283 lines
35 KiB
XML
283 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1191.114021" ID-Pensoft-Pub="1313-2970-1191-287" ID-Pensoft-UUID="5911E5DEBF3E5E249F39C0323B591149" ID-ZooBank="F412786C54D1408D8BC7D630A05F086F" ModsDocID="1313-2970-1191-287" checkinTime="1708002480185" checkinUser="pensoft" docAuthor="Li, Chun-Lin & Wang, Chuan-Chan" docDate="2024" docId="4203E0EB4F025F3D935674590CF2005F" docLanguage="en" docName="ZooKeys 1191: 287-305" docOrigin="ZooKeys 1191" docPubDate="2024-02-14" docSource="http://dx.doi.org/10.3897/zookeys.1191.114021" docTitle="Bolbelasmus yutangi Li & Wang 2024, sp. nov." docType="treatment" docUuid="A57879F9-F8A6-4D40-B922-DF3429C8BA4D" docUuidSource="ZooBank" docVersion="1" id="5911E5DEBF3E5E249F39C0323B591149" lastPageNumber="287" masterDocId="5911E5DEBF3E5E249F39C0323B591149" masterDocTitle="A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas" masterLastPageNumber="305" masterPageNumber="287" pageNumber="287" updateTime="1708002480185" updateUser="pensoft">
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<mods:mods id="3673BC6B5708362D89858662AEE46DDE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4533DDFA7B87E05D11926E99DCCC0650">
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<mods:title id="7F811AE7B5D853A36400434AE2BEAAD6">A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas</mods:title>
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</mods:titleInfo>
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<mods:name id="F2C0858AFBB86B582E25825A056833D5" type="personal">
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<mods:role id="C63C6B77BF1B5FC6F8EFD939D2BAAF0A">
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<mods:roleTerm id="03A6938CA7C3640D8EA4F81811C00A8C">Author</mods:roleTerm>
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<mods:namePart id="842888A1357A6B069D9DF91B64D66761">Li, Chun-Lin</mods:namePart>
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<mods:affiliation id="B7DE076211F368BF98E42E9B757D06DD">The Experimental Forest, National Taiwan University, Chushan, Nantou County 557, Taiwan</mods:affiliation>
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</mods:name>
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<mods:name id="25D048E5DBCFDB2C43F4C2B267534F11" type="personal">
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<mods:roleTerm id="74FD4A343BEE739645A79011D7A6ABE4">Author</mods:roleTerm>
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<mods:namePart id="5D5FCD83AAFD674FC5609C7B9D74985C">Wang, Chuan-Chan</mods:namePart>
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<mods:affiliation id="4233C5560862288C9EEFD4480E92B2DD">Department of Life Science, Fu Jen Catholic University, Hsinchuang, New Taipei City 24205, Taiwan</mods:affiliation>
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<mods:nameIdentifier id="6DBC54E1EB511BDFF037E1DFDC7D6213" type="email">gwennywang@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="A6734E489A08AAA481C0424B62625225">text</mods:typeOfResource>
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<mods:titleInfo id="763CA6924A8BD73A6A87B104104A9D56">
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<mods:title id="BE8A8F269532CBE78FBA9E81BF14469C">ZooKeys</mods:title>
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<mods:part id="BED70684ACC97B001F70719F7DED1912">
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<mods:date id="CC4DB86B6C4A81798DC7AEA3FE4C7DBC">2024</mods:date>
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<mods:number id="71B57AF1C0F7F91DEEDFB35F8DEBF1DC">2024-02-14</mods:number>
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<mods:number id="BE55F057DBB6BEF98578357D8FDA28C5">1191</mods:number>
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<mods:url id="257B1E908A176FCFFB5CC15207853232">http://dx.doi.org/10.3897/zookeys.1191.114021</mods:url>
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<mods:classification id="D2525C75D0765B07113B46E160F037D7">journal article</mods:classification>
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<mods:identifier id="2885B63FA149479BD5A55E19D21C8FBD" type="DOI">http://dx.doi.org/10.3897/zookeys.1191.114021</mods:identifier>
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<treatment id="4203E0EB4F025F3D935674590CF2005F" LSID="urn:lsid:zoobank.org:act:A57879F9-F8A6-4D40-B922-DF3429C8BA4D" httpUri="http://treatment.plazi.org/id/4203E0EB4F025F3D935674590CF2005F" lastPageNumber="287" pageId="0" pageNumber="287">
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<subSubSection id="EBFF248A34B8A632C29F9B56A8940F6A" pageId="0" pageNumber="287" type="nomenclature">
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<paragraph id="D2F4B1C2542C26C8BD04FC4651ECAC40" pageId="0" pageNumber="287">
