484 lines
71 KiB
XML
484 lines
71 KiB
XML
<document id="D6ECF0824555675524BE8D835E285E15" ID-CLB-Dataset="299409" ID-DOI="10.5281/zenodo.11067293" ID-GBIF-Dataset="9235db45-04c4-4570-8987-cb84765763a4" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067293" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052207303" checkinUser="felipe" docAuthor="Kavanaugh, David H., Cueva-Dabkoski, Mollie & Liang, Hongbin" docDate="2023" docId="644DE316FF87FFB4D34FDCBCFDF06134" docLanguage="en" docName="ProcCalifAcadSci.67.20.451-491.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (20)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="462" masterDocId="98749B6EFF8FFFB8D334D867FFCA6027" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species." masterLastPageNumber="491" masterPageNumber="451" pageNumber="459" updateTime="1719954462017" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="7642E4D9E5A4389A5457037EB8348344">
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<mods:title id="15F2599190D8A461264167897D714360">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:name id="48C6C9FA444349C368BFC076BCBD07D2" type="personal">
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<mods:roleTerm id="5756079C2615F573532FA17B106C7235">Author</mods:roleTerm>
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<mods:namePart id="9EDD184CD458D96079B564495AEF2089">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="F666A8288519B8D3EC20ED4F906F889C">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:name id="63B533D8A07397452479B929A6EADC88" type="personal">
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<mods:roleTerm id="FFA47F560C321C13C2744614B548B813">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="01A91B975C9A3DBB9D7AA24DB4560250">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="7B49EF2EA3ADC57691BC5B35100D24F9">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:name id="6608BF269FE6D9203727FA9AE640F864" type="personal">
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<mods:roleTerm id="A53088A66461020B92F343233FD90529">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A301694581CD315D7146331360F23E16">Liang, Hongbin</mods:namePart>
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<mods:affiliation id="D2B55D6FF7FFA5810EEC73CC61F2AEA5">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
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<mods:typeOfResource id="2F4C040A767CD6058840A5A2D7E0C799">text</mods:typeOfResource>
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<mods:title id="E4D83C963588F362780EE7A5F8875F99">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="CB5A915BF1FE082094C26BD4D47C9F67">
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<mods:date id="FFDD1BD1F8CED7EE89EA77A3B8E7C9AA">2023</mods:date>
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<mods:number id="1E69A40A64F57CEDE352DF4DD40C5ACF">2023-02-15</mods:number>
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<mods:number id="3E64DAB3EC806E63E6E74F6AB679A922">67</mods:number>
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<treatment id="644DE316FF87FFB4D34FDCBCFDF06134" ID-DOI="http://doi.org/10.5281/zenodo.12628248" ID-GBIF-Taxon="234412263" ID-Zenodo-Dep="12628248" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134" httpUri="http://treatment.plazi.org/id/644DE316FF87FFB4D34FDCBCFDF06134" lastPageId="12" lastPageNumber="462" pageId="8" pageNumber="459">
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<subSubSection id="A4FE018BFF87FFB0D34FDCBCFD556536" pageId="8" pageNumber="459" type="nomenclature">
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<paragraph id="EC5B5200FF87FFB0D34FDCBCFCBF64D4" blockId="8.[123,885,1243,1297]" box="[123,885,1243,1267]" pageId="8" pageNumber="459">
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<heading id="B713E56CFF87FFB0D34FDCBCFCBF64D4" box="[123,885,1243,1267]" fontSize="10" level="2" pageId="8" pageNumber="459" reason="4">
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<emphasis id="DE908E12FF87FFB0D34FDCBCFCBF64D4" bold="true" box="[123,885,1243,1267]" pageId="8" pageNumber="459">
