treatments-xml/data/D9/75/87/D97587D9D11BFFEA83AAFF32FC2DA193.xml

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<mods:title>A new species of Fusigobius (Teleostei: Gobiidae) from the Red Sea and Gulf of Aden</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bogorodsky, Sergey V.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kovačić, Marcelo</mods:namePart>
<mods:affiliation>Natural History Museum Rijeka, Lorenzov prolaz 1, HR- 51000 Rijeka, Croatia.</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mal, Ahmad O.</mods:namePart>
<mods:affiliation>Marine Biology Department, Faculty of Marine Sciences, King Abdulaziz University, Jeddah, Saudi Arabia.</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Alpermann, Tilman J.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8723-4576</mods:nameIdentifier>
<mods:affiliation>Tilman. Alpermann @ senckenberg. de; https: // orcid. org / 0000 - 0002 - 8723 - 4576</mods:affiliation>
<mods:nameIdentifier type="email">lpermann@senckenberg.de</mods:nameIdentifier>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-17</mods:number>
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<mods:number>5256</mods:number>
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<heading bold="true" box="[151,982,150,178]" fontSize="11" level="1" pageId="5" pageNumber="106" reason="1">
<emphasis bold="true" box="[151,982,150,178]" pageId="5" pageNumber="106">
<taxonomicName authority="Kovacic, Bogorodsky &amp; Alpermann" authorityName="Kovacic, Bogorodsky &amp; Alpermann" authorityYear="2023" box="[151,880,150,178]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis bold="true" box="[151,428,151,178]" italics="true" pageId="5" pageNumber="106">Fusigobius humerosus</emphasis>
Kovačić, Bogorodsky &amp; Alpermann
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<taxonomicNameLabel box="[888,982,151,177]" pageId="5" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph blockId="5.[151,982,150,250]" box="[151,518,187,213]" pageId="5" pageNumber="106">
<collectingRegion box="[151,243,187,213]" country="Egypt" name="Al Bahr al Ahmar" pageId="5" pageNumber="106">Red Sea</collectingRegion>
Shoulderspot Sandgoby
</paragraph>
</subSubSection>
<subSubSection box="[158,507,223,250]" pageId="5" pageNumber="106" type="description">
<paragraph blockId="5.[151,982,150,250]" box="[158,507,223,250]" pageId="5" pageNumber="106">
(
<figureCitation box="[166,303,223,250]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,250,1864,1888]" captionStartId-1="7.[151,250,1090,1114]" captionStartId-2="9.[151,250,1868,1892]" captionStartId-3="10.[151,250,1927,1951]" captionTargetBox-0="[225,1363,309,1840]" captionTargetBox-1="[211,1436,215,1019]" captionTargetBox-2="[151,1436,201,1843]" captionTargetBox-3="[151,1436,181,1903]" captionTargetId-0="figure-107@6.[225,1363,309,1840]" captionTargetId-1="figure-20@7.[151,1436,181,1067]" captionTargetId-2="figure-20@9.[151,1436,201,1844]" captionTargetId-3="figure-26@10.[151,1436,181,1903]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 2. Fusigobius humerosus sp. nov. Paratype, KAUMM 199, female, 27.6 + 7.9 mm, Al Lith, Saudi Arabia, Red Sea: A: the origin of posterior nostril about equal to orbit and to the origin of anterior nostril, the origin of nostrils and the edge of orbit outlined; B: 1–2 rows of cycloid scales extends above opercle. C: Paratype, PMR VP5272, male, 24.1 + 6.7 mm, Al Lith, Saudi Arabia, Red Sea, the scales ending anterior at line between upper pectoral-fin base and first dorsal fin origin. The anteriormost scales marked with arrow. Photos by M. Kovačić." captionText-1="FIGURE 3. Cephalic sensory pores system and papillae of Fusigobius humerosus sp. nov. Paratype, KAUMM 199, female, 27.6 + 7.9 mm, Al Lith, Saudi Arabia, Red Sea. Terminology in text. Drawing by M. Kovačić." captionText-2="FIGURE 4. Fusigobius humerosus sp. nov. holotype, SMF 35901, male, 28.7 + 8.0 mm, northern side of Farasan Island, Saudi Arabia, Red Sea, A: live specimen, B: freshly collected, C: preserved specimen. Photos by S.V. Bogorodsky (A &amp; B) and M. Kovačić (C)." captionText-3="FIGURE 5. Fusigobius humerosus sp. nov., underwater photographs of uncollected individuals. A: Dahab, Egypt, Red Sea; B: Al Wajh, Saudi Arabia, Red Sea; C: Mangrove Bay, El Quseir, Egypt, Red Sea. Photos by S.V. Bogorodsky." figureDoi-0="http://doi.org/10.5281/zenodo.7745453" figureDoi-1="http://doi.org/10.5281/zenodo.7745457" figureDoi-2="http://doi.org/10.5281/zenodo.7745459" figureDoi-3="http://doi.org/10.5281/zenodo.7745461" httpUri-0="https://zenodo.org/record/7745453/files/figure.png" httpUri-1="https://zenodo.org/record/7745457/files/figure.png" httpUri-2="https://zenodo.org/record/7745459/files/figure.png" httpUri-3="https://zenodo.org/record/7745461/files/figure.png" pageId="5" pageNumber="106">Figures 2–5</figureCitation>
, 
<figureCitation box="[315,404,223,249]" captionStart="FIGURE 6" captionStartId="11.[151,250,1156,1180]" captionTargetBox="[151,1436,181,1131]" captionTargetId="figure-20@11.[151,1436,181,1131]" captionTargetPageId="11" captionText="FIGURE 6. A and C: Fusigobius humerosus sp. nov., PMR VP5271, paratype, male, 26.2 + 7.3 mm, northern side of Farasan Island, Saudi Arabia, Red Sea. B and D: F. humeralis, BPBM 36627, paratype, female, 29.5 + 7.8 mm, Manuk Island, Banda Sea, Indonesia. Humeral region just above base of pectoral fin. A: round black spot is smaller than or equal to pupil in F. humerosus sp. nov., B: dark spot larger than pupil in F. humeralis; midbase of caudal fin: C: oval black spot covers less than 1/3 of caudal base height in F. humerosus sp. nov., D: irregular spot at midbase of caudal fin larger than 1/3 of caudal base height in F. humeralis." figureDoi="http://doi.org/10.5281/zenodo.7745463" httpUri="https://zenodo.org/record/7745463/files/figure.png" pageId="5" pageNumber="106">6A &amp; C</figureCitation>
, 
<tableCitation box="[415,496,223,249]" captionStart="TABLE 1" captionStartId="16.[151,239,476,500]" captionTargetPageId="16" captionText="TABLE 1. Morphometric characters (% SL) of Fusigobius humerosus sp. nov. Characters are sorted in alphabetic order." httpUri="http://table.plazi.org/id/05A36647D10EFFF783AAFE79FF1FA456" pageId="5" pageNumber="106" tableUuid="05A36647D10EFFF783AAFE79FF1FA456">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="106" type="reference_group">
<paragraph blockId="5.[151,1436,291,408]" pageId="5" pageNumber="106">
<treatmentCitationGroup pageId="5" pageNumber="106">
<taxonomicName ID-CoL="6K63K" baseAuthorityName="Randall" baseAuthorityYear="2001" box="[151,372,292,316]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[151,372,292,316]" italics="true" pageId="5" pageNumber="106">Fusigobius humeralis</emphasis>
</taxonomicName>
(non Randall) — 
<treatmentCitation author="Randall, J. E." box="[558,746,292,316]" page="212" pageId="5" pageNumber="106" year="2001">
<bibRefCitation author="Randall, J. E." box="[558,746,292,316]" pageId="5" pageNumber="106" pagination="206 - 225" refId="ref16951" refString="Randall, J. E. (2001) Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zoological Studies, 40 (3), 206 - 225. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall 2001: 212</bibRefCitation>
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(in part, 
<collectingRegion box="[840,925,292,316]" country="Egypt" name="Al Bahr al Ahmar" pageId="5" pageNumber="106">Red Sea</collectingRegion>
&amp; Gulf of Aden); 
<treatmentCitation author="Ryanskiy, A." box="[1115,1321,291,316]" page="105" pageId="5" pageNumber="106" year="2022">
<bibRefCitation author="Ryanskiy, A." box="[1115,1321,291,316]" pageId="5" pageNumber="106" refId="ref17035" refString="Ryanskiy, A. (2022) Red Sea marine life. Andrey Ryanskiy publisher, 200 pp." type="book" year="2022">Ryanskiy 2022: 105</bibRefCitation>
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, line 2, left (Sharm el Naga, 
<collectingCountry box="[371,436,322,346]" name="Egypt" pageId="5" pageNumber="106">Egypt</collectingCountry>
).
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</paragraph>
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<treatmentCitationGroup box="[151,926,353,377]" pageId="5" pageNumber="106">
<taxonomicName ID-CoL="6K63J" authorityName="Goren" authorityYear="1978" box="[151,387,353,377]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="longispinus">
<emphasis box="[151,387,353,377]" italics="true" pageId="5" pageNumber="106">Fusigobius longispinus</emphasis>
</taxonomicName>
(non Goren) — 
<treatmentCitation author="Lips, J. &amp; Lips, B. &amp; Roux, J. - M." box="[556,753,353,377]" page="96" pageId="5" pageNumber="106" year="2016">
<bibRefCitation author="Lips, J. &amp; Lips, B. &amp; Roux, J. - M." box="[556,753,353,377]" pageId="5" pageNumber="106" refId="ref16510" refString="Lips, J., Lips, B. &amp; Roux, J. - M. (2016) Poissons de Djibouti. Universite de Djibouti, 248 pp." type="book" year="2016">
Lips 
<emphasis box="[607,658,353,377]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
2016: 96
</bibRefCitation>
</treatmentCitation>
(
<collectingRegion box="[767,816,353,377]" country="Djibouti" name="Arta" pageId="5" pageNumber="106">Arta</collectingRegion>
, 
<collectingCountry box="[826,915,353,377]" name="Djibouti" pageId="5" pageNumber="106">Djibouti</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="5.[151,1436,291,408]" box="[151,946,383,408]" pageId="5" pageNumber="106">
<treatmentCitationGroup box="[151,946,383,408]" pageId="5" pageNumber="106">
<taxonomicName ID-CoL="6JTRQ" baseAuthorityName="Randall" baseAuthorityYear="2001" box="[151,360,384,408]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="maximus">
<emphasis box="[151,360,384,408]" italics="true" pageId="5" pageNumber="106">Fusigobius maximus</emphasis>
</taxonomicName>
(non Randall) — 
<treatmentCitation author="Ryanskiy, A." box="[545,750,383,408]" page="105" pageId="5" pageNumber="106" year="2022">
<bibRefCitation author="Ryanskiy, A." box="[545,750,383,408]" pageId="5" pageNumber="106" refId="ref17035" refString="Ryanskiy, A. (2022) Red Sea marine life. Andrey Ryanskiy publisher, 200 pp." type="book" year="2022">Ryanskiy 2022: 105</bibRefCitation>
</treatmentCitation>
, line 3, left (
<collectingRegion box="[877,934,384,408]" country="Egypt" name="Shamal Sina'" pageId="5" pageNumber="106">Sinai</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="106" type="materials_examined">
<paragraph blockId="5.[151,1437,454,2028]" pageId="5" pageNumber="106">
<materialsCitation collectingDate="2017-02-09" collectionCode="T, V" collectorName="T. J. Alpermann &amp; S. V. Bogorodsky &amp; M. Sonnewald" country="Saudi Arabia" elevation="5" latitude="16.91315" location="Red Sea" longLatPrecision="1" longitude="41.841568" municipality="Farasan Island" pageId="5" pageNumber="106" specimenCode="SMF 35901" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[151,260,454,480]" pageId="5" pageNumber="106">
<typeStatus box="[151,260,454,480]" pageId="5" pageNumber="106">Holotype</typeStatus>
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. 
<specimenCode box="[273,405,454,481]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35901</specimenCode>
(sample number KAU17-167), male, 28.7 + 8.0 mm, 
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, 
<collectingCountry box="[1104,1251,454,480]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, northern side of 
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, a small sand patch at wreck, 
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, 
<geoCoordinate box="[816,962,490,517]" degrees="41" direction="east" minutes="50.494" orientation="longitude" pageId="5" pageNumber="106" precision="1" value="41.841568">41°50.494’E</geoCoordinate>
, 
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<elevation box="[973,1047,490,517]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="106" unit="m" value="4.5" valueMax="5.0" valueMin="4.0">4–5 m</elevation>
</quantity>
, 
<collectorName box="[1059,1234,490,516]" pageId="5" pageNumber="106">T.J. Alpermann</collectorName>
, 
<collectorName box="[1245,1436,490,516]" pageId="5" pageNumber="106">S.V. Bogorodsky</collectorName>
&amp; 
<collectorName box="[180,346,526,552]" pageId="5" pageNumber="106">M. Sonnewald</collectorName>
, 
<date box="[357,560,526,552]" pageId="5" pageNumber="106" value="2017-02-09">
<collectingDate box="[357,560,526,552]" pageId="5" pageNumber="106" value="2017-02-09">09 February 2017</collectingDate>
</date>
.
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</paragraph>
<paragraph blockId="5.[151,1437,454,2028]" pageId="5" pageNumber="106">
<emphasis bold="true" box="[199,576,561,588]" pageId="5" pageNumber="106">
<typeStatus box="[199,320,562,588]" pageId="5" pageNumber="106">Paratypes</typeStatus>
, all from the Red Sea
</emphasis>
. 
<materialsCitation collectionCode="KAUMM" country="Saudi Arabia" elevation="8" latitude="20.245417" location="Saudi Arabia" longLatPrecision="1" longitude="40.00095" pageId="5" pageNumber="106" specimenCode="KAUMM 199" specimenCount="1" typeStatus="paratype">
<specimenCode box="[590,751,562,588]" collectionCode="KAUMM" pageId="5" pageNumber="106">KAUMM 199</specimenCode>
, female, 27.6 + 
<quantity box="[931,1019,562,588]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="5" pageNumber="106" unit="mm" value="7.9">7.9 mm</quantity>
, 
<collectingCountry box="[1030,1179,562,588]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, Al Lith, 
<geoCoordinate box="[1283,1431,562,589]" degrees="20" direction="north" minutes="14.725" orientation="latitude" pageId="5" pageNumber="106" precision="1" value="20.245417">20°14.725’N</geoCoordinate>
, 
<geoCoordinate box="[151,310,598,625]" degrees="040" direction="east" minutes="00.057" orientation="longitude" pageId="5" pageNumber="106" precision="1" value="40.00095">040°00.057’E</geoCoordinate>
, 
<quantity box="[323,369,598,624]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="106" unit="m" value="8.0">
<elevation box="[323,369,598,624]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="106" unit="m" value="8.0">8 m</elevation>
</quantity>
, fringing reef, coll
</materialsCitation>
. 
<materialsCitation box="[596,822,598,624]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[596,616,598,624]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE281BFFDF3FCC9A43E" box="[642,768,598,624]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[826,1166,598,624]" collectingDate="2013-06-19" collectionCode="V" country="Saudi Arabia" location="Bogorodsky" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[826,849,598,624]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28060FDF3FC2EA43E" box="[861,999,598,624]" country="Saudi Arabia" name="Bogorodsky" pageId="5" pageNumber="106">Bogorodsky</location>
, 
<date box="[1011,1166,598,624]" pageId="5" pageNumber="106" value="2013-06-19">
<collectingDate box="[1011,1166,598,624]" pageId="5" pageNumber="106" value="2013-06-19">19 June 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="KAUMM" country="Saudi Arabia" elevation="7" latitude="16.598984" location="Farasan Isl." longLatPrecision="1" longitude="42.3346" pageId="5" pageNumber="106" specimenCode="KAUMM 463" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1179,1339,598,624]" collectionCode="KAUMM" pageId="5" pageNumber="106">KAUMM 463</specimenCode>
(sample number KAU12-91), male, 31.2 + 
<quantity box="[550,641,634,660]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="5" pageNumber="106" unit="mm" value="8.2">8.2 mm</quantity>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE281B3FDDFFCDAA4DA" box="[654,787,634,660]" country="Saudi Arabia" latitude="16.598984" longLatPrecision="1" longitude="42.3346" name="Farasan Isl." pageId="5" pageNumber="106">Farasan Isl.</location>
, Mazaghef, 
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, 
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, 
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</quantity>
, coll
</materialsCitation>
. 
<materialsCitation collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[1396,1416,634,660]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
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&amp; S
</materialsCitation>
. 
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<collectionCode box="[335,358,670,696]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
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, 
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<collectingDate box="[521,727,670,696]" pageId="5" pageNumber="106" value="2012-02-19">19 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="PMR" country="Saudi Arabia" elevation="5" latitude="16.91315" location="Farasan Island" longLatPrecision="1" longitude="41.841568" pageId="5" pageNumber="106" specimenCode="VP5271" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
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, 
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, 25.2 + 6.3, and 25.0 + 
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and male, 26.4 + 
<quantity box="[302,391,706,732]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="106" unit="mm" value="7.3">7.3 mm</quantity>
, 
<collectingCountry box="[404,554,706,732]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, northern side of 
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, 
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, 
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, 
<quantity box="[1252,1326,706,733]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="106" unit="m" value="4.5" valueMax="5.0" valueMin="4.0">
<elevation box="[1252,1326,706,733]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="106" unit="m" value="4.5" valueMax="5.0" valueMin="4.0">4–5 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[1396,1416,706,732]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE283AAFD43FEDCA54E" box="[151,277,742,768]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[337,732,742,768]" collectingDate="2017-02-09" collectionCode="V" country="Saudi Arabia" location="Bogorodsky" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
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. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2824BFD43FDC9A54E" box="[374,512,742,768]" country="Saudi Arabia" name="Bogorodsky" pageId="5" pageNumber="106">Bogorodsky</location>
, 
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<collectingDate box="[525,732,742,768]" pageId="5" pageNumber="106" value="2017-02-09">09 February 2017</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="PMR" country="Saudi Arabia" elevation="7" latitude="20.13235" location="Al Lith" longLatPrecision="1" longitude="40.09878" pageId="5" pageNumber="106" specimenCode="VP5272" specimenCount="1" typeStatus="paratype">
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, male, 24.1 + 
<quantity box="[1078,1168,742,768]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="106" unit="mm" value="6.7">6.7 mm</quantity>
, 
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, 
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, 
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, 
<geoCoordinate box="[309,454,778,805]" degrees="40" direction="east" minutes="05.927" orientation="longitude" pageId="5" pageNumber="106" precision="1" value="40.09878">40°05.927’E</geoCoordinate>
, 
<quantity box="[463,534,778,804]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">
<elevation box="[463,534,778,804]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">6–8 m</elevation>
</quantity>
, a small isolated coral reef, coll
</materialsCitation>
. 
<materialsCitation box="[887,1103,778,804]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[887,907,778,804]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2809FFCAFFBE9A56A" box="[930,1056,778,804]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation collectingDate="2014-11-16" collectionCode="V" country="Saudi Arabia" location="Bogorodsky" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[1107,1130,778,804]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2874FFCAFFB35A56A" box="[1138,1276,778,804]" country="Saudi Arabia" name="Bogorodsky" pageId="5" pageNumber="106">Bogorodsky</location>
, 
<date pageId="5" pageNumber="106" value="2014-11-16">
<collectingDate pageId="5" pageNumber="106" value="2014-11-16">16 November 2014</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="SMF" country="Saudi Arabia" elevation="14" latitude="27.095089" location="Saudi Arabia" longLatPrecision="1" longitude="35.778442" pageId="5" pageNumber="106" specimenCode="SMF 35224" specimenCount="1" typeStatus="paratype">
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, female, 23.2 + 
<quantity box="[549,638,814,840]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="106" unit="mm" value="6.4">6.4 mm</quantity>
, 
<collectingCountry box="[652,802,814,840]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, Duba, 
<geoCoordinate box="[891,1066,814,841]" degrees="27" direction="north" minutes="05" orientation="latitude" pageId="5" pageNumber="106" precision="1" seconds="42.32" value="27.095089">27°05’42.32”N</geoCoordinate>
, 
<geoCoordinate box="[1079,1251,814,841]" degrees="35" direction="east" minutes="46" orientation="longitude" pageId="5" pageNumber="106" precision="1" seconds="42.39" value="35.778442">35°46’42.39”E</geoCoordinate>
, 
<quantity box="[1264,1367,814,841]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="5" pageNumber="106" unit="m" value="14.0" valueMax="15.0" valueMin="13.0">
<elevation box="[1264,1367,814,841]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="5" pageNumber="106" unit="m" value="14.0" valueMax="15.0" valueMin="13.0">13–15 m</elevation>
</quantity>
, coral block on steep slope of island, coll. S
</materialsCitation>
. 
<materialsCitation box="[570,904,850,876]" collectingDate="2013-06-19" collectionCode="V" country="Saudi Arabia" location="Bogorodsky" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[570,593,850,876]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28166FCF7FD2CA522" box="[603,741,850,876]" country="Saudi Arabia" name="Bogorodsky" pageId="5" pageNumber="106">Bogorodsky</location>
, 
<date box="[752,904,850,876]" pageId="5" pageNumber="106" value="2013-06-19">
<collectingDate box="[752,904,850,876]" pageId="5" pageNumber="106" value="2013-06-19">19 June 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="SMF" collectorName="Abkar I." country="Saudi Arabia" elevation="7" latitude="16.618717" location="Farasan Isl." longLatPrecision="1" longitude="41.933968" pageId="5" pageNumber="106" specimenCode="SMF 35902" specimenCount="1" typeStatus="paratype">
<specimenCode box="[915,1051,850,877]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35902</specimenCode>
, female, 22.1 + 
<quantity box="[1229,1317,850,876]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="5" pageNumber="106" unit="mm" value="6.2">6.2 mm</quantity>
, and male, 25.1 + 
<quantity box="[231,320,886,912]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="106" unit="mm" value="7.4">7.4 mm</quantity>
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<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28270FCD3FE19A5DE" box="[333,464,886,912]" country="Saudi Arabia" latitude="16.618717" longLatPrecision="1" longitude="41.933968" name="Farasan Isl." pageId="5" pageNumber="106">Farasan Isl.</location>
, 
<collectorName box="[476,571,886,912]" pageId="5" pageNumber="106">Abkar I.</collectorName>
, 
<geoCoordinate box="[584,732,886,912]" degrees="16" direction="north" minutes="37.123" orientation="latitude" pageId="5" pageNumber="106" precision="1" value="16.618717">16°37.123’N</geoCoordinate>
, 
<geoCoordinate box="[745,904,886,913]" degrees="041" direction="east" minutes="56.038" orientation="longitude" pageId="5" pageNumber="106" precision="1" value="41.933968">041°56.038’E</geoCoordinate>
, 
<quantity box="[916,962,886,912]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="106" unit="m" value="7.0">
<elevation box="[916,962,886,912]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="106" unit="m" value="7.0">7 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation box="[1031,1255,886,912]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[1031,1051,886,912]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28708FCD3FB7AA5DE" box="[1077,1203,886,912]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<collectionCode box="[1259,1282,886,912]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. Bogorodsky, 
<date box="[151,350,922,948]" pageId="5" pageNumber="106" value="2012-02-27">27 February 2012</date>
(used only to study morphological characters in the mouth cavity and gill chamber); 
<materialsCitation collectionCode="SMF" country="Saudi Arabia" elevation="7" latitude="16.598984" location="Farasan Isl." longLatPrecision="1" longitude="42.3346" pageId="5" pageNumber="106" specimenCode="SMF 35904" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1303,1436,922,949]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35904</specimenCode>
(sample number KAU12-40), female, 28.2 + 7.0 mm, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE281C3FC1BFC4AA596" box="[766,899,958,984]" country="Saudi Arabia" latitude="16.598984" longLatPrecision="1" longitude="42.3346" name="Farasan Isl." pageId="5" pageNumber="106">Farasan Isl.</location>
, Mazaghef, 
<geoCoordinate box="[1039,1187,958,985]" degrees="16" direction="north" minutes="35.939" orientation="latitude" pageId="5" pageNumber="106" precision="1" value="16.598984">16°35.939’N</geoCoordinate>
, 
<geoCoordinate box="[1200,1345,958,984]" degrees="42" direction="east" minutes="20.076" orientation="longitude" pageId="5" pageNumber="106" precision="1" value="42.3346">42°20.076’E</geoCoordinate>
, 
<quantity box="[1357,1431,958,984]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">
<elevation box="[1357,1431,958,984]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">6–8 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation box="[209,439,994,1020]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[209,229,994,1020]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2823CFC47FEB6A5B2" box="[257,383,994,1020]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[444,840,994,1020]" collectingDate="2012-02-19" collectionCode="V" country="Saudi Arabia" location="Bogorodsky" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[444,467,994,1020]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE282DDFC47FDA3A5B2" box="[480,618,994,1020]" country="Saudi Arabia" name="Bogorodsky" pageId="5" pageNumber="106">Bogorodsky</location>
, 
<date box="[632,840,994,1020]" pageId="5" pageNumber="106" value="2012-02-19">
<collectingDate box="[632,840,994,1020]" pageId="5" pageNumber="106" value="2012-02-19">19 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="SMF" country="Saudi Arabia" elevation="7" latitude="16.91555" location="Farasan Isl." longLatPrecision="1" longitude="41.846016" pageId="5" pageNumber="106" specimenCode="SMF 35905" specimenCount="1" typeStatus="paratype">
<specimenCode box="[863,1002,994,1021]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35905</specimenCode>
, female, 20.9 + 
<quantity box="[1193,1283,994,1021]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="5" pageNumber="106" unit="mm" value="5.7">5.7 mm</quantity>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2862FFC47FA5EA5B2" box="[1298,1431,994,1020]" country="Saudi Arabia" latitude="16.91555" longLatPrecision="1" longitude="41.846016" name="Farasan Isl." pageId="5" pageNumber="106">Farasan Isl.</location>
, 
<geoCoordinate box="[151,299,1030,1057]" degrees="16" direction="north" minutes="54.933" orientation="latitude" pageId="5" pageNumber="106" precision="1" value="16.91555">16°54.933’N</geoCoordinate>
, 
<geoCoordinate box="[311,470,1030,1057]" degrees="041" direction="east" minutes="50.761" orientation="longitude" pageId="5" pageNumber="106" precision="1" value="41.846016">041°50.761’E</geoCoordinate>
, 
<quantity box="[482,527,1030,1056]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="106" unit="m" value="7.0">
<elevation box="[482,527,1030,1056]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="106" unit="m" value="7.0">7 m</elevation>
</quantity>
, isolated coral block, coll
</materialsCitation>
. 
<materialsCitation box="[827,1049,1030,1056]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[827,847,1030,1056]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28055FBA3FC2FA26E" box="[872,998,1030,1056]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[1053,1394,1030,1056]" collectingDate="2011-04-02" collectionCode="V" country="Saudi Arabia" location="Bogorodsky" pageId="5" pageNumber="106" specimenCount="1" typeStatus="paratype">
<collectionCode box="[1053,1076,1030,1056]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28702FBA3FB00A26E" box="[1087,1225,1030,1056]" country="Saudi Arabia" name="Bogorodsky" pageId="5" pageNumber="106">Bogorodsky</location>
, 
<date box="[1236,1394,1030,1056]" pageId="5" pageNumber="106" value="2011-04-02">
<collectingDate box="[1236,1394,1030,1056]" pageId="5" pageNumber="106" value="2011-04-02">02 April 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="5.[151,1437,454,2028]" pageId="5" pageNumber="106">
<emphasis bold="true" box="[199,420,1065,1092]" pageId="5" pageNumber="106">Non-type material</emphasis>
. 
<emphasis bold="true" box="[437,546,1065,1092]" pageId="5" pageNumber="106">Red Sea:</emphasis>
<materialsCitation box="[556,1432,1066,1093]" collectionCode="BPBM" country="Israel" location="Gulf of Aqaba" pageId="5" pageNumber="106" specimenCode="BPBM 18272" specimenCount="1">
<specimenCode box="[556,717,1066,1092]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="106" type="Museum">BPBM 18272</specimenCode>
, female, 
<quantity box="[823,928,1066,1093]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="5" pageNumber="106" unit="mm" value="23.5">23.5 mm</quantity>
, caudal fin damaged, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2879DFB8FFA81A20A" box="[1184,1352,1066,1092]" country="Israel" name="Gulf of Aqaba" pageId="5" pageNumber="106">Gulf of Aqaba</location>
, 
<collectingCountry box="[1366,1432,1066,1092]" name="Israel" pageId="5" pageNumber="106">Israel</collectingCountry>
</materialsCitation>
; 
<materialsCitation box="[151,847,1102,1129]" collectionCode="BPBM" country="Yemen" location="Hanish" pageId="5" pageNumber="106" specimenCode="BPBM 35714" specimenCount="1">
<specimenCode box="[151,310,1102,1129]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="106" type="Museum">BPBM 35714</specimenCode>
, female, 25.6 + 
<quantity box="[495,585,1102,1128]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="106" unit="mm" value="6.9">6.9 mm</quantity>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2816BFBEBFD6CA226" box="[598,677,1102,1128]" country="Yemen" name="Hanish" pageId="5" pageNumber="106">Hanish</location>
group, 
<collectingCountry box="[765,847,1102,1128]" name="Yemen" pageId="5" pageNumber="106">Yemen</collectingCountry>
</materialsCitation>
; 
<materialsCitation collectionCode="KAUMM" country="Saudi Arabia" elevation="7" location="Amaq" pageId="5" pageNumber="106" specimenCode="KAUMM 464" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCode box="[859,1023,1102,1128]" collectionCode="KAUMM" pageId="5" pageNumber="106">KAUMM 464</specimenCode>
, 
<specimenCount box="[1035,1125,1102,1128]" pageId="5" pageNumber="106" type="male">2 males</specimenCount>
, 24.7 + 
<quantity box="[1219,1306,1102,1129]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="106" unit="mm" value="7.5">7.5 mm</quantity>
and 24.4 + 
<quantity box="[151,241,1138,1164]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="106" unit="mm" value="7.4">7.4 mm</quantity>
, and 
<specimenCount box="[304,417,1138,1164]" pageId="5" pageNumber="106" type="female">2 females</specimenCount>
, 20.9 + 
<quantity box="[512,600,1138,1165]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="106" unit="mm" value="6.5">6.5 mm</quantity>
and 
<quantity box="[658,760,1138,1164]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="5" pageNumber="106" unit="mm" value="18.7">18.7 mm</quantity>
+ caudal fin damaged, 
<collectingCountry box="[1032,1182,1138,1164]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28797FBD7FB38A2C2" box="[1194,1265,1138,1164]" country="Saudi Arabia" name="Amaq" pageId="5" pageNumber="106">Amaq</location>
, 
<quantity box="[1278,1325,1138,1164]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="106" unit="m" value="7.0">
<elevation box="[1278,1325,1138,1164]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="106" unit="m" value="7.0">7 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[1396,1416,1138,1164]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE283AAFB33FEDCA2FE" box="[151,277,1174,1200]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[332,688,1174,1200]" collectingDate="2011-03-31" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[332,355,1174,1200]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[366,504,1174,1200]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
<date box="[515,688,1174,1200]" pageId="5" pageNumber="106" value="2011-03-31">
<collectingDate box="[515,688,1174,1200]" pageId="5" pageNumber="106" value="2011-03-31">31 March 2011</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="KAUMM" country="Saudi Arabia" elevation="8" location="Al Lith" pageId="5" pageNumber="106" specimenCode="KAUMM 465" specimenCount="1">
<specimenCode box="[699,862,1174,1201]" collectionCode="KAUMM" pageId="5" pageNumber="106">KAUMM 465</specimenCode>
, female, 20.2 + 
<quantity box="[1041,1129,1174,1200]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="5" pageNumber="106" unit="mm" value="6.3">6.3 mm</quantity>
, 
<collectingCountry box="[1141,1289,1174,1200]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2862EFB33FAAFA2FE" box="[1299,1382,1174,1200]" country="Saudi Arabia" name="Al Lith" pageId="5" pageNumber="106">Al Lith</location>
, 
<quantity metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.5" valueMax="9.0" valueMin="6.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.5" valueMax="9.0" valueMin="6.0">6–9 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation box="[249,480,1210,1236]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[249,269,1210,1236]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28214FB1FFE6EA29A" box="[297,423,1210,1236]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[484,899,1210,1236]" collectingDate="2014-11-18" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[484,507,1210,1236]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[521,659,1210,1236]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
<date box="[673,899,1210,1236]" pageId="5" pageNumber="106" value="2014-11-18">
<collectingDate box="[673,899,1210,1236]" pageId="5" pageNumber="106" value="2014-11-18">18 November 2014</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectingDate="2012-02-26" collectionCode="KAUMM" country="Saudi Arabia" location="Farasan Arch." pageId="5" pageNumber="106" specimenCode="KAUMM 466" specimenCount="1">
<specimenCode box="[913,1076,1210,1236]" collectionCode="KAUMM" pageId="5" pageNumber="106">KAUMM 466</specimenCode>
(sample number KAU12-443), male, 26.2 + 7.0 mm, 
<collectingCountry box="[393,540,1246,1272]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2811AFB7BFD0DA2B6" box="[551,708,1246,1272]" country="Saudi Arabia" name="Farasan Arch." pageId="5" pageNumber="106">Farasan Arch.</location>
, 
<date box="[718,919,1246,1272]" pageId="5" pageNumber="106" value="2012-02-26">
<collectingDate box="[718,919,1246,1272]" pageId="5" pageNumber="106" value="2012-02-26">26 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="KAUMM" country="Saudi Arabia" elevation="7" location="Farasan Arch." pageId="5" pageNumber="106" specimenCode="KAUMM 467" specimenCount="1">
<specimenCode box="[929,1088,1246,1272]" collectionCode="KAUMM" pageId="5" pageNumber="106">KAUMM 467</specimenCode>
(sample number KAU12-445), female, 26.6 + 
<quantity box="[320,409,1282,1308]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="5" pageNumber="106" unit="mm" value="7.8">7.8 mm</quantity>
, 
<collectingCountry box="[420,569,1282,1308]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28179FAA7FD2AA352" box="[580,739,1282,1308]" country="Saudi Arabia" name="Farasan Arch." pageId="5" pageNumber="106">Farasan Arch.</location>
, 
<quantity box="[750,823,1282,1308]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">
<elevation box="[750,823,1282,1308]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="106" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">6–8 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation box="[890,1113,1282,1308]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[890,910,1282,1308]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28095FAA7FBEFA352" box="[936,1062,1282,1308]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation collectingDate="2012-02-26" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[1117,1140,1282,1308]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[1151,1289,1282,1308]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
<date pageId="5" pageNumber="106" value="2012-02-26">
<collectingDate pageId="5" pageNumber="106" value="2012-02-26">26 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation box="[225,1155,1318,1345]" collectionCode="SMF" country="Saudi Arabia" elevation="10" location="Al Khoreybah" pageId="5" pageNumber="106" specimenCode="SMF 35207" specimenCount="1">
<specimenCode box="[225,363,1318,1345]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35207</specimenCode>
, male, 19.0 + 
<quantity box="[531,622,1318,1345]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="106" unit="mm" value="5.2">5.2 mm</quantity>
, 
<collectingCountry box="[636,787,1318,1344]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28022FA83FC0DA30E" box="[799,964,1318,1344]" country="Saudi Arabia" name="Al Khoreybah" pageId="5" pageNumber="106">Al Khoreybah</location>
, 
<quantity box="[978,1067,1318,1344]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="5" pageNumber="106" unit="m" value="10.0" valueMax="12.0" valueMin="8.0">
<elevation box="[978,1067,1318,1344]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="5" pageNumber="106" unit="m" value="10.0" valueMax="12.0" valueMin="8.0">8–12 m</elevation>
</quantity>
, coll. S
</materialsCitation>
. 
<materialsCitation collectingDate="2013-06-23" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[1159,1182,1318,1344]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[1196,1334,1318,1344]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
<date pageId="5" pageNumber="106" value="2013-06-23">
<collectingDate pageId="5" pageNumber="106" value="2013-06-23">23 June 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="SMF" country="Saudi Arabia" elevation="8" location="Al Lith" pageId="5" pageNumber="106" specimenCode="SMF 35906" specimenCount="2" specimenCount-male="2">
<specimenCode box="[222,359,1354,1381]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35906</specimenCode>
, 
<specimenCount box="[370,459,1354,1380]" pageId="5" pageNumber="106" type="male">2 males</specimenCount>
, 31.5 + 
<quantity box="[550,637,1354,1380]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="5" pageNumber="106" unit="mm" value="8.9">8.9 mm</quantity>
and 23.4 + 
<quantity box="[771,859,1354,1380]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="106" unit="mm" value="7.3">7.3 mm</quantity>
, and female, 21.9 + 
<quantity box="[1088,1177,1354,1380]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="106" unit="mm" value="6.8">6.8 mm</quantity>
, 
<collectingCountry box="[1189,1337,1354,1380]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2867EFAEFFA5EA32A" box="[1347,1431,1354,1380]" country="Saudi Arabia" name="Al Lith" pageId="5" pageNumber="106">Al Lith</location>
, 
<quantity box="[151,197,1390,1416]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="106" unit="m" value="8.0">
<elevation box="[151,197,1390,1416]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="106" unit="m" value="8.0">8 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation box="[265,489,1390,1416]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[265,285,1390,1416]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2820AFACBFE7CA3C6" box="[311,437,1390,1416]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[493,853,1390,1417]" collectingDate="2012-03-05" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[493,516,1390,1416]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[527,665,1390,1416]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
<date box="[677,853,1390,1417]" pageId="5" pageNumber="106" value="2012-03-05">
<collectingDate box="[677,853,1390,1417]" pageId="5" pageNumber="106" value="2012-03-05">05 March 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="SMF" country="Saudi Arabia" elevation="10" location="Rabigh" pageId="5" pageNumber="106" specimenCode="SMF 35907" specimenCount="1">
<specimenCode box="[864,1001,1390,1417]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35907</specimenCode>
, female, 22.1 + 
<quantity box="[1183,1271,1390,1416]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="106" unit="mm" value="6.9">6.9 mm</quantity>
, 
<collectingCountry box="[1283,1432,1390,1416]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE283AAFA37FF22A3E2" box="[151,235,1426,1452]" country="Saudi Arabia" name="Rabigh" pageId="5" pageNumber="106">Rabigh</location>
, 
<quantity box="[246,305,1426,1452]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="106" unit="m" value="10.0">
<elevation box="[246,305,1426,1452]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="106" unit="m" value="10.0">10 m</elevation>
</quantity>
, coll
</materialsCitation>
. 
<materialsCitation box="[372,595,1426,1452]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[372,392,1426,1452]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2829FFA37FDE9A3E2" box="[418,544,1426,1452]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[599,940,1426,1452]" collectingDate="2011-04-08" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[599,622,1426,1452]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[633,771,1426,1452]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
<date box="[782,940,1426,1452]" pageId="5" pageNumber="106" value="2011-04-08">
<collectingDate box="[782,940,1426,1452]" pageId="5" pageNumber="106" value="2011-04-08">08 April 2011</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectionCode="SMF" country="Saudi Arabia" elevation="7" location="Farasan Arch." pageId="5" pageNumber="106" specimenCode="SMF 35908" specimenCount="1">
<specimenCode box="[952,1085,1426,1453]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35908</specimenCode>
(sample number KAU12-442), male, 29.8 + 
<quantity box="[303,393,1462,1488]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="5" pageNumber="106" unit="mm" value="7.8">7.8 mm</quantity>
, 
<collectingCountry box="[406,556,1462,1488]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28105FA13FD10A39E" box="[568,729,1462,1488]" country="Saudi Arabia" name="Farasan Arch." pageId="5" pageNumber="106">Farasan Arch.</location>
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. 
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.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2809DFA13FBD7A39E" box="[928,1054,1462,1488]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
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. 
<materialsCitation collectingDate="2012-02-26" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[1112,1135,1462,1488]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[1148,1286,1462,1488]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
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<collectingDate pageId="5" pageNumber="106" value="2012-02-26">26 February 2012</collectingDate>
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; 
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<specimenCode box="[223,357,1498,1525]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="106">SMF 35909</specimenCode>
(sample number KAU12-444), male, 24.3 + 
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, 
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, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE28757FA7FFAC0A3BA" box="[1130,1289,1498,1524]" country="Saudi Arabia" name="Farasan Arch." pageId="5" pageNumber="106">Farasan Arch.</location>
, 
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, coll
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. 
<materialsCitation box="[151,375,1534,1560]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[151,171,1534,1560]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
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&amp; S
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. 
<materialsCitation box="[379,768,1534,1560]" collectingDate="2012-02-26" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[379,402,1534,1560]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[414,552,1534,1560]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
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<collectingDate box="[563,768,1534,1560]" pageId="5" pageNumber="106" value="2012-02-26">26 February 2012</collectingDate>
</date>
</materialsCitation>
; 
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(sample number KAU12-446), female, 27.4 + 
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, 
<collectingCountry box="[252,401,1570,1596]" name="Saudi Arabia" pageId="5" pageNumber="106">Saudi Arabia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE282A1F987FDF2A072" box="[412,571,1570,1596]" country="Saudi Arabia" name="Farasan Arch." pageId="5" pageNumber="106">Farasan Arch.</location>
, 
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, coll
</materialsCitation>
. 
<materialsCitation box="[724,948,1570,1596]" collectionCode="T" country="Saudi Arabia" location="Alpermann" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[724,744,1570,1596]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="106">T</collectionCode>
.J. 
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2803FF987FC49A072" box="[770,896,1570,1596]" country="Saudi Arabia" name="Alpermann" pageId="5" pageNumber="106">Alpermann</location>
&amp; S
</materialsCitation>
. 
<materialsCitation box="[952,1340,1570,1596]" collectingDate="2012-02-26" collectionCode="V" collectorName="Bogorodsky" country="Saudi Arabia" pageId="5" pageNumber="106" specimenCount="1">
<collectionCode box="[952,975,1570,1596]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="106" type="Museum">V</collectionCode>
. 
<collectorName box="[987,1125,1570,1596]" pageId="5" pageNumber="106">Bogorodsky</collectorName>
, 
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<collectingDate box="[1136,1340,1570,1596]" pageId="5" pageNumber="106" value="2012-02-26">26 February 2012</collectingDate>
</date>
</materialsCitation>
. 
<emphasis bold="true" pageId="5" pageNumber="106">
<materialsCitation country="Saudi Arabia" location="Gulf of Aden" pageId="5" pageNumber="106" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:D97587D9D11BFFEA83AAFF32FC2DA193:54036014D11BFFE2867AF984FF12A011" country="Saudi Arabia" name="Gulf of Aden" pageId="5" pageNumber="106">Gulf of Aden</location>
</materialsCitation>
:
</emphasis>
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<specimenCode box="[231,389,1606,1633]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="106" type="Museum">BPBM 35726</specimenCode>
, male, 28.2 + 
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, 
<collectingCountry box="[647,744,1606,1632]" name="Djibouti" pageId="5" pageNumber="106">Djibouti</collectingCountry>
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.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="107" pageId="5" pageNumber="106" type="diagnosis">
<paragraph blockId="5.[151,1437,454,2028]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="107" pageId="5" pageNumber="106">
<emphasis bold="true" box="[199,314,1641,1667]" pageId="5" pageNumber="106">Diagnosis</emphasis>
. Dorsal-fin rays VI+I,8–9; anal-fin rays I,8; pectoral-fin rays 17–18; first dorsal-fin spine longest, 
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head length, 19.9–23.6% SL, subsequent spines progressively shorter; first and second dorsal-fin spines not elongate or filamentous; first dorsal fin slightly higher than second in females, about equal in males; anal-fin spine length 
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head length; pelvic fins noticeably joined medially by membrane, fifth ray branched, shortest branch of fifth pelvic-fin soft ray 83–94% of longest branch of fourth ray, 18.1-22.1% SL; pelvic frenum present, its width at midline about 1/4–1/5 of pelvic-spine length; body depth 
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SL; snout short, its length 4.0– 
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head length, 
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SL; posterior nostril about halfway between orbit and anterior nostril (
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); eye diameter 3.0– 
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head length; upper-jaw length 
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head length; gill openings extending forward to about half of opercle width from posterior edge of preopercle; longitudinal scale series 26–28; head and predorsal area naked, scales on side of nape usually ending just anterior at line between upper pectoral-fin base and origin of first dorsal fin (
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) or 1–2 rows of cycloid scales extending forward to above preopercular edge (
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); body translucent with numerous small dusky orange-yellow spots on head and body, series of melanophores within and adjacent of each spot; a round black spot smaller than or as large as pupil in humeral region just above base of pectoral fin and equal or slightly larger than pupil a black spot at midbase of caudal fin, covering less than 1/3 height of caudal-fin base; first dorsal fin with oblique dusky line from middle of first spine to anterior base of third spine.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7745453" ID-Zenodo-Dep="7745453" httpUri="https://zenodo.org/record/7745453/files/figure.png" pageId="6" pageNumber="107" startId="6.[151,250,1864,1888]" targetBox="[225,1363,309,1840]" targetPageId="6">
<paragraph blockId="6.[151,1437,1864,2032]" pageId="6" pageNumber="107">
<emphasis bold="true" box="[151,274,1864,1888]" pageId="6" pageNumber="107">FIGURE 2.</emphasis>
<taxonomicName authority="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann, 2023" authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[279,506,1864,1888]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis box="[279,506,1864,1888]" italics="true" pageId="6" pageNumber="107">Fusigobius humerosus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[512,591,1864,1888]" pageId="6" pageNumber="107">
<taxonomicNameLabel box="[512,591,1864,1888]" pageId="6" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype, KAUMM 199, female, 27.6 + 7.9 mm, Al Lith, Saudi Arabia, Red Sea: 
<emphasis bold="true" box="[1411,1429,1864,1888]" pageId="6" pageNumber="107">A</emphasis>
: the origin of posterior nostril about equal to orbit and to the origin of anterior nostril, the origin of nostrils and the edge of orbit outlined; 
<emphasis bold="true" box="[247,264,1936,1960]" pageId="6" pageNumber="107">B</emphasis>
: 1–2 rows of cycloid scales extends above opercle. 
<emphasis bold="true" box="[787,805,1936,1960]" pageId="6" pageNumber="107">C</emphasis>
: Paratype, PMR VP5272, male, 24.1 + 6.7 mm, Al Lith, Saudi Arabia, Red Sea, the scales ending anterior at line between upper pectoral-fin base and first dorsal fin origin. The anteriormost scales marked with arrow. Photos by M. Kovačić.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7745457" ID-Zenodo-Dep="7745457" httpUri="https://zenodo.org/record/7745457/files/figure.png" pageId="7" pageNumber="108" startId="7.[151,250,1090,1114]" targetBox="[211,1436,215,1019]" targetPageId="7">
<paragraph blockId="7.[151,1437,1090,1151]" pageId="7" pageNumber="108">
<emphasis bold="true" box="[151,276,1090,1115]" pageId="7" pageNumber="108">FIGURE 3.</emphasis>
Cephalic sensory pores system and papillae of 
<taxonomicName authority="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann, 2023" authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[766,995,1091,1115]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="108" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis box="[766,995,1091,1115]" italics="true" pageId="7" pageNumber="108">Fusigobius humerosus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1002,1084,1091,1115]" pageId="7" pageNumber="108">
<taxonomicNameLabel box="[1002,1084,1091,1115]" pageId="7" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype, KAUMM 199, female, 27.6 + 7.9 mm, Al Lith, Saudi Arabia, Red Sea. Terminology in text. Drawing by M. Kovačić.
</paragraph>
</caption>
<subSubSection lastPageId="12" lastPageNumber="113" pageId="7" pageNumber="108" type="description">
<paragraph blockId="7.[151,1437,1197,2016]" pageId="7" pageNumber="108">
<emphasis bold="true" box="[199,337,1197,1223]" pageId="7" pageNumber="108">Description</emphasis>
. Body moderately elongate, body depth 
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SL, and laterally compressed, width 
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depth, caudal peduncle narrow, caudal peduncle depth 
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SL, 
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head, caudal peduncle length 
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SL, 
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head. Head large, length 
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SL, ventral part and breast broad; head triangular from dorsal view. Head triangular in cross section. Crest or ridge of head absent. Snout moderately short and steep, its length 4.0– 
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head length, 
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SL. Anterior nostril a nasal tube without process from rim, at level of lower edge of pupil, about equally distant between front of snout at base of upper lip and anterior edge of orbit, reaching to or nearly reaching upper lip. Posterior nostril with a slightly erected rim, dorsoposterior to anterior nostril and about equally distant to orbit and to anterior nostril. Eyes dorsolateral, moderately large, eye diameter is 3.0– 
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head length, orbit projecting slightly above dorsal profile. Interorbital very narrow, 
<quantity box="[1235,1376,1485,1512]" metricMagnitude="0" metricUnit="m" metricValue="1.51003" metricValueMax="1.7170399999999997" metricValueMin="1.3030199999999998" pageId="7" pageNumber="108" unit="in" value="59.449999999999996" valueMax="67.6" valueMin="51.3">51.3–67.6 in</quantity>
head length. Mouth terminal, oblique. Mouth moderately large, upper jaw 
<quantity box="[914,1026,1521,1548]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" metricValueMax="8.89" metricValueMin="8.382" pageId="7" pageNumber="108" unit="in" value="3.4" valueMax="3.5" valueMin="3.3">3.3–3.5 in</quantity>
head length, angle of jaws extending posteriorly to between a vertical of anterior edge of eye and of vertical at anterior edge of pupil. Cheek broad.
</paragraph>
<paragraph blockId="7.[151,1437,1197,2016]" pageId="7" pageNumber="108">Upper jaw with outer row of long, slender, well-spaced, incurved conical teeth continuing backwards on side of jaw, middle band of small conical teeth in a few irregular rows, narrowing backwards on side of jaw and inner row of slender, strongly incurved, conical teeth of medium size. Lower jaw with 10–12 well-spaced, long, slender, slightly incurved, conical teeth restricted to a front of jaw; a middle band of small conical teeth in a few irregular rows narrowing and decreasing in number of rows backwards on side of jaw, and an inner row of long, slender, curved teeth. Tongue well developed, with anterior edge rounded and anterior part free from floor of mouth. Branchiostegal membranes fused to isthmus, gill openings extending forward to about half of opercle width from posterior edge of preopercle. Gill rakers 1–2 + 6, pseudobranchial filaments 6. Branchiostegal rays 5. Genital papilla of adult males 3/5 of orbit diameter, of adult females 1/5 to 2/5 of orbit diameter.</paragraph>
<paragraph blockId="7.[151,1437,1197,2016]" lastBlockId="8.[151,1437,150,2013]" lastPageId="8" lastPageNumber="109" pageId="7" pageNumber="108">
<emphasis box="[199,249,1918,1943]" italics="true" pageId="7" pageNumber="108">Fins</emphasis>
. Dorsal-fin rays 
<collectionCode box="[445,477,1917,1943]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="108">VI</collectionCode>
+I,8–9 (
<typeStatus box="[566,666,1917,1943]" pageId="7" pageNumber="108">holotype</typeStatus>
8, 
<typeStatus box="[705,812,1917,1943]" pageId="7" pageNumber="108">paratypes</typeStatus>
8: 2, 9: 7); anal-fin rays I,8; branched caudal-fin rays 12–13 (
<typeStatus box="[238,338,1953,1979]" pageId="7" pageNumber="108">holotype</typeStatus>
12, 
<typeStatus box="[391,498,1953,1979]" pageId="7" pageNumber="108">paratypes</typeStatus>
12: 8, 13: 1); upper unbranched caudal-fin rays 7–8 (
<typeStatus box="[1128,1228,1953,1979]" pageId="7" pageNumber="108">holotype</typeStatus>
8, 
<typeStatus box="[1268,1375,1953,1979]" pageId="7" pageNumber="108">paratypes</typeStatus>
7: 1, 8: 8), posterior 2-3 segmented (
<typeStatus box="[514,614,1989,2015]" pageId="7" pageNumber="108">holotype</typeStatus>
3, 
<typeStatus box="[651,758,1989,2015]" pageId="7" pageNumber="108">paratypes</typeStatus>
2: 3, 3: 6); lower unbranched caudal-fin rays 5–7 (
<typeStatus box="[1336,1436,1989,2015]" pageId="7" pageNumber="108">holotype</typeStatus>
5, 
<typeStatus box="[181,288,151,177]" pageId="8" pageNumber="109">paratypes</typeStatus>
5: 4, 6: 3, 7: 2), posterior 1–2 segmented (
<typeStatus box="[784,884,151,177]" pageId="8" pageNumber="109">holotype</typeStatus>
2, 
<typeStatus box="[923,1030,151,177]" pageId="8" pageNumber="109">paratypes</typeStatus>
1: 2, 2: 7). 
<collectorName box="[1170,1302,151,177]" pageId="8" pageNumber="109">Pectoral-fin</collectorName>
rays 17–18 (
<typeStatus box="[159,259,187,213]" pageId="8" pageNumber="109">holotype</typeStatus>
18: 2, 
<typeStatus box="[339,446,187,213]" pageId="8" pageNumber="109">paratypes</typeStatus>
17: 14, 18: 4), upper 2–5 and lowermost 1–2 unbranched. 
<collectorName box="[1113,1183,187,213]" pageId="8" pageNumber="109">Pelvic</collectorName>
fins I,5+5, 
<collectorName box="[1309,1405,186,213]" pageId="8" pageNumber="109">I. Origin</collectorName>
of first dorsal fin slightly behind upper base of pectoral fin, predorsal distance 
<quantity box="[1026,1141,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.366" metricValueMin="6.604" pageId="8" pageNumber="109" unit="in" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 in</quantity>
SL. 
<collectorName box="[1199,1250,223,249]" pageId="8" pageNumber="109">First</collectorName>
dorsal-fin spine longest, 4.2–5.0 in SL, 
<quantity box="[413,526,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.5559999999999996" pageId="8" pageNumber="109" unit="in" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 in</quantity>
head length, first and second dorsal-fin spine not elongate or filamentous in both sexes, second to sixth spine progressively diminishing in length. 
<collectorName box="[875,911,295,321]" pageId="8" pageNumber="109">All</collectorName>
spines of first dorsal fin barely reaching origin of second dorsal fin when folded down. 
<collectorName box="[612,660,331,357]" pageId="8" pageNumber="109">Last</collectorName>
membrane of first dorsal fin separated from origin of second dorsal fin by a space about equal to space of first membrane of second dorsal fin. 
<collectorName box="[1015,1079,367,393]" pageId="8" pageNumber="109">Spine</collectorName>
of second dorsal fin 1.8–2.0 in head. 
<collectorName box="[219,270,403,429]" pageId="8" pageNumber="109">First</collectorName>
dorsal fin slightly higher than second in females, about equal in males, although longest dorsal-fin soft ray longer in males, 
<quantity box="[382,495,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.5559999999999996" pageId="8" pageNumber="109" unit="in" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 in</quantity>
head length, than in females, 
<quantity box="[833,946,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="4.571999999999999" metricValueMax="4.826" metricValueMin="4.318" pageId="8" pageNumber="109" unit="in" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7–1.9 in</quantity>
head. 
<collectorName box="[1021,1094,439,465]" pageId="8" pageNumber="109">Origin</collectorName>
of anal fin between vertical at base of first soft ray and vertical of second soft ray of second dorsal fin, preanal length 
<quantity box="[1119,1231,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="8" pageNumber="109" unit="in" value="1.65" valueMax="1.7" valueMin="1.6">1.6–1.7 in</quantity>
SL, anal-fin spine 
<quantity box="[151,265,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="9.270999999999999" metricValueMax="9.906" metricValueMin="8.636" pageId="8" pageNumber="109" unit="in" value="3.65" valueMax="3.9" valueMin="3.4">3.4–3.9 in</quantity>
head length, longest anal-fin soft ray longer in males, 
<quantity box="[881,994,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="3.9369999999999994" metricValueMax="4.318" metricValueMin="3.5559999999999996" pageId="8" pageNumber="109" unit="in" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 in</quantity>
head, than in females, 
<quantity box="[1256,1369,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="5.588" metricValueMin="4.572" pageId="8" pageNumber="109" unit="in" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 in</quantity>
head. 
<collectorName box="[151,230,547,573]" pageId="8" pageNumber="109">Caudal</collectorName>
fin rounded, shorter than head, 
<quantity box="[603,718,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="8" pageNumber="109" unit="in" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 in</quantity>
head length, 3.5–4.0 in SL. 
<collectorName box="[1050,1142,547,573]" pageId="8" pageNumber="109">Pectoral</collectorName>
fins pointed, middle rays longest, 
<quantity box="[248,363,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="8.763" metricValueMax="9.398" metricValueMin="8.128" pageId="8" pageNumber="109" unit="in" value="3.45" valueMax="3.7" valueMin="3.2">3.2–3.7 in</quantity>
SL. 
<collectorName box="[419,453,583,609]" pageId="8" pageNumber="109">No</collectorName>
free pectoral-fin rays. 
<collectorName box="[716,808,583,609]" pageId="8" pageNumber="109">Pectoral</collectorName>
fins extending posteriorly well below origin of second dorsal fin. 
<collectorName box="[276,346,619,645]" pageId="8" pageNumber="109">Pelvic</collectorName>
fins joined medially by membrane at least 2/3 of pelvic fin length (membrane connection was broken in specimens so exact extend of connection along fin was not determined, but membrane was still visible on rays from origin up to 2/3 of pelvic fin length) possibly to 9/10 of pelvic fin length (length of fifth pelvic-fin soft ray to tip of shortest branch of fifth 83–94% of length of fourth pelvic-fin soft ray to tip). 
<collectorName box="[1117,1153,727,753]" pageId="8" pageNumber="109">All</collectorName>
pelvic-fin rays including fifth ray branched. 
<collectorName box="[370,440,763,789]" pageId="8" pageNumber="109">Pelvic</collectorName>
frenum present, width at midline about 1/4–1/5 of pelvic-spine length. 
<collectorName box="[1255,1328,763,789]" pageId="8" pageNumber="109">Origin</collectorName>
of pelvic fins below base of pectoral fins, prepelvic length 2.8–3.0 in SL, pelvic-fin spine 
<quantity box="[1056,1169,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="1.1683999999999999" metricValueMax="1.3462" metricValueMin="0.9906" pageId="8" pageNumber="109" unit="in" value="4.6" valueMax="5.3" valueMin="3.9">3.9–5.3 in</quantity>
head, fourth pelvic soft ray longest, 
<quantity box="[290,404,835,861]" metricMagnitude="-1" metricUnit="m" metricValue="1.1176" metricValueMax="1.2446" metricValueMin="0.9906" pageId="8" pageNumber="109" unit="in" value="4.4" valueMax="4.9" valueMin="3.9">3.9–4.9 in</quantity>
SL, longer in males, 
<quantity box="[648,762,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="3.3019999999999996" metricValueMax="3.5559999999999996" metricValueMin="3.048" pageId="8" pageNumber="109" unit="in" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4 in</quantity>
head length, than in females, 
<quantity box="[1103,1216,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.5559999999999996" pageId="8" pageNumber="109" unit="in" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 in</quantity>
head, and reaching to origin of anal fin.
</paragraph>
<paragraph blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="109">
<emphasis box="[199,333,907,933]" italics="true" pageId="8" pageNumber="109">Squamation</emphasis>
. Body covered with ctenoid scales, cycloid on breast, prepectoral area, above base of pelvic fins and on belly, anterior part of belly naked. Head, including cheek and operculum, and predorsal area naked, scales on side of nape usually ending anterior at line between upper pectoral-fin base and origin of first dorsal fin or 1–2 rows of cycloid scales extending forward to above opercle. No scales on fins except of a few on base of caudal fin, most of which are smaller than scales on body. Longitudinal scale series 26–28 (
<typeStatus box="[1063,1163,1051,1077]" pageId="8" pageNumber="109">holotype</typeStatus>
26, right side damaged, 
<typeStatus box="[151,258,1087,1113]" pageId="8" pageNumber="109">paratypes</typeStatus>
26: 3, 27: 12, 28: 1, right sides of 
<specimenCode box="[637,770,1086,1113]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="109">SMF 35904</specimenCode>
and 
<specimenCode box="[823,981,1087,1113]" collectionCode="KAUMM" pageId="8" pageNumber="109">KAUMM 463</specimenCode>
too damaged for count), transverse scale series 6–7 (
<typeStatus box="[279,379,1123,1149]" pageId="8" pageNumber="109">holotype</typeStatus>
6, 
<typeStatus box="[414,521,1123,1149]" pageId="8" pageNumber="109">paratypes</typeStatus>
6: 6, 7: 10, right sides of 
<specimenCode box="[811,944,1122,1149]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="109">SMF 35904</specimenCode>
and 
<specimenCode box="[999,1158,1123,1149]" collectionCode="KAUMM" pageId="8" pageNumber="109">KAUMM 463</specimenCode>
too damaged for count); circumpeduncular scales 10–12 (
<typeStatus box="[529,629,1159,1185]" pageId="8" pageNumber="109">holotype</typeStatus>
11, right side damaged, 
<typeStatus box="[907,1014,1159,1185]" pageId="8" pageNumber="109">paratypes</typeStatus>
10: 1, 11: 5, 12: 3); breast with three scales in longitudinal series and prepectoral area with single vertical row of scales.
</paragraph>
<paragraph blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="109">
<emphasis box="[199,486,1231,1257]" italics="true" pageId="8" pageNumber="109">Cephalic sensory systems</emphasis>
(
<figureCitation box="[501,569,1231,1257]" captionStart="FIGURE 3" captionStartId="7.[151,250,1090,1114]" captionTargetBox="[211,1436,215,1019]" captionTargetId="figure-20@7.[151,1436,181,1067]" captionTargetPageId="7" captionText="FIGURE 3. Cephalic sensory pores system and papillae of Fusigobius humerosus sp. nov. Paratype, KAUMM 199, female, 27.6 + 7.9 mm, Al Lith, Saudi Arabia, Red Sea. Terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.7745457" httpUri="https://zenodo.org/record/7745457/files/figure.png" pageId="8" pageNumber="109">Fig. 3</figureCitation>
). All three head canals present: anterior oculoscapular canal with pores B’, C (single), D (single), E, F, G, and H’; posterior oculoscapular canal with pores K’ and L’; preopercular canal with pores M’, N, and O’. Anterior oculoscapular head canal with short side branch to pore E below eye. Rows and number of sensory papillae as follows: (1) 
<emphasis box="[503,616,1339,1365]" italics="true" pageId="8" pageNumber="109">preorbital</emphasis>
: snout with three rows in median preorbital series, superior row 
<emphasis box="[1328,1339,1339,1364]" italics="true" pageId="8" pageNumber="109">r</emphasis>
close on inner side to pore B (1–2), inferior row 
<emphasis box="[592,603,1375,1400]" italics="true" pageId="8" pageNumber="109">s</emphasis>
with two sections, 
<emphasis box="[818,829,1375,1400]" italics="true" pageId="8" pageNumber="109">s</emphasis>
<superScript attach="left" box="[828,836,1374,1388]" fontSize="6" pageId="8" pageNumber="109">1</superScript>
(1) below pore B, close to nostrils, 
<emphasis box="[1234,1245,1375,1400]" italics="true" pageId="8" pageNumber="109">s</emphasis>
<superScript attach="left" box="[1245,1253,1374,1388]" fontSize="6" pageId="8" pageNumber="109">3</superScript>
(1) above upper lip. Lateral series 
<emphasis box="[355,367,1411,1436]" italics="true" pageId="8" pageNumber="109">c</emphasis>
in four parts: superior 
<emphasis box="[630,642,1411,1436]" italics="true" pageId="8" pageNumber="109">c</emphasis>
<superScript attach="left" box="[642,650,1410,1424]" fontSize="6" pageId="8" pageNumber="109">2</superScript>
(2–3) between anterior and posterior nostrils, middle 
<emphasis box="[1263,1275,1411,1436]" italics="true" pageId="8" pageNumber="109">c</emphasis>
<superScript attach="left" box="[1275,1283,1410,1424]" fontSize="6" pageId="8" pageNumber="109">1</superScript>
(1–2) behind and bellow anterior nostril and inferior upper 
<emphasis box="[672,684,1447,1472]" italics="true" pageId="8" pageNumber="109">c</emphasis>
<subScript attach="left" box="[685,693,1465,1479]" fontSize="6" pageId="8" pageNumber="109">2</subScript>
(1–2) and lower 
<emphasis box="[891,903,1447,1472]" italics="true" pageId="8" pageNumber="109">c</emphasis>
<subScript attach="left" box="[903,911,1465,1479]" fontSize="6" pageId="8" pageNumber="109">1</subScript>
(2) above upper lip. (2) 
<emphasis box="[1193,1308,1447,1473]" italics="true" pageId="8" pageNumber="109">suborbital</emphasis>
: suborbital rows without transverse proliferations and uniserial. Row 
<emphasis box="[803,817,1483,1508]" italics="true" pageId="8" pageNumber="109">a</emphasis>
(3) longitudinal without transverse proliferation below rear part of eye; row 
<emphasis box="[385,397,1519,1544]" italics="true" pageId="8" pageNumber="109">c</emphasis>
(4) longitudinal without transverse proliferation below anterior and middle part of eye, more distant from eye than row 
<emphasis box="[454,468,1555,1580]" italics="true" pageId="8" pageNumber="109">a</emphasis>
, starting anteriorly at about vertical of front border of eye, posteriorly ending before vertical of row 
<emphasis box="[319,340,1591,1616]" italics="true" pageId="8" pageNumber="109">a.</emphasis>
Two 
<emphasis box="[401,427,1591,1616]" italics="true" pageId="8" pageNumber="109">cp</emphasis>
as papillae below first papilla of row 
<emphasis box="[841,855,1591,1617]" italics="true" pageId="8" pageNumber="109">b</emphasis>
, lower papilla nearly below upper one. Longitudinal row 
<emphasis box="[205,219,1627,1653]" italics="true" pageId="8" pageNumber="109">b</emphasis>
(6–11) below rear border of eye. Longitudinal row 
<emphasis box="[829,843,1627,1653]" italics="true" pageId="8" pageNumber="109">d</emphasis>
continuous from anteriorly above posterior part of upper lip along cheek posterior nearly to a vertical at rear edge of eye (17–19) or divided in supralabial part and part on cheek (6+3–9+7). (3) 
<emphasis box="[491,766,1699,1725]" italics="true" pageId="8" pageNumber="109">preoperculo-mandibular</emphasis>
: external row 
<emphasis box="[933,945,1699,1724]" italics="true" pageId="8" pageNumber="109">e</emphasis>
and internal row 
<emphasis box="[1150,1158,1699,1724]" italics="true" pageId="8" pageNumber="109">i</emphasis>
divided into anterior (
<emphasis box="[1416,1428,1699,1724]" italics="true" pageId="8" pageNumber="109">e</emphasis>
: 11–16, 
<emphasis box="[233,241,1735,1760]" italics="true" pageId="8" pageNumber="109">i</emphasis>
: 6–8), and posterior sections (
<emphasis box="[578,590,1735,1760]" italics="true" pageId="8" pageNumber="109">e</emphasis>
: 14–20, 
<emphasis box="[687,695,1735,1760]" italics="true" pageId="8" pageNumber="109">i</emphasis>
: 6–7); mental row 
<emphasis box="[907,915,1735,1761]" italics="true" pageId="8" pageNumber="109">f</emphasis>
longitudinal (1–4). (4) 
<emphasis box="[1175,1335,1735,1761]" italics="true" pageId="8" pageNumber="109">oculoscapular</emphasis>
: anterior longitudinal row 
<emphasis box="[343,355,1771,1796]" italics="true" pageId="8" pageNumber="109">x</emphasis>
<superScript attach="left" box="[355,363,1770,1784]" fontSize="6" pageId="8" pageNumber="109">1</superScript>
(7–12) continuous from above pore G to above halfway between pores H’ and K’, posterior row 
<emphasis box="[151,163,1807,1832]" italics="true" pageId="8" pageNumber="109">x</emphasis>
<superScript attach="left" box="[164,172,1806,1820]" fontSize="6" pageId="8" pageNumber="109">2</superScript>
(1–4) above pore L’; row 
<emphasis box="[475,486,1807,1832]" italics="true" pageId="8" pageNumber="109">z</emphasis>
(3–5) from behind pore G downwards to behind pore M’, row 
<emphasis box="[1210,1224,1807,1832]" italics="true" pageId="8" pageNumber="109">q</emphasis>
(1–3) behind pore H’, row 
<emphasis box="[249,261,1843,1868]" italics="true" pageId="8" pageNumber="109">y</emphasis>
(1–2) behind L’. Axillary rows 
<emphasis box="[629,654,1843,1868]" italics="true" pageId="8" pageNumber="109">as</emphasis>
<superScript attach="left" box="[654,662,1842,1856]" fontSize="6" pageId="8" pageNumber="109">1</superScript>
(5–8), 
<emphasis box="[744,769,1843,1868]" italics="true" pageId="8" pageNumber="109">as</emphasis>
<superScript attach="left" box="[769,777,1842,1856]" fontSize="6" pageId="8" pageNumber="109">2</superScript>
(5–9), 
<emphasis box="[859,884,1843,1868]" italics="true" pageId="8" pageNumber="109">as</emphasis>
<superScript attach="left" box="[883,891,1842,1856]" fontSize="6" pageId="8" pageNumber="109">3</superScript>
(6–8) not visible in every specimen, 
<emphasis box="[1318,1340,1843,1869]" italics="true" pageId="8" pageNumber="109">la</emphasis>
<superScript attach="left" box="[1340,1348,1842,1856]" fontSize="6" pageId="8" pageNumber="109">1</superScript>
and 
<emphasis box="[1406,1428,1843,1869]" italics="true" pageId="8" pageNumber="109">la</emphasis>
<superScript attach="left" box="[1428,1436,1842,1856]" fontSize="6" pageId="8" pageNumber="109">2</superScript>
absent. (5) 
<emphasis box="[279,388,1879,1905]" italics="true" pageId="8" pageNumber="109">opercular</emphasis>
: transverse row 
<emphasis box="[578,600,1879,1904]" italics="true" pageId="8" pageNumber="109">ot</emphasis>
(12-18); superior longitudinal row 
<emphasis box="[1004,1029,1879,1904]" italics="true" pageId="8" pageNumber="109">os</emphasis>
(6–11); inferior longitudinal row 
<emphasis box="[1414,1436,1879,1904]" italics="true" pageId="8" pageNumber="109">oi</emphasis>
(5–8) anteriorly starts close to row 
<emphasis box="[542,564,1915,1940]" italics="true" pageId="8" pageNumber="109">ot</emphasis>
. (6) 
<emphasis box="[617,787,1915,1941]" italics="true" pageId="8" pageNumber="109">anterior dorsal</emphasis>
: transverse row 
<emphasis box="[971,985,1915,1940]" italics="true" pageId="8" pageNumber="109">n</emphasis>
behind pore 
<emphasis box="[1131,1148,1915,1940]" italics="true" pageId="8" pageNumber="109">E</emphasis>
(1); transverse rows 
<emphasis box="[1383,1397,1915,1940]" italics="true" pageId="8" pageNumber="109">o</emphasis>
(1) divided from each other; longitudinal row 
<emphasis box="[627,641,1951,1976]" italics="true" pageId="8" pageNumber="109">g</emphasis>
(3–4) ends posteriorly laterally and slightly behind row 
<emphasis box="[1273,1287,1951,1976]" italics="true" pageId="8" pageNumber="109">o</emphasis>
, longitudinal row 
<emphasis box="[202,222,1987,2012]" italics="true" pageId="8" pageNumber="109">m</emphasis>
(2–3) behind and below of row 
<emphasis box="[584,598,1987,2012]" italics="true" pageId="8" pageNumber="109">g</emphasis>
; longitudinal row 
<emphasis box="[805,819,1987,2013]" italics="true" pageId="8" pageNumber="109">h</emphasis>
discontinuous (2+3–3+3) in front of first dorsal fin.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7745459" ID-Zenodo-Dep="7745459" httpUri="https://zenodo.org/record/7745459/files/figure.png" pageId="9" pageNumber="110" startId="9.[151,250,1868,1892]" targetBox="[151,1436,201,1843]" targetPageId="9">
<paragraph blockId="9.[151,1437,1868,1964]" pageId="9" pageNumber="110">
<emphasis bold="true" box="[151,274,1868,1892]" pageId="9" pageNumber="110">FIGURE 4.</emphasis>
<taxonomicName authority="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann, 2023" authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[280,508,1868,1892]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="110" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis box="[280,508,1868,1892]" italics="true" pageId="9" pageNumber="110">Fusigobius humerosus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[514,594,1868,1892]" pageId="9" pageNumber="110">
<taxonomicNameLabel box="[514,594,1868,1892]" pageId="9" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype, SMF 35901, male, 28.7 + 8.0 mm, northern side of Farasan Island, Saudi Arabia, Red Sea, 
<emphasis bold="true" box="[338,356,1904,1928]" pageId="9" pageNumber="110">A</emphasis>
: live specimen, 
<emphasis bold="true" box="[525,542,1904,1928]" pageId="9" pageNumber="110">B</emphasis>
: freshly collected, 
<emphasis bold="true" box="[738,756,1904,1928]" pageId="9" pageNumber="110">C</emphasis>
: preserved specimen. Photos by S.V. Bogorodsky (A &amp; B) and M. Kovačić (C).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7745461" ID-Zenodo-Dep="7745461" httpUri="https://zenodo.org/record/7745461/files/figure.png" pageId="10" pageNumber="111" startId="10.[151,250,1927,1951]" targetBox="[151,1436,181,1903]" targetPageId="10">
<paragraph blockId="10.[151,1436,1927,1987]" pageId="10" pageNumber="111">
<emphasis bold="true" box="[151,275,1927,1951]" pageId="10" pageNumber="111">FIGURE 5.</emphasis>
<taxonomicName authority="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann, 2023" authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[281,508,1927,1951]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis box="[281,508,1927,1951]" italics="true" pageId="10" pageNumber="111">Fusigobius humerosus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[514,594,1927,1951]" pageId="10" pageNumber="111">
<taxonomicNameLabel box="[514,594,1927,1951]" pageId="10" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, underwater photographs of uncollected individuals. 
<emphasis bold="true" box="[1133,1151,1927,1951]" pageId="10" pageNumber="111">A</emphasis>
: Dahab, Egypt, Red Sea; 
<emphasis bold="true" box="[1412,1429,1927,1951]" pageId="10" pageNumber="111">B</emphasis>
: Al Wajh, Saudi Arabia, Red Sea; 
<emphasis bold="true" box="[489,507,1963,1987]" pageId="10" pageNumber="111">C</emphasis>
: Mangrove Bay, El Quseir, Egypt, Red Sea. Photos by S.V. Bogorodsky.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7745463" ID-Zenodo-Dep="7745463" httpUri="https://zenodo.org/record/7745463/files/figure.png" pageId="11" pageNumber="112" startId="11.[151,250,1156,1180]" targetBox="[151,1436,181,1131]" targetPageId="11">
<paragraph blockId="11.[151,1437,1156,1360]" pageId="11" pageNumber="112">
<emphasis bold="true" box="[151,299,1156,1180]" pageId="11" pageNumber="112">FIGURE 6. A</emphasis>
and 
<emphasis bold="true" box="[349,367,1156,1180]" pageId="11" pageNumber="112">C</emphasis>
: 
<taxonomicName authority="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann, 2023" authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[381,609,1156,1180]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis box="[381,609,1156,1180]" italics="true" pageId="11" pageNumber="112">Fusigobius humerosus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[615,696,1156,1180]" pageId="11" pageNumber="112">
<taxonomicNameLabel box="[615,696,1156,1180]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, PMR VP5271, paratype, male, 26.2 + 7.3 mm, northern side of Farasan Island, Saudi Arabia, Red Sea. 
<emphasis bold="true" box="[475,492,1192,1216]" pageId="11" pageNumber="112">B</emphasis>
and 
<emphasis bold="true" box="[542,560,1192,1216]" pageId="11" pageNumber="112">D</emphasis>
: 
<taxonomicName authority=", BPBM" authorityName="BPBM" box="[574,783,1192,1216]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[574,700,1192,1216]" italics="true" pageId="11" pageNumber="112">F. humeralis</emphasis>
, BPBM
</taxonomicName>
36627, paratype, female, 29.5 + 7.8 mm, Manuk Island, Banda Sea, Indonesia. Humeral region just above base of pectoral fin. 
<emphasis bold="true" box="[823,841,1228,1252]" pageId="11" pageNumber="112">A</emphasis>
: round black spot is smaller than or equal to pupil in 
<taxonomicName authority="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann, 2023" authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis italics="true" pageId="11" pageNumber="112">F. humerosus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[266,346,1264,1288]" pageId="11" pageNumber="112">
<taxonomicNameLabel box="[266,346,1264,1288]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis bold="true" box="[358,375,1264,1288]" pageId="11" pageNumber="112">B</emphasis>
: dark spot larger than pupil in 
<taxonomicName box="[683,808,1264,1288]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[683,808,1264,1288]" italics="true" pageId="11" pageNumber="112">F. humeralis</emphasis>
</taxonomicName>
; midbase of caudal fin: 
<emphasis bold="true" box="[1050,1068,1264,1288]" pageId="11" pageNumber="112">C</emphasis>
: oval black spot covers less than 1/3 of caudal base height in 
<taxonomicName authority="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann, 2023" authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[401,535,1300,1324]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="humerosus" status="sp. nov.">
<emphasis box="[401,535,1300,1324]" italics="true" pageId="11" pageNumber="112">F. humerosus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[542,623,1300,1324]" pageId="11" pageNumber="112">
<taxonomicNameLabel box="[542,623,1300,1324]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis bold="true" box="[637,655,1300,1324]" pageId="11" pageNumber="112">D</emphasis>
: irregular spot at midbase of caudal fin larger than 1/3 of caudal base height in 
<taxonomicName box="[177,303,1336,1360]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[177,303,1336,1360]" italics="true" pageId="11" pageNumber="112">F. humeralis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="11.[151,1437,1406,2009]" lastBlockId="12.[151,1437,151,2013]" lastPageId="12" lastPageNumber="113" pageId="11" pageNumber="112">
<emphasis box="[199,668,1406,1432]" italics="true" pageId="11" pageNumber="112">Color in life and immediately after death</emphasis>
(
<figureCitation box="[685,832,1406,1433]" captionStart="FIGURE 4" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[151,1436,201,1843]" captionTargetId="figure-20@9.[151,1436,201,1844]" captionTargetPageId="9" captionText="FIGURE 4. Fusigobius humerosus sp. nov. holotype, SMF 35901, male, 28.7 + 8.0 mm, northern side of Farasan Island, Saudi Arabia, Red Sea, A: live specimen, B: freshly collected, C: preserved specimen. Photos by S.V. Bogorodsky (A &amp; B) and M. Kovačić (C)." figureDoi="http://doi.org/10.5281/zenodo.7745459" httpUri="https://zenodo.org/record/7745459/files/figure.png" pageId="11" pageNumber="112">Fig. 4A &amp; B</figureCitation>
, 
<figureCitation box="[846,915,1406,1433]" captionStart="FIGURE 5" captionStartId="10.[151,250,1927,1951]" captionTargetBox="[151,1436,181,1903]" captionTargetId="figure-26@10.[151,1436,181,1903]" captionTargetPageId="10" captionText="FIGURE 5. Fusigobius humerosus sp. nov., underwater photographs of uncollected individuals. A: Dahab, Egypt, Red Sea; B: Al Wajh, Saudi Arabia, Red Sea; C: Mangrove Bay, El Quseir, Egypt, Red Sea. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.7745461" httpUri="https://zenodo.org/record/7745461/files/figure.png" pageId="11" pageNumber="112">Fig. 5</figureCitation>
). Body translucent in live specimens, slightly murky in fresh specimens, with numerous small dusky orange-yellow spots on head and body, series of melanophores within and adjacent of each spot, spots on head irregularly arranged in oblique rows going posterodorsally and posteriorly. Body covered with small silver or bluish reflective markings. Internal organs well visible, especially in live specimens, gills red and peritoneum silvery; body with an internal series of six horizontally-elongate, dark brown blotches along vertebral column and several, irregularly scattered, oval dark brown blotches in lower part of body. A round black spot smaller than or as large as pupil in humeral region just above base of pectoral fin (
<figureCitation box="[1334,1423,1622,1648]" captionStart="FIGURE 6" captionStartId="11.[151,250,1156,1180]" captionTargetBox="[151,1436,181,1131]" captionTargetId="figure-20@11.[151,1436,181,1131]" captionTargetPageId="11" captionText="FIGURE 6. A and C: Fusigobius humerosus sp. nov., PMR VP5271, paratype, male, 26.2 + 7.3 mm, northern side of Farasan Island, Saudi Arabia, Red Sea. B and D: F. humeralis, BPBM 36627, paratype, female, 29.5 + 7.8 mm, Manuk Island, Banda Sea, Indonesia. Humeral region just above base of pectoral fin. A: round black spot is smaller than or equal to pupil in F. humerosus sp. nov., B: dark spot larger than pupil in F. humeralis; midbase of caudal fin: C: oval black spot covers less than 1/3 of caudal base height in F. humerosus sp. nov., D: irregular spot at midbase of caudal fin larger than 1/3 of caudal base height in F. humeralis." figureDoi="http://doi.org/10.5281/zenodo.7745463" httpUri="https://zenodo.org/record/7745463/files/figure.png" pageId="11" pageNumber="112">Fig. 6A</figureCitation>
), a second subequal or slightly larger black spot at midbase of caudal fin, barely visible underwater, more apparent in freshly dead material, covers less than 1/3 of caudal-fin base height (
<figureCitation box="[960,1049,1694,1720]" captionStart="FIGURE 6" captionStartId="11.[151,250,1156,1180]" captionTargetBox="[151,1436,181,1131]" captionTargetId="figure-20@11.[151,1436,181,1131]" captionTargetPageId="11" captionText="FIGURE 6. A and C: Fusigobius humerosus sp. nov., PMR VP5271, paratype, male, 26.2 + 7.3 mm, northern side of Farasan Island, Saudi Arabia, Red Sea. B and D: F. humeralis, BPBM 36627, paratype, female, 29.5 + 7.8 mm, Manuk Island, Banda Sea, Indonesia. Humeral region just above base of pectoral fin. A: round black spot is smaller than or equal to pupil in F. humerosus sp. nov., B: dark spot larger than pupil in F. humeralis; midbase of caudal fin: C: oval black spot covers less than 1/3 of caudal base height in F. humerosus sp. nov., D: irregular spot at midbase of caudal fin larger than 1/3 of caudal base height in F. humeralis." figureDoi="http://doi.org/10.5281/zenodo.7745463" httpUri="https://zenodo.org/record/7745463/files/figure.png" pageId="11" pageNumber="112">Fig. 6C</figureCitation>
). Seven faint, brown dorsal marks on back, two in front of the first dorsal fin, two more below first dorsal fin, remaining three marks below second dorsal fin, marks extending onto basal part of both fins and more visible in freshly dead material. Upper pectoralfin base with white streak. An oblique blackish bar extending from anteroventral edge of eye across suborbital area ending on upper lip visible in freshly dead specimens but not visible in life. Eyes with uniform brown-orange iris, sometimes suffused with yellow, and green pupil in life, pupil black immediately after death. Sometimes an orange-brown oblique dash starts at the end of jaw extending backwards and up on cheek. Dorsal and caudal fins with orange and yellow dots, except for unspotted ventral one-fourth of caudal fin, the first dorsal fin with variably distinct oblique dusky line from the middle of first spine to anterior base of third spine. Pectoral fins transparent with a white longitudinal stripe at the midline of pectoral-fin base extending into middle of pectoral fin and nearly reaching fin margin. Anal and pelvic fins transparent with snow-white area: origin of anal fin and center of pelvic fin, respectively.
</paragraph>
<paragraph blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="113">
<emphasis box="[199,381,259,285]" italics="true" pageId="12" pageNumber="113">Color preserved</emphasis>
(
<figureCitation box="[398,488,259,285]" captionStart="FIGURE 4" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[151,1436,201,1843]" captionTargetId="figure-20@9.[151,1436,201,1844]" captionTargetPageId="9" captionText="FIGURE 4. Fusigobius humerosus sp. nov. holotype, SMF 35901, male, 28.7 + 8.0 mm, northern side of Farasan Island, Saudi Arabia, Red Sea, A: live specimen, B: freshly collected, C: preserved specimen. Photos by S.V. Bogorodsky (A &amp; B) and M. Kovačić (C)." figureDoi="http://doi.org/10.5281/zenodo.7745459" httpUri="https://zenodo.org/record/7745459/files/figure.png" pageId="12" pageNumber="113">Fig. 4C</figureCitation>
). Body opaque fawn, orange-yellow coloration lost. The pigmented pattern of small spots on head and body brown. A round black spot smaller or as large as pupil in humeral region just above base of pectoral fin (
<figureCitation box="[296,387,331,357]" captionStart="FIGURE 6" captionStartId="11.[151,250,1156,1180]" captionTargetBox="[151,1436,181,1131]" captionTargetId="figure-20@11.[151,1436,181,1131]" captionTargetPageId="11" captionText="FIGURE 6. A and C: Fusigobius humerosus sp. nov., PMR VP5271, paratype, male, 26.2 + 7.3 mm, northern side of Farasan Island, Saudi Arabia, Red Sea. B and D: F. humeralis, BPBM 36627, paratype, female, 29.5 + 7.8 mm, Manuk Island, Banda Sea, Indonesia. Humeral region just above base of pectoral fin. A: round black spot is smaller than or equal to pupil in F. humerosus sp. nov., B: dark spot larger than pupil in F. humeralis; midbase of caudal fin: C: oval black spot covers less than 1/3 of caudal base height in F. humerosus sp. nov., D: irregular spot at midbase of caudal fin larger than 1/3 of caudal base height in F. humeralis." figureDoi="http://doi.org/10.5281/zenodo.7745463" httpUri="https://zenodo.org/record/7745463/files/figure.png" pageId="12" pageNumber="113">Fig. 6A</figureCitation>
), a second about equal black spot at midbase of caudal fin, covers less than 1/3 of caudal-fin base height, both well visible to less vivid in some specimens (
<figureCitation box="[852,940,367,393]" captionStart="FIGURE 6" captionStartId="11.[151,250,1156,1180]" captionTargetBox="[151,1436,181,1131]" captionTargetId="figure-20@11.[151,1436,181,1131]" captionTargetPageId="11" captionText="FIGURE 6. A and C: Fusigobius humerosus sp. nov., PMR VP5271, paratype, male, 26.2 + 7.3 mm, northern side of Farasan Island, Saudi Arabia, Red Sea. B and D: F. humeralis, BPBM 36627, paratype, female, 29.5 + 7.8 mm, Manuk Island, Banda Sea, Indonesia. Humeral region just above base of pectoral fin. A: round black spot is smaller than or equal to pupil in F. humerosus sp. nov., B: dark spot larger than pupil in F. humeralis; midbase of caudal fin: C: oval black spot covers less than 1/3 of caudal base height in F. humerosus sp. nov., D: irregular spot at midbase of caudal fin larger than 1/3 of caudal base height in F. humeralis." figureDoi="http://doi.org/10.5281/zenodo.7745463" httpUri="https://zenodo.org/record/7745463/files/figure.png" pageId="12" pageNumber="113">Fig. 6C</figureCitation>
). Eyes dark, including pupil. Oblique brown bar from eye to upper lip, parallel oblique brown bar from end of jaw backwards and up onto cheek. Dorsal fins transparent with dark pigmented pattern extending from the base, more intense on the first dorsal fin, a dark lines extends from the origin of the third spine upward to the upper part of the first spine. Anal fin darkly pigmented in males, poorly pigmented in females. Caudal fin, pectoral fins and pelvic fins transparent, poorly pigmented except for the black spot at origin of caudal fin that extends slightly onto caudal rays.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="113" type="etymology">
<paragraph blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="113">
<emphasis bold="true" box="[199,328,583,609]" pageId="12" pageNumber="113">Etymology</emphasis>
. The species name 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[551,671,583,609]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[551,671,583,609]" italics="true" pageId="12" pageNumber="113">humerosus</emphasis>
</taxonomicName>
in reference to resemblance to the sister 
<taxonomicName box="[1138,1278,583,609]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[1138,1278,583,609]" italics="true" pageId="12" pageNumber="113">F. humeralis</emphasis>
</taxonomicName>
. Both species share the presence of a distinct black spot in the humeral area above pectoral-fin base.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="113" type="distribution">
<paragraph blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="113">
<emphasis bold="true" box="[199,501,654,681]" pageId="12" pageNumber="113">Distribution and Habitat</emphasis>
. At present 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[643,896,655,681]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[643,896,655,681]" italics="true" pageId="12" pageNumber="113">Fusigobius humerosus</emphasis>
</taxonomicName>
is known from the 
<collectingRegion box="[1126,1224,655,681]" country="Egypt" name="Al Bahr al Ahmar" pageId="12" pageNumber="113">Red Sea</collectingRegion>
, including Gulf of Aqaba, south to the inner Gulf of 
<collectingRegion box="[528,592,691,717]" country="Yemen" name="'Adan" pageId="12" pageNumber="113">Aden</collectingRegion>
. Found on sand and rubble near or within coral reefs, at depths of 
<quantity box="[1333,1421,690,717]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="12" pageNumber="113" unit="m" value="9.0" valueMax="15.0" valueMin="3.0">
3– 
<specimenCount box="[1362,1421,690,717]" pageId="12" pageNumber="113" type="generic">15 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113" type="discussion">
<paragraph blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="113">
<emphasis bold="true" box="[199,308,727,753]" pageId="12" pageNumber="113">Remarks</emphasis>
. 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[328,585,727,753]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[328,585,727,753]" italics="true" pageId="12" pageNumber="113">Fusigobius humerosus</emphasis>
</taxonomicName>
differs from all other 
<taxonomicName box="[861,1079,727,753]" pageId="12" pageNumber="113">
<emphasis box="[861,985,727,753]" italics="true" pageId="12" pageNumber="113">Fusigobius</emphasis>
species
</taxonomicName>
by a combination of external morphological characters and coloration, except from 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[759,873,763,789]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[759,873,763,789]" italics="true" pageId="12" pageNumber="113">F. gracilis</emphasis>
</taxonomicName>
and 
<taxonomicName box="[927,1066,763,789]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[927,1066,763,789]" italics="true" pageId="12" pageNumber="113">F. humeralis</emphasis>
</taxonomicName>
(e.g., 
<bibRefCitation author="Randall, J. E." box="[1137,1290,763,789]" pageId="12" pageNumber="113" pagination="206 - 225" refId="ref16951" refString="Randall, J. E. (2001) Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zoological Studies, 40 (3), 206 - 225. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall 2001</bibRefCitation>
, 
<figureCitation box="[1302,1374,763,789]" captionStart="FIGURE 1" captionStartId="4.[152,255,1824,1848]" captionTargetBox="[304,1282,180,1789]" captionTargetId="figure-26@4.[304,1284,179,1798]" captionTargetPageId="4" captionText="FIGURE 1. Maximum likelihood tree estimated from the barcoding region of the mitochondrial COI gene displaying phylogenetic relationships among haplotypes from Fusigobius species.The holotype of the new species F. humerosus sp. nov. is printed in bold. Branch length is proportional to the estimated number of average nucleotide substitutions. Numbers at branches represent bootstrap proportions in percentages. Values &lt;50% are not shown. Results from species delimitation analyses based on COI gene sequences using the ABGD and bPTP algorithms are displayed as vertical bars that link terminal taxa in the tree if assigned to the same hypothetical species." figureDoi="http://doi.org/10.5281/zenodo.7745449" httpUri="https://zenodo.org/record/7745449/files/figure.png" pageId="12" pageNumber="113">Figs. 1</figureCitation>
–18). The new species is compared with each species below by the set of the most useful characters. Comparison is ordered as in diagnosis, the species descriptions may be found in the following publications (
<bibRefCitation author="Gunther, A." box="[1194,1354,835,861]" pageId="12" pageNumber="113" pagination="169 - 216" refId="ref15637" refString="Gunther, A. (1877) Andrew Garrett's Fische der Sudsee. Band II, Heft. VI. Journal des Museum Godeffroy, 4 (13), 169 - 216." type="journal article" year="1877">Günther 1877</bibRefCitation>
; 
<bibRefCitation author="Goren, M." pageId="12" pageNumber="113" pagination="191 - 203" refId="ref15604" refString="Goren, M. (1978) A new gobiid genus and seven new species from Sinai coasts (Pisces: Gobiidae). Senckenbergiana Biologica, 59, 191 - 203." type="journal article" year="1978">Goren 1978</bibRefCitation>
; 
<bibRefCitation author="Hoese, D. H. &amp; Reader, S." box="[224,482,870,897]" pageId="12" pageNumber="113" pagination="1 - 9" refId="ref15769" refString="Hoese, D. H. &amp; Reader, S. (1985) A new gobiid fish, Fusigobius duospilus, from the tropical Indo-pacific. Special Publication, J. L. B. Smith Institute of Ichthyology, 36, 1 - 9." type="journal article" year="1985">Hoese &amp; Reader 1985</bibRefCitation>
; 
<bibRefCitation author="Winterbottom, R. &amp; Emery, A. R." box="[496,836,871,897]" pageId="12" pageNumber="113" pagination="1 - 82" refId="ref17727" refString="Winterbottom, R. &amp; Emery, A. R. (1986) Review of the Gobioid fishes of the Chagos Archipelago, central Indian Ocean. Royal Ontario Museum Life Science Contributions, 142, 1 - 82. https: // doi. org / 10.5962 / bhl. title. 52226" type="journal article" year="1986">Winterbottom &amp; Emery 1986</bibRefCitation>
; 
<bibRefCitation author="Hoese, D. H. &amp; Obika, Y." box="[850,1097,871,897]" pageId="12" pageNumber="113" pagination="282 - 288" refId="ref15719" refString="Hoese, D. H. &amp; Obika, Y. (1988) A new gobiid fish, Fusigobius signipinnis, from the western tropical Pacific. Japanese Journal of Ichthyology, 35, 282 - 288. https: // doi. org / 10.1007 / BF 02938428" type="journal article" year="1988">Hoese &amp; Obika 1988</bibRefCitation>
; 
<bibRefCitation author="Chen, I. - S. &amp; Shao, K. - T." box="[1111,1335,871,897]" pageId="12" pageNumber="113" pagination="87 - 92" refId="ref15121" refString="Chen, I. - S. &amp; Shao, K. - T. (1997) Fusigobius aureus, a new species of gobiid fish (Perciformes, Gobiidae) from Flores Island, Indonesia. Acta Zoologica Taiwanica, 8 (2), 87 - 92." type="journal article" year="1997">Chen &amp; Shao 1997</bibRefCitation>
; 
<bibRefCitation author="Randall, J. E." pageId="12" pageNumber="113" pagination="317 - 340" refId="ref16877" refString="Randall, J. E. (1994) A new genus and six new gobiid fishes (Perciformes: Gobiidae) from Arabian waters. Fauna of Saudi Arabia, 14, 317 - 340." type="journal article" year="1994">Randall 1994</bibRefCitation>
, 
<bibRefCitation author="Randall, J. E." box="[221,280,907,933]" pageId="12" pageNumber="113" pagination="206 - 225" refId="ref16951" refString="Randall, J. E. (2001) Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zoological Studies, 40 (3), 206 - 225. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">2001</bibRefCitation>
, 
<bibRefCitation author="Randall, J. E." box="[291,350,906,933]" pageId="12" pageNumber="113" refId="ref16999" refString="Randall, J. E. (2005) Reef and shore fishes of the South Pacific: New Caledonia to Tahiti and the Pitcairn Islands. University of Hawai'i, Hawaii, 707 pp." type="book" year="2005">2005</bibRefCitation>
; 
<bibRefCitation author="Allen, G. R. &amp; Erdmann, M. V." box="[360,628,907,933]" pageId="12" pageNumber="113" refId="ref14765" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef Fishes of the East Indies, Volume 3. Tropical Reef Research, Perth, pp. 857 - 1260." type="book" year="2012">Allen &amp; Erdmann 2012</bibRefCitation>
; 
<bibRefCitation author="Larson, H. K." box="[639,785,907,933]" pageId="12" pageNumber="113" pagination="18 - 202" refId="ref16435" refString="Larson, H. K. (2022) Suborder Gobioidei. In: Heemstra, P. C., Heemstra, E., Ebert, D. A., Holleman, W. &amp; Randall, J. E. (Eds.), Coastal fishes of the Western Indian Ocean, volume 5. South African Institute for Aquatic Biodiversity, Makhanda, pp. 18 - 202." type="book chapter" year="2022">Larson 2022</bibRefCitation>
).
</paragraph>
<paragraph blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113">
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[199,453,943,969]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[199,453,943,969]" italics="true" pageId="12" pageNumber="113">Fusigobius humerosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName authorityName="Chen &amp; Shao" authorityYear="1997" box="[607,712,943,968]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="aureus">
<emphasis box="[607,712,943,968]" italics="true" pageId="12" pageNumber="113">F. aureus</emphasis>
</taxonomicName>
by pelvic fins joined by membrane, pelvic frenum present (vs. pelvic fins separated, except for the narrow connection, pelvic frenum absent), snout length 4.0– 
<quantity box="[1235,1299,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.2446" pageId="12" pageNumber="113" unit="in" value="4.9">4.9 in</quantity>
head length (vs. 
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head length) and longitudinal scale series 26–28 (vs. 23–24). The new species differs from 
<taxonomicName authorityName="Hoese &amp; Reader" authorityYear="1985" box="[1304,1436,1015,1041]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="duospilus">
<emphasis box="[1304,1436,1015,1041]" italics="true" pageId="12" pageNumber="113">F. duospilus</emphasis>
</taxonomicName>
by first dorsal-fin spine longest, longer than second and third (vs. first three spines subequal in length), the shortest branch of fifth pelvic-fin soft ray 83–94% of the longest branch of fourth ray (vs. 2/3 of the longest branch of fourth ray) and longitudinal scale series 26–28 (vs. 22–25, however the method applied by 
<bibRefCitation author="Hoese, D. H. &amp; Reader, S." box="[1170,1436,1122,1149]" pageId="12" pageNumber="113" pagination="1 - 9" refId="ref15769" refString="Hoese, D. H. &amp; Reader, S. (1985) A new gobiid fish, Fusigobius duospilus, from the tropical Indo-pacific. Special Publication, J. L. B. Smith Institute of Ichthyology, 36, 1 - 9." type="journal article" year="1985">Hoese &amp; Reader (1985)</bibRefCitation>
could count 2–3 fewer scales than the present method following 
<bibRefCitation author="Randall, J. E." box="[898,1070,1159,1185]" pageId="12" pageNumber="113" pagination="206 - 225" refId="ref16951" refString="Randall, J. E. (2001) Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zoological Studies, 40 (3), 206 - 225. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall (2001)</bibRefCitation>
and, counts for the two species may overlap). The new species differs from 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[655,770,1195,1221]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[655,770,1195,1221]" italics="true" pageId="12" pageNumber="113">F. gracilis</emphasis>
</taxonomicName>
by pelvic frenum present (vs. pelvic frenum absent), body depth 
<quantity box="[220,332,1231,1257]" metricMagnitude="-1" metricUnit="m" metricValue="1.1811000000000003" metricValueMax="1.2446000000000002" metricValueMin="1.1176000000000001" pageId="12" pageNumber="113" unit="in" value="4.65" valueMax="4.9" valueMin="4.4">4.4–4.9 in</quantity>
SL (vs. body slender, its depth 
<quantity box="[679,792,1230,1257]" metricMagnitude="-1" metricUnit="m" metricValue="1.4477999999999998" metricValueMax="1.4731999999999998" metricValueMin="1.4223999999999999" pageId="12" pageNumber="113" unit="in" value="5.699999999999999" valueMax="5.8" valueMin="5.6">5.6–5.8 in</quantity>
SL) and longitudinal scale series 26–28 (vs. 25). The new species differs from 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="1994" box="[387,587,1267,1293]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="inframaculatus">
<emphasis box="[387,587,1267,1293]" italics="true" pageId="12" pageNumber="113">F. inframaculatus</emphasis>
</taxonomicName>
by pectoral-fin rays 17–18 (vs. 19), first and second dorsal-fin spines not elongate or filamentous (vs. first and second dorsal-fin spines elongate and first usually filamentous), scales on side of nape not extending forward to above preopercular edge (vs. scales on side of nape extending forward nearly to eye). 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[214,466,1375,1401]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[214,466,1375,1401]" italics="true" pageId="12" pageNumber="113">Fusigobius humerosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName authorityName="Goren" authorityYear="1978" box="[616,773,1375,1401]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="longispinus">
<emphasis box="[616,773,1375,1401]" italics="true" pageId="12" pageNumber="113">F. longispinus</emphasis>
</taxonomicName>
by first and second dorsal-fin spines not elongate (vs. first dorsal-fin spine very long, reaching to the rear base of the second dorsal fin), gill openings extending forward to about half of opercle width from posterior edge of preopercle (vs. gill openings reaches only to below pectoral-fin base), longitudinal scale series 26–28 (vs. 25). The new species differs from 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[1031,1161,1483,1508]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="maximus">
<emphasis box="[1031,1161,1483,1508]" italics="true" pageId="12" pageNumber="113">F. maximus</emphasis>
</taxonomicName>
by first dorsal-fin spine longest, remaining spines progressively shorter (vs. second dorsal-fin spine longest, first to fifth spines only slightly shorter than second), first dorsal fin slightly higher than second in females, about equal in males (vs. first dorsal fin lower than second in females, about equal in males), scales on side of nape usually ending anterior at line between upper pectoral-base and first dorsal-fin origin or rarely 1–2 rows of cycloid scales extending to above opercle (vs. scales on side of nape extending forward to above upper end of preopercular margin). The new species differs from 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[213,347,1699,1725]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="melacron">
<emphasis box="[213,347,1699,1725]" italics="true" pageId="12" pageNumber="113">F. melacron</emphasis>
</taxonomicName>
by dorsal-fin rays VI+I,8–9 (vs. VI+I,10), anal-fin rays I,8 (vs. I,9), pectoral-fin rays 17–18 (vs. 19–21), first dorsal fin slightly higher than second in females, about equal in males (vs. first dorsal fin distinctly higher than second dorsal fin). The new species differs from 
<taxonomicName authorityName="Gunther" authorityYear="1877" box="[841,983,1771,1797]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="neophytus">
<emphasis box="[841,983,1771,1797]" italics="true" pageId="12" pageNumber="113">F. neophytus</emphasis>
</taxonomicName>
by eye diameter, 3.0– 
<quantity box="[1234,1298,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="12" pageNumber="113" unit="in" value="3.2">3.2 in</quantity>
head length (vs. 3.2–3.7), longitudinal scale series 26–28 (vs. 23–25), scales on side of nape ending anterior at line between upper pectoral-base and first dorsal-fin origin or rarely 1–2 rows of cycloid scales extending above opercle (vs. scales extending forward nearly to eye), operculum naked (vs. operculum rarely with a single scale on the dorsal half). The new species differs from 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[547,664,1915,1941]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="pallidus">
<emphasis box="[547,664,1915,1941]" italics="true" pageId="12" pageNumber="113">F. pallidus</emphasis>
</taxonomicName>
by pectoral-fin rays 17–18 (vs. 19–20), scales on side of nape ending anterior at line between upper pectoral-base and first dorsal-fin origin or rarely 1–2 rows of cycloid scales extends above opercle (vs. scales on side of nape extending forward to above upper end of preopercular margin). The new species differs from 
<taxonomicName authorityName="Hoese &amp; Obika" authorityYear="1988" box="[380,531,151,176]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="signipinnis">
<emphasis box="[380,531,151,176]" italics="true" pageId="13" pageNumber="114">F. signipinnis</emphasis>
</taxonomicName>
by pelvic fins joined by membrane, pelvic frenum present (vs. pelvic fins nearly fully separated, pelvic frenum absent), longitudinal scale series 26–28 (vs. 21–25, however the method applied by 
<bibRefCitation author="Hoese, D. H. &amp; Obika, Y." box="[151,407,223,249]" pageId="13" pageNumber="114" pagination="282 - 288" refId="ref15719" refString="Hoese, D. H. &amp; Obika, Y. (1988) A new gobiid fish, Fusigobius signipinnis, from the western tropical Pacific. Japanese Journal of Ichthyology, 35, 282 - 288. https: // doi. org / 10.1007 / BF 02938428" type="journal article" year="1988">Hoese &amp; Obika (1988)</bibRefCitation>
could count 2–3 fewer scales than the present method following 
<bibRefCitation author="Randall, J. E." box="[1140,1308,223,249]" pageId="13" pageNumber="114" pagination="206 - 225" refId="ref16951" refString="Randall, J. E. (2001) Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zoological Studies, 40 (3), 206 - 225. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall (2001)</bibRefCitation>
and counts for the two species may overlap).
</paragraph>
<paragraph blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="114">
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[199,420,295,321]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[199,420,295,321]" italics="true" pageId="13" pageNumber="114">Fusigobius gracilis</emphasis>
</taxonomicName>
, 
<taxonomicName box="[436,579,295,321]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[436,579,295,321]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[639,789,295,321]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[639,789,295,321]" italics="true" pageId="13" pageNumber="114">F. humerosus</emphasis>
</taxonomicName>
are very similar in coloration and possess a translucent body with numerous small dusky orange-yellow spots on the head and body combined with a round black spot smaller than or as large as pupil in humeral region just above base of pectoral fin and another slightly larger black spot at midbase of caudal fin. 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[492,744,403,429]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[492,744,403,429]" italics="true" pageId="13" pageNumber="114">Fusigobius humerosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[894,1008,403,429]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[894,1008,403,429]" italics="true" pageId="13" pageNumber="114">F. gracilis</emphasis>
</taxonomicName>
by having dusky orange-yellow spots with a series of melanophores within and adjacent in each spot vs. numerous very small orange-yellow spots that cover head and body, each containing a single melanophore except for ventral half of the posterior part of body with a longitudinal series of orange-yellow spots surrounded by a series of melanophores, and a distinct patch of melanophores on side of abdomen in 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[566,679,547,573]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[566,679,547,573]" italics="true" pageId="13" pageNumber="114">F. gracilis</emphasis>
</taxonomicName>
. Also iris is usually unicolor in 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[1031,1281,547,573]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[1031,1281,547,573]" italics="true" pageId="13" pageNumber="114">Fusigobius humerosus</emphasis>
</taxonomicName>
vs. dorsal iris and dorsal surface of orbit with tiny orange-brown spots in 
<taxonomicName baseAuthorityName="Randall" baseAuthorityYear="2001" box="[817,930,583,609]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[817,930,583,609]" italics="true" pageId="13" pageNumber="114">F. gracilis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="114">
The new species differs from the similarly colored 
<taxonomicName box="[781,921,619,645]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[781,921,619,645]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
(
<figureCitation box="[938,1006,619,645]" captionStart="FIGURE 7" captionStartId="14.[151,250,1752,1776]" captionTargetBox="[151,1436,181,1728]" captionTargetId="figure-26@14.[151,1436,181,1728]" captionTargetPageId="14" captionText="FIGURE 7. Fusigobius humeralis. A: BPBM 32955, holotype, 28 mm SL, South MaleAtoll, Maldives; B: USNM 439586, 25.7 mm SL, Moorea, Society Islands; C: Ngulu Atoll, Micronesia; D: Raja Ampat Is., West Papua, Indonesia. Photos by J.E. Randall (A), J.T. Williams (B) and G.R. Allen (C &amp; D)." figureDoi="http://doi.org/10.5281/zenodo.7745465" httpUri="https://zenodo.org/record/7745465/files/figure.png" pageId="13" pageNumber="114">Fig. 7</figureCitation>
) in minor details of coloration of live individuals: a unicolor iris, rarely suffused with yellow vs. iris with orange-red marks anteriorly and dorsally around eye in 
<taxonomicName box="[225,365,691,717]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[225,365,691,717]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
. The new species also differs from 
<taxonomicName box="[762,901,691,717]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[762,901,691,717]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
in the following morphological characters: first dorsal-fin spine longest, 
<quantity box="[432,547,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.5559999999999996" pageId="13" pageNumber="114" unit="in" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 in</quantity>
head length, 4.2–5.0 in SL, remaining spines progressively shorter vs. second and third dorsal-fin spines longest and subequal, 
<quantity box="[717,832,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="5.460999999999999" metricValueMax="6.35" metricValueMin="4.572" pageId="13" pageNumber="114" unit="in" value="2.15" valueMax="2.5" valueMin="1.8">1.8–2.5 in</quantity>
head length, 
<quantity box="[990,1106,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.5112999999999999" metricValueMax="1.7271999999999998" metricValueMin="1.2953999999999999" pageId="13" pageNumber="114" unit="in" value="5.949999999999999" valueMax="6.8" valueMin="5.1">5.1–6.8 in</quantity>
SL; anal-fin spine short, the length 
<quantity box="[231,346,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="9.270999999999999" metricValueMax="9.906" metricValueMin="8.636" pageId="13" pageNumber="114" unit="in" value="3.65" valueMax="3.9" valueMin="3.4">3.4–3.9 in</quantity>
head length vs. 
<quantity box="[539,654,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="7.239" metricValueMax="8.128" metricValueMin="6.35" pageId="13" pageNumber="114" unit="in" value="2.85" valueMax="3.2" valueMin="2.5">2.5–3.2 in</quantity>
head length; the shortest branch of fifth pelvic-fin ray short, length 18.1–22.1% SL vs. 19.4–27.5% SL (single overlapping value of 19.4% SL was found only in ROM 60914, 
<collectingCountry name="French Polynesia" pageId="13" pageNumber="114">French Polynesia</collectingCountry>
); posterior nostril about halfway between anterior margin of orbit and anterior nostril vs. posterior nostril closer to orbit than to anterior nostril; upper jaw short, the length 
<quantity box="[884,997,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" metricValueMax="8.89" metricValueMin="8.382" pageId="13" pageNumber="114" unit="in" value="3.4" valueMax="3.5" valueMin="3.3">3.3–3.5 in</quantity>
head length vs. 
<quantity box="[1179,1292,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="7.239" metricValueMax="8.128" metricValueMin="6.35" pageId="13" pageNumber="114" unit="in" value="2.85" valueMax="3.2" valueMin="2.5">2.5–3.2 in</quantity>
head length; longitudinal scale series 26–28 vs. 23–25; head and predorsal naked, scales on side of nape ending usually anterior at line between upper pectoral-base and first dorsal-fin origin or rarely 1–2 rows of cycloid scales extending above opercle vs. scales on side of nape variably extending forward nearly to eye, to between eye and above preopercular edge or to above preopercular edge. The two species are also delimited by their geographic ranges, 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[1287,1436,1051,1077]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[1287,1436,1051,1077]" italics="true" pageId="13" pageNumber="114">F. humerosus</emphasis>
</taxonomicName>
is known from the Red Sea and the Gulf of Aden, while 
<taxonomicName box="[804,945,1087,1113]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[804,945,1087,1113]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
is widespread from the 
<collectingCountry box="[1224,1330,1087,1113]" name="Comoros" pageId="13" pageNumber="114">Comoros</collectingCountry>
, 
<collectingCountry name="Maldives" pageId="13" pageNumber="114">Maldive Islands</collectingCountry>
and Chagos Archipelago in the western Indian Ocean to 
<collectingCountry box="[892,1088,1123,1149]" name="French Polynesia" pageId="13" pageNumber="114">French Polynesia</collectingCountry>
in the Pacific Ocean (
<bibRefCitation author="Randall, J. E." pageId="13" pageNumber="114" pagination="206 - 225" refId="ref16951" refString="Randall, J. E. (2001) Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zoological Studies, 40 (3), 206 - 225. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall 2001</bibRefCitation>
, present research).
</paragraph>
<paragraph blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="114">
Size of the spot in humeral area and spot at caudal-fin base may be useful characters to differentiate 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[1290,1436,1195,1221]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[1290,1436,1195,1221]" italics="true" pageId="13" pageNumber="114">F. humerosus</emphasis>
</taxonomicName>
from 
<taxonomicName box="[215,356,1231,1257]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[215,356,1231,1257]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
, with exceptions of some specimens of the latter species showing overlapping characters. The round black spot in humeral region just above the pectoral-fin base is smaller or as large as pupil in 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[1287,1436,1267,1293]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[1287,1436,1267,1293]" italics="true" pageId="13" pageNumber="114">F. humerosus</emphasis>
</taxonomicName>
vs. dark spot usually larger than pupil in 
<taxonomicName box="[616,757,1303,1329]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[616,757,1303,1329]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
(with the exception of some comparative specimens having dark spot size of pupil or slightly smaller: ROM 85028, 
<collectingCountry box="[805,918,1339,1365]" name="Indonesia" pageId="13" pageNumber="114">Indonesia</collectingCountry>
; ROM 81801 and ROM 81686, 
<collectingCountry box="[1293,1359,1339,1365]" name="Palau" pageId="13" pageNumber="114">Palau</collectingCountry>
; ROM 64085, 
<collectingCountry box="[235,412,1375,1401]" name="New Caledonia" pageId="13" pageNumber="114">New Caledonia</collectingCountry>
; and ROM 37240, ROM 37241, ROM 37243, ROM 37244, Chagos Archipelago). The oval black spot at midbase of caudal fin covers less than 1/3 of caudal-fin base height in 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[1092,1239,1411,1437]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[1092,1239,1411,1437]" italics="true" pageId="13" pageNumber="114">F. humerosus</emphasis>
</taxonomicName>
vs. irregular spot at midbase of caudal fin larger than 1/3 of caudal-fin base height in 
<taxonomicName box="[905,1044,1447,1473]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[905,1044,1447,1473]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
(except for some specimens where it is equal to but not less than 1/3 of caudal-fin base height: ROM 85028, 
<collectingCountry box="[971,1084,1483,1509]" name="Indonesia" pageId="13" pageNumber="114">Indonesia</collectingCountry>
; ROM 81801 and ROM 81686, 
<collectingCountry box="[151,217,1519,1545]" name="Palau" pageId="13" pageNumber="114">Palau</collectingCountry>
; 
<figureCitation box="[228,372,1519,1545]" captionStart="FIGURE 6" captionStartId="11.[151,250,1156,1180]" captionTargetBox="[151,1436,181,1131]" captionTargetId="figure-20@11.[151,1436,181,1131]" captionTargetPageId="11" captionText="FIGURE 6. A and C: Fusigobius humerosus sp. nov., PMR VP5271, paratype, male, 26.2 + 7.3 mm, northern side of Farasan Island, Saudi Arabia, Red Sea. B and D: F. humeralis, BPBM 36627, paratype, female, 29.5 + 7.8 mm, Manuk Island, Banda Sea, Indonesia. Humeral region just above base of pectoral fin. A: round black spot is smaller than or equal to pupil in F. humerosus sp. nov., B: dark spot larger than pupil in F. humeralis; midbase of caudal fin: C: oval black spot covers less than 1/3 of caudal base height in F. humerosus sp. nov., D: irregular spot at midbase of caudal fin larger than 1/3 of caudal base height in F. humeralis." figureDoi="http://doi.org/10.5281/zenodo.7745463" httpUri="https://zenodo.org/record/7745463/files/figure.png" pageId="13" pageNumber="114">Fig. 6B &amp; D</figureCitation>
).
</paragraph>
<paragraph blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="114">
<bibRefCitation author="Randall, J. E." box="[199,368,1555,1581]" pageId="13" pageNumber="114" pagination="206 - 225" refId="ref16951" refString="Randall, J. E. (2001) Five new Indo-Pacific gobiid fishes of the genus Coryphopterus. Zoological Studies, 40 (3), 206 - 225. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall (2001)</bibRefCitation>
noted that specimens from the 
<collectingRegion box="[723,816,1555,1581]" country="Egypt" name="Al Bahr al Ahmar" pageId="13" pageNumber="114">Red Sea</collectingRegion>
and Gulf of 
<collectingRegion box="[960,1023,1555,1581]" country="Yemen" name="'Adan" pageId="13" pageNumber="114">Aden</collectingRegion>
, which were described as non-types, have 17 pectoral-fin rays whereas specimens from the Indo-West Pacific usually have 18 rays. Re-examination of Randall’s material and additional 
<collectingRegion box="[525,616,1627,1653]" country="Egypt" name="Al Bahr al Ahmar" pageId="13" pageNumber="114">Red Sea</collectingRegion>
material of the new species confirmed possessing usually of 17 rays. This material was added as non-type material of 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[640,788,1663,1689]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[640,788,1663,1689]" italics="true" pageId="13" pageNumber="114">F. humerosus</emphasis>
</taxonomicName>
in the present new species description.
</paragraph>
<paragraph blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="114">
Specimens of 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" box="[358,610,1699,1725]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis box="[358,610,1699,1725]" italics="true" pageId="13" pageNumber="114">Fusigobius humerosus</emphasis>
</taxonomicName>
from the Red Sea and Gulf of Aden have been incorrectly identified as 
<taxonomicName class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
. Along with the above morphological evidence, both molecular approaches for the species delimitation (i.e. ABGD and PTP) recognized the new species as an independent lineage at species level (see 
<figureCitation box="[1297,1365,1771,1797]" captionStart="FIGURE 1" captionStartId="4.[152,255,1824,1848]" captionTargetBox="[304,1282,180,1789]" captionTargetId="figure-26@4.[304,1284,179,1798]" captionTargetPageId="4" captionText="FIGURE 1. Maximum likelihood tree estimated from the barcoding region of the mitochondrial COI gene displaying phylogenetic relationships among haplotypes from Fusigobius species.The holotype of the new species F. humerosus sp. nov. is printed in bold. Branch length is proportional to the estimated number of average nucleotide substitutions. Numbers at branches represent bootstrap proportions in percentages. Values &lt;50% are not shown. Results from species delimitation analyses based on COI gene sequences using the ABGD and bPTP algorithms are displayed as vertical bars that link terminal taxa in the tree if assigned to the same hypothetical species." figureDoi="http://doi.org/10.5281/zenodo.7745449" httpUri="https://zenodo.org/record/7745449/files/figure.png" pageId="13" pageNumber="114">Fig. 1</figureCitation>
). The phylogenetic analysis of mitochondrial COI showed that sequences generated from the 
<typeStatus box="[1109,1157,1807,1833]" pageId="13" pageNumber="114">type</typeStatus>
specimens of 
<taxonomicName authorityName="Bogorodsky &amp; Kovačić &amp; Mal &amp; Alpermann" authorityYear="2023" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humerosus">
<emphasis italics="true" pageId="13" pageNumber="114">Fusigobius humerosus</emphasis>
</taxonomicName>
formed a monophyletic clade with other sequences of Red Sea origin. The resulting clade was sister to a clade composed of 
<taxonomicName box="[367,610,1879,1905]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[367,610,1879,1905]" italics="true" pageId="13" pageNumber="114">Fusigobius humeralis</emphasis>
</taxonomicName>
from 
<collectingCountry box="[678,871,1879,1905]" name="French Polynesia" pageId="13" pageNumber="114">French Polynesia</collectingCountry>
and 
<collectingCountry box="[925,1011,1879,1905]" name="Taiwan" pageId="13" pageNumber="114">Taiwan</collectingCountry>
. Unfortunately, no tissue material was available from or close to the 
<typeStatus box="[488,536,1915,1941]" pageId="13" pageNumber="114">type</typeStatus>
locality of 
<taxonomicName box="[666,806,1915,1941]" class="Actinopterygii" family="Gobiidae" genus="Fusigobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="humeralis">
<emphasis box="[666,806,1915,1941]" italics="true" pageId="13" pageNumber="114">F. humeralis</emphasis>
</taxonomicName>
(i.e. 
<collectingCountry box="[864,973,1915,1941]" name="Maldives" pageId="13" pageNumber="114">Maldives</collectingCountry>
). However, the identities of the two most distant specimens (from 
<collectingCountry box="[426,621,1951,1977]" name="French Polynesia" pageId="13" pageNumber="114">French Polynesia</collectingCountry>
) were confirmed by images in BOLD and by the original identifier based on distinguishing characters as defined herein (J.T. Williams, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>