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<taxonomicName id="9755AF31E97F94F1D0602B8D6E21E3B1" LSID="https://zoobank.org/A57879F9-F8A6-4D40-B922-DF3429C8BA4D" authority="Li & Wang" authorityName="Li & Wang" authorityYear="2024" class="Insecta" family="Geotrupidae" genus="Bolbelasmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolbelasmus yutangi" order="Coleoptera" pageId="0" pageNumber="287" phylum="Arthropoda" rank="species" species="yutangi" status="sp. nov.">Bolbelasmus yutangi Li & Wang</taxonomicName>
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<taxonomicNameLabel id="9A143181367991CCF4B51A0C560284DF" pageId="0" pageNumber="287">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="35CE929598B38DC7EB64672A937F7875" pageId="0" pageNumber="287" type="description">
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<paragraph id="96C91D1BC0900157A26D129C9F6166F1" pageId="0" pageNumber="287">
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<figureCitation id="3A418738B0A89E8A211F904977E588F5" captionStart="Figures 7–10" captionStartId="F2" captionText="Figures 7 - 10. Dorsal and lateral views of male Bolbelasmus spp. 7, 8 B. korshunovi 9, 10 B. yutangi sp. nov., holotype." figureDoi="10.3897/zookeys.1191.114021.figures7-10" httpUri="https://binary.pensoft.net/fig/984307" pageId="0" pageNumber="287">Figs 9</figureCitation>
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<figureCitation id="2BF8D47938CC06699C53DA57D09C02CC" captionStart="Figures 7–10" captionStartId="F2" captionText="Figures 7 - 10. Dorsal and lateral views of male Bolbelasmus spp. 7, 8 B. korshunovi 9, 10 B. yutangi sp. nov., holotype." figureDoi="10.3897/zookeys.1191.114021.figures7-10" httpUri="https://binary.pensoft.net/fig/984307" pageId="0" pageNumber="287">, 10</figureCitation>
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<figureCitation id="FC20C149D0960F0FCC56397D8FEC5C88" captionStart="Figures 11–16" captionStartId="F3" captionText="Figures 11 - 16. Scanning electron micrographs of heads of male Bolbelasmus spp. 11 B. concavisuturalis sp. nov. 12 B. chifengi sp. nov. 13 B. meridionalis 14 B. korshunovi 15 B. yutangi sp. nov. 16 B. coreanus. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1191.114021.figures11-16" httpUri="https://binary.pensoft.net/fig/984308" pageId="0" pageNumber="287">, 15</figureCitation>
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<figureCitation id="95B2699F0C4F0638C2B394F98237E4DB" captionStart="Figures 17–22" captionStartId="F4" captionText="Figures 17 - 22. Scanning electron micrographs of pronota of male Bolbelasmus spp. 17 B. concavisuturalis sp. nov. 18 B. chifengi sp. nov. 19 B. meridionalis 20 B. korshunovi 21 B. yutangi sp. nov. 22 B. coreanus. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1191.114021.figures17-22" httpUri="https://binary.pensoft.net/fig/984309" pageId="0" pageNumber="287">, 21</figureCitation>
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<figureCitation id="EC8E12BF7AC7131763B5F8FD94926522" captionStart="Figures 23–28" captionStartId="F5" captionText="Figures 23 - 28. Scanning electron micrographs of elytra of male Bolbelasmus spp. 23 B. concavisuturalis sp. nov. 24 B. chifengi sp. nov. 25 B. meridionalis 26 B. korshunovi 27 B. yutangi sp. nov. 28 B. coreanus. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1191.114021.figures23-28" httpUri="https://binary.pensoft.net/fig/984310" pageId="0" pageNumber="287">, 27</figureCitation>
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<figureCitation id="D5A323C56F08CDF8C77D314F0A0752C2" captionStart="Figures 29–38" captionStartId="F6" captionText="Figures 29 - 38. Male genitalia of Bolbelasmus spp. (29, 31, 33, 35, 37, dorsal views; 30, 32, 34, 36, 38, ventral views) 29, 30 B. concavisuturalis sp. nov. 31, 32 B. chifengi sp. nov. 33, 34 B. meridionalis 35, 36 B. korshunovi 37, 38 B. yutangi sp. nov. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1191.114021.figures29-38" httpUri="https://binary.pensoft.net/fig/984311" pageId="0" pageNumber="287">, 37</figureCitation>
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<figureCitation id="D5CAEEBA4C574F170F6224ABB0D468E1" captionStart="Figures 29–38" captionStartId="F6" captionText="Figures 29 - 38. Male genitalia of Bolbelasmus spp. (29, 31, 33, 35, 37, dorsal views; 30, 32, 34, 36, 38, ventral views) 29, 30 B. concavisuturalis sp. nov. 31, 32 B. chifengi sp. nov. 33, 34 B. meridionalis 35, 36 B. korshunovi 37, 38 B. yutangi sp. nov. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1191.114021.figures29-38" httpUri="https://binary.pensoft.net/fig/984311" pageId="0" pageNumber="287">, 38</figureCitation>
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<figureCitation id="82F8E7EACB0F5CD105668E13257F6B45" captionStart="Figures 39–43" captionStartId="F7" captionText="Figures 39 - 43. Male genitalia of Bolbelasmus spp., lateral views 39 B. concavisuturalis sp. nov. 40 B. chifengi sp. nov. 41 B. meridionalis 42 B. korshunovi 43 B. yutangi sp. nov. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1191.114021.figures39-43" httpUri="https://binary.pensoft.net/fig/984312" pageId="0" pageNumber="287">, 43</figureCitation>
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<figureCitation id="09E242362D347C92F5EC5E9975856267" captionStart="Figures 44–51" captionStartId="F8" captionText="Figures 44 - 51. Dorsal and lateral views of female Bolbelasmus spp. 44, 45 B. concavisuturalis sp. nov., paratype 46, 47 B. chifengi sp. nov., paratype 48, 49 B. meridionalis 50, 51 B. yutangi sp. nov., paratype." figureDoi="10.3897/zookeys.1191.114021.figures44-51" httpUri="https://binary.pensoft.net/fig/984313" pageId="0" pageNumber="287">, 50</figureCitation>
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<figureCitation id="E845289BA1C5ED3B5615274D0003D7FE" captionStart="Figures 44–51" captionStartId="F8" captionText="Figures 44 - 51. Dorsal and lateral views of female Bolbelasmus spp. 44, 45 B. concavisuturalis sp. nov., paratype 46, 47 B. chifengi sp. nov., paratype 48, 49 B. meridionalis 50, 51 B. yutangi sp. nov., paratype." figureDoi="10.3897/zookeys.1191.114021.figures44-51" httpUri="https://binary.pensoft.net/fig/984313" pageId="0" pageNumber="287">, 51</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="F253F62A261FFCD8F358684C46A47A8B" pageId="0" pageNumber="287" type="type material">
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<paragraph id="4816DF220D13BEDA9B4387387DA115BF" pageId="0" pageNumber="287">Type materials.</paragraph>
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<paragraph id="F5BE76FCF369E33F648A7C820DB9FCAC" pageId="0" pageNumber="287">
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<emphasis id="9819608151F3524D04B4FC40E750DA26" bold="true" pageId="0" pageNumber="287">
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<emphasis id="84CC3E490D7F3BD0E81CFC9E7A355AC7" italics="true" pageId="0" pageNumber="287">Holotype</emphasis>
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male.
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</emphasis>
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"Myanmar: Bago Region, Moe Yin Gyi, CCCC, 21.V.2017. leg. Y.-T. Wang." (glued on label, TARI).
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<emphasis id="A3F0AB3F5D337BF9141D010961AF4B6C" bold="true" pageId="0" pageNumber="287">
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<emphasis id="5F329BFC20319363358E7E9E58D39639" italics="true" pageId="0" pageNumber="287">Paratypes</emphasis>
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.
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</emphasis>
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1♀(TARI). data as the holotype.
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<emphasis id="C8A70B144386F2BD9FED0396D74BFF5D" bold="true" pageId="0" pageNumber="287">5</emphasis>
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♂♂ (TARI). "China: Yunnan, Wudian (武甸), 17.IX.2014. leg. Y.-T. Wang". 1♀ (CCLI) "China: Yunnan, Banggunjianshan (邦棍尖山), 18.IX.2015. leg. Y.-T. Wang". 6♂♂ (CCLI). "China: Yunnan, Bangdashan (邦達山), 01.IX.2015. leg. Y.-T. Wang". 1♂ (CCLI). "China: Yunnan, Ruili (瑞麗), 15.IX.2014. leg. Y.-T. Wang". 1♂ (NSMT). "Doi Saket, Chiang Mai, Thailand, 12-X-1989, Y. MANIT leg". 1♂ (NSMT). "Doi Mon Unggate, Samoeng Distr., Chiang Mai Prov., Thailand, 18-VII-1989, Y. MANIT leg".
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</paragraph>
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</subSubSection>
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<subSubSection id="61D83BBF29C406356BE288A4CB03A238" pageId="0" pageNumber="287" type="description">
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<paragraph id="B9D12F5C1DD23FA055F4787E3F6DF690" pageId="0" pageNumber="287">Description.</paragraph>
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<paragraph id="6F2523C0BAC259EE7897871867549500" pageId="0" pageNumber="287">
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<emphasis id="01401165DAC633767EF467346A6208C9" bold="true" pageId="0" pageNumber="287">Holotype male</emphasis>
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(Figs
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<figureCitation id="426F29C40F90E81D780BCA5C8BE1BCB1" captionStart="Figures 7–10" captionStartId="F2" captionText="Figures 7 - 10. Dorsal and lateral views of male Bolbelasmus spp. 7, 8 B. korshunovi 9, 10 B. yutangi sp. nov., holotype." figureDoi="10.3897/zookeys.1191.114021.figures7-10" httpUri="https://binary.pensoft.net/fig/984307" pageId="0" pageNumber="287">9</figureCitation>
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,
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<figureCitation id="2A704791DEB147700414DC66BC4E7797" captionStart="Figures 7–10" captionStartId="F2" captionText="Figures 7 - 10. Dorsal and lateral views of male Bolbelasmus spp. 7, 8 B. korshunovi 9, 10 B. yutangi sp. nov., holotype." figureDoi="10.3897/zookeys.1191.114021.figures7-10" httpUri="https://binary.pensoft.net/fig/984307" pageId="0" pageNumber="287">10</figureCitation>
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). Body length 9.3 mm; width across humeri 5.9 mm. Dorsum moderately shiny. Head, pronotum and scutellum dark brown with elytra reddish brown.