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<taxonomicName id="2BE42983FF87FFB0D34FDCBCFCD864D4" authority="Kavanaugh and Cueva-Dabkoski" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[123,786,1243,1267]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
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<emphasis id="DE908E12FF87FFB0D34FDCBCFE4464D4" bold="true" box="[123,398,1243,1267]" italics="true" pageId="8" pageNumber="459">Cyclosomus acutangulus</emphasis>
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Kavanaugh and Cueva-Dabkoski
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</taxonomicName>
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,
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<taxonomicNameLabel id="C5A33369FF87FFB0D02BDCBBFCBF64D4" box="[799,885,1244,1267]" pageId="8" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="EC5B5200FF87FFB0D34FDC9CFD556536" blockId="8.[123,885,1243,1297]" box="[123,671,1275,1297]" pageId="8" pageNumber="459">
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<figureCitation id="74DF4E85FF87FFB0D34FDC9CFF236536" box="[123,233,1275,1297]" captionStart="FIGURE" captionStartId="5.[686,698,839,858]" captionTargetBox="[666,1158,216,826]" captionTargetId="figure-473@5.[652,1176,216,826]" captionText="FIGURE 5. Left hind tibial medial spur and tarsomere 1, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12628237" httpUri="https://zenodo.org/record/12628237/files/figure.png" pageId="8" pageNumber="459">Figures 5A</figureCitation>
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, 7,
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<figureCitation id="74DF4E85FF87FFB0D238DC9CFED36536" box="[268,281,1275,1297]" captionStart="FIGURE" captionStartId="11.[687,699,975,994]" captionTargetBox="[657,1156,383,944]" captionTargetId="figure-502@11.[652,1176,357,962]" captionText="FIGURE 8. Male genitalia, Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012642, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Dorsal aspect; B. Left lateral aspect; scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12628241" httpUri="https://zenodo.org/record/12628241/files/figure.png" pageId="8" pageNumber="459">8</figureCitation>
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, 13A, 14A, 15A, 16A, 17A, 21, and 23.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A4FE018BFF87FFB1D39FDD42FE8A6174" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="materials_examined">
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<paragraph id="EC5B5200FF87FFB1D39FDD42FCD26134" blockId="8.[123,1174,1316,1725]" lastBlockId="9.[123,1174,219,1725]" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
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<emphasis id="DE908E12FF87FFB0D39FDD42FE9A651B" bold="true" box="[171,336,1317,1340]" pageId="8" pageNumber="459">Type material</emphasis>
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.
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<materialsCitation id="5C8C585DFF87FFB0D251DD43FE32659B" ID-GBIF-Occurrence="4902827327" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ" collectorName="Liang H. - B & Shi X. - C" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1015370" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan Province" typeStatus="holotype">
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||
<typeStatus id="335FECA2FF87FFB0D251DD43FE3F651B" box="[357,501,1316,1340]" pageId="8" pageNumber="459" type="holotype">HOLOTYPE</typeStatus>
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||
, a
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<specimenCount id="FAE29989FF87FFB0D115DD43FD9D651B" box="[545,599,1316,1340]" pageId="8" pageNumber="459" type="male">male</specimenCount>
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||
, deposited in
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||
<collectionCode id="8AF5CAC5FF87FFB0D034DD43FCE5651B" box="[768,815,1316,1340]" pageId="8" pageNumber="459">IOZ</collectionCode>
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||
, labeled: “
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||
<specimenCode id="BC42FA7BFF87FFB0D099DD43FB4E651B" box="[941,1156,1316,1340]" pageId="8" pageNumber="459">CASENT 1015370</specimenCode>
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”/ “
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||
<collectingCountry id="94F31290FF87FFB0D3B0DD23FF17657B" box="[132,221,1348,1372]" name="China" pageId="8" pageNumber="459">CHINA</collectingCountry>
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||
,
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||
<collectingRegion id="2E209CE2FF87FFB0D3DEDD22FE6D657B" box="[234,423,1348,1372]" country="China" name="Yunnan" pageId="8" pageNumber="459">Yunnan Province</collectingRegion>
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,
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<collectingCounty id="053A2A8CFF87FFB0D287DD23FD91657B" box="[435,603,1348,1372]" pageId="8" pageNumber="459">Tengchong Co.</collectingCounty>
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||
,
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<collectingMunicipality id="0C3FC87AFF87FFB0D15CDD23FCD1657B" box="[616,795,1348,1372]" pageId="8" pageNumber="459">Wuhe Township</collectingMunicipality>
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,
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<location id="E93B04DBFF87FFB0D01CDD23FF79655B" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D01CDD23FF79655B" country="China" county="Tengchong Co." latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" name="Longjiang Bridge on Longchuanjiang" pageId="8" pageNumber="459" stateProvince="Yunnan Province">Longjiang Bridge on Longchuanjiang</location>
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,
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<quantity id="2B1CFFE5FF87FFB0D38ADD03FED8655B" box="[190,274,1380,1404]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="8" pageNumber="459" unit="m" value="1215.0">
|
||
<elevation id="67C9B533FF87FFB0D38ADD03FED8655B" box="[190,274,1380,1404]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="8" pageNumber="459" unit="m" value="1215.0">1215 m</elevation>
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</quantity>
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,”/ “
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<geoCoordinate id="89D034C7FF87FFB0D20EDD03FE77655B" box="[314,445,1380,1404]" degrees="24.89889" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="1" value="24.89889">N24.89889°</geoCoordinate>
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/
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||
<geoCoordinate id="89D034C7FF87FFB0D2F1DD03FD98655B" box="[453,594,1380,1404]" degrees="098.66667" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="1" value="98.66667">E098.66667°</geoCoordinate>
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<date id="985A74C0FF87FFB0D16EDD03FCDB655B" box="[602,785,1380,1404]" pageId="8" pageNumber="459" value="2003-10-30">
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<collectingDate id="881E8D28FF87FFB0D16EDD03FCDB655B" box="[602,785,1380,1404]" pageId="8" pageNumber="459" value="2003-10-30">30 October 2003</collectingDate>
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</date>
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, on beach,
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<collectorName id="411137D6FF87FFB0D0BFDD03FBC2655B" box="[907,1032,1380,1404]" pageId="8" pageNumber="459">Liang H.-B</collectorName>
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. &
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<collectorName id="411137D6FF87FFB0D71BDD03FB5B655B" box="[1071,1169,1380,1404]" pageId="8" pageNumber="459">Shi X.-C</collectorName>
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. collectors, LHB03-42”/ “HOLOTYPE
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<taxonomicName id="2BE42983FF87FFB0D118DDE3FCF165BB" authority="Kavanaugh & Cueva-Dabkoski & Liang, 2023" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[556,827,1412,1436]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
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<emphasis id="DE908E12FF87FFB0D118DDE3FCF165BB" box="[556,827,1412,1436]" italics="true" pageId="8" pageNumber="459">Cyclosomus acutangulus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C5A33369FF87FFB0D073DDE2FC4C65BB" box="[839,902,1413,1436]" pageId="8" pageNumber="459" rank="species">sp. n.</taxonomicNameLabel>
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D.H. Kavanaugh & M. Cueva-Dabkoski 2022” [red label].
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</materialsCitation>
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<materialsCitation id="5C8C585DFF87FFB0D2CBDDC3FECB65DB" ID-GBIF-Occurrence="4902827302" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="CAS" collectorName="D. H. Kavanaugh & M. Cueva-Dabkoski & P. Pacholatko & L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1012641"" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Yunnan" typeStatus="holotype">
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A total of 28 paratypes:
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<specimenCount id="FAE29989FF87FFB0D03FDDC3FCB1659B" box="[779,891,1444,1468]" count="5" pageId="8" pageNumber="459" type="male">five males</specimenCount>
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||
and
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<specimenCount id="FAE29989FF87FFB0D087DDC3FBFB659B" box="[947,1073,1444,1468]" count="6" pageId="8" pageNumber="459" type="female">six females</specimenCount>
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, in
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||
<collectionCode id="8AF5CAC5FF87FFB0D76FDDC3FB5B659B" box="[1115,1169,1444,1468]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="459" type="Museum">CAS</collectionCode>
|
||
, same label data as holotype, except first label [unique identifier label] different (“
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<specimenCode id="BC42FA7BFF87FFB0D71EDDA3FF2E65DA" pageId="8" pageNumber="459">CASENT 1012641”</specimenCode>
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||
to
|
||
</materialsCitation>
|
||
<materialsCitation id="5C8C585DFF87FFB0D23DDD82FDD665DA" ID-GBIF-Occurrence="4902827317" box="[265,540,1509,1533]" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ, CAS, R, NHMUK, BMNH" collectorName="D. H. Kavanaugh & M. Cueva-Dabkoski & P. Pacholatko & L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1012646" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan" typeStatus="holotype">
|
||
“
|
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<specimenCode id="BC42FA7BFF87FFB0D226DD82FE2E65DA" box="[274,484,1509,1533]" pageId="8" pageNumber="459">CASENT 1012646</specimenCode>
|
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” and
|
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</materialsCitation>
|
||
<materialsCitation id="5C8C585DFF87FFB0D110DD82FCEF65DB" ID-GBIF-Occurrence="4902827325" box="[548,805,1509,1533]" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ, CAS, R, NHMUK, BMNH" collectorName="D. H. Kavanaugh & M. Cueva-Dabkoski & P. Pacholatko & L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1015371" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan" typeStatus="holotype">
|
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“
|
||
<specimenCode id="BC42FA7BFF87FFB0D119DD82FD3465DA" box="[557,766,1509,1533]" pageId="8" pageNumber="459">CASENT 1015371</specimenCode>
|
||
” to
|
||
</materialsCitation>
|
||
<materialsCitation id="5C8C585DFF87FFB0D019DD82FF6A663A" ID-GBIF-Occurrence="4902827303" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ, CAS, R, NHMUK, BMNH" collectorName="D. H. Kavanaugh & M. Cueva-Dabkoski & P. Pacholatko & L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1015375" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan" typeStatus="holotype">
|
||
“
|
||
<specimenCode id="BC42FA7BFF87FFB0D003DD82FBC065DA" box="[823,1034,1509,1533]" pageId="8" pageNumber="459">CASENT 1015375</specimenCode>
|
||
”, respectively);
|
||
</materialsCitation>
|
||
<materialsCitation id="5C8C585DFF87FFB0D392DE62FBA2661A" ID-GBIF-Occurrence="4902827313" collectingDate="2015-03-22" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ" collectorName="Liu Y & Shi H. L" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longchuan river" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCount="13" specimenCount-female="7" specimenCount-male="6" stateProvince="Yunnan" typeStatus="holotype">
|
||
<specimenCount id="FAE29989FF87FFB0D392DE62FEC0663A" box="[166,266,1541,1565]" count="6" pageId="8" pageNumber="459" type="male">six males</specimenCount>
|
||
and
|
||
<specimenCount id="FAE29989FF87FFB0D20BDE62FE12663A" box="[319,472,1541,1565]" count="7" pageId="8" pageNumber="459" type="female">seven females</specimenCount>
|
||
, in
|
||
<collectionCode id="8AF5CAC5FF87FFB0D2CADE62FDE4663A" box="[510,558,1541,1565]" pageId="8" pageNumber="459">IOZ</collectionCode>
|
||
. labeled: “
|
||
<collectingCountry id="94F31290FF87FFB0D1A9DE62FD2B663A" box="[669,737,1541,1565]" name="China" pageId="8" pageNumber="459">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="2E209CE2FF87FFB0D1DFDE62FCF5663B" box="[747,831,1541,1564]" country="China" name="Yunnan" pageId="8" pageNumber="459">Yunnan</collectingRegion>
|
||
, Tengchong, Wuhe, on beach of
|
||
<location id="E93B04DBFF87FFB0D34FDE42FEE4661A" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D34FDE42FEE4661A" box="[123,302,1573,1597]" country="China" county="Tengchong Co." latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" name="Longchuan river" pageId="8" pageNumber="459" stateProvince="Yunnan">Longchuan river</location>
|
||
,
|
||
<geoCoordinate id="89D034C7FF87FFB0D20CDE42FE6F661A" box="[312,421,1573,1597]" degrees="24.8941" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="5" value="24.8941">N24.8941</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="89D034C7FF87FFB0D284DE42FDD6661A" box="[432,540,1573,1597]" degrees="98.6750" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="5" value="98.675">E98.6750</geoCoordinate>
|
||
,” / “
|
||
<quantity id="2B1CFFE5FF87FFB0D17DDE42FD54661A" box="[585,670,1573,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="8" pageNumber="459" unit="m" value="1215.0">
|
||
<elevation id="67C9B533FF87FFB0D17DDE42FD54661A" box="[585,670,1573,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="8" pageNumber="459" unit="m" value="1215.0">1215 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="985A74C0FF87FFB0D19DDE42FCDE661A" box="[681,788,1573,1597]" pageId="8" pageNumber="459" value="2015-03-22">
|
||
<collectingDate id="881E8D28FF87FFB0D19DDE42FCDE661A" box="[681,788,1573,1597]" pageId="8" pageNumber="459" value="2015-03-22">2015.3.22</collectingDate>
|
||
</date>
|
||
N,
|
||
<collectorName id="411137D6FF87FFB0D008DE42FCB6661A" box="[828,892,1573,1597]" pageId="8" pageNumber="459">Liu Y</collectorName>
|
||
.,
|
||
<collectorName id="411137D6FF87FFB0D0BFDE42FC28661A" box="[907,994,1573,1597]" pageId="8" pageNumber="459">Shi H.L</collectorName>
|
||
. collectors”;
|
||
</materialsCitation>
|
||
<materialsCitation id="5C8C585DFF87FFB0D75BDE41FBA2665A" ID-GBIF-Occurrence="4902827316" collectingDate="2006-06-03" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="CAS, IOZ" collectorName="Kavanaugh D & Brett R." country="China" county="Wuhe" elevation="1230" latitude="24.89889" location="Longjiangqiao" longLatPrecision="1" longitude="98.66667" municipality="Tengchong" pageId="8" pageNumber="459" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan Prov." typeStatus="holotype">
|
||
<specimenCount id="FAE29989FF87FFB0D75BDE41FF65667A" count="1" pageId="8" pageNumber="459" type="male">one male</specimenCount>
|
||
and
|
||
<specimenCount id="FAE29989FF87FFB0D3C4DE21FEBA667A" box="[240,368,1605,1629]" count="1" pageId="8" pageNumber="459" type="female">one female</specimenCount>
|
||
, in
|
||
<collectionCode id="8AF5CAC5FF87FFB0D290DE22FE12667A" box="[420,472,1605,1629]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="459" type="Museum">CAS</collectionCode>
|
||
and
|
||
<collectionCode id="8AF5CAC5FF87FFB0D12EDE22FD83667A" box="[538,585,1605,1629]" pageId="8" pageNumber="459">IOZ</collectionCode>
|
||
, labeled: “
|
||
<collectingCountry id="94F31290FF87FFB0D1F3DE22FCC1667A" box="[711,779,1605,1629]" name="China" pageId="8" pageNumber="459">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="2E209CE2FF87FFB0D028DE21FC7F667A" box="[796,949,1606,1629]" country="China" name="Yunnan" pageId="8" pageNumber="459">Yunnan Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="0C3FC87AFF87FFB0D0F2DE22FBF5667A" box="[966,1087,1605,1629]" pageId="8" pageNumber="459">Tengchong</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="053A2A8CFF87FFB0D764DE22FB5B667A" box="[1104,1169,1605,1629]" pageId="8" pageNumber="459">Wuhe</collectingCounty>
|
||
,
|
||
<location id="E93B04DBFF87FFB0D34FDE02FED3665A" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D34FDE02FED3665A" box="[123,281,1637,1661]" country="China" county="Wuhe" latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Tengchong" name="Longjiangqiao" pageId="8" pageNumber="459" stateProvince="Yunnan Prov.">Longjiangqiao</location>
|
||
, beach,
|
||
<geoCoordinate id="89D034C7FF87FFB0D244DE02FE38665A" box="[368,498,1637,1661]" degrees="24.89176" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="1" value="24.89176">24.89176°N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="89D034C7FF87FFB0D2CFDE02FDB2665A" box="[507,632,1637,1661]" degrees="98.67551" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="1" value="98.67551">98.67551°E</geoCoordinate>
|
||
/
|
||
<quantity id="2B1CFFE5FF87FFB0D1BADE02FD2B665A" box="[654,737,1637,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="8" pageNumber="459" unit="m" value="1230.0">
|
||
<elevation id="67C9B533FF87FFB0D1BADE02FD2B665A" box="[654,737,1637,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="8" pageNumber="459" unit="m" value="1230.0">1230 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="985A74C0FF87FFB0D1D9DE02FC87665A" box="[749,845,1637,1661]" pageId="8" pageNumber="459" value="2006-06-03">
|
||
<collectingDate id="881E8D28FF87FFB0D1D9DE02FC87665A" box="[749,845,1637,1661]" pageId="8" pageNumber="459" value="2006-06-03">2006.6.3</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="411137D6FF87FFB0D06CDE02FC3A665A" box="[856,1008,1637,1661]" pageId="8" pageNumber="459">Kavanaugh D</collectorName>
|
||
.,
|
||
<collectorName id="411137D6FF87FFB0D734DE01FB9D665A" box="[1024,1111,1637,1661]" pageId="8" pageNumber="459">Brett R.</collectorName>
|
||
”;
|
||
</materialsCitation>
|
||
<materialsCitation id="5C8C585DFF87FFB1D75BDE01FCD26134" ID-GBIF-Occurrence="4902827314" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="NHMUK, BMNH" collectorName="P. Pacholatko & L. Dembicky" country="India" county="Tengchong Co." elevation="150" lastPageId="9" lastPageNumber="460" latitude="24.89889" location="Bhalukpong" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="ASSAM" typeStatus="holotype">
|
||
<specimenCount id="FAE29989FF87FFB0D75BDE01FF6566BA" count="1" pageId="8" pageNumber="459" type="male">one male</specimenCount>
|
||
and
|
||
<specimenCount id="FAE29989FF87FFB0D3D5DEE1FE9266BA" box="[225,344,1669,1693]" count="1" pageId="8" pageNumber="459" type="female">one female</specimenCount>
|
||
, both in
|
||
<collectionCode id="8AF5CAC5FF87FFB0D287DEE1FDD166BA" box="[435,539,1670,1693]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="459" type="Museum">NHMUK</collectionCode>
|
||
, labeled: “NE
|
||
<collectingCountry id="94F31290FF87FFB0D180DEE2FD2766BA" box="[692,749,1669,1693]" name="India" pageId="8" pageNumber="459">India</collectingCountry>
|
||
,
|
||
<collectingRegion id="2E209CE2FF87FFB0D1C2DEE2FC9F66BA" box="[758,853,1669,1693]" country="India" name="Assam" pageId="8" pageNumber="459">ASSAM</collectingRegion>
|
||
,
|
||
<location id="E93B04DBFF87FFB0D06ADEE2FC2966BA" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D06ADEE2FC2966BA" box="[862,995,1669,1693]" country="India" county="Tengchong Co." latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" name="Bhalukpong" pageId="8" pageNumber="459" stateProvince="ASSAM">Bhalukpong</location>
|
||
,
|
||
<date id="985A74C0FF87FFB0D0D9DEE2FB5866BA" box="[1005,1170,1669,1693]" pageId="8" pageNumber="459" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">26.v.-3.vi.2006</date>
|
||
.
|
||
<geoCoordinate id="89D034C7FF87FFB0D34FDEC2FF14669A" box="[123,222,1701,1725]" degrees="27" direction="north" minutes="02" orientation="latitude" pageId="8" pageNumber="459" precision="925" value="27.033333">27° 02’N</geoCoordinate>
|
||
<geoCoordinate id="89D034C7FF87FFB0D3D2DEC2FE82669A" box="[230,328,1701,1725]" degrees="92" direction="east" minutes="35" orientation="longitude" pageId="8" pageNumber="459" precision="925" value="92.583336">92° 35’E</geoCoordinate>
|
||
,
|
||
<quantity id="2B1CFFE5FF87FFB0D261DEC2FE5E669A" box="[341,404,1701,1725]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="459" unit="m" value="150.0">
|
||
<elevation id="67C9B533FF87FFB0D261DEC2FE5E669A" box="[341,404,1701,1725]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="459" unit="m" value="150.0">150m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="411137D6FF87FFB0D2ABDEC1FDFA669A" box="[415,560,1701,1725]" pageId="8" pageNumber="459">P. Pacholátko</collectorName>
|
||
leg.”/ “
|
||
<collectorName id="411137D6FF87FFB0D1B1DEC2FCD9669A" box="[645,787,1701,1725]" pageId="8" pageNumber="459">L. Dembicky</collectorName>
|
||
&
|
||
<collectorName id="411137D6FF87FFB0D002DEC1FC0D669A" box="[822,967,1701,1725]" pageId="8" pageNumber="459">P. Pacholátko</collectorName>
|
||
<collectionCode id="8AF5CAC5FF87FFB0D0FADEC1FBD4669A" box="[974,1054,1702,1725]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="8" pageNumber="459">BMNH</collectionCode>
|
||
{E} 2006- 48” Each paratype also bears the following label: “PARATYPE
|
||
<taxonomicName id="2BE42983FF86FFB1D076D8BCFB8560D4" authority="Kavanaugh & Cueva-Dabkoski & Liang, 2023" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[834,1103,219,243]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
|
||
<emphasis id="DE908E12FF86FFB1D076D8BCFB8560D4" box="[834,1103,219,243]" italics="true" pageId="9" pageNumber="460">Cyclosomus acutangulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C5A33369FF86FFB1D76DD8BBFB5F60D4" box="[1113,1173,220,243]" pageId="9" pageNumber="460" rank="species">sp. n.</taxonomicNameLabel>
|
||
D.H. Kavanaugh & M. Cueva-Dabkoski 2022” [yellow label].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF86FFB1D39FD97BFE8A6174" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
|
||
<emphasis id="DE908E12FF86FFB1D39FD97BFE826114" bold="true" box="[171,328,284,307]" pageId="9" pageNumber="460">
|
||
<typeStatus id="335FECA2FF86FFB1D39FD97BFF296114" box="[171,227,284,307]" pageId="9" pageNumber="460">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="94F31290FF86FFB1D260D97CFE5C6114" box="[340,406,283,307]" name="China" pageId="9" pageNumber="460">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="2E209CE2FF86FFB1D293D97BFE316114" box="[423,507,284,307]" country="China" name="Yunnan" pageId="9" pageNumber="460">Yunnan</collectingRegion>
|
||
,
|
||
<collectingCounty id="053A2A8CFF86FFB1D13FD97CFD146114" box="[523,734,283,307]" pageId="9" pageNumber="460">Tengchong County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="0C3FC87AFF86FFB1D1DAD97CFC6F6114" box="[750,933,283,307]" pageId="9" pageNumber="460">Wuhe Township</collectingMunicipality>
|
||
,
|
||
<location id="E93B04DBFF86FFB1D081D97CFBBF6114" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF86FFB1D081D97CFBBF6114" box="[949,1141,283,307]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longchuan River" pageId="9" pageNumber="460" stateProvince="Yunnan">Longchuan River</location>
|
||
at
|
||
<location id="E93B04DBFF86FFB1D34FD95CFEF66174" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF86FFB1D34FD95CFEF66174" box="[123,316,315,339]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longjiang Bridge" pageId="9" pageNumber="460" stateProvince="Yunnan">Longjiang Bridge</location>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4FE018BFF86FFB1D39FD93BFDEF61F3" pageId="9" pageNumber="460" type="etymology">
|
||
<paragraph id="EC5B5200FF86FFB1D39FD93BFDEF61F3" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
|
||
<emphasis id="DE908E12FF86FFB1D39FD93BFEE16154" bold="true" box="[171,299,348,371]" pageId="9" pageNumber="460">Etymology.</emphasis>
|
||
The species epithet,
|
||
<taxonomicName id="2BE42983FF86FFB1D126D93BFD5E6153" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[530,660,348,372]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="DE908E12FF86FFB1D126D93BFD5E6153" box="[530,660,348,372]" italics="true" pageId="9" pageNumber="460">acutangulus</emphasis>
|
||
</taxonomicName>
|
||
, is a combination of the Latin adjective,
|
||
<emphasis id="DE908E12FF86FFB1D77DD93AFB456153" box="[1097,1167,349,372]" italics="true" pageId="9" pