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<document id="E04036537AE157957F8CF6E3D03A829E" ID-CLB-Dataset="37404" ID-DOI="10.11646/zootaxa.4205.4.1" ID-GBIF-Dataset="eb5dfbad-e024-4318-b58e-6d9b9e19eac3" ID-ISSN="1175-5326" ID-Zenodo-Dep="208241" ID-ZooBank="BAC8F107-21D6-49FE-BAC7-BF4EE6C3E6A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1482105213028" checkinUser="plazi" docAuthor="Kolesik, Peter &amp; Gagné, Raymond J." docDate="2016" docId="D759878B0E77EE595BF6FF4D287BFE9F" docLanguage="en" docName="Zootaxa.4205.4.301-338" docOrigin="Zootaxa 4205 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brachineura australis Skuse 1890" docType="treatment" docVersion="12" lastPageNumber="309" masterDocId="2B60FFF30E7FEE505B61FFDA2A35FFDF" masterDocTitle="Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae)" masterLastPageNumber="338" masterPageNumber="301" pageNumber="309" updateTime="1698434883341" updateUser="ExternalLinkService">
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<mods:title id="8031394F5C270133CE8CB86991976A53">Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae)</mods:title>
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<mods:namePart id="5BED1F52018FE3832B11D53923CDFBC8">Kolesik, Peter</mods:namePart>
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<mods:namePart id="C958D7F38B45650DE5BFFFF45BB2C8D2">Gagné, Raymond J.</mods:namePart>
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<mods:title id="1D310B613992383D22D40D284B6B4E9E">Zootaxa</mods:title>
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<mods:date id="51197501267B6F62330893A285E5693B">2016</mods:date>
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<treatment id="D759878B0E77EE595BF6FF4D287BFE9F" ID-DOI="http://doi.org/10.5281/zenodo.4752501" ID-GBIF-Taxon="125140064" ID-Zenodo-Dep="4752501" LSID="urn:lsid:plazi:treatment:D759878B0E77EE595BF6FF4D287BFE9F" httpUri="http://treatment.plazi.org/id/D759878B0E77EE595BF6FF4D287BFE9F" lastPageId="9" lastPageNumber="309" pageId="8" pageNumber="309">
<subSubSection id="17EA65160E77EE585BF6FF4D2ADEFF0C" pageId="8" pageNumber="309" type="nomenclature">
<paragraph id="5F4F369D0E77EE585BF6FF4D287BFF6D" blockId="8.[151,590,151,211]" box="[151,590,151,178]" pageId="8" pageNumber="309">
<heading id="040781F10E77EE585BF6FF4D287BFF6D" bold="true" box="[151,590,151,178]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<taxonomicName id="98F04D1E0E77EE585BF6FF4D287BFF6D" ID-CoL="MSVS" authority="Skuse, 1890" authorityName="Skuse" authorityYear="1890" box="[151,590,151,178]" class="Insecta" family="Cecidomyiidae" genus="Brachineura" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="6D84EA8F0E77EE585BF6FF4D287BFF6D" bold="true" box="[151,590,151,178]" pageId="8" pageNumber="309">
<emphasis id="6D84EA8F0E77EE585BF6FF4D2B94FF6E" bold="true" box="[151,417,151,177]" italics="true" pageId="8" pageNumber="309">Brachineura australis</emphasis>
(
<bibRefCitation id="3B614B6C0E77EE585AD1FF422873FF6D" author="Skuse" box="[432,582,152,178]" pageId="8" pageNumber="309" refString="Skuse, F. A. A. (1890) Diptera of Australia. Nematocera. Supplement I. Proceedings of the Linnean Society of New South Wales, 5, 373 - 412. [pl. XVI.]" type="journal article" year="1890">Skuse, 1890</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="5F4F369D0E77EE585BF6FF602ADEFF0C" blockId="8.[151,590,151,211]" box="[151,235,186,211]" pageId="8" pageNumber="309">
[
<figureCitation id="C7CB2A180E77EE585BFFFF602AD1FF0C" box="[158,228,186,211]" captionStart="FIGURE 4" captionStartId="8.[151,250,1370,1392]" captionTargetBox="[185,1388,409,1309]" captionTargetId="figure@8.[151,1436,361,1349]" captionTargetPageId="8" captionText="FIGURE 4. Brachineura australis (Skuse). Male. (a) wing (part missing), (b) second flagellomere, (c) last tarsomere of foreleg with claws and empodium, (d) terminalia in dorsolateral view (substantially compressed, mediobasal lobes not recognisable). Figure a from Kolesik (2014)." httpUri="https://zenodo.org/record/208245/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
]
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E77EE585BF6FED9291FFEE8" pageId="8" pageNumber="309" type="reference_group">
<paragraph id="5F4F369D0E77EE585BF6FED92813FEC7" blockId="8.[151,810,258,311]" box="[151,550,258,280]" pageId="8" pageNumber="309">
<treatmentCitationGroup id="7FE011B30E77EE585BF6FED92813FEC7" box="[151,550,258,280]" pageId="8" pageNumber="309">
<taxonomicName id="98F04D1E0E77EE585BF6FED92814FEC7" ID-CoL="8F3DY" authority="Skuse, 1890: 385" authorityName="Skuse" authorityPageNumber="385" authorityYear="1890" box="[151,545,258,280]" class="Insecta" family="Cecidomyiidae" genus="Spaniocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="6D84EA8F0E77EE585BF6FED92B5CFEC7" box="[151,361,258,280]" italics="true" pageId="8" pageNumber="309">Spaniocera australis</emphasis>
<treatmentCitation id="DE51108C0E77EE585A0EFED82814FEC7" author="Skuse" box="[367,545,258,280]" page="385" pageId="8" pageNumber="309" year="1890">
<bibRefCitation id="3B614B6C0E77EE585A0EFED82BDAFEC7" author="Skuse" box="[367,495,258,280]" pageId="8" pageNumber="309" refString="Skuse, F. A. A. (1890) Diptera of Australia. Nematocera. Supplement I. Proceedings of the Linnean Society of New South Wales, 5, 373 - 412. [pl. XVI.]" type="journal article" year="1890">Skuse, 1890</bibRefCitation>
: 385
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="5F4F369D0E77EE585BF6FEFB291FFEE8" blockId="8.[151,810,258,311]" box="[151,810,289,311]" pageId="8" pageNumber="309">
<treatmentCitationGroup id="7FE011B30E77EE585BF6FEFB291FFEE8" box="[151,810,289,311]" pageId="8" pageNumber="309">
<taxonomicName id="98F04D1E0E77EE585BF6FEFB2B43FEE8" ID-CoL="MSVS" box="[151,374,289,311]" class="Insecta" family="Cecidomyiidae" genus="Brachineura" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="australis" status="comb. nov.">
<emphasis id="6D84EA8F0E77EE585BF6FEFB2B43FEE8" box="[151,374,289,311]" italics="true" pageId="8" pageNumber="309">Brachineura australis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="76B757F40E77EE585AE3FEF82804FEE8" box="[386,561,289,311]" pageId="8" pageNumber="309">new combination</taxonomicNameLabel>
by
<treatmentCitation id="DE51108C0E77EE585936FEFB2913FEE8" author="Gagne" box="[599,806,289,311]" page="156" pageId="8" pageNumber="309" year="1989">
<bibRefCitation id="3B614B6C0E77EE585936FEFB2913FEE8" author="Gagne" box="[599,806,289,311]" pageId="8" pageNumber="309" refString="Gagne, R. J. (1989 b) Family Cecidomyiidae. In: Evenhuis, N. L. (Ed.), &quot; Catalog of the Diptera of the Australasian and Oceanian Regions &quot; (Bishop Museum Press and E. J. Brill, Honolulu), pp. 152 - 163." type="book chapter" year="1989" yearSuffix="b">Gagné (1989b: 156)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="0B8F66150E77EE585BF6FA802868FA72" httpUri="https://zenodo.org/record/208245/files/figure.png" pageId="8" pageNumber="309" targetBox="[185,1388,409,1309]" targetPageId="8">
<paragraph id="5F4F369D0E77EE585BF6FA802868FA72" blockId="8.[151,1435,1370,1453]" pageId="8" pageNumber="309">
<emphasis id="6D84EA8F0E77EE585BF6FA802B24FAAF" bold="true" box="[151,273,1370,1392]" pageId="8" pageNumber="309">FIGURE 4</emphasis>
.
<taxonomicName id="98F04D1E0E77EE585A40FA80286AFAAF" authority="Skuse" authorityName="Skuse" box="[289,607,1370,1392]" class="Insecta" family="Cecidomyiidae" genus="Brachineura" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="6D84EA8F0E77EE585A40FA802830FAAF" box="[289,517,1370,1392]" italics="true" pageId="8" pageNumber="309">Brachineura australis</emphasis>
(Skuse)
</taxonomicName>
. Male. (
<emphasis id="6D84EA8F0E77EE5859DBFA8028F3FAAF" bold="true" box="[698,710,1370,1392]" pageId="8" pageNumber="309">a</emphasis>
) wing (part missing), (
<emphasis id="6D84EA8F0E77EE5858A3FA8029E5FAAF" bold="true" box="[962,976,1370,1392]" pageId="8" pageNumber="309">b</emphasis>
) second flagellomere, (
<emphasis id="6D84EA8F0E77EE585FADFA802EE2FAAF" bold="true" box="[1228,1239,1370,1392]" pageId="8" pageNumber="309">c</emphasis>
) last tarsomere of foreleg with claws and empodium, (
<emphasis id="6D84EA8F0E77EE585942FAA32804FA50" bold="true" box="[547,561,1401,1423]" pageId="8" pageNumber="309">d</emphasis>
) terminalia in dorsolateral view (substantially compressed, mediobasal lobes not recognisable). Figure a from Kolesik (2014).
</paragraph>
</caption>
<subSubSection id="17EA65160E77EE585BF6FA0F2B98F9A1" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="5F4F369D0E77EE585BF6FA0F2B98F9A1" blockId="8.[151,1436,1493,2024]" pageId="8" pageNumber="309">
<materialsCitation id="EF983CC00E77EE585BF6FA0F2B98F9A1" ID-GBIF-Occurrence="1416805656" collectionCode="ANIC" collectorName="Gosford, N. S. W." county="The" location="Hogan's Brush" municipality="August" pageId="8" pageNumber="309" specimenCode="ANIC 29-38480" specimenCount="2" typeStatus="holotype">
<emphasis id="6D84EA8F0E77EE585BF6FA0F2BD6FA31" bold="true" box="[151,483,1493,1518]" pageId="8" pageNumber="309">
Material studied.
<typeStatus id="804B883F0E77EE585A17FA0F2BD6FA31" box="[374,483,1493,1518]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
</emphasis>
male
<typeStatus id="804B883F0E77EE585953FA0F28A1FA31" box="[562,660,1493,1518]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
(
<specimenCode id="0F569EE60E77EE5859C9FA0C295CFA31" box="[680,873,1494,1518]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="8" pageNumber="309">
<collectionCode id="39E1AE580E77EE5859C9FA0C28DBFA31" box="[680,750,1494,1518]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="8" pageNumber="309">ANIC</collectionCode>
29-38480
</specimenCode>
), “
<location id="5A2F60460E77EE5858EDFA0C2E08FA31" LSID="urn:lsid:plazi:treatment:D759878B0E77EE595BF6FF4D287BFE9F:5A2F60460E77EE5858EDFA0C2E08FA31" box="[908,1085,1493,1518]" county="The" municipality="August" name="Hogan's Brush" pageId="8" pageNumber="309">Hogans Brush</location>
,
<location id="5A2F60460E77EE585F2FFA0C2ED9FA31" LSID="urn:lsid:plazi:treatment:D759878B0E77EE595BF6FF4D287BFE9F:5A2F60460E77EE585F2FFA0C2ED9FA31" box="[1102,1260,1493,1518]" county="The" municipality="August" name="Narara Creek" pageId="8" pageNumber="309">Narara Creek</location>
, near
<collectorName id="F205534B0E77EE585E58FA0F2ADDF9CE" pageId="8" pageNumber="309">Gosford, N.S.W.</collectorName>
(Skuse).
<location id="5A2F60460E77EE585A30FA232BB5F9CE" LSID="urn:lsid:plazi:treatment:D759878B0E77EE595BF6FF4D287BFE9F:5A2F60460E77EE585A30FA232BB5F9CE" box="[337,384,1529,1553]" county="The" municipality="August" name="One" pageId="8" pageNumber="309">One</location>
specimen in
<collectingMunicipality id="BF2BACE70E77EE585973FA232853F9CE" box="[530,614,1529,1553]" pageId="8" pageNumber="309">August</collectingMunicipality>
, in dense bush”.
<collectingCounty id="B62E4E110E77EE585840FA232978F9CD" box="[801,845,1529,1554]" pageId="8" pageNumber="309">The</collectingCounty>
mounted has retained most of one wing, the first two and three flagellomeres [remaining ones missing originally (
<bibRefCitation id="3B614B6C0E77EE58582DF9C729EDF9E9" author="Skuse" box="[844,984,1565,1590]" pageId="8" pageNumber="309" refString="Skuse, F. A. A. (1890) Diptera of Australia. Nematocera. Supplement I. Proceedings of the Linnean Society of New South Wales, 5, 373 - 412. [pl. XVI.]" type="journal article" year="1890">Skuse 1890</bibRefCitation>
)], a shrivelled head with missing palpi, thorax with parts of legs including one entire foreleg, its claws partially obscured, and the substantially compressed abdomen and terminalia.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E77EE585BA6F953294AF87F" pageId="8" pageNumber="309" type="description">
<paragraph id="5F4F369D0E77EE585BA6F953294AF87F" blockId="8.[151,1436,1493,2024]" pageId="8" pageNumber="309">
<emphasis id="6D84EA8F0E77EE585BA6F9532B6CF97D" bold="true" box="[199,345,1673,1698]" pageId="8" pageNumber="309">Description.</emphasis>
Wing
<quantity id="98089B780E77EE585AC5F9532BCFF97E" box="[420,506,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="309" unit="mm" value="1.4">1.4 mm</quantity>
long,
<quantity id="98089B780E77EE585923F95328ADF97E" box="[578,664,1673,1698]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="309" unit="mm" value="0.6">0.6 mm</quantity>
wide, R4+5 4/5 wing length, Cu simple, fading apically, wingfold not apparent. Antennal flagellomeres cylindrical, second
<date id="2B4E105D0E77EE585990F96A2938F917" box="[753,781,1712,1736]" pageId="8" pageNumber="309">3x</date>
longer than wide, with short necks; circumfila appressed, consisting of a continuous sinuous band along length of flagellomere and a separate, apical transverse band. Claws on foreleg apparently simple, thin, curved near midlength, empodium shorter than claws. Last abdominal segments with scales, the sclerites obscured on mount. Terminalia: gonocoxite with acute apicoventral process; gonostylus wide and microtrichose on basal half, narrow and carinate on distal half, with comb-like tooth apically; aedeagus long, strongly recurved, tapering apically; cerci large, elliptical; hypoproct broad, slightly shorter than cerci, shallowly concave distally, with apical seta on each resulting lobe.
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E77EE595BA6F876287BFE9F" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" type="discussion">
<paragraph id="5F4F369D0E77EE595BA6F876287BFE9F" blockId="8.[151,1436,1493,2024]" lastBlockId="9.[151,1436,151,320]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
<emphasis id="6D84EA8F0E77EE585BA6F8762B0EF81A" bold="true" box="[199,315,1964,1989]" pageId="8" pageNumber="309">Remarks.</emphasis>
The terminalia are distorted on the slide mount but are nevertheless tantalizingly odd. The curved, acute apex of the gonocoxite is reminiscent of the shape of the ventrodistal lobe in
<taxonomicName id="98F04D1E0E77EE585F2FF80A2E9EF838" box="[1102,1195,2000,2023]" class="Insecta" family="Cecidomyiidae" genus="Epimyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="6D84EA8F0E77EE585F2FF80A2E9EF838" box="[1102,1195,2000,2023]" italics="true" pageId="8" pageNumber="309">Epimyia</emphasis>
</taxonomicName>
. A generally similar, greatly recurved aedeagus is also found in
<taxonomicName id="98F04D1E0E76EE595913FF4228FAFF70" box="[626,719,152,175]" class="Insecta" family="Cecidomyiidae" genus="Epimyia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="6D84EA8F0E76EE595913FF4228FAFF70" box="[626,719,152,175]" italics="true" pageId="9" pageNumber="310">Epimyia</emphasis>
</taxonomicName>
, but the cerci and hypoproct in the present species are shaped as in
<taxonomicName id="98F04D1E0E76EE595BD4FF662B77FF0B" box="[181,322,188,212]" class="Insecta" family="Cecidomyiidae" genus="Brachineura" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="6D84EA8F0E76EE595BD4FF662B77FF0B" box="[181,322,188,212]" italics="true" pageId="9" pageNumber="310">Brachineura</emphasis>
</taxonomicName>
. The bizarre shape of the terminalia in this species insures a species identification should another specimen become available. The tarsal claws appear to lack the basal tooth, but the acromere of the only available leg is not well displayed and partially obscured on the mount. If the claws are truly simple, this species will be the first to lack toothed claws in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>