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<emphasis id="3F8F1B93FCCA5E26304099136BC9ECE8" bold="true" italics="true" pageId="0" pageNumber="287">Head</emphasis>
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(Fig.
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<figureCitation id="8F9C17B9E76397A80706A3FCA0A69A00" captionStart="Figures 11–16" captionStartId="F3" captionText="Figures 11 - 16. Scanning electron micrographs of heads of male Bolbelasmus spp. 11 B. concavisuturalis sp. nov. 12 B. chifengi sp. nov. 13 B. meridionalis 14 B. korshunovi 15 B. yutangi sp. nov. 16 B. coreanus. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1191.114021.figures11-16" httpUri="https://binary.pensoft.net/fig/984308" pageId="0" pageNumber="287">15</figureCitation>
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): labrum with anterior margin crenulate, disc transversally rugose. Clypeus trapezoidal, surface densely rugopunctate; protrusion at basal angle moderately developed. Clypeofrontal suture well defined, slightly curved in front of frontal tubercle. Frons with surface moderately punctate, punctures coarse, frontal tubercle vertically located at center of disc, triangular when viewed laterally. Eye prominent, canthus simple, not exceeding eye.
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<emphasis id="D5F52FD103BE16A91F3C47DBB75CE523" bold="true" italics="true" pageId="0" pageNumber="287">Thorax</emphasis>
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: pronotum (Fig.
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<figureCitation id="F4DF03CBD3FAF0C3490F50069C959A30" captionStart="Figures 17–22" captionStartId="F4" captionText="Figures 17 - 22. Scanning electron micrographs of pronota of male Bolbelasmus spp. 17 B. concavisuturalis sp. nov. 18 B. chifengi sp. nov. 19 B. meridionalis 20 B. korshunovi 21 B. yutangi sp. nov. 22 B. coreanus. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1191.114021.figures17-22" httpUri="https://binary.pensoft.net/fig/984309" pageId="0" pageNumber="287">21</figureCitation>
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) quadrituberculate, tubercles placed in a line, lateral tubercle greatly reduced in size; anterior face of median tubercles almost perpendicular to plane of pronotum; primary punctures coarse, dense on disc with small impunctate area in front of middle of posterior margin, punctures between lateral margins of pronotum and fovea bigger and denser, scattered coarse punctures distributed along posterior margin with seven punctures in front of scutellum, secondary punctures tiny, evenly scattered on surface of pronotum; frontal and lateral margins beaded, posterior margin beaded only in front of elytral humeri. Scutellum elongate, fine punctures sparsely distributed on surface.
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<emphasis id="3744CBC44534EA2B33384E2EFCE38BB5" bold="true" italics="true" pageId="0" pageNumber="287">Elytron</emphasis>
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(Fig.
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<figureCitation id="09A49BD7E5C5E55B2E2E8E592140763A" captionStart="Figures 23–28" captionStartId="F5" captionText="Figures 23 - 28. Scanning electron micrographs of elytra of male Bolbelasmus spp. 23 B. concavisuturalis sp. nov. 24 B. chifengi sp. nov. 25 B. meridionalis 26 B. korshunovi 27 B. yutangi sp. nov. 28 B. coreanus. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1191.114021.figures23-28" httpUri="https://binary.pensoft.net/fig/984310" pageId="0" pageNumber="287">27</figureCitation>
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): elytral striae shallowly impressed, punctures mostly spaced by 1-3 times diameters of punctures. Intervals slightly convex, including sutural interval, surface with scattered secondary punctures. Male genitalia. Figs
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<figureCitation id="481419615603613D854E7A1F71E0826C" captionStart="Figures 29–38" captionStartId="F6" captionText="Figures 29 - 38. Male genitalia of Bolbelasmus spp. (29, 31, 33, 35, 37, dorsal views; 30, 32, 34, 36, 38, ventral views) 29, 30 B. concavisuturalis sp. nov. 31, 32 B. chifengi sp. nov. 33, 34 B. meridionalis 35, 36 B. korshunovi 37, 38 B. yutangi sp. nov. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1191.114021.figures29-38" httpUri="https://binary.pensoft.net/fig/984311" pageId="0" pageNumber="287">37</figureCitation>
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,
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<figureCitation id="09E6FA6830FFA6576ED72F436394CFEE" captionStart="Figures 29–38" captionStartId="F6" captionText="Figures 29 - 38. Male genitalia of Bolbelasmus spp. (29, 31, 33, 35, 37, dorsal views; 30, 32, 34, 36, 38, ventral views) 29, 30 B. concavisuturalis sp. nov. 31, 32 B. chifengi sp. nov. 33, 34 B. meridionalis 35, 36 B. korshunovi 37, 38 B. yutangi sp. nov. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1191.114021.figures29-38" httpUri="https://binary.pensoft.net/fig/984311" pageId="0" pageNumber="287">38</figureCitation>
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,
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<figureCitation id="C67C5B934EFA13DBC29BC13E76E50A8B" captionStart="Figures 39–43" captionStartId="F7" captionText="Figures 39 - 43. Male genitalia of Bolbelasmus spp., lateral views 39 B. concavisuturalis sp. nov. 40 B. chifengi sp. nov. 41 B. meridionalis 42 B. korshunovi 43 B. yutangi sp. nov. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1191.114021.figures39-43" httpUri="https://binary.pensoft.net/fig/984312" pageId="0" pageNumber="287">43</figureCitation>
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.
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</paragraph>
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<caption id="1BB41F686F8654A1EBF3733DFE876370" doi="10.3897/zookeys.1191.114021.figures29-38" httpUri="https://binary.pensoft.net/fig/984311" pageId="0" pageNumber="287" start="Figures 29–38" startId="F6">
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<paragraph id="1BFD16769915B787CB1AA35A666427D2" pageId="0" pageNumber="287">
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<emphasis id="215C5322E05AB36754D1D774E7D14920" bold="true" pageId="0" pageNumber="287">Figures 29-38.</emphasis>
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Male genitalia of
|
||
<taxonomicName id="0DAB9EB49FBD64134BB8BDB9EB2A427D" authorityName="Boucomont" authorityYear="1910" class="Insecta" family="Geotrupidae" genus="Bolbelasmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolbelasmus" order="Coleoptera" pageId="0" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1906F9DDAD53C91DDEB437317901F507" italics="true" pageId="0" pageNumber="287">Bolbelasmus</emphasis>
|
||
</taxonomicName>
|
||
spp. (29, 31, 33, 35, 37, dorsal views; 30, 32, 34, 36, 38, ventral views)
|
||
<emphasis id="D09FE1FFA105254992ABA219AE82E2E3" bold="true" pageId="0" pageNumber="287">29, 30</emphasis>
|
||
<taxonomicName id="A5F1451185D80DE4495ED5637551813C" family="Geotrupidae" kingdom="Animalia" lsidName="B. concavisuturalis" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="concavisuturalis">
|
||
<emphasis id="E4879E7964D7262B5EDCDCB11F71CC5A" italics="true" pageId="0" pageNumber="287">B. concavisuturalis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="BF6CEA59D7ACA48944020686C9797DC0" bold="true" pageId="0" pageNumber="287">31, 32</emphasis>
|
||
<taxonomicName id="70E88EC04EC3AD0E609CCFEB2005F85E" family="Geotrupidae" kingdom="Animalia" lsidName="B. chifengi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="chifengi">
|
||
<emphasis id="65AEE067C3682D3195AE8C9D50D65BC5" italics="true" pageId="0" pageNumber="287">B. chifengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="FDA5B402D165B42D92F6DDB964208BC4" bold="true" pageId="0" pageNumber="287">33, 34</emphasis>
|
||
<taxonomicName id="AA8F062758DC68F725248EC50D0A60F5" family="Geotrupidae" kingdom="Animalia" lsidName="B. meridionalis" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="meridionalis">
|
||
<emphasis id="C041D855388B00CA07CA4FD9DDF1391B" italics="true" pageId="0" pageNumber="287">B. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="430F902882220164E2A7AE1BEF29903B" bold="true" pageId="0" pageNumber="287">35, 36</emphasis>
|
||
<taxonomicName id="5AE6D087AD30D0A39720F20A56865174" family="Geotrupidae" kingdom="Animalia" lsidName="B. korshunovi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="korshunovi">
|
||
<emphasis id="2C921195158EB5086F7D7607660E0717" italics="true" pageId="0" pageNumber="287">B. korshunovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7B8192084AC27FB85C215839067E064F" bold="true" pageId="0" pageNumber="287">37, 38</emphasis>
|
||
<taxonomicName id="3C4A685ABCA480D6FDE2A8DEBD051619" family="Geotrupidae" kingdom="Animalia" lsidName="B. yutangi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="yutangi">
|
||
<emphasis id="A2446BDBF24C67969355E859013CE7C2" italics="true" pageId="0" pageNumber="287">B. yutangi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Scale bar: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FD94849A72DEFF5E710AE281BAA61572" doi="10.3897/zookeys.1191.114021.figures39-43" httpUri="https://binary.pensoft.net/fig/984312" pageId="0" pageNumber="287" start="Figures 39–43" startId="F7">
|
||
<paragraph id="4379708FB082292CE76B71D7D7E8899F" pageId="0" pageNumber="287">
|
||
<emphasis id="E18FD447011C60F7B56CCA4D5C612123" bold="true" pageId="0" pageNumber="287">Figures 39-43.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="2AECF31969E9FDB20EBB3A6D91A52A7C" authorityName="Boucomont" authorityYear="1910" class="Insecta" family="Geotrupidae" genus="Bolbelasmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolbelasmus" order="Coleoptera" pageId="0" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75E88A3C4FC11D0262930C7344B3C26D" italics="true" pageId="0" pageNumber="287">Bolbelasmus</emphasis>
|
||
</taxonomicName>
|
||
spp., lateral views
|
||
<emphasis id="1378643DFBD4D45D0686F443652B10EB" bold="true" pageId="0" pageNumber="287">39</emphasis>
|
||
<taxonomicName id="1A8EDFFEF4A2920DC1A8DE08C483EA74" family="Geotrupidae" kingdom="Animalia" lsidName="B. concavisuturalis" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="concavisuturalis">
|
||
<emphasis id="3C90A927D608DC6652D546826FE3EDF6" italics="true" pageId="0" pageNumber="287">B. concavisuturalis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="7F0973A1D7F239548F1002FE51BB1671" bold="true" pageId="0" pageNumber="287">40</emphasis>
|
||
<taxonomicName id="FB5A3F755E871B6F87C25CE758FF38EC" family="Geotrupidae" kingdom="Animalia" lsidName="B. chifengi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="chifengi">
|
||
<emphasis id="80D553ACBEA4E41FFABEA4135475BF5A" italics="true" pageId="0" pageNumber="287">B. chifengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="F8C3E75962934C3334D57FF843466AC0" bold="true" pageId="0" pageNumber="287">41</emphasis>
|
||
<taxonomicName id="D1A820EB9C829FE249706AB0D1DA9BD3" family="Geotrupidae" kingdom="Animalia" lsidName="B. meridionalis" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="meridionalis">
|
||
<emphasis id="EC213785282E17CDF755625013DA27ED" italics="true" pageId="0" pageNumber="287">B. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B6717F8BC21EA557893C8D503843383D" bold="true" pageId="0" pageNumber="287">42</emphasis>
|
||
<taxonomicName id="C39659758A055F1B56323596EF5FDC4D" family="Geotrupidae" kingdom="Animalia" lsidName="B. korshunovi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="korshunovi">
|
||
<emphasis id="3506CAFF76DFD0B7F59E46C248511B6D" italics="true" pageId="0" pageNumber="287">B. korshunovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A8F2B4859E36790890D158D1461D23CD" bold="true" pageId="0" pageNumber="287">43</emphasis>
|
||
<taxonomicName id="70D6C7C82390393E8EEAC262C5628EBC" family="Geotrupidae" kingdom="Animalia" lsidName="B. yutangi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="yutangi">
|
||
<emphasis id="3DD5F78BD47CEC1CD3FAD15614145E88" italics="true" pageId="0" pageNumber="287">B. yutangi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Scale bar: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6B182951059181D9D1B48910CF11782B" pageId="0" pageNumber="287">
|
||
<emphasis id="308CB0A3C7A81DF396270FE03C8B8A37" bold="true" pageId="0" pageNumber="287">Female</emphasis>
|
||
(Figs
|
||
<figureCitation id="888A0B217F86F36130FCFA1831645CBA" captionStart="Figures 44–51" captionStartId="F8" captionText="Figures 44 - 51. Dorsal and lateral views of female Bolbelasmus spp. 44, 45 B. concavisuturalis sp. nov., paratype 46, 47 B. chifengi sp. nov., paratype 48, 49 B. meridionalis 50, 51 B. yutangi sp. nov., paratype." figureDoi="10.3897/zookeys.1191.114021.figures44-51" httpUri="https://binary.pensoft.net/fig/984313" pageId="0" pageNumber="287">50</figureCitation>
|
||
,
|
||
<figureCitation id="49FBA9842E0E91F912115BEF62A33A6D" captionStart="Figures 44–51" captionStartId="F8" captionText="Figures 44 - 51. Dorsal and lateral views of female Bolbelasmus spp. 44, 45 B. concavisuturalis sp. nov., paratype 46, 47 B. chifengi sp. nov., paratype 48, 49 B. meridionalis 50, 51 B. yutangi sp. nov., paratype." figureDoi="10.3897/zookeys.1191.114021.figures44-51" httpUri="https://binary.pensoft.net/fig/984313" pageId="0" pageNumber="287">51</figureCitation>
|
||
). Body length 8.4-8.7 mm; width across humeri 5.1-5.3 mm. Similar to male with minor differences in the form of strongly wrinkled surface of clypeus, transverse frontal carina trilobed, central lobe more prominent than lateral lobes, punctures on frons and vertex rugose, transverse pronotal carina feebly bilobed, with lobes broadly developed to reduced, bigger punctures on pronotal disc denser than those of males and scutellum with 1 or 2 bigger punctures.
|
||
</paragraph>
|
||
<caption id="5B5686D5679A95C29DC8368ACF40C5AB" doi="10.3897/zookeys.1191.114021.figures44-51" httpUri="https://binary.pensoft.net/fig/984313" pageId="0" pageNumber="287" start="Figures 44–51" startId="F8">
|
||
<paragraph id="4206EF94F1F38E601DC07A21C3C96473" pageId="0" pageNumber="287">
|
||
<emphasis id="5694E3FD13754816E53D0EE535397EEC" bold="true" pageId="0" pageNumber="287">Figures 44-51.</emphasis>
|
||
Dorsal and lateral views of female
|
||
<taxonomicName id="B687E9553798D7107F3648A74AA19C5C" authorityName="Boucomont" authorityYear="1910" class="Insecta" family="Geotrupidae" genus="Bolbelasmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolbelasmus" order="Coleoptera" pageId="0" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FC422F56ADD4978C91D48578E7350B7F" italics="true" pageId="0" pageNumber="287">Bolbelasmus</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="A7FDB3671E541B20C4258FBDDA051BB5" bold="true" pageId="0" pageNumber="287">44, 45</emphasis>
|
||
<taxonomicName id="4280DA5DE956B1C7C8C835B8039B3872" family="Geotrupidae" kingdom="Animalia" lsidName="B. concavisuturalis" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="concavisuturalis">
|
||
<emphasis id="D56B9F00BEC3CD567C4A32E80AB01310" italics="true" pageId="0" pageNumber="287">B. concavisuturalis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype
|
||
<emphasis id="0C35B3CC8518ADF8EF78F458A4DD907F" bold="true" pageId="0" pageNumber="287">46, 47</emphasis>
|
||
<taxonomicName id="214BB0E1962597276F9D3746E8C9A4EA" family="Geotrupidae" kingdom="Animalia" lsidName="B. chifengi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="chifengi">
|
||
<emphasis id="F79A86DF6B701B0604A4DA9DAA10FF93" italics="true" pageId="0" pageNumber="287">B. chifengi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype
|
||
<emphasis id="9A02640B012929213955090FA95BA2BD" bold="true" pageId="0" pageNumber="287">48, 49</emphasis>
|
||
<taxonomicName id="27BE58B9803616EBF413D06A64B11157" family="Geotrupidae" kingdom="Animalia" lsidName="B. meridionalis" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="meridionalis">
|
||
<emphasis id="CF8789EA556EAEEC2F16C7CAF5E48C96" italics="true" pageId="0" pageNumber="287">B. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="33F29F90B13CF237E4BC02B07C08204E" bold="true" pageId="0" pageNumber="287">50, 51</emphasis>
|
||
<taxonomicName id="53A0B948ADF2F1266F4CFB09C57E1766" family="Geotrupidae" kingdom="Animalia" lsidName="B. yutangi" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="yutangi">
|
||
<emphasis id="D00301E76A39B2AC496BF1AA75564765" italics="true" pageId="0" pageNumber="287">B. yutangi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4754618BB15CDA77C4332C79F222A42E" pageId="0" pageNumber="287">
|
||
<emphasis id="6CAF5877E950E1222A0C91262F211A51" bold="true" pageId="0" pageNumber="287">Variation in male.</emphasis>
|
||
Male paratypes differ from the holotype in the following respects: smaller body size, 6.6 mm in body length and 5.2 mm in width across humeri, frontal tubercle less developed and not in junction of clypeofrontal suture, pronotal tubercle feebly convex, reduced, and number of coarse punctures along pronotal posterior margin variable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5E66DB22E2BA317499F2BDF88CB829CD" pageId="0" pageNumber="287" type="diagnosis">
|
||
<paragraph id="17D92D1B9B6F153DBE1461B27058F3B2" pageId="0" pageNumber="287">Diagnosis.</paragraph>
|
||
<paragraph id="A25249456EB00F9278A6F304E6EB93A4" pageId="0" pageNumber="287">
|
||
<taxonomicName id="F2909FF7E697CD390DB46C366DA67620" authorityName="Li & Wang" authorityYear="2024" class="Insecta" family="Geotrupidae" genus="Bolbelasmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolbelasmus yutangi" order="Coleoptera" pageId="0" pageNumber="287" phylum="Arthropoda" rank="species" species="yutangi">
|
||
<emphasis id="EE01DF8200E321E58A4A7AD0149D6D21" italics="true" pageId="0" pageNumber="287">Bolbelasmus yutangi</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="DD25A17D20642D672572D0E92AD245EA" family="Geotrupidae" kingdom="Animalia" lsidName="B. nativus" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="nativus">
|
||
<emphasis id="8CD7224313CCDB97935BC73179ED020A" italics="true" pageId="0" pageNumber="287">B. nativus</emphasis>
|
||
</taxonomicName>
|
||
in sharing the frontal tubercle location at the center of the frons as well as having smaller parameres. It can be distinguished from the latter by the weakly convex basal angle of the clypeus (distinctly bulging in
|
||
<taxonomicName id="3B528F886261CDA570788D19BA508678" family="Geotrupidae" kingdom="Animalia" lsidName="B. nativus" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="nativus">
|
||
<emphasis id="827BFF92321D2028FB2BCA32247A1B71" italics="true" pageId="0" pageNumber="287">B. nativus</emphasis>
|
||
</taxonomicName>
|
||
), primary punctures densely distributed on the disc (almost impunctate along the middle of disc in
|
||
<taxonomicName id="9417EB2198CE958F989BA7D3EF139DBD" family="Geotrupidae" kingdom="Animalia" lsidName="B. nativus" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="nativus">
|
||
<emphasis id="B7C0B3A0492B363D4BE4FA7DBA891B77" italics="true" pageId="0" pageNumber="287">B. nativus</emphasis>
|
||
</taxonomicName>
|
||
), sutural interval convex, similar to discal intervals (distinctly more convex than discal intervals in
|
||
<taxonomicName id="3B075F23DF7A33E1EF22966115A50E1B" family="Geotrupidae" kingdom="Animalia" lsidName="B. nativus" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="nativus">
|
||
<emphasis id="D9845F4FA0DDA1197E8C547083B0E081" italics="true" pageId="0" pageNumber="287">B. nativus</emphasis>
|
||
</taxonomicName>
|
||
) and parameres with the tips tapered (parameres broader at tips in
|
||
<taxonomicName id="3C0B6DD2C85856A112A2E982D1D8C72D" family="Geotrupidae" kingdom="Animalia" lsidName="B. nativus" order="Coleoptera" pageId="0" pageNumber="287" rank="species" species="nativus">
|
||
<emphasis id="D633448B6A0265868EA9C21A4A472EC2" italics="true" pageId="0" pageNumber="287">B. nativus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="444737A6DD9F0BBF86E20836DDDAA6C9" pageId="0" pageNumber="287" type="distribution">
|
||
<paragraph id="5024F377530F404B2DE21FABBD75A393" pageId="0" pageNumber="287">Distribution.</paragraph>
|
||
<paragraph id="EB6CC0F435E78837053920D64B47AAAE" pageId="0" pageNumber="287">
|
||
Eastern Myanmar, northern Thailand and western Yunnan, China (Fig.
|
||
<figureCitation id="61517599BCAA3B15C7ECEF4905439FC5" captionStart="Figure 52" captionStartId="F9" captionText="Figure 52. Distribution map of the eastern and southeastern Asian Bolbelasmus species. Question marks indicate doubtful distributional records." figureDoi="10.3897/zookeys.1191.114021.figure52" httpUri="https://binary.pensoft.net/fig/984314" pageId="0" pageNumber="287">52</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="5B2EFDF6158C39F74FBF1C3747F093B9" doi="10.3897/zookeys.1191.114021.figure52" httpUri="https://binary.pensoft.net/fig/984314" pageId="0" pageNumber="287" start="Figure 52" startId="F9">
|
||
<paragraph id="31CEC5B56D45DE5C86A4B824B69EC905" pageId="0" pageNumber="287">
|
||
<emphasis id="E9C20086A3187D52D8B68AEA73196CB0" bold="true" pageId="0" pageNumber="287">Figure 52.</emphasis>
|
||
Distribution map of the eastern and southeastern Asian
|
||
<taxonomicName id="DCE68FE1845FE63940BFA6E139DB8653" authorityName="Boucomont" authorityYear="1910" class="Insecta" family="Geotrupidae" genus="Bolbelasmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolbelasmus" order="Coleoptera" pageId="0" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EA4CF7D2840433847FE9B6973C8B76A" italics="true" pageId="0" pageNumber="287">Bolbelasmus</emphasis>
|
||
</taxonomicName>
|
||
species. Question marks indicate doubtful distributional records.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="6D8921331BB9FA909E8B78677721E34B" pageId="0" pageNumber="287" type="etymology">
|
||
<paragraph id="C1C6A36DAC1AC5840ADF9D90AAC4C02A" pageId="0" pageNumber="287">Etymology.</paragraph>
|
||
<paragraph id="D5445BCC5A882364730C69051F69DD88" pageId="0" pageNumber="287">
|
||
<taxonomicName id="0BE5D78CF19FA366F449CFA4880B6D50" authorityName="Li & Wang" authorityYear="2024" class="Insecta" family="Geotrupidae" genus="Bolbelasmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolbelasmus yutangi" order="Coleoptera" pageId="0" pageNumber="287" phylum="Arthropoda" rank="species" species="yutangi">
|
||
<emphasis id="6D4C31D3D35B5ACC156A3CE928D199B4" italics="true" pageId="0" pageNumber="287">Bolbelasmus yutangi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is named after Mr Yu-tang Wang, a beetle enthusiast of Taiwan, who collected most of the material used in this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FC2AA97D05094EB7C048576BF5B8F480" pageId="0" pageNumber="287" type="species with doubtful locality record">
|
||
<paragraph id="422B92728C3E3167E7CE3C1ECEABFE7A" pageId="0" pageNumber="287">Species with doubtful locality record</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |