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<document id="143C3ED2D599DF1E87159F8EE946B303" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E45FFC994FD4ABCED99E9D3" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) griseus Panzer 1796" docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="9" updateTime="1698562432667" updateUser="plazi">
<mods:mods id="EFF09D346D7690321D072A6D064FCE35" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0C5CD10A28943042B970E516220712E3">
<mods:title id="65A786D285A6C014EB20F41963747F2F">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="E71FF7AA21781DDB96968BBC40CC0914" type="personal">
<mods:role id="3111B81F3C0035916A55E8D374302D89">
<mods:roleTerm id="C5BD2AB79DEA83373E480735AF238C10">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D70EDF942976D435486953520922DD1E">Kataev, Boris M.</mods:namePart>
</mods:name>
<mods:name id="BA0534CF3E83E947505702B196DF49DE" type="personal">
<mods:role id="F7FC9D775F1F96A7F281B551F113AC1F">
<mods:roleTerm id="112B0F4B2EF9447293138037B2EF8592">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47213995FD8B3D64D7D087A621918D8F">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:typeOfResource id="310C51CD87DB0E735F79DDBA3F098D0C">text</mods:typeOfResource>
<mods:relatedItem id="6F0B74DB842655F1F47C708284E1E086" type="host">
<mods:titleInfo id="8CE5D629D6DEC529A80870BD63215FFA">
<mods:title id="07DC37D605B8C3A119B2C0D06A499D38">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="767D71BA9AC81783AF58A9ADB7C16125">
<mods:date id="04961F16AEE932D1A7EF49B6D5A910C0">2015</mods:date>
<mods:detail id="D0C7E513BFDE495BC35AEFD1C0262AF9" type="volume">
<mods:number id="3C4E26744BEE1FADB084DB402B6A60AE">3920</mods:number>
</mods:detail>
<mods:detail id="E8F5631A69F897609255A6EA7740E05B" type="issue">
<mods:number id="F45DF43FBA345980B342FD2DF5C04F59">1</mods:number>
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<mods:start id="9083854973342B786D64B2DAC043F9F7">1</mods:start>
<mods:end id="26941CBE75B5366415A1F25D2D5F965C">39</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CDF63ADC9D4994006EA85B5C46D68976">journal article</mods:classification>
<mods:identifier id="D9A4551F5158D449D0AE49EB790ADE1C" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
<mods:identifier id="2E718B8F4FA43392699CFEB674EC0F1B" type="GBIF-Dataset">7289c86d-0ed1-4e93-b7b3-bc596042d854</mods:identifier>
<mods:identifier id="F52453A6A922001202075782FA71B726" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7830D803F655DC1074259932F21305E6" type="Zenodo-Dep">287886</mods:identifier>
<mods:identifier id="317B29FCB6A523B675B7D8A646837E2F" type="ZooBank">025F9918-F9F6-4049-B7CD-5AC632094F5E</mods:identifier>
</mods:mods>
<treatment id="039787812E45FFC994FD4ABCED99E9D3" ID-DOI="http://doi.org/10.5281/zenodo.6114246" ID-GBIF-Taxon="127671659" ID-Zenodo-Dep="6114246" LSID="urn:lsid:plazi:treatment:039787812E45FFC994FD4ABCED99E9D3" httpUri="http://treatment.plazi.org/id/039787812E45FFC994FD4ABCED99E9D3" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="C324651C2E45FFC994FD4ABCE801EFC9" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B8136972E45FFC994FD4ABCEBDDEFA8" blockId="8.[151,762,151,211]" box="[151,762,151,178]" pageId="8" pageNumber="9">
<heading id="D0C981FB2E45FFC994FD4ABCEBDDEFA8" bold="true" box="[151,762,151,178]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="4C3E4D142E45FFC994FD4ABCEBDDEFA8" ID-CoL="3JSTS" authority="Panzer, 1796" authorityName="Panzer" authorityYear="1796" box="[151,762,151,178]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="griseus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E45FFC994FD4ABCEBDDEFA8" bold="true" box="[151,762,151,178]" pageId="8" pageNumber="9">
<emphasis id="B94AEA852E45FFC994FD4ABCE82EEFAB" bold="true" box="[151,265,151,177]" italics="true" pageId="8" pageNumber="9">Harpalus</emphasis>
(
<emphasis id="B94AEA852E45FFC995704ABCE8FDEFAB" bold="true" box="[282,474,151,177]" italics="true" pageId="8" pageNumber="9">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E45FFC995814ABCEB66EFAB" bold="true" box="[491,577,151,177]" italics="true" pageId="8" pageNumber="9">griseus</emphasis>
(
<bibRefCitation id="EFAF4B662E45FFC9963A4AB3EBD5EFA8" author="Panzer" box="[592,754,152,178]" pageId="8" pageNumber="9" refString="Panzer, G. W. F. (1796) Faunae Insectorum Germaniae initia oder Deutschlands Insecten gesammelt und herausgegeben. 38. Felseckerscken Buchhandlung, Nurnberg, 24 pp. + 24 pls." type="book" year="1796">Panzer, 1796</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E45FFC994FD4A91E801EFC9" blockId="8.[151,762,151,211]" box="[151,294,186,211]" pageId="8" pageNumber="9">
(
<figureCitation id="13052A122E45FFC994F54A91E838EFC9" box="[159,287,186,211]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,263,1483,1506]" captionTargetBox="[228,1353,642,1453]" captionTargetId="figure@9.[220,1366,628,1463]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Harpalus, median lobe of aedeagus. 15 – 16, H. griseus (Liaoning); 17 – 18, H. jureceki (Heilongjiang). 15, 17, view from the left side; 16, 18, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287888/files/figure.png" pageId="8" pageNumber="9">Figs 15–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E45FFC994FD4B28ECACEE6E" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8B8136972E45FFC994FD4B28EAB8EE02" blockId="8.[151,1419,258,372]" box="[151,927,258,280]" pageId="8" pageNumber="9">
<taxonomicName id="4C3E4D142E45FFC994FD4B28E8ECEE02" ID-CoL="68YN2" authority="Panzer, 1796" authorityName="Panzer" authorityYear="1796" box="[151,459,258,280]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="B94AEA852E45FFC994FD4B28E81AEE02" box="[151,317,259,280]" italics="true" pageId="8" pageNumber="9">Carabus griseus</emphasis>
<bibRefCitation id="EFAF4B662E45FFC9952E4B28E8ECEE02" author="Panzer" box="[324,459,258,280]" pageId="8" pageNumber="9" refString="Panzer, G. W. F. (1796) Faunae Insectorum Germaniae initia oder Deutschlands Insecten gesammelt und herausgegeben. 38. Felseckerscken Buchhandlung, Nurnberg, 24 pp. + 24 pls." type="book" year="1796">Panzer, 1796</bibRefCitation>
</taxonomicName>
: no 1.
<typeStatus id="548588352E45FFC996674B28EB18EE02" box="[525,575,259,280]" pageId="8" pageNumber="9">Type</typeStatus>
locality: &quot;Brunsvigiae&quot;,
<collectingCountry id="F32976072E45FFC997504B29EABDEE02" box="[826,922,258,280]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
.
</paragraph>
<paragraph id="8B8136972E45FFC994FD4B0AEAE7EE2D" blockId="8.[151,1419,258,372]" box="[151,960,289,311]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB2E11B92E45FFC994FD4B0AEAE7EE2D" box="[151,960,289,311]" pageId="8" pageNumber="9">
<taxonomicName id="4C3E4D142E45FFC994FD4B0AEB31EE2D" ID-CoL="8L5MN" authority="Marsham, 1802: 436" authorityName="Marsham" authorityPageNumber="436" authorityYear="1802" box="[151,534,289,311]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B94AEA852E45FFC994FD4B0AE81AEE2C" box="[151,317,289,310]" italics="true" pageId="8" pageNumber="9">Carabus bicolor</emphasis>
<treatmentCitation id="0A9F10862E45FFC9952E4B0AEB31EE2D" author="Marsham" box="[324,534,289,311]" page="436" pageId="8" pageNumber="9" year="1802">
<bibRefCitation id="EFAF4B662E45FFC9952E4B0AE8C1EE2D" author="Marsham" box="[324,486,289,311]" pageId="8" pageNumber="9" refString="Marsham, Th. (1802) Entomologia Britannica, sistens Insecta Britanniae indigena, secundum, methodum Linnaeanam disposita. Tomus I. Coleoptera. White, Londini, xxxi + 547 + 1 pp." type="book" year="1802">Marsham, 1802</bibRefCitation>
: 436
</treatmentCitation>
</taxonomicName>
(non Drury, 1770).
<typeStatus id="548588352E45FFC9968A4B09EA35EE2D" box="[736,786,290,311]" pageId="8" pageNumber="9">Type</typeStatus>
locality:
<collectingCountry id="F32976072E45FFC997184B0AEA9BEE2D" box="[882,956,289,311]" name="United Kingdom" pageId="8" pageNumber="9">Britain</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E45FFC994FD4B6BEAACEE4C" blockId="8.[151,1419,258,372]" box="[151,907,320,342]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB2E11B92E45FFC994FD4B6BEAACEE4C" box="[151,907,320,342]" pageId="8" pageNumber="9">
<taxonomicName id="4C3E4D142E45FFC994FD4B6BE8EAEE4C" ID-CoL="8KWCG" authority="Reichei Desbrochers" authorityName="Reichei Desbrochers" box="[151,461,320,342]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E45FFC994FD4B6BE86EEE4F" box="[151,329,320,341]" italics="true" pageId="8" pageNumber="9">Harpalus Reichei</emphasis>
Desbrochers
</taxonomicName>
des
<treatmentCitation id="0A9F10862E45FFC995964B6AEB85EE4C" author="Loges" box="[508,674,320,342]" page="42" pageId="8" pageNumber="9" year="1867">
<bibRefCitation id="EFAF4B662E45FFC995964B6AEB5BEE4C" author="Loges" box="[508,636,320,342]" pageId="8" pageNumber="9" refString="Desbrochers des Loges (1867) Notice sur l'entomologie du Bourbonnais. Suivie de la description de trois especes nouvelles. Assises scientifiques du Bourbonnais, 44 pp." type="book" year="1867">Loges, 1867</bibRefCitation>
: 42
</treatmentCitation>
.
<typeStatus id="548588352E45FFC996C64B6AEBF9EE4C" box="[684,734,321,342]" pageId="8" pageNumber="9">Type</typeStatus>
locality:
<collectingCountry id="F32976072E45FFC997544B6AEAA1EE4C" box="[830,902,321,342]" name="France" pageId="8" pageNumber="9">France</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E45FFC994FD4B74ECACEE6E" blockId="8.[151,1419,258,372]" box="[151,1419,350,372]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB2E11B92E45FFC994FD4B74ECACEE6E" box="[151,1419,350,372]" pageId="8" pageNumber="9">
<taxonomicName id="4C3E4D142E45FFC994FD4B74EA4CEE6E" ID-CoL="8LG2C" authority="Huang, Hu &amp; Sun, 1994: 263" authorityName="Huang, Hu &amp; Sun" authorityPageNumber="263" authorityYear="1994" box="[151,875,350,372]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xinjiangensis" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E45FFC994FD4B74E9D0EE6E" box="[151,247,351,372]" italics="true" pageId="8" pageNumber="9">Harpalus</emphasis>
(
<emphasis id="B94AEA852E45FFC9956F4B74E882EE6E" box="[261,421,351,372]" italics="true" pageId="8" pageNumber="9">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E45FFC995DE4B74EB1DEE6E" box="[436,570,351,372]" italics="true" pageId="8" pageNumber="9">xinjiangensis</emphasis>
<treatmentCitation id="0A9F10862E45FFC9962A4B74EA4CEE6E" author="Huang" box="[576,875,350,372]" page="263" pageId="8" pageNumber="9" year="1994">
<bibRefCitation id="EFAF4B662E45FFC9962A4B74EA1EEE6E" author="Huang" box="[576,825,350,372]" pageId="8" pageNumber="9" refString="Huang, T., Hu, J. &amp; Sun, D. (1994) Four new species of the genus Harpalus Latreille (Coleoptera: Carabidae) from China. Entomotaxonomia, 16 (4), 263 - 268." type="journal article" year="1994">Huang, Hu &amp; Sun, 1994</bibRefCitation>
: 263
</treatmentCitation>
</taxonomicName>
, 267,
<emphasis id="B94AEA852E45FFC997C34B74EAC9EE6E" bold="true" box="[937,1006,351,372]" pageId="8" pageNumber="9">syn. n.</emphasis>
<typeStatus id="548588352E45FFC9979F4B74ED00EE6E" box="[1013,1063,351,372]" pageId="8" pageNumber="9">Type</typeStatus>
locality: Urumqi, Xinjiang,
<collectingCountry id="F32976072E45FFC9912D4B75ECA0EE6E" box="[1351,1415,350,372]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E45FFC994FD4B8AEA27ECCC" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8B8136972E45FFC994FD4B8AEA27ECCC" blockId="8.[151,1437,417,1738]" pageId="8" pageNumber="9">
<emphasis id="B94AEA852E45FFC994FD4B8AE85EEEA0" bold="true" box="[151,377,417,442]" pageId="8" pageNumber="9">Material examined</emphasis>
. More than 2700 specimens (more than 2150 from
<collectingCountry id="F32976072E45FFC997D44B8AED22EEA0" box="[958,1029,417,442]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
), including the following specimens from Xinjiang, Hebei, Sichuan and Guizhou.
<collectingCountry id="F32976072E45FFC996F24BEEEBC6EEC4" box="[664,737,453,478]" name="China" pageId="8" pageNumber="9">
<emphasis id="B94AEA852E45FFC996F24BEEEBC6EEC4" bold="true" box="[664,737,453,478]" pageId="8" pageNumber="9">China</emphasis>
</collectingCountry>
. XINJIANG:
<specimenCount id="9D38FD1E2E45FFC997E84BEDEA88EEC7" box="[898,943,454,478]" pageId="8" pageNumber="9" type="generic">1 ♂</specimenCount>
, 2 ♀, &quot;Kachgarie, oasis de Koutchar, Dr. L. Vaillant 1909&quot; (
<collectionCode id="ED2FAE522E45FFC995224BC2E883ED1B" LSID="urn:lsid:biocol.org:col:34988" box="[328,420,489,513]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="9">MNHN</collectionCode>
); 1 ♀, &quot;Kuldzha A. Regel. 1878&quot; (
<collectionCode id="ED2FAE522E45FFC997424BC2EA7BED1B" box="[808,860,489,513]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E45FFC9971A4BC2EABBED1B" box="[880,924,489,513]" pageId="8" pageNumber="9" type="generic">1 ♂</specimenCount>
, &quot;Tien Shan to Taushkan-darya 20.V.
<date id="FF8010572E45FFC9912D4BC2ECBBED1B" box="[1351,1436,489,513]" pageId="8" pageNumber="9" value="1989-06-03">3.VI.89</date>
Pevtsov&quot; (
<collectionCode id="ED2FAE522E45FFC995604825E819ED3C" box="[266,318,526,550]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
); 1 ♀, &quot;Kashgaria, foothills of Kara-Teke, Yark.[end] -Dar.[ya]
<date id="FF8010572E45FFC990784825ED85ED3C" box="[1042,1186,526,550]" pageId="8" pageNumber="9" value="1989-06-10" valueMax="1989-06-22" valueMin="1989-06-10">10–22.VI.89</date>
, Pevtsov&quot; (
<collectionCode id="ED2FAE522E45FFC991484825EC71ED3C" box="[1314,1366,526,550]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E45FFC991014825ECB0ED3F" box="[1387,1431,526,550]" pageId="8" pageNumber="9" type="generic">1 ♂</specimenCount>
, &quot;Tat-..uia
<date id="FF8010572E45FFC9956C481AE842ED53" box="[262,357,561,585]" pageId="8" pageNumber="9" value="1990-07-01">1.VII.90</date>
Gr.[abczewski]&quot; (
<collectionCode id="ED2FAE522E45FFC99658481AEB41ED53" box="[562,614,561,585]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E45FFC99610481AEB81ED53" box="[634,678,561,585]" pageId="8" pageNumber="9" type="generic">1 ♂</specimenCount>
, &quot;Sandjac
<date id="FF8010572E45FFC99774481AEA5BED53" box="[798,892,561,585]" pageId="8" pageNumber="9" value="1990-07-04">4.VII.90</date>
Gr.[abczewski]&quot; (
<collectionCode id="ED2FAE522E45FFC99023481AED5AED53" box="[1097,1149,561,585]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
); 1 ♀, &quot;Ilsmash
<date id="FF8010572E45FFC9915E481AECBBED50" box="[1332,1436,561,586]" pageId="8" pageNumber="9" value="1990-08-06">6.VIII.90</date>
Gr.[abczewski]&quot; (
<collectionCode id="ED2FAE522E45FFC99531487DE8A8ED74" box="[347,399,598,622]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
); 1 ♀, &quot;Kul
<date id="FF8010572E45FFC9967F487EEBACED74" box="[533,651,597,622]" pageId="8" pageNumber="9" value="1990-08-15">15.VIII.90</date>
Gr.[abczewski]&quot; (
<collectionCode id="ED2FAE522E45FFC9973C487DEAADED74" box="[854,906,598,622]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
); 1 ♀, &quot;Arlalyn
<date id="FF8010572E45FFC99051487DEDB6ED74" box="[1083,1169,598,622]" pageId="8" pageNumber="9" value="1990-09-04">4.IX.90</date>
Gr.[abczewski]&quot; (
<collectionCode id="ED2FAE522E45FFC99131487DECA8ED74" box="[1371,1423,598,622]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E45FFC994FD4851E9EEED8B" box="[151,201,634,658]" pageId="8" pageNumber="9" type="generic">1 ♂</specimenCount>
, &quot;Takrakhan
<date id="FF8010572E45FFC995004851E891ED88" box="[362,438,634,658]" pageId="8" pageNumber="9" value="1990-10-03">3.X.90</date>
Gr.[abczewski]&quot; (
<collectionCode id="ED2FAE522E45FFC996E54851EBE4ED88" box="[655,707,634,658]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E45FFC996B74851EA28ED8B" box="[733,783,634,658]" pageId="8" pageNumber="9" type="generic">2 ♂</specimenCount>
, N Tien Shan, Koeksu Riv., S of Tekes,
<geoCoordinate id="EE0A50502E45FFC991614851ECBBED88" box="[1291,1436,634,658]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="42.784443">42°47' 04&quot;N</geoCoordinate>
<geoCoordinate id="EE0A50502E45FFC994FD48B6E834EDAC" box="[151,275,669,694]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="81.90305">81°54'11&quot;E</geoCoordinate>
–
<geoCoordinate id="EE0A50502E45FFC9954948B6E88EEDAC" box="[291,425,669,694]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="42.753887">42°45'14&quot;N</geoCoordinate>
<geoCoordinate id="EE0A50502E45FFC995D848B6EB12EDAC" box="[434,565,669,694]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="81.92972">81°55'47&quot;E</geoCoordinate>
,
<quantity id="4CC69B722E45FFC9962948B6EBC5EDAF" box="[579,738,669,694]" metricMagnitude="3" metricUnit="m" metricValue="1.5825" metricValueMax="1.6149999999999998" metricValueMin="1.55" pageId="8" pageNumber="9" unit="m" value="1582.5" valueMax="1615.0" valueMin="1550.0">1550–1615 m</quantity>
,
<date id="FF8010572E45FFC9969A48B5EA49EDAC" box="[752,878,670,694]" pageId="8" pageNumber="9" value="2012-07-07">7.VII.2012</date>
, I.I. Kabak leg. (cIB&amp;IK);
<specimenCount id="9D38FD1E2E45FFC990C048B5EC1EEDAC" box="[1194,1337,669,694]" pageId="8" pageNumber="9" type="generic">2 specimens</specimenCount>
, Koeksu Vall., Dunsala Riv., S of Tekes,
<geoCoordinate id="EE0A50502E45FFC9966348EAEBABEDC0" box="[521,652,705,730]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="42.7375">42°44'15&quot;N</geoCoordinate>
<geoCoordinate id="EE0A50502E45FFC996FC48EAEA35EDC0" box="[662,786,705,730]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="81.928055">81°55'41&quot;E</geoCoordinate>
–
<geoCoordinate id="EE0A50502E45FFC9974848EAEA8FEDC0" box="[802,936,705,730]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="42.719166">42°43'09&quot;N</geoCoordinate>
<geoCoordinate id="EE0A50502E45FFC997D848EAED12EDC0" box="[946,1077,705,730]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="81.86472">81°51'53&quot;E</geoCoordinate>
,
<quantity id="4CC69B722E45FFC9902948EAEDC5EDC3" box="[1091,1250,705,730]" metricMagnitude="3" metricUnit="m" metricValue="2.1125" metricValueMax="2.555" metricValueMin="1.67" pageId="8" pageNumber="9" unit="m" value="2112.5" valueMax="2555.0" valueMin="1670.0">1670–2555 m</quantity>
,
<date id="FF8010572E45FFC9909A48E9EC4AEDC0" box="[1264,1389,706,730]" pageId="8" pageNumber="9" value="2012-07-07">7.VII.2012</date>
, I.I. Kabak leg. (cIB&amp;IK);
<specimenCount id="9D38FD1E2E45FFC995F048CDE8EFEDE7" box="[410,456,742,766]" pageId="8" pageNumber="9" type="generic">2 ♂</specimenCount>
, Kunges Riv., N Kunges,
<geoCoordinate id="EE0A50502E45FFC9969C48CDEA5DEDE4" box="[758,890,742,766]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="43.52">43°31'12&quot;N</geoCoordinate>
<geoCoordinate id="EE0A50502E45FFC997E948CEED21EDE4" box="[899,1030,741,766]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="83.25611">83°15'22&quot;E</geoCoordinate>
, H=
<quantity id="4CC69B722E45FFC9905D48CDEDA5EDE7" box="[1079,1154,742,766]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="9" unit="m" value="840.0">840 m</quantity>
,
<date id="FF8010572E45FFC990FB48CEEC3BEDE4" box="[1169,1308,741,766]" pageId="8" pageNumber="9" value="2012-07-15">15.VII.2012</date>
, I.I. Kabak leg. (cIB&amp;IK); 1 ♀, S slope of East Tien Shan, Kucha R., N of Kucha Vill.,
<quantity id="4CC69B722E45FFC990644921ED40EC3B" box="[1038,1127,778,802]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FF8010572E45FFC9901F4921EDCFEC38" box="[1141,1256,778,802]" pageId="8" pageNumber="9" value="2001-06-07">7.VI.2001</date>
, I.I. Kabak leg. (cIB&amp;IK).
<collectionCode id="ED2FAE522E45FFC9957D4905E849EC5F" box="[279,366,814,837]" pageId="8" pageNumber="9">HEBEI</collectionCode>
:
<specimenCount id="9D38FD1E2E45FFC995164905EB3EEC5C" box="[380,537,813,838]" pageId="8" pageNumber="9" type="generic">18 specimens</specimenCount>
(♂ and ♀), Zhuolu County, Nan-Jiakou,
<quantity id="4CC69B722E45FFC9906E4906ED82EC5F" box="[1028,1189,813,838]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="8" pageNumber="9" unit="m" value="1400.0" valueMax="1600.0" valueMin="1200.0">1200–1600 m</quantity>
,
<date id="FF8010572E45FFC990DF4906EC53EC5C" box="[1205,1396,813,838]" pageId="8" pageNumber="9" value="2005-08-21" valueMax="2005-08-22" valueMin="2005-08-21">21–22.VIII.2005</date>
, Li Xueyan leg. (
<collectionCode id="ED2FAE522E45FFC995594979E840EC70" box="[307,359,850,874]" pageId="8" pageNumber="9">IOZ</collectionCode>
). SICHUAN:
<specimenCount id="9D38FD1E2E45FFC996614979EB1EEC73" box="[523,569,850,874]" pageId="8" pageNumber="9" type="generic">1 ♂</specimenCount>
, 1 ♀, Batang, Zhubalong, Sanjiacun, Jinshajiang,
<geoCoordinate id="EE0A50502E45FFC990144979EC21EC70" box="[1150,1286,850,874]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="29.84109">29.84109°N</geoCoordinate>
<geoCoordinate id="EE0A50502E45FFC9917A4979ECBFEC70" box="[1296,1432,850,874]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="99.0239">99.02390°E</geoCoordinate>
,
<quantity id="4CC69B722E45FFC994FD495DE9D4EC97" box="[151,243,886,910]" metricMagnitude="3" metricUnit="m" metricValue="2.48" pageId="8" pageNumber="9" unit="m" value="2480.0">2480 m</quantity>
,
<date id="FF8010572E45FFC9956E495DE85FEC94" box="[260,376,886,910]" pageId="8" pageNumber="9" value="2009-06-01">1.VI.2009</date>
, day, H.B. Liang leg. (
<collectionCode id="ED2FAE522E45FFC996FA495DEBE3EC94" box="[656,708,886,910]" pageId="8" pageNumber="9">IOZ</collectionCode>
); 1 ♀,
<quantity id="4CC69B722E45FFC9974A495DEA4BEC94" box="[800,876,885,910]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="9" unit="km" value="70.0">70 km</quantity>
W Chengdu, Qingchenghoushan Mts,
<quantity id="4CC69B722E45FFC99151495DECB0EC97" box="[1339,1431,886,910]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="9" unit="m" value="1400.0">1400 m</quantity>
,
<date id="FF8010572E45FFC994FD49B2E872ECA8" box="[151,341,921,946]" pageId="8" pageNumber="9" value="2005-08-16" valueMax="2005-08-25" valueMin="2005-08-16">16–25.VIII.2005</date>
, S. Murzin leg. (
<collectionCode id="ED2FAE522E45FFC9967249B1EB6BECA8" box="[536,588,922,946]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9">ZIN</collectionCode>
). GUIZHOU:
<specimenCount id="9D38FD1E2E45FFC9969B49B1EA38ECAB" box="[753,799,922,946]" pageId="8" pageNumber="9" type="generic">1 ♂</specimenCount>
, &quot;Guizhou, Guiyang / 1981.X&quot; [in
<taxonomicName id="4C3E4D142E45FFC990D949B2EC34ECA8" box="[1203,1299,921,946]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="8" pageNumber="37" rank="genus">Chinese</taxonomicName>
], &quot;
<taxonomicName id="4C3E4D142E45FFC9915849B2E80CECCF" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xinjiangensis">
<emphasis id="B94AEA852E45FFC9915849B2E80CECCF" italics="true" pageId="8" pageNumber="9">Harpalus xinjiangensis</emphasis>
</taxonomicName>
H. T. L. (det by Huang Tongling)&quot; (
<collectionCode id="ED2FAE522E45FFC996AA4995EBD3ECCC" box="[704,756,958,982]" pageId="8" pageNumber="9">IOZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E45FFC994AD49CAEDC4EB93" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8B8136972E45FFC994AD49CAEDC4EB93" blockId="8.[151,1437,417,1738]" pageId="8" pageNumber="9">
<emphasis id="B94AEA852E45FFC994AD49CAE81DECE0" bold="true" box="[199,314,993,1018]" pageId="8" pageNumber="9">Diagnosis</emphasis>
. The most distinctive characters are: clypeus with one seta on each side anteriorly; pronotum with one lateral seta on each side and with basal angles blunt at apex; pronotal disc in central portion impunctate; elytra densely punctate and pubescent on all intervals, without row of discal pores on intervals 1, 3, 5 and 7; and legs reddish brown or brownish yellow. In addition, labial basal palpomere is not carinate, and median lobe of aedeagus (
<figureCitation id="13052A122E45FFC994F54E5AE839EB90" box="[159,286,1137,1162]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,263,1483,1506]" captionTargetBox="[228,1353,642,1453]" captionTargetId="figure@9.[220,1366,628,1463]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Harpalus, median lobe of aedeagus. 15 – 16, H. griseus (Liaoning); 17 – 18, H. jureceki (Heilongjiang). 15, 17, view from the left side; 16, 18, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287888/files/figure.png" pageId="8" pageNumber="9">Figs 15–16</figureCitation>
) lacks any distinct sclerotized elements in the internal sac. Body length
<quantity id="4CC69B722E45FFC9902D4E5AEDF8EB93" box="[1095,1247,1137,1162]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" metricValueMax="1.25" metricValueMin="0.83" pageId="8" pageNumber="9" unit="mm" value="10.4" valueMax="12.5" valueMin="8.3">8.3–12.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E45FFC994AD4EBEE83EEAB0" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8B8136972E45FFC994AD4EBEE8C7EA24" blockId="8.[151,1437,417,1738]" pageId="8" pageNumber="9">
<emphasis id="B94AEA852E45FFC994AD4EBEE87EEBB4" bold="true" box="[199,345,1173,1198]" pageId="8" pageNumber="9">Distribution</emphasis>
.
<collectingCountry id="F32976072E45FFC995034EBEE897EBB4" box="[361,432,1173,1198]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
: the north-western and eastern parts. It was reported from Xinjiang, Heilongjiang, Jilin, Liaoning, Shandong, Jiangsu, Gansu, Shanxi, Shaanxi, Hubei, Zhejiang, Yunnan (
<bibRefCitation id="EFAF4B662E45FFC990374E91EC67EBC8" author="Kataev" box="[1117,1344,1209,1234]" pageId="8" pageNumber="9" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E45FFC990D04E91EDD6EBCB" box="[1210,1265,1209,1233]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2003
</bibRefCitation>
), Hebei (Schauberger 1938), Nei Mongol, Hubei, Henan, Anhui, Jiangxi, Sichuan, Guizhou, Fujian,
<collectingCountry id="F32976072E45FFC990DB4EF6EC21EBEC" box="[1201,1286,1245,1270]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
, Guangdong, Hunan, Guangxi, and Xizang (
<bibRefCitation id="EFAF4B662E45FFC995954F29EB50EA00" author="Hua" box="[511,631,1282,1306]" pageId="8" pageNumber="9" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua 2002</bibRefCitation>
). We provided here the additional examined material from Xinjiang, Hebei, Sichuan and Guizhou.
</paragraph>
<paragraph id="8B8136972E45FFC994AD4F62E83EEAB0" blockId="8.[151,1437,417,1738]" pageId="8" pageNumber="9">
General distribution includes the Palaearctic from North-Western Africa, Azores and
<collectingCountry id="F32976072E45FFC990C34F62EC2FEA78" box="[1193,1288,1353,1378]" name="Portugal" pageId="8" pageNumber="9">Portugal</collectingCountry>
to Sakhalin, southern Kurils,
<collectingCountry id="F32976072E45FFC9953E4F45E8B0EA9F" box="[340,407,1390,1413]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
,
<collectingCountry id="F32976072E45FFC995C84F45E8C1EA9F" box="[418,486,1390,1413]" name="South Korea" pageId="8" pageNumber="9">Korea</collectingCountry>
including Jejudo, and northern
<collectingCountry id="F32976072E45FFC997244F46EA8AEA9C" box="[846,941,1389,1414]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
(
<bibRefCitation id="EFAF4B662E45FFC997D64F45ED77EA9C" author="Kataev" box="[956,1104,1390,1414]" pageId="8" pageNumber="9" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E45FFC990374F45EC17EA9C" author="Kataev" box="[1117,1328,1389,1414]" pageId="8" pageNumber="9" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E45FFC990DE4F45EDC2EA9F" box="[1204,1253,1389,1413]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2003
</bibRefCitation>
; Moon &amp;
<bibRefCitation id="EFAF4B662E45FFC994FD4FBAE82AEAB0" author="Paik" box="[151,269,1425,1450]" pageId="8" pageNumber="9" refString="Moon, Ch. - S. &amp; Paik, J. - Ch. (2006) Notes on Harpalini (Coleoptera, Carabidae) from Korea (1 - 3). Korean Journal of Soil Zoology, 11 (1 - 2), 30 - 84." type="journal article" year="2006">Paik 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E45FFC994AD4F9EED99E9D3" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8B8136972E45FFC994AD4F9EED99E9D3" blockId="8.[151,1437,417,1738]" pageId="8" pageNumber="9">
<emphasis id="B94AEA852E45FFC994AD4F9EE813EAD4" bold="true" box="[199,308,1461,1486]" pageId="8" pageNumber="9">Remarks</emphasis>
.
<taxonomicName id="4C3E4D142E45FFC995284F9EEB60EAD7" box="[322,583,1461,1485]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xinjiangensis">
<emphasis id="B94AEA852E45FFC995284F9EEB60EAD7" box="[322,583,1461,1485]" italics="true" pageId="8" pageNumber="9">Harpalus xinjiangensis</emphasis>
</taxonomicName>
was described on the basis of six specimens from Urumqi, Xinjiang,
<collectingCountry id="F32976072E45FFC991384F9EECBEEAD4" box="[1362,1433,1461,1486]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
. According to the original description (
<bibRefCitation id="EFAF4B662E45FFC996084FF1EA64EAE8" author="Huang" box="[610,835,1497,1522]" pageId="8" pageNumber="9" refString="Huang, T., Hu, J. &amp; Sun, D. (1994) Four new species of the genus Harpalus Latreille (Coleoptera: Carabidae) from China. Entomotaxonomia, 16 (4), 263 - 268." type="journal article" year="1994">
Huang
<emphasis id="B94AEA852E45FFC996D64FF1EBD4EAEB" box="[700,755,1497,1521]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1994
</bibRefCitation>
), this species is similar to
<taxonomicName id="4C3E4D142E45FFC990E54FF1EC2FEAEB" box="[1167,1288,1498,1521]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="B94AEA852E45FFC990E54FF1EC2FEAEB" box="[1167,1288,1498,1521]" italics="true" pageId="8" pageNumber="9">H. griseus</emphasis>
</taxonomicName>
, but can be distinguished by its frons &quot;with a shallow foveae and punctures&quot;, and metafemur &quot;with 10–14 setae in two rows at hind margin&quot;. We could not study the
<typeStatus id="548588352E45FFC996294C09EB54E920" box="[579,627,1570,1594]" pageId="8" pageNumber="9">type</typeStatus>
material, however the examination of the specimens of
<taxonomicName id="4C3E4D142E45FFC990814C0AECBBE923" box="[1259,1436,1569,1593]" class="Insecta" family="Carabidae" genus="Pseudoophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E45FFC990814C0AECBBE923" box="[1259,1436,1569,1593]" italics="true" pageId="8" pageNumber="9">Pseudoophonus</emphasis>
</taxonomicName>
from Xinjiang and the specimens from Guizhou, determined as
<taxonomicName id="4C3E4D142E45FFC9970F4C6DED3BE947" box="[869,1052,1606,1629]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xinjiangensis">
<emphasis id="B94AEA852E45FFC9970F4C6DED3BE947" box="[869,1052,1606,1629]" italics="true" pageId="8" pageNumber="9">H. xinjiangensis</emphasis>
</taxonomicName>
by Huang Tongling, revealed that this description is based on members of
<taxonomicName id="4C3E4D142E45FFC9961A4C41EBC1E99B" box="[624,742,1642,1665]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="B94AEA852E45FFC9961A4C41EBC1E99B" box="[624,742,1642,1665]" italics="true" pageId="8" pageNumber="9">H. griseus</emphasis>
</taxonomicName>
which is rather variable in number of setae along posterior margin of metafemur (three to five long setae and usually numerous additional short setae); head also has often very fine punctures on frons. Thus we treat
<taxonomicName id="4C3E4D142E45FFC996EA4C99EA11E9D3" box="[640,822,1714,1737]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xinjiangensis">
<emphasis id="B94AEA852E45FFC996EA4C99EA11E9D3" box="[640,822,1714,1737]" italics="true" pageId="8" pageNumber="9">H. xinjiangensis</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C3E4D142E45FFC9902E4C99ED9FE9D3" box="[1092,1208,1714,1737]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="B94AEA852E45FFC9902E4C99ED9FE9D3" box="[1092,1208,1714,1737]" italics="true" pageId="8" pageNumber="9">H. griseus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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