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92 KiB
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507 lines
92 KiB
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<document id="82C9BAC6F78DFEA3DCC2A508493BBD06" ID-CLB-Dataset="299409" ID-DOI="10.5281/zenodo.11067293" ID-GBIF-Dataset="9235db45-04c4-4570-8987-cb84765763a4" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067293" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052207303" checkinUser="felipe" docAuthor="Kavanaugh, David H., Cueva-Dabkoski, Mollie & Liang, Hongbin" docDate="2023" docId="644DE316FF9AFFA0D34FD95BFF6861B4" docLanguage="en" docName="ProcCalifAcadSci.67.20.451-491.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (20)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Tetragonoderus punctatus" docType="treatment" docVersion="5" lastPageNumber="474" masterDocId="98749B6EFF8FFFB8D334D867FFCA6027" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species." masterLastPageNumber="491" masterPageNumber="451" pageNumber="472" updateTime="1719954462017" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="EF48BA9625A924057FB7A0F450451A32">
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<mods:title id="28D87BF05AE3BAF6F33773A22D3DB679">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:name id="C2B017D203B946897F0174E49C86BEC3" type="personal">
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<mods:roleTerm id="4CDA0E4D70184BEB97F75E7FA1FF0BF2">Author</mods:roleTerm>
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<mods:namePart id="041190F3E8057D6A6CA8EC1EF96FF37C">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="B38AE4C29BC043228F83F1B6A1139057">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:name id="873FE29EAFA4B7BE0A8DE195E30C88FC" type="personal">
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<mods:roleTerm id="A804C29C492F94C4CF3415A1FD9AA2AE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E80914F49D94B526F27ACD9ED0641279">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="0E2BC40726278D3A4ABDAAEE51D8DA55">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:name id="428D5BD909D0B604018F929194F06D39" type="personal">
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<mods:roleTerm id="5A4A837FEF48135CB636ADD57DBB7CFA">Author</mods:roleTerm>
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<mods:namePart id="1EFE5D923A4273F2A07B45AAC1AF32D8">Liang, Hongbin</mods:namePart>
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<mods:affiliation id="8E089FB3CE94E9C5302DBE4C76C68583">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
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<mods:typeOfResource id="D087DB0AF5B466D84E2A06DAE6DAF924">text</mods:typeOfResource>
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<mods:title id="D767221329E002CAEDD1E566C9483CB3">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="0A73DBA7540359CF4A911B862197E830">
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<mods:date id="699F0FB0E4388A66A48D468923E3F61F">2023</mods:date>
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<mods:detail id="CB61DFBCA6DDF220A4DF0677791EAD22" type="pubDate">
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<mods:number id="97B716763493681DD44581D7973636AB">2023-02-15</mods:number>
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<mods:detail id="4F5287CD3965827A3BD5D59DFAFF7A8E" type="volume">
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<mods:number id="358D65422927C76A7B004CD65CF1D9C6">67</mods:number>
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<mods:number id="2296A9A9D0AD7D6FC602B0FF36FDAB27">20</mods:number>
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<mods:start id="B2F560BD3102D9D2FC2F32059A61A9E6">451</mods:start>
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<mods:identifier id="58A14612465451839C705B88DADB11EC" type="DOI">10.5281/zenodo.11067293</mods:identifier>
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<treatment id="644DE316FF9AFFA0D34FD95BFF6861B4" ID-DOI="http://doi.org/10.5281/zenodo.12628252" ID-GBIF-Taxon="234412270" ID-Zenodo-Dep="12628252" LSID="urn:lsid:plazi:treatment:644DE316FF9AFFA0D34FD95BFF6861B4" httpUri="http://treatment.plazi.org/id/644DE316FF9AFFA0D34FD95BFF6861B4" lastPageId="24" lastPageNumber="474" pageId="21" pageNumber="472">
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<subSubSection id="A4FE018BFF9AFFADD34FD95BFDF76174" box="[123,573,316,340]" pageId="21" pageNumber="472" type="nomenclature">
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<paragraph id="EC5B5200FF9AFFADD34FD95BFDF76174" blockId="21.[123,681,316,370]" box="[123,573,316,340]" pageId="21" pageNumber="472">
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<heading id="B713E56CFF9AFFADD34FD95BFDF76174" box="[123,573,316,340]" fontSize="10" level="2" pageId="21" pageNumber="472" reason="4">
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<taxonomicName id="2BE42983FF9AFFADD34FD95BFDF76174" ID-CoL="55N24" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[123,573,316,340]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
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<emphasis id="DE908E12FF9AFFADD34FD95BFDF76174" bold="true" box="[123,573,316,340]" pageId="21" pageNumber="472">
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<emphasis id="DE908E12FF9AFFADD34FD95BFE506173" bold="true" box="[123,410,316,340]" italics="true" pageId="21" pageNumber="472">Tetragonoderus punctatus</emphasis>
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(Wiedemann)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A4FE018BFF9AFFADD34FD93BFD636155" box="[123,681,348,370]" pageId="21" pageNumber="472" type="description">
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<paragraph id="EC5B5200FF9AFFADD34FD93BFD636155" blockId="21.[123,681,316,370]" box="[123,681,348,370]" pageId="21" pageNumber="472">
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<figureCitation id="74DF4E85FF9AFFADD34FD93BFF296155" box="[123,227,348,370]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="21" pageNumber="472">
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Figures
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<geoCoordinate id="89D034C7FF9AFFADD3FDD93BFF296155" box="[201,227,348,370]" degrees="12, 13" direction="east" orientation="longitude" pageId="21" pageNumber="472" precision="555" value="12.13">12</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD3DAD93BFEDD6155" box="[238,279,348,370]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="21" pageNumber="472">13E</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD215D93BFE836155" box="[289,329,348,370]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="21" pageNumber="472">14E</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD260D93BFEB76155" box="[340,381,348,370]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="21" pageNumber="472">15E</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD2BCD93BFE7B6155" box="[392,433,348,370]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">16E</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD28FD93BFE2E6155" box="[443,484,348,370]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="21" pageNumber="472">17E</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD2DAD93BFDCF6155" box="[494,517,348,370]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="21" pageNumber="472">19</figureCitation>
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C-D, 21, and 23.
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</paragraph>
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</subSubSection>
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<subSubSection id="A4FE018BFF9AFFADD34FD9E1FD3E6284" pageId="21" pageNumber="472" type="reference_group">
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<paragraph id="EC5B5200FF9AFFADD34FD9E1FE4E624E" blockId="21.[123,1175,390,675]" pageId="21" pageNumber="472">
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<treatmentCitationGroup id="CCF4752EFF9AFFADD34FD9E1FE4E624E" pageId="21" pageNumber="472">
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<treatmentCitation id="6D457411FF9AFFADD34FD9E1FDFF61BB" author="WIEDEMANN, C. R. W." box="[123,565,390,412]" page="61" pageId="21" pageNumber="472" year="1823">
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<taxonomicName id="2BE42983FF9AFFADD34FD9E1FDFF61BB" ID-CoL="LG8X" authority="Wiedemann, 1823: 61" authorityName="Wiedemann" authorityPageNumber="61" authorityYear="1823" box="[123,565,390,412]" class="Insecta" family="Carabidae" genus="Bembidium" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatum">
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<emphasis id="DE908E12FF9AFFADD34FD9E1FE9D61BB" box="[123,343,390,412]" italics="true" pageId="21" pageNumber="472">Bembidium punctatum</emphasis>
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<bibRefCitation id="88752FF1FF9AFFADD255D9E1FDFF61BB" author="WIEDEMANN, C. R. W." box="[353,565,390,412]" pageId="21" pageNumber="472" pagination="162 - 164" refId="ref18095" refString="WIEDEMANN, C. R. W. 1823. Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." type="journal article" year="1823">Wiedemann, 1823:61</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="335FECA2FF9AFFADD176D9E1FD0761BB" box="[578,717,390,412]" pageId="21" pageNumber="472" type="lectotype">LECTOTYPE</typeStatus>
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, here designated, a male, in ZMUC, labeled: “TYPE” [red label]/ “Bengala Westermann
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<taxonomicName id="2BE42983FF9AFFADD172D9C3FD2F619E" ID-CoL="55N24" authority="Wied." authorityName="Wied." box="[582,741,419,441]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">Punctatus Wied.</taxonomicName>
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” [handwritten]/ “ZMUC 00027869”/ “
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<typeStatus id="335FECA2FF9AFFADD76BD9C4FF3D61F0" pageId="21" pageNumber="472" type="lectotype">LECTOTYPE</typeStatus>
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<taxonomicName id="2BE42983FF9AFFADD3CBD9A6FD5961F0" ID-CoL="LG8X" authority="Wiedemann, 1823" authorityName="Wiedemann" authorityYear="1823" box="[255,659,449,471]" class="Insecta" family="Carabidae" genus="Bembidium" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatum">
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<emphasis id="DE908E12FF9AFFADD3CBD9A6FE1061F0" box="[255,474,449,471]" italics="true" pageId="21" pageNumber="472">Bembidium punctatum</emphasis>
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<bibRefCitation id="88752FF1FF9AFFADD2D6D9A6FD5961F0" author="WIEDEMANN, C. R. W." box="[482,659,449,471]" pageId="21" pageNumber="472" pagination="162 - 164" refId="ref18095" refString="WIEDEMANN, C. R. W. 1823. Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
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</taxonomicName>
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design. by D.H. Kavanaugh & M. Cueva-Dabkoski 2022” [red label].
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<typeStatus id="335FECA2FF9AFFADD278D9B9FE1261D3" box="[332,472,478,500]" pageId="21" pageNumber="472" type="paralectotype">Paralectotypes</typeStatus>
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examined:
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<specimenCount id="FAE29989FF9AFFADD17CD9B9FD5461D3" box="[584,670,478,500]" count="1" pageId="21" pageNumber="472" type="female">1 female</specimenCount>
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, in ZMUC, labeled: “Mus. Westerm.”/ “TYPE” [red label]/ “Bengal Maj. 1808.
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<taxonomicName id="2BE42983FF9AFFADD286D99BFD9E6236" ID-CoL="55N24" authority="Wied." authorityName="Wied." box="[434,596,507,529]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">Punctatus Wied.</taxonomicName>
|
|
” [handwritten]/ “ZMUC 00027868” [label upside down]/ “
|
|
<typeStatus id="335FECA2FF9AFFADD390DA7FFEA76209" box="[164,365,536,558]" pageId="21" pageNumber="472" type="paralectotype">PARALECTOTYPE</typeStatus>
|
|
<taxonomicName id="2BE42983FF9AFFADD247DA7FFCCA6209" ID-CoL="LG8X" authority="Wiedemann, 1823" authorityName="Wiedemann" authorityYear="1823" box="[371,768,536,558]" class="Insecta" family="Carabidae" genus="Bembidium" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatum">
|
|
<emphasis id="DE908E12FF9AFFADD247DA7FFD816209" box="[371,587,536,558]" italics="true" pageId="21" pageNumber="472">Bembidium punctatum</emphasis>
|
|
<bibRefCitation id="88752FF1FF9AFFADD165DA7FFCCA6209" author="WIEDEMANN, C. R. W." box="[593,768,536,558]" pageId="21" pageNumber="472" pagination="162 - 164" refId="ref18095" refString="WIEDEMANN, C. R. W. 1823. Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
|
|
</taxonomicName>
|
|
design. by D.H. Kavanaugh & M. Cueva-Dabkoski 2022” [yellow label]. Type locality: “Bengalia” [area which now includes
|
|
<collectingRegion id="2E209CE2FF9AFFADD0FDDA51FB89626B" box="[969,1091,566,588]" country="India" name="West Bengal" pageId="21" pageNumber="472">West Bengal</collectingRegion>
|
|
(eastern
|
|
<collectingCountry id="94F31290FF9AFFADD3AFDA34FF04624E" box="[155,206,595,617]" name="India" pageId="21" pageNumber="472">India</collectingCountry>
|
|
) and
|
|
<collectingCountry id="94F31290FF9AFFADD230DA34FEB0624E" box="[260,378,595,617]" name="Bangladesh" pageId="21" pageNumber="472">Bangladesh</collectingCountry>
|
|
].
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF9AFFADD34FDA17FD3E6284" blockId="21.[123,1175,390,675]" pageId="21" pageNumber="472">
|
|
<treatmentCitationGroup id="CCF4752EFF9AFFADD34FDA17FD3E6284" pageId="21" pageNumber="472">
|
|
<taxonomicName id="2BE42983FF9AFFADD34FDA17FDCB62A1" ID-CoL="55N24" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[123,513,624,646]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF9AFFADD34FDA17FEBF62A1" box="[123,373,624,646]" italics="true" pageId="21" pageNumber="472">Tetragonoderus punctatus</emphasis>
|
|
(Wiedemann)
|
|
</taxonomicName>
|
|
,
|
|
<treatmentCitation id="6D457411FF9AFFADD13FDA17FD0862A1" author="DEJEAN, P. F. M. A." box="[523,706,624,646]" page="505" pageId="21" pageNumber="472" year="1829">
|
|
<bibRefCitation id="88752FF1FF9AFFADD13FDA17FD0862A1" author="DEJEAN, P. F. M. A." box="[523,706,624,646]" pageId="21" pageNumber="472" refId="ref16379" refString="DEJEAN, P. F. M. A. 1829. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean (1829:505)</bibRefCitation>
|
|
</treatmentCitation>
|
|
;
|
|
<treatmentCitation id="6D457411FF9AFFADD1F8DA17FC0C62A1" author="SCHMIDT-GOBEL, M." box="[716,966,624,646]" page="92" pageId="21" pageNumber="472" year="1846">
|
|
<bibRefCitation id="88752FF1FF9AFFADD1F8DA17FC0C62A1" author="SCHMIDT-GOBEL, M." box="[716,966,624,646]" pageId="21" pageNumber="472" pagination="1 - 3" refId="ref17793" refString="SCHMIDT-GOBEL, M. 1846. Faunula coleopterorum Birmaniae, adjectis nonnulis Bengaliae indigenis. Med. Dr. Johann Wilhelm Helfer's hinterlassene Sammlungen aus Vorder- und Hinter-Indien. Nach seinem Tode im Auftrage des bohm. National-Museums unter Mitwirkung Mehrerer bearbeitet und herausgegeben. G. Haase Sohne, Prague, viii + 94 pp. + plates 1 - 3." type="book chapter" year="1846">Schmidt-Göbel (1846:92)</bibRefCitation>
|
|
</treatmentCitation>
|
|
;
|
|
<treatmentCitation id="6D457411FF9AFFADD0E5DA17FB5B62A1" author="CHAUDOIR, M." box="[977,1169,624,646]" page="48" pageId="21" pageNumber="472" year="1876">
|
|
<bibRefCitation id="88752FF1FF9AFFADD0E5DA17FB5B62A1" author="CHAUDOIR, M." box="[977,1169,624,646]" pageId="21" pageNumber="472" pagination="1 - 84" refId="ref16317" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir (1876:48)</bibRefCitation>
|
|
</treatmentCitation>
|
|
;
|
|
<treatmentCitation id="6D457411FF9AFFADD3AFDAEAFEBA6284" author="ANDREWES, H. E." box="[155,368,653,675]" page="174" pageId="21" pageNumber="472" year="1921">
|
|
<bibRefCitation id="88752FF1FF9AFFADD3AFDAEAFEBA6284" author="ANDREWES, H. E." box="[155,368,653,675]" pageId="21" pageNumber="472" pagination="145 - 195" refId="ref15718" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:174)</bibRefCitation>
|
|
</treatmentCitation>
|
|
;
|
|
<treatmentCitation id="6D457411FF9AFFADD24FDAEAFDE46284" author="CSIKI, E." box="[379,558,653,675]" page="1298" pageId="21" pageNumber="472" year="1932">
|
|
<bibRefCitation id="88752FF1FF9AFFADD24FDAEAFDE46284" author="CSIKI, E." box="[379,558,653,675]" pageId="21" pageNumber="472" pagination="1279 - 1598" refId="ref16350" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1298)</bibRefCitation>
|
|
</treatmentCitation>
|
|
;
|
|
<treatmentCitation id="6D457411FF9AFFADD10CDAE9FD3A6284" author="LORENZ, W." box="[568,752,653,675]" page="453" pageId="21" pageNumber="472" year="2005">
|
|
<bibRefCitation id="88752FF1FF9AFFADD10CDAE9FD3A6284" author="LORENZ, W." box="[568,752,653,675]" pageId="21" pageNumber="472" refId="ref17529" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005:453)</bibRefCitation>
|
|
</treatmentCitation>
|
|
.
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A4FE018BFF9AFFA0D39FDADFFF6861B4" lastPageId="24" lastPageNumber="475" pageId="21" pageNumber="472" type="diagnosis">
|
|
<paragraph id="EC5B5200FF9AFFADD39FDADFFD6164B2" blockId="21.[123,1174,695,1725]" pageId="21" pageNumber="472">
|
|
<emphasis id="DE908E12FF9AFFADD39FDADFFED562E8" bold="true" box="[171,287,696,719]" pageId="21" pageNumber="472">Diagnosis.</emphasis>
|
|
Adults of
|
|
<taxonomicName id="2BE42983FF9AFFADD294DADFFDE262E8" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[416,552,696,719]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF9AFFADD294DADFFDE262E8" box="[416,552,696,719]" italics="true" pageId="21" pageNumber="472">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
can be distinguished from those of other
|
|
<emphasis id="DE908E12FF9AFFADD726DAD0FB5C62E8" box="[1042,1174,695,719]" italics="true" pageId="21" pageNumber="472">cyclosomine</emphasis>
|
|
species in the study area by the following combination of character states: Body size medium for genus, BL males =
|
|
<quantity id="2B1CFFE5FF9AFFADD273DA9FFE146337" box="[327,478,760,784]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="21" pageNumber="472" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5 to 4.8 mm</quantity>
|
|
, females =
|
|
<quantity id="2B1CFFE5FF9AFFADD163DA9FFD246337" box="[599,750,760,784]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.4" metricValueMin="4.6" pageId="21" pageNumber="472" unit="mm" value="5.0" valueMax="5.4" valueMin="4.6">4.6 to 5.4 mm</quantity>
|
|
; pronotum piceous or rufopiceous, with or without faint greenish metallic reflection; elytral color pattern as in
|
|
<figureCitation id="74DF4E85FF9AFFADD04BDB7FFC286317" box="[895,994,792,816]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="21" pageNumber="472">Fig. 12A</figureCitation>
|
|
, with dark areas less expansive, connection between medial portion of the basal dark band and humeral portion absent or very narrow, less than one-fifth the length of humeral portion, middle discal band with portion between anterior and posterior dark edges not or only slightly darker than remainder of pale elytral areas; femora pale tan; pronotum (
|
|
<figureCitation id="74DF4E85FF9AFFADD10EDBFDFD536395" box="[570,665,922,946]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="21" pageNumber="472">Fig. 13E</figureCitation>
|
|
) widest near mid-length and posterior to insertion of midlateral seta, not or only very slightly narrowed basally (ratio PWB/PWA = 1.15 to 1.25), lateral margins slightly and evenly convex or straight or very slightly sinuate anterior to basal angles; elytra slightly to distinctly truncate apically; front tarsomeres 1 to 3 without lateral expansions (
|
|
<figureCitation id="74DF4E85FF9AFFADD3F6DC7BFEE76413" box="[194,301,1052,1076]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="21" pageNumber="472">Figs. 14E</figureCitation>
|
|
,
|
|
<figureCitation id="74DF4E85FF9AFFADD20FDC7BFEA36413" box="[315,361,1052,1076]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="21" pageNumber="472">15E</figureCitation>
|
|
); male with middle mesotarsomeres (
|
|
<figureCitation id="74DF4E85FF9AFFADD037DC7BFCA96413" box="[771,867,1052,1076]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">Fig. 16E</figureCitation>
|
|
) only slightly wider than in female and with only tarsomeres 1 to 3 with pads of adhesive setae ventrally (i.e., tarsomere 4 without adhesive setae ventrally) (
|
|
<figureCitation id="74DF4E85FF9AFFADD2F4DC3AFDEB6452" box="[448,545,1117,1141]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="21" pageNumber="472">Fig. 17E</figureCitation>
|
|
); median lobe of male genitalia (
|
|
<figureCitation id="74DF4E85FF9AFFADD0BDDC3AFBC46452" box="[905,1038,1117,1141]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="21" pageNumber="472">Figs. 19C,D</figureCitation>
|
|
) with apical lamella short and broad, internal sac without spines.
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF9AFFAFD39FDCF8FED16193" blockId="21.[123,1174,695,1725]" lastBlockId="23.[123,1174,219,1725]" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472">
|
|
In the study area, specimens of
|
|
<taxonomicName id="2BE42983FF9AFFADD2CEDCF8FDB16491" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[506,635,1183,1206]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF9AFFADD2CEDCF8FDB16491" box="[506,635,1183,1206]" italics="true" pageId="21" pageNumber="472">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="74DF4E85FF9AFFADD1BDDCF9FD206491" box="[649,746,1182,1206]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="21" pageNumber="472">Fig. 12A</figureCitation>
|
|
) might be confused with those of
|
|
<taxonomicName id="2BE42983FF9AFFADD760DCF8FF6764F1" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9AFFADD760DCF8FF6764F1" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="74DF4E85FF9AFFADD389DCD9FED564F1" box="[189,287,1214,1238]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="21" pageNumber="472">Fig. 10A</figureCitation>
|
|
) and also with those of
|
|
<taxonomicName id="2BE42983FF9AFFADD112DCD8FD0E64F1" authorityName="Jian & Tian" authorityYear="2009" box="[550,708,1214,1238]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF9AFFADD112DCD8FD0E64F1" box="[550,708,1214,1238]" italics="true" pageId="21" pageNumber="472">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="74DF4E85FF9AFFADD1E0DCD9FCF164F1" box="[724,827,1214,1238]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="21" pageNumber="472">Figs. 13F</figureCitation>
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,
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|
<figureCitation id="74DF4E85FF9AFFADD07CDCD9FCB864F1" box="[840,882,1214,1238]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="21" pageNumber="472">14F</figureCitation>
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,
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|
<figureCitation id="74DF4E85FF9AFFADD04ADCD9FC6D64F1" box="[894,935,1214,1238]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="21" pageNumber="472">15F</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD080DCD9FC1764F1" box="[948,989,1214,1238]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">16F</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD0DEDCD9FBD964F1" box="[1002,1043,1214,1238]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="21" pageNumber="472">17F</figureCitation>
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,
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<figureCitation id="74DF4E85FF9AFFADD714DCD9FBF364F1" box="[1056,1081,1214,1238]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="21" pageNumber="472">19</figureCitation>
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E-F, and 20), although the latter has not yet been recorded from the Gaoligong
|
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<collectingRegion id="2E209CE2FF9AFFADD055DCB8FC2E64D0" box="[865,996,1247,1271]" country="Myanmar" name="Shan" pageId="21" pageNumber="472">Shan region</collectingRegion>
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.
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<taxonomicName id="2BE42983FF9AFFADD0D9DCB8FF296530" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
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<emphasis id="DE908E12FF9AFFADD0D9DCB8FF296530" italics="true" pageId="21" pageNumber="472">Tetragonoderus punctatus</emphasis>
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</taxonomicName>
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|
members differ from those of
|
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<taxonomicName id="2BE42983FF9AFFADD11ADD67FD506530" box="[558,666,1279,1303]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="DE908E12FF9AFFADD11ADD67FD506530" box="[558,666,1279,1303]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
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|
</taxonomicName>
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|
in having the forebody slightly lighter in color (i.e., less heavily pigmented) and the elytral pattern of dark areas less extensive, with connection between medial portion of the basal dark band and humeral portion absent or very narrow, less than one-fifth the length of the humeral portion (at least one-fourth the length of the humeral portion in
|
|
<taxonomicName id="2BE42983FF9AFFADD34FDDE5FF2D65BE" box="[123,231,1409,1433]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9AFFADD34FDDE5FF2D65BE" box="[123,231,1409,1433]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
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|
</taxonomicName>
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|
), the middle discal band with the area between anterior and posterior dark edges not or only very slightly darker than remainder of pale elytral areas (distinctly darker tan-orange in
|
|
<taxonomicName id="2BE42983FF9AFFADD34FDDA4FF2265FD" box="[123,232,1474,1498]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
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|
<emphasis id="DE908E12FF9AFFADD34FDDA4FF2265FD" box="[123,232,1474,1498]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
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</taxonomicName>
|
|
), and with a less distinct and discontinuous preapical dark band. Also the pronotum is proportionally slightly wider (ratio PWM/EW = 0.55 to 0.57; in
|
|
<taxonomicName id="2BE42983FF9AFFADD01BDD84FC5165DD" box="[815,923,1506,1530]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9AFFADD01BDD84FC5165DD" box="[815,923,1506,1530]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
= 0.51 to 0.55), widest at or near midlength, posterior to the insertion of the midlateral seta (widest more anteriad and at or near the insertion of the midlateral seta in
|
|
<taxonomicName id="2BE42983FF9AFFADD147DE43FD2B661C" box="[627,737,1571,1595]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9AFFADD147DE43FD2B661C" box="[627,737,1571,1595]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
), less distinctly narrowed basally (ratio PWM/PWB = 1.06 to 1.13; in
|
|
<taxonomicName id="2BE42983FF9AFFADD2FDDE22FDFC667B" box="[457,566,1604,1628]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9AFFADD2FDDE22FDFC667B" box="[457,566,1604,1628]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
= 1.13 to 1.20), and the lateral margins are not at all or only very slightly sinuate anterior to the basal angles (distinctly sinuate in
|
|
<taxonomicName id="2BE42983FF9AFFADD0FDDE02FBF3665B" box="[969,1081,1636,1660]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9AFFADD0FDDE02FBF3665B" box="[969,1081,1636,1660]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
). Males differ in several additional features. In
|
|
<taxonomicName id="2BE42983FF9AFFADD12ADEE1FD6B66BA" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[542,673,1670,1693]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF9AFFADD12ADEE1FD6B66BA" box="[542,673,1670,1693]" italics="true" pageId="21" pageNumber="472">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
males, the middle tarsi are only slightly wider than those in females and middle tarsomere 4 lacks a pad of adhesive setae ventrally (
|
|
<figureCitation id="74DF4E85FF9AFFADD712DEC2FB5B669A" box="[1062,1169,1701,1725]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">Figs. 16E</figureCitation>
|
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,
|
|
<figureCitation id="74DF4E85FF98FFAFD34FD8BCFF6360D4" box="[123,169,219,243]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="23" pageNumber="474">17E</figureCitation>
|
|
), whereas middle tarsomeres 1 to 4 are distinctly wider than in females and bear adhesive setae ventrally in
|
|
<taxonomicName id="2BE42983FF98FFAFD3CFD89BFEAC6134" box="[251,358,251,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF98FFAFD3CFD89BFEAC6134" box="[251,358,251,275]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
males (
|
|
<figureCitation id="74DF4E85FF98FFAFD28ED89CFDEE6134" box="[442,548,251,275]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="23" pageNumber="474">Figs. 16C</figureCitation>
|
|
,
|
|
<figureCitation id="74DF4E85FF98FFAFD11BD89CFD956134" box="[559,607,251,275]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="23" pageNumber="474">17C</figureCitation>
|
|
). Males also differ in the form of their genitalia, with the median lobe (
|
|
<figureCitation id="74DF4E85FF98FFAFD200D97CFE7D6114" box="[308,439,283,307]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="23" pageNumber="474">Figs. 19C,D</figureCitation>
|
|
) slightly shorter and distinctly thicker, the apical lamella shorter and broader, and the internal sac without distinct spines in
|
|
<taxonomicName id="2BE42983FF98FFAFD1F4D95BFC886174" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[704,834,316,339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF98FFAFD1F4D95BFC886174" box="[704,834,316,339]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
. In contrast, the median lobe of
|
|
<taxonomicName id="2BE42983FF98FFAFD34FD93AFF206153" box="[123,234,348,372]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF98FFAFD34FD93AFF206153" box="[123,234,348,372]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
males (
|
|
<figureCitation id="74DF4E85FF98FFAFD272D93BFE076153" box="[326,461,348,372]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="23" pageNumber="474">Figs. 19A,B</figureCitation>
|
|
) is longer and more slender, the apical lamella is much longer, narrower, and distinctly curved ventrad, and the internal sac is armed with five or more large and distinct spines.
|
|
</paragraph>
|
|
<caption id="B89B0288FF99FFAED3AFDE54FBC8665F" ID-DOI="http://doi.org/10.5281/zenodo.11067313" ID-Zenodo-Dep="11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="22" pageNumber="473" startId="22.[155,167,1587,1606]" targetBox="[128,1176,216,1576]" targetPageId="22" targetType="figure">
|
|
<paragraph id="EC5B5200FF99FFAED3AFDE54FBC8665F" blockId="22.[123,1174,1586,1657]" pageId="22" pageNumber="473">
|
|
<smallCapsWord id="EABDC4DCFF99FFAED3AFDE54FF126662" baselines="1601,1601" box="[155,216,1587,1606]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="22" pageNumber="473">FIGURE</smallCapsWord>
|
|
12.
|
|
<taxonomicName id="2BE42983FF99FFAED23CDE55FDA56662" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[264,623,1586,1605]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF99FFAED23CDE55FE266662" box="[264,492,1586,1605]" italics="true" pageId="22" pageNumber="473">Tetragonoderus punctatus</emphasis>
|
|
(Wiedemann)
|
|
</taxonomicName>
|
|
. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="EC5B5200FF98FFAFD39FD9DBFE0663DF" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
|
|
Members of
|
|
<taxonomicName id="2BE42983FF98FFAFD209D9DAFE0961F3" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[317,451,445,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF98FFAFD209D9DAFE0961F3" box="[317,451,445,468]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
are extremely similar to those of
|
|
<taxonomicName id="2BE42983FF98FFAFD078D9DAFC2461F3" authorityName="Jian & Tian" authorityYear="2009" box="[844,1006,444,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF98FFAFD078D9DAFC2461F3" box="[844,1006,444,468]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
and these two species are no doubt confused in collections. Most specimens of
|
|
<taxonomicName id="2BE42983FF98FFAFD011D9BAFC6C61D3" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[805,934,477,500]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF98FFAFD011D9BAFC6C61D3" box="[805,934,477,500]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
are slightly larger (BL males =
|
|
<quantity id="2B1CFFE5FF98FFAFD3E3D99AFEBA6233" box="[215,368,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="23" pageNumber="474" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5 to 4.8 mm</quantity>
|
|
, females =
|
|
<quantity id="2B1CFFE5FF98FFAFD2D8D99AFD4F6232" box="[492,645,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.4" metricValueMin="4.6" pageId="23" pageNumber="474" unit="mm" value="5.0" valueMax="5.4" valueMin="4.6">4.6 to 5.4 mm</quantity>
|
|
) than those of
|
|
<taxonomicName id="2BE42983FF98FFAFD012D999FC0E6232" authorityName="Jian & Tian" authorityYear="2009" box="[806,964,509,533]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF98FFAFD012D999FC0E6232" box="[806,964,509,533]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
(BL males =
|
|
<quantity id="2B1CFFE5FF98FFAFD76CD99AFF1A6212" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.6" metricValueMin="3.9" pageId="23" pageNumber="474" unit="mm" value="4.25" valueMax="4.6" valueMin="3.9">3.9 to 4.6 mm</quantity>
|
|
, females =
|
|
<quantity id="2B1CFFE5FF98FFAFD279DA7AFE236212" box="[333,489,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.7" metricValueMin="4.2" pageId="23" pageNumber="474" unit="mm" value="4.45" valueMax="4.7" valueMin="4.2">4.2 to 4.7 mm</quantity>
|
|
). They have the pronotum (
|
|
<figureCitation id="74DF4E85FF98FFAFD027DA7AFCBB6212" box="[787,881,541,565]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="23" pageNumber="474">Fig. 13E</figureCitation>
|
|
) slightly shorter and wider (ratio PWM/PL = 1.52 to 1.70), widest at or near midlength and posterior to insertion of midlateral seta and not or only very slightly narrowed basally (ratio PWB/PWA = 1.15 to 1.25), and with the lateral margins slightly and evenly convex or straight or very slightly sinuate anterior to the basal angles. In
|
|
<taxonomicName id="2BE42983FF98FFAFD206DAF8FE196291" authorityName="Jian & Tian" authorityYear="2009" box="[306,467,670,694]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF98FFAFD206DAF8FE196291" box="[306,467,670,694]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
, the pronotum (
|
|
<figureCitation id="74DF4E85FF98FFAFD1BDDAF9FD236291" box="[649,745,670,694]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="23" pageNumber="474">Fig. 13F</figureCitation>
|
|
) is slightly longer and narrower (ratio PWM/PL = 1.48 to 1.57), widest anterior to mid-length at or near insertion of midlateral seta, slightly to moderately narrowed basally (ratio PWB/PWA = 1.04 to 1.14), and with the lateral margins slightly to moderately sinuate anterior to the basal angles. Unfortunately, the ranges in size and pronotal proportions (except for ratio PWB/PWA) seen among specimens of these two species overlap slightly, so measurements alone cannot distinguish all specimens of the two. The male genitalia are also very similar, except that the apical lamella of the median lobe is slightly shorter and broader in
|
|
<taxonomicName id="2BE42983FF98FFAFD215DBE7FE6F63B0" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[289,421,896,919]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF98FFAFD215DBE7FE6F63B0" box="[289,421,896,919]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
males (
|
|
<figureCitation id="74DF4E85FF98FFAFD2C8DB18FD4B63B0" box="[508,641,895,919]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="23" pageNumber="474">Figs. 19C,D</figureCitation>
|
|
) than in
|
|
<taxonomicName id="2BE42983FF98FFAFD1D2DBE7FC4F63B0" authorityName="Jian & Tian" authorityYear="2009" box="[742,901,895,919]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF98FFAFD1D2DBE7FC4F63B0" box="[742,901,895,919]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
(Figs. E,F) males. There are also subtle differences in the convexity of the elytral intervals (flat in
|
|
<taxonomicName id="2BE42983FF98FFAFD0BADBC7FBDB6390" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[910,1041,928,951]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF98FFAFD0BADBC7FBDB6390" box="[910,1041,928,951]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
and slightly convex in
|
|
<taxonomicName id="2BE42983FF98FFAFD3D8DBA6FE4063FF" authorityName="Jian & Tian" authorityYear="2009" box="[236,394,960,984]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF98FFAFD3D8DBA6FE4063FF" box="[236,394,960,984]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
) and depth of impression of the striae (moderately deep in
|
|
<taxonomicName id="2BE42983FF98FFAFD738DBA6FB4563FF" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[1036,1167,961,984]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF98FFAFD738DBA6FB4563FF" box="[1036,1167,961,984]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
, even deeper in
|
|
<taxonomicName id="2BE42983FF98FFAFD22BDB86FE7763DF" authorityName="Jian & Tian" authorityYear="2009" box="[287,445,992,1016]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF98FFAFD22BDB86FE7763DF" box="[287,445,992,1016]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF98FFAFD39FDC66FD8864BE" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
|
|
<emphasis id="DE908E12FF98FFAFD39FDC66FE5D643F" bold="true" box="[171,407,1025,1048]" pageId="23" pageNumber="474">Habitat distribution.</emphasis>
|
|
Members of this species were found at night, running on the surface of the open sandy shores of the Longchuan River (
|
|
<figureCitation id="74DF4E85FF98FFAFD1B2DC47FD1C641F" box="[646,726,1056,1080]" captionStart="FIGURE" captionStartId="38.[155,167,1623,1642]" captionTargetBox="[136,1161,216,1607]" captionTargetId="figure-10@38.[136,1161,216,1607]" captionTargetPageId="38" captionText="FIGURE 21. Photographs of habitat for cyclosomine species, Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov., Tetragonoderus elegans Andrewes, T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., and T. punctatus Wiedemann at Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China. A. lower river bank; B. sandy upper bank with sparse vegetation. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067337" httpUri="https://zenodo.org/record/11067337/files/figure.png" pageId="23" pageNumber="474">Fig. 21</figureCitation>
|
|
), a medium-sized stream, at two sites (at elevations of 1215 and
|
|
<quantity id="2B1CFFE5FF98FFAFD241DC27FE02647F" box="[373,456,1088,1112]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="23" pageNumber="474" unit="m" value="1230.0">1230 m</quantity>
|
|
, respectively). They were running on the same beaches with adults of
|
|
<taxonomicName id="2BE42983FF98FFAFD3ADDC05FECC645E" box="[153,262,1121,1145]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF98FFAFD3ADDC05FECC645E" box="[153,262,1121,1145]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
, which were much more abundant. They appear to prefer upper beach areas, where zones of slightly moist and dry sand meet.
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF98FFAFD39FDCC5FC0E657D" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
|
|
<emphasis id="DE908E12FF98FFAFD39FDCC5FD18649E" bold="true" box="[171,722,1186,1209]" pageId="23" pageNumber="474">Geographical distribution within the Gaoligong</emphasis>
|
|
<emphasis id="DE908E12FF98FFAFD1EFDCC5FCD6649E" bold="true" box="[731,796,1186,1209]" pageId="23" pageNumber="474">
|
|
<collectingRegion id="2E209CE2FF98FFAFD1EFDCC5FCDD649E" box="[731,791,1186,1209]" country="Myanmar" name="Shan" pageId="23" pageNumber="474">Shan</collectingRegion>
|
|
.
|
|
</emphasis>
|
|
<figureCitation id="74DF4E85FF98FFAFD011DCC6FC42649E" box="[805,904,1185,1209]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="23" pageNumber="474">Fig. 12B</figureCitation>
|
|
.
|
|
<materialsCitation id="5C8C585DFF98FFAFD0A0DCC5FBFB653D" ID-GBIF-Occurrence="4902827310" collectingDate="2003-10-28" collectingDateMax="2006-06-03" collectingDateMin="2003-10-28" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh & L. Brett & H. B. Liang & D. Z. Dong & Longchuan Jiang & Longjiang Bridge & X. C. Shi" county="Tengchong County" elevation="1230" location="Wuhe Township" municipality="We" pageId="23" pageNumber="474" specimenCount="15" specimenCount-female="8" specimenCount-male="2" stateProvince="Shan">
|
|
<collectingMunicipality id="0C3FC87AFF98FFAFD0A0DCC5FC7D649E" box="[916,951,1186,1209]" pageId="23" pageNumber="474">We</collectingMunicipality>
|
|
examined a total of
|
|
<specimenCount id="FAE29989FF98FFAFD34FDCA6FED764FE" box="[123,285,1217,1241]" count="5" pageId="23" pageNumber="474" type="generic">five specimens</specimenCount>
|
|
(
|
|
<specimenCount id="FAE29989FF98FFAFD21BDCA6FE5F64FE" box="[303,405,1217,1241]" count="1" pageId="23" pageNumber="474" type="male">one male</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF98FFAFD2E4DCA6FD9164FE" box="[464,603,1217,1241]" count="4" pageId="23" pageNumber="474" type="female">four females</specimenCount>
|
|
) from the following localities:
|
|
<emphasis id="DE908E12FF98FFAFD08CDCA5FB5F64FD" bold="true" box="[952,1173,1218,1242]" italics="true" pageId="23" pageNumber="474">
|
|
<collectingCounty id="053A2A8CFF98FFAFD08CDCA5FB4464FD" box="[952,1166,1218,1242]" pageId="23" pageNumber="474">Tengchong County</collectingCounty>
|
|
:
|
|
</emphasis>
|
|
<location id="E93B04DBFF98FFAFD34FDC86FEE164DE" LSID="urn:lsid:plazi:treatment:644DE316FF9AFFA0D34FD95BFF6861B4:E93B04DBFF98FFAFD34FDC86FEE164DE" box="[123,299,1249,1273]" county="Tengchong County" municipality="We" name="Wuhe Township" pageId="23" pageNumber="474" stateProvince="Shan">Wuhe Township</location>
|
|
(
|
|
<location id="E93B04DBFF98FFAFD20FDC86FE3C64DE" LSID="urn:lsid:plazi:treatment:644DE316FF9AFFA0D34FD95BFF6861B4:E93B04DBFF98FFAFD20FDC86FE3C64DE" box="[315,502,1249,1273]" county="Tengchong County" municipality="We" name="Longchuan Jiang" pageId="23" pageNumber="474" stateProvince="Shan">Longchuan Jiang</location>
|
|
just below bridge at
|
|
<location id="E93B04DBFF98FFAFD1D6DC86FC5764DE" LSID="urn:lsid:plazi:treatment:644DE316FF9AFFA0D34FD95BFF6861B4:E93B04DBFF98FFAFD1D6DC86FC5764DE" box="[738,925,1249,1273]" county="Tengchong County" municipality="We" name="Menglian village" pageId="23" pageNumber="474" stateProvince="Shan">Menglian village</location>
|
|
, 24.89176°/98.67551°,
|
|
<quantity id="2B1CFFE5FF98FFAFD34FDD65FF04653D" box="[123,206,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="23" pageNumber="474" unit="m" value="1230.0">
|
|
<elevation id="67C9B533FF98FFAFD34FDD65FF04653D" box="[123,206,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="23" pageNumber="474" unit="m" value="1230.0">1230 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF98FFAFD3EEDD65FE95653D" box="[218,351,1282,1306]" pageId="23" pageNumber="474" value="2006-06-03">
|
|
<collectingDate id="881E8D28FF98FFAFD3EEDD65FE95653D" box="[218,351,1282,1306]" pageId="23" pageNumber="474" value="2006-06-03">3 June 2006</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF98FFAFD25EDD64FDEB653D" box="[362,545,1282,1306]" pageId="23" pageNumber="474">D.H. Kavanaugh</collectorName>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF98FFAFD118DD64FD8B653D" box="[556,577,1283,1306]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="474">R</collectionCode>
|
|
.
|
|
<collectorName id="411137D6FF98FFAFD171DD64FD51653D" box="[581,667,1283,1306]" pageId="23" pageNumber="474">L. Brett</collectorName>
|
|
,
|
|
<collectorName id="411137D6FF98FFAFD192DD64FCD4653D" box="[678,798,1282,1306]" pageId="23" pageNumber="474">H.B. Liang</collectorName>
|
|
&
|
|
<collectorName id="411137D6FF98FFAFD075DD64FC7E653C" box="[833,948,1283,1307]" pageId="23" pageNumber="474">D.Z. Dong</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF98FFAFD705DD64FCDF657D" ID-GBIF-Occurrence="4902827331" collectingDate="2003-10-28" collectingDateMax="2006-06-03" collectingDateMin="2003-10-28" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh & L. Brett & H. B. Liang & D. Z. Dong & Longchuan Jiang & Longjiang Bridge & X. C. Shi" county="Tengchong County" elevation="1230" location="Wuhe Township" municipality="We" pageId="23" pageNumber="474" specimenCount="15" specimenCount-female="8" specimenCount-male="2" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF98FFAFD705DD64FB5C653D" box="[1073,1174,1282,1306]" count="1" pageId="23" pageNumber="474" type="male">one male</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF98FFAFD398DD45FE89651D" box="[172,323,1314,1338]" count="3" pageId="23" pageNumber="474" type="female">three females</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF98FFAFD267DD45FE43651D" box="[339,393,1314,1338]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="474" type="Museum">CAS</collectionCode>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF98FFAFD2ACDD45FE00651D" box="[408,458,1314,1338]" pageId="23" pageNumber="474">IOZ</collectionCode>
|
|
]), (
|
|
<collectorName id="411137D6FF98FFAFD2C4DD45FD67651D" box="[496,685,1314,1338]" pageId="23" pageNumber="474">Longchuan Jiang</collectorName>
|
|
at
|
|
<collectorName id="411137D6FF98FFAFD1E3DD45FC50651D" box="[727,922,1314,1338]" pageId="23" pageNumber="474">Longjiang Bridge</collectorName>
|
|
, 24.89889°/98.66667°,
|
|
<quantity id="2B1CFFE5FF98FFAFD34FDD25FF04657D" box="[123,206,1346,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="23" pageNumber="474" unit="m" value="1215.0">
|
|
<elevation id="67C9B533FF98FFAFD34FDD25FF04657D" box="[123,206,1346,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="23" pageNumber="474" unit="m" value="1215.0">1215 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF98FFAFD3EEDD25FE5B657D" box="[218,401,1346,1370]" pageId="23" pageNumber="474" value="2003-10-28">
|
|
<collectingDate id="881E8D28FF98FFAFD3EEDD25FE5B657D" box="[218,401,1346,1370]" pageId="23" pageNumber="474" value="2003-10-28">28 October 2003</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF98FFAFD2A9DD24FDDE657D" box="[413,532,1346,1370]" pageId="23" pageNumber="474">H.B. Liang</collectorName>
|
|
&
|
|
<collectorName id="411137D6FF98FFAFD10CDD25FD5C657D" box="[568,662,1346,1370]" pageId="23" pageNumber="474">X.C. Shi</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF98FFAFD021DD25FC0E657D" ID-GBIF-Occurrence="4902827315" box="[789,964,1346,1370]" collectingDate="2003-10-28" collectingDateMax="2006-06-03" collectingDateMin="2003-10-28" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh & L. Brett & H. B. Liang & D. Z. Dong & Longchuan Jiang & Longjiang Bridge & X. C. Shi" county="Tengchong County" elevation="1230" location="Wuhe Township" municipality="We" pageId="23" pageNumber="474" specimenCount="15" specimenCount-female="8" specimenCount-male="2" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF98FFAFD021DD25FCBF657D" box="[789,885,1346,1370]" count="1" pageId="23" pageNumber="474" type="female">1 female</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF98FFAFD0B4DD25FC73657D" box="[896,953,1346,1370]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="474" type="Museum">CAS</collectionCode>
|
|
].
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF98FFAFD39FDD05FEDF65BC" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">At present, this species has been recorded only from the southwestern part of the study area (Core Area 6).</paragraph>
|
|
<paragraph id="EC5B5200FF98FFAFD39FDDC3FE7B665B" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
|
|
<emphasis id="DE908E12FF98FFAFD39FDDC3FDE7659C" bold="true" box="[171,557,1444,1467]" pageId="23" pageNumber="474">Overall geographical distribution.</emphasis>
|
|
<figureCitation id="74DF4E85FF98FFAFD106DDC4FD4A659C" box="[562,640,1443,1467]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="23" pageNumber="474">Fig. 23</figureCitation>
|
|
. We have examined specimens of this species only from “Bengala” (i.e.,
|
|
<collectingRegion id="2E209CE2FF98FFAFD251DDA3FE2465FC" box="[357,494,1475,1499]" country="India" name="West Bengal" pageId="23" pageNumber="474">West Bengal</collectingRegion>
|
|
(
|
|
<collectingCountry id="94F31290FF98FFAFD2C9DDA4FDFD65FC" box="[509,567,1475,1499]" name="India" pageId="23" pageNumber="474">India</collectingCountry>
|
|
) and/or
|
|
<collectingCountry id="94F31290FF98FFAFD1A6DDA4FCDA65FC" box="[658,784,1475,1499]" name="Bangladesh" pageId="23" pageNumber="474">Bangladesh</collectingCountry>
|
|
) and the localities in the Gaoligong Shan as listed above. Its occurrence in the study area represents the eastern limit of its known geographical range and also a first record for
|
|
<collectingCountry id="94F31290FF98FFAFD14CDE64FD71663C" box="[632,699,1539,1563]" name="China" pageId="23" pageNumber="474">China</collectingCountry>
|
|
. It is likely that this species occurs also in northern
|
|
<collectingCountry id="94F31290FF98FFAFD3EFDE43FE89661B" box="[219,323,1572,1596]" name="Myanmar" pageId="23" pageNumber="474">Myanmar</collectingCountry>
|
|
and northeastern
|
|
<collectingCountry id="94F31290FF98FFAFD135DE43FDF2661B" box="[513,568,1572,1596]" name="India" pageId="23" pageNumber="474">India</collectingCountry>
|
|
along rivers draining the western slope of the Gaoligong Shan and the southern slope of the eastern part of the Himalaya Mountains, respectively, at low elevations (
|
|
<quantity id="2B1CFFE5FF98FFAFD3C2DE03FE6E665B" box="[246,420,1636,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="23" pageNumber="474" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">1000 to 1300 m</quantity>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF98FFA0D39FDEE2FF6861B4" blockId="23.[123,1174,219,1725]" lastBlockId="24.[123,1174,219,403]" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474">
|
|
<emphasis id="DE908E12FF98FFAFD39FDEE2FCB766BB" bold="true" box="[171,893,1669,1693]" pageId="23" pageNumber="474">
|
|
Geographical relationships with other
|
|
<taxonomicName id="2BE42983FF98FFAFD144DEE2FCB766BB" box="[624,893,1669,1693]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="undetermined">
|
|
<emphasis id="DE908E12FF98FFAFD144DEE2FCD666BA" bold="true" box="[624,796,1669,1693]" italics="true" pageId="23" pageNumber="474">Tetragonoderus</emphasis>
|
|
species.
|
|
</taxonomicName>
|
|
</emphasis>
|
|
Members of this species have been found synotopic with those of
|
|
<taxonomicName id="2BE42983FF98FFAFD102DEC1FD68669B" box="[566,674,1700,1725]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF98FFAFD102DEC1FD68669B" box="[566,674,1700,1725]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="2BE42983FF98FFAFD1ECDEC1FCAE669B" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[728,868,1700,1725]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="parviculus">
|
|
<emphasis id="DE908E12FF98FFAFD1ECDEC1FCAE669B" box="[728,868,1700,1725]" italics="true" pageId="23" pageNumber="474">T. parviculus</emphasis>
|
|
</taxonomicName>
|
|
in the study area. The range of this species also overlaps that of
|
|
<taxonomicName id="2BE42983FF97FFA0D2C0D8BBFDA060D4" box="[500,618,220,243]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="arcuatus">
|
|
<emphasis id="DE908E12FF97FFA0D2C0D8BBFDA060D4" box="[500,618,220,243]" italics="true" pageId="24" pageNumber="475">T. arcuatus</emphasis>
|
|
</taxonomicName>
|
|
in the study area, but these two species have not yet been found syntopic. Also, the range of
|
|
<taxonomicName id="2BE42983FF97FFA0D11BD89BFD046134" authorityName="Jian & Tian" authorityYear="2009" box="[559,718,251,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF97FFA0D11BD89BFD046134" box="[559,718,251,275]" italics="true" pageId="24" pageNumber="475">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
broadly overlaps that of
|
|
<taxonomicName id="2BE42983FF97FFA0D0D5D89BFBAF6134" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[993,1125,252,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF97FFA0D0D5D89BFBAF6134" box="[993,1125,252,275]" italics="true" pageId="24" pageNumber="475">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
, but the former species has not yet been recorded from the study area. We note that all current records of
|
|
<taxonomicName id="2BE42983FF97FFA0D3A3D95BFEFE6174" authorityName="Jian & Tian" authorityYear="2009" box="[151,308,315,339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF97FFA0D3A3D95BFEFE6174" box="[151,308,315,339]" italics="true" pageId="24" pageNumber="475">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
are from localities well below
|
|
<quantity id="2B1CFFE5FF97FFA0D148D95CFD066174" box="[636,716,315,339]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="475" unit="m" value="1000.0">1000 m</quantity>
|
|
in elevation and such elevations are absent from all but the extreme southwesternmost part of the study area, which we were unable to sample.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |