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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.7.e29130" ID-GBIF-Dataset="709abdc2-9cad-4c9b-9786-7816ac10a9fb" ID-PMC="PMC6477854" ID-Pensoft-Pub="1314-2828--29130" ID-PubMed="31048976" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2828--e29130" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="A new species of Trismegistomya Reinhard (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica" checkinTime="1554922601376" checkinUser="pensoft" docAuthor="Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Dapkey, Tanya, Hallwachs, Winnie &amp; Janzen, Daniel H" docDate="2019" docId="60654EF40978C8E31999079FC314904A" docLanguage="en" docName="BiodivDatJour 7: e29130" docOrigin="Biodiversity Data Journal 7" docSource="http://dx.doi.org/10.3897/BDJ.7.e29130" docTitle="Trismegistomya jimoharai Fleming &amp; Wood, sp. n." docType="treatment" docVersion="5" lastPageNumber="29130" masterDocId="CC33CC6A59727543190EED182C40FFD2" masterDocTitle="A new species of Trismegistomya Reinhard (Diptera: Tachinidae) from Area de Conservacion Guanacaste in northwestern Costa Rica" masterLastPageNumber="29130" masterPageNumber="29130" pageNumber="29130" updateTime="1668124899685" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Trismegistomya Reinhard (Diptera: Tachinidae) from Area de Conservacion Guanacaste in northwestern Costa Rica</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fleming, AJ</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wood, D. Monty</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, M. Alex</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dapkey, Tanya</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hallwachs, Winnie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Janzen, Daniel H</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>29130</mods:start>
<mods:end>29130</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.7.e29130</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.7.e29130</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--29130</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="156146698" LSID="urn:lsid:zoobank.org:act:790573DE-5676-4D55-B62B-78CDBDE101C7" httpUri="http://treatment.plazi.org/id/60654EF40978C8E31999079FC314904A" lastPageNumber="29130" pageId="0" pageNumber="29130">
<subSubSection pageId="0" pageNumber="29130" type="nomenclature">
<paragraph pageId="0" pageNumber="29130">
<taxonomicName LSID="urn:lsid:zoobank.org:act:790573DE-5676-4D55-B62B-78CDBDE101C7" authority="Fleming &amp; Wood" class="Insecta" family="Tachinidae" genus="Trismegistomya" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trismegistomya jimoharai" order="Diptera" pageId="0" pageNumber="29130" phylum="Arthropoda" rank="species" species="jimoharai">Trismegistomya jimoharai Fleming &amp; Wood</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="29130">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29130" type="materials_examined">
<paragraph pageId="0" pageNumber="29130">Materials</paragraph>
<paragraph pageId="0" pageNumber="29130">
<materialsCitation ID-GBIF-Occurrence="2235579005" collectingDate="1984-05-25" collectingMethod="Reared from Erebidae moth larva, Melipotisjanuaris" collectionCode="CNC, Insects" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" latitude="10.802" location="Area de Conservacion Guanacaste" longitude="-85.654" pageId="0" pageNumber="29130" specimenCode="DHJPAR 0018620" specimenCount="1" specimenCount-male="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="29130">Holotype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="29130">DHJPAR0018620</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="29130">D.H. Janzen &amp; W. Hallwachs</collectorName>
; individualID: DHJPAR0018620; individualCount:
<specimenCount pageId="0" pageNumber="29130">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="29130">male</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="29130">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 84-SRNP-154.3,BOLD:AAW7939,ASTAI1267-07; Taxon: scientificName: Trismegistomyajimoharai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Trismegistomya; specificEpithet: jimoharai; scientificNameAuthorship: Fleming &amp; Wood, 2015; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="29130">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality:
<location LSID="urn:lsid:plazi:treatment:60654EF40978C8E31999079FC314904A:B7592D21BAA4C10E8EFD55DEB1A465C6" country="Costa Rica" latitude="10.802" longitude="-85.654" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="29130">
Area de
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
Guanacaste
</location>
; verbatimLocality: Vado Nisperal; verbatimElevation:
<elevation pageId="0" pageNumber="29130">10</elevation>
; verbatimLatitude: 10.802; verbatimLongitude: -85.654; verbatimCoordinateSystem: Decimal; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="29130" value="10.802">10.802</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="29130" value="-85.654">-85.654</geoCoordinate>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="29130">AJ Fleming</determinerName>
; dateIdentified: 2015; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="29130">Reared from Erebidae moth larva, Melipotisjanuaris</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="29130" value="1984-05-25">05/25/1984</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="29130">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="29130">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2235579006" collectingDate="1984-05-25" collectingMethod="Reared from Erebidae moth larva, Melipotisjanuaris" collectionCode="CNC, Insects" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" latitude="10.802" location="Area de Conservacion Guanacaste" longitude="-85.654" pageId="0" pageNumber="29130" specimenCode="DHJPAR 0018621" specimenCount="1" specimenCount-male="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="29130">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="29130">DHJPAR0018621</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="29130">D.H. Janzen &amp; W. Hallwachs</collectorName>
; individualID: DHJPAR0018621; individualCount:
<specimenCount pageId="0" pageNumber="29130">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="29130">male</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="29130">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 84-SRNP-154,ASTAI1268-07; Taxon: scientificName: Trismegistomyajimoharai; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Trismegistomya; specificEpithet: jimoharai; scientificNameAuthorship: Fleming &amp; Wood, 2015; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="29130">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality:
<location LSID="urn:lsid:plazi:treatment:60654EF40978C8E31999079FC314904A:8123093C60C7F48EE6EA898D62744D21" country="Costa Rica" latitude="10.802" longitude="-85.654" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="29130">
Area de
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
Guanacaste
</location>
; verbatimLocality: Vado Nisperal; verbatimElevation:
<elevation pageId="0" pageNumber="29130">10</elevation>
; verbatimLatitude: 10.802; verbatimLongitude: -85.654; verbatimCoordinateSystem: Decimal; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="29130" value="10.802">10.802</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="29130" value="-85.654">-85.654</geoCoordinate>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="29130">AJ Fleming</determinerName>
; dateIdentified: 2015; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="29130">Reared from Erebidae moth larva, Melipotisjanuaris</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="29130" value="1984-05-25">05/25/1984</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="29130">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="29130">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29130" type="description">
<paragraph pageId="0" pageNumber="29130">Description</paragraph>
<paragraph pageId="0" pageNumber="29130">
Male (Fig. 2). Length: 3 mm. Head (Fig. 2b): postpedicel orange basally, directly adjacent to pedicel; arista orange on apical half; ocellar setae well-developed, proclinate and divergent, arising behind anterior ocellus; fronto-orbital plate with two regular rows of short setulae outside of frontal setae; frontal setae appearing as continuous with parafacial setulae; two pairs of proclinate orbital setae, one pair of reclinate upper orbital setae. Thorax (Fig. 2a, c): three postsutural supra-alar setae, middle seta twice as long as outer two; two postsutural intra-alar setae; four postsutural dorsocentral setae; scutellum with one pair of basal scutellar setae, two pairs of lateral scutellars and one pair of weaker divergent apical scutellar setae; scutellum bearing one pair of weak but evident discal setae slightly wider than apical scutellars. Legs (Fig. 2c): entirely glabrous black, as in generic description. Wing (Fig. 2a): hyaline, very slightly darkening basally; basicosta dark brownr; wing vein R4+5 bearing 5-6 setulae extending 0.75
<normalizedToken originalValue="×">x</normalizedToken>
distance from node to crossvein r-m on dorsal surface; ventral surface bearing at most 0-2 setulae; calypters pale white translucent. Abdomen (Fig. 2a, c): ground color of abdomen glabrous maroon or reddish-black; very slight silver tomentum only visible when observed under different angles of light, appearing as a silver sheen on dorsal surface of tergites; complete row of marginal setae present on both T4 and T5; median discal setae absent on all tergites except row on T5.
</paragraph>
<paragraph pageId="0" pageNumber="29130">Female: unknown at this time, assumed to be similar to male as is the case with the type species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29130" type="diagnosis">
<paragraph pageId="0" pageNumber="29130">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="29130">
<taxonomicName class="Insecta" family="Tachinidae" genus="Trismegistomya" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trismegistomya jimoharai" order="Diptera" pageId="0" pageNumber="29130" phylum="Arthropoda" rank="species" species="jimoharai">Trismegistomya jimoharai</taxonomicName>
sp. n. is easily differentiated from its only congener, the type species
<taxonomicName lsidName="T. pumilis" pageId="0" pageNumber="29130" rank="species" species="pumilis">T. pumilis</taxonomicName>
Reinhard, by the following combination of traits: basal portion of postpedicel distinctly orange; arista orange apically; and four postsutural dorsocentral setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29130" type="etymology">
<paragraph pageId="0" pageNumber="29130">Etymology</paragraph>
<paragraph pageId="0" pageNumber="29130">
<taxonomicName class="Insecta" family="Tachinidae" genus="Trismegistomya" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trismegistomya jimoharai" order="Diptera" pageId="0" pageNumber="29130" phylum="Arthropoda" rank="species" species="jimoharai">Trismegistomya jimoharai</taxonomicName>
sp. n. is named in honor of Dr. James
<normalizedToken originalValue="OHara">O'Hara</normalizedToken>
of Ottawa, Canada in recognition of his many years of support to the curation, taxonomy and administrative logistics of the on-going effort to inventory the caterpillar-attacking species of
<taxonomicName family="Tachinidae" lsidName="" pageId="0" pageNumber="29130" rank="family">Tachinidae</taxonomicName>
of ACG and secure the residence of their voucher specimens in the Canadian National Collection.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29130" type="distribution">
<paragraph pageId="0" pageNumber="29130">Distribution</paragraph>
<paragraph pageId="0" pageNumber="29130">Costa Rica, ACG, Prov. Guanacaste, coastal margin dry forest, 10 m elevation.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29130" type="ecology">
<paragraph pageId="0" pageNumber="29130">Ecology</paragraph>
<paragraph pageId="0" pageNumber="29130">
<taxonomicName lsidName="T. jimoharai" pageId="0" pageNumber="29130" rank="species" species="jimoharai">T. jimoharai</taxonomicName>
has been reared two times, from only four larvae of
<taxonomicName class="Insecta" family="Noctuidae" genus="Melipotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melipotis januaris" order="Lepidoptera" pageId="0" pageNumber="29130" phylum="Arthropoda" rank="species" species="januaris">Melipotis januaris</taxonomicName>
(
<normalizedToken originalValue="Guenée">Guenee</normalizedToken>
, 1852) (
<taxonomicName lsidName="Noctuoidea" order="Lepidoptera" pageId="0" pageNumber="29130" rank="order">Lepidoptera</taxonomicName>
:
<taxonomicName family="Erebidae" lsidName="" pageId="0" pageNumber="29130" rank="family">Erebidae</taxonomicName>
) reared by the inventory to date and collected while feeding on
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pithecellobium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pithecellobium oblongum" order="Fabales" pageId="0" pageNumber="29130" phylum="Tracheophyta" rank="species" species="oblongum">Pithecellobium oblongum</taxonomicName>
Benth. (
<taxonomicName family="Fabaceae" lsidName="Noctuoidea" pageId="0" pageNumber="29130" rank="family">Fabaceae</taxonomicName>
) in ACG dry forest. It should be noted that
<taxonomicName class="Insecta" family="Tachinidae" genus="Trismegistomya" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trismegistomya" order="Diptera" pageId="0" pageNumber="29130" phylum="Arthropoda" rank="genus">Trismegistomya</taxonomicName>
has not been noted out of the other 808 rearings of
<taxonomicName class="Insecta" family="Noctuidae" genus="Melipotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melipotis" order="Lepidoptera" pageId="0" pageNumber="29130" phylum="Arthropoda" rank="genus">Melipotis</taxonomicName>
spp. reared by the inventory.
</paragraph>
</subSubSection>
</treatment>
</document>