName id="26EBA6AA1351532CFCD6F9F4FC22F9AB" box="[807,932,1550,1576]" pageId="25" pageNumber="26">E. Moreno</collectorName>
leg.;
<collectionCode id="ED0F5BB91351532CFC17F9F7FBBCF9A4" box="[998,1082,1549,1575]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="25" pageNumber="26" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C1351532CFF1BF9B5FD90F9EA" blockId="25.[189,1399,1613,1746]" box="[234,534,1613,1641]" pageId="25" pageNumber="26">
<emphasis id="B96A1F6E1351532CFF1BF9B5FEE1F9EA" bold="true" box="[234,359,1615,1641]" pageId="25" pageNumber="26">
<typeStatus id="54A57DDE1351532CFF1BF9B5FEE1F9EA" box="[234,359,1615,1641]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
(2 ♂♂,
<specimenCount id="9D1808F51351532CFE3BF9B5FD8CF9EA" box="[458,522,1615,1641]" pageId="25" pageNumber="26" type="female">2 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CF988FE23F951" blockId="25.[189,1399,1613,1746]" pageId="25" pageNumber="26">
<materialsCitation id="3B76C9211351532CFF4CF988FC42F90F" ID-GBIF-Occurrence="2620303385" box="[189,964,1648,1676]" collectingDate="2016-08-04" collectionCode="MNHN" collectorName="P. Moret &amp; S. Aguirre &amp; E. Moreno" country="Ecuador" elevation="3240" latitude="-4.711861" location="ECUADOR" longLatPrecision="1" longitude="-79.44035" municipality="Parque Nacional Yacuri" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<collectingCountry id="F30983EC1351532CFF4CF988FECDF90F" box="[189,331,1650,1676]" name="Ecuador" pageId="25" pageNumber="26">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F51351532CFE99F988FE11F90F" box="[360,407,1648,1676]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB91351532CFC9BF988FC42F90F" box="[874,964,1650,1676]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="26">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211351532CFC11F988FEF4F92C" ID-GBIF-Occurrence="2620303388" collectingDate="2016-08-04" collectionCode="CPM" collectorName="P. Moret &amp; S. Aguirre &amp; E. Moreno" country="Ecuador" elevation="3240" latitude="-4.711861" location="Waypoint" longLatPrecision="1" longitude="-79.44035" municipality="Parque Nacional Yacuri" pageId="25" pageNumber="26" specimenCount="2" specimenCount-female="2" stateProvince="Loja">
<specimenCount id="9D1808F51351532CFC11F988FBA3F90F" box="[992,1061,1650,1676]" pageId="25" pageNumber="26" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB91351532CFEC5F96FFEF4F92C" box="[308,370,1685,1711]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="25" pageNumber="26">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211351532CFE7DF96FFE27F951" ID-GBIF-Occurrence="2620303363" collectingDate="2016-08-04" collectionCode="CPM" collectorName="P. Moret &amp; S. Aguirre &amp; E. Moreno" country="Ecuador" elevation="3240" latitude="-4.711861" location="Waypoint" longLatPrecision="1" longitude="-79.44035" municipality="Parque Nacional Yacuri" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F51351532CFE7DF96FFE3CF92C" box="[396,442,1683,1711]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
; same collection data as for holotype; COI voucher PM167-09, BOLD sequence SUM061-18;
<collectionCode id="ED0F5BB91351532CFEAEF942FE27F951" box="[351,417,1720,1746]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="25" pageNumber="26">CPM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CF903FE52F890" blockId="25.[189,468,1785,1811]" box="[189,468,1785,1811]" pageId="25" pageNumber="26">
<heading id="D0E974101351532CFF4CF903FE52F890" bold="true" box="[189,468,1785,1811]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B96A1F6E1351532CFF4CF903FE52F890" bold="true" box="[189,468,1785,1811]" pageId="25" pageNumber="26">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1351532FFF4CF8D2FD05FE7B" blockId="25.[189,1398,1832,1999]" lastBlockId="26.[189,1399,267,505]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Habitus:
<figureCitation id="1325DFF91351532CFED5F8D2FEF0F8C1" box="[292,374,1832,1859]" captionStart="Figs 19–24" captionStartId="24.[189,241,1879,1905]" captionTargetBox="[306,1275,267,1823]" captionTargetId="figure@24.[269,1319,262,1835]" captionTargetPageId="24" captionText="Figs 19–24. Dyscolus spp., habitus (19–21) and aedeagus, median lobe in lateral view (22–24). 19. D. rugitarsis Moret sp. nov., male holotype (QACZ). 20. D. sulcipedis Moret sp. nov., male holotype (QACZ). 21. D. marini Moret sp. nov., female paratype. 22. D. rugitarsis Moret sp. nov. 23. D. sulcipedis Moret sp. nov. 24. D. marini Moret sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829694" httpUri="https://zenodo.org/record/3829694/files/figure.png" pageId="25" pageNumber="26">Fig.20</figureCitation>
.Wingless.Body length:
<quantity id="4CE66E991351532CFD7AF8D2FCB2F8C0" box="[651,820,1832,1859]" metricMagnitude="-2" metricUnit="m" metricValue="1.105" metricValueMax="1.15" metricValueMin="1.06" pageId="25" pageNumber="26" unit="mm" value="11.05" valueMax="11.5" valueMin="10.6">10.6–11.5mm</quantity>
.Body colour variable,from brunneous to brownish black; femora and tibiae brownish; tarsi, antennae and mouthparts reddish brown. Elytral microsculpture transverse, weakly impressed. Head moderately broad, constricted basally, eyes moderately bulging, genae flat. Pronotum subcordiform, transverse, sides arcuate anterad, markedly sinuate posterad; hind angles slightly obtuse or almost right angled; two pairs of lateral setae. Elytra elongate-ovoid, slightly convex; striae entire, well impressed; intervals variable, flat to slightly convex in basal half, convex at apex. Third elytral interval with three setae. Last visible abdominal ventrite with two pairs (♂) or four pairs (
<specimenCount id="9D1808F51352532FFEFAFEA8FE98FEEF" box="[267,286,338,364]" pageId="26" pageNumber="27" type="female">♀</specimenCount>
) of setae along its apical margin. Legs: tarsi smooth dorsally, metatarsomeres 1–3 bisulcate, fourth metatarsomere with one pair of dorsolateral subapical setae, apical lobes moderately long, the external lobe 1.5 times longer than the inner lobe; fifth metatarsomere convex dorsally, asetose ventrally.
<emphasis id="B96A1F6E1352532FFF4CFE41FEEEFE56" box="[189,360,443,469]" italics="true" pageId="26" pageNumber="27">Male genitalia</emphasis>
:
<figureCitation id="1325DFF91352532FFE86FE41FE4BFE56" box="[375,461,443,469]" captionStart="Figs 19–24" captionStartId="24.[189,241,1879,1905]" captionTargetBox="[306,1275,267,1823]" captionTargetId="figure@24.[269,1319,262,1835]" captionTargetPageId="24" captionText="Figs 19–24. Dyscolus spp., habitus (19–21) and aedeagus, median lobe in lateral view (22–24). 19. D. rugitarsis Moret sp. nov., male holotype (QACZ). 20. D. sulcipedis Moret sp. nov., male holotype (QACZ). 21. D. marini Moret sp. nov., female paratype. 22. D. rugitarsis Moret sp. nov. 23. D. sulcipedis Moret sp. nov. 24. D. marini Moret sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829694" httpUri="https://zenodo.org/record/3829694/files/figure.png" pageId="26" pageNumber="27">Fig. 23</figureCitation>
. Median lobe weakly arcuate, apex short, acute, endophallus without sclerotized structure.
<emphasis id="B96A1F6E1352532FFEC3FE25FE7CFE7A" box="[306,506,479,505]" italics="true" pageId="26" pageNumber="27">Female genitalia</emphasis>
: unstudied.
</paragraph>
<paragraph id="8BA1C37C1352532FFF4CFDDAFEE5FDB9" blockId="26.[189,355,544,570]" box="[189,355,544,570]" pageId="26" pageNumber="27">
<heading id="D0E974101352532FFF4CFDDAFEE5FDB9" bold="true" box="[189,355,544,570]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96A1F6E1352532FFF4CFDDAFEE5FDB9" bold="true" box="[189,355,544,570]" pageId="26" pageNumber="27">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1352532FFF4CFDB5FC6FFCDC" blockId="26.[189,1399,590,863]" pageId="26" pageNumber="27">
<taxonomicName id="4C1EB8FF1352532FFF4CFDB5FE7FFDEA" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,505,591,617]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sulcipedis" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFF4CFDB5FE21FDEA" box="[189,423,591,617]" italics="true" pageId="26" pageNumber="27">Dyscolus sulcipedis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFDF3FDB5FDD9FDEA" box="[514,607,591,617]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C1EB8FF1352532FFCEAFDB5FCEFFDEB" authorityName="Moret" authorityYear="2020" box="[795,873,591,616]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="marini">
<emphasis id="B96A1F6E1352532FFCEAFDB5FCEFFDEB" box="[795,873,591,616]" italics="true" pageId="26" pageNumber="27">marini</emphasis>
</taxonomicName>
clade along with
<taxonomicName id="4C1EB8FF1352532FFBB2FDB5FA8FFDEA" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1091,1289,591,617]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFBB2FDB5FB31FDEB" box="[1091,1207,591,616]" italics="true" pageId="26" pageNumber="27">D. marini</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFAE2FDB5FAF4FDEA" box="[1299,1394,591,617]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C1EB8FF1352532FFF4CFD89FEEFFD0F" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,361,626,652]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ruizi" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFF4CFD89FE9EFD0F" box="[189,280,627,652]" italics="true" pageId="26" pageNumber="27">D. ruizi</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFE80FD88FE56FD0F" box="[369,464,626,652]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C1EB8FF1352532FFE2CFD89FD54FD0F" authority="Moret" authorityName="Moret" authorityYear="2020" box="[477,722,626,652]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="arborarius" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFE2CFD89FD07FD0F" box="[477,641,626,652]" italics="true" pageId="26" pageNumber="27">D. arborarius</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFD2AFD88FCBFFD0F" box="[731,825,626,652]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C1EB8FF1352532FFCB7FD89FC68FD0F" authorityName="Perrault" authorityYear="1993" box="[838,1006,627,652]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="moretianus">
<emphasis id="B96A1F6E1352532FFCB7FD89FC68FD0F" box="[838,1006,627,652]" italics="true" pageId="26" pageNumber="27">D. moretianus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1EB8FF1352532FFBDBFD89FA94FD0F" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1066,1298,626,652]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFBDBFD89FB44FD0F" box="[1066,1218,627,652]" italics="true" pageId="26" pageNumber="27">D. rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFAEAFD88FAF1FD0F" box="[1307,1399,626,652]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
Based on external morphology (not on molecular data), its closest relative is
<taxonomicName id="4C1EB8FF1352532FFBDEFD6CFA97FD2C" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1071,1297,661,687]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFBDEFD6CFB45FD2C" box="[1071,1219,662,687]" italics="true" pageId="26" pageNumber="27">D. rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFAE6FD6FFAF4FD2C" box="[1303,1394,661,687]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, from which it differs mainly in the following characters: elytral microsculpture transverse, weakly impressed (coarser and mostly isodiametric in
<taxonomicName id="4C1EB8FF1352532FFD03FD26FC5BFD75" authority="Moret" authorityName="Moret" authorityYear="2020" box="[754,989,732,758]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFD03FD26FC0DFD76" box="[754,907,732,757]" italics="true" pageId="26" pageNumber="27">D. rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFC16FD26FBCFFD75" box="[999,1097,732,758]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
); head slightly narrower; latero-posterior sinuation of the pronotum deeper; last visible abdominal ventrite with two pairs (♂) or four pairs (
<specimenCount id="9D1808F51352532FFEB3FCD8FED3FCBF" box="[322,341,802,828]" pageId="26" pageNumber="27" type="female">♀</specimenCount>
) of apical setae (one and two pairs in
<taxonomicName id="4C1EB8FF1352532FFCEDFCD9FB85FCBF" authority="Moret" authorityName="Moret" authorityYear="2020" box="[796,1027,802,828]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1352532FFCEDFCD9FC35FCBF" box="[796,947,803,828]" italics="true" pageId="26" pageNumber="27">D. rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151352532FFBFBFCD8FBECFCBF" box="[1034,1130,802,828]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
); tarsi not rugose; fifth metatarsomere plesiomorphic, not broadened and dorsally depressed.
</paragraph>
<paragraph id="8BA1C37C1352532FFF4CFC7CFE9BFC23" blockId="26.[189,285,902,928]" box="[189,285,902,928]" pageId="26" pageNumber="27">
<heading id="D0E974101352532FFF4CFC7CFE9BFC23" bold="true" box="[189,285,902,928]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96A1F6E1352532FFF4CFC7CFE9BFC23" bold="true" box="[189,285,902,928]" pageId="26" pageNumber="27">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1352532FFF4CFC4FFD60FC71" blockId="26.[189,1399,949,1011]" pageId="26" pageNumber="27">
Upper montane forest, at around
<quantity id="4CE66E991352532FFDB0FC4FFD1CFC4C" box="[577,666,949,975]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="26" pageNumber="27" unit="m" value="3240.0">3240 m</quantity>
a.s.l. Active at the beginning of the night on the surface of the leaf litter and on tree trunks
<quantity id="4CE66E991352532FFDF8FC22FDB0FC70" box="[521,566,984,1011]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="m" value="2.0">2 m</quantity>
above ground.
</paragraph>
<paragraph id="8BA1C37C1352532FFF4CFBE3FE6DFBB0" blockId="26.[189,491,1049,1075]" box="[189,491,1049,1075]" pageId="26" pageNumber="27">
<heading id="D0E974101352532FFF4CFBE3FE6DFBB0" bold="true" box="[189,491,1049,1075]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96A1F6E1352532FFF4CFBE3FE6DFBB0" bold="true" box="[189,491,1049,1075]" pageId="26" pageNumber="27">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1352532FFF4CFBB2FEEBFB06" blockId="26.[189,1399,1096,1157]" pageId="26" pageNumber="27">
Only known from the
<typeStatus id="54A57DDE1352532FFE15FBB3FD91FBE0" box="[484,535,1097,1123]" pageId="26" pageNumber="27">type</typeStatus>
locality in Southern
<collectingCountry id="F30983EC1352532FFCC1FBB2FC12FBE1" box="[816,916,1096,1122]" name="Ecuador" pageId="26" pageNumber="27">Ecuador</collectingCountry>
, in Parque Nacional Yacuri. Probably microendemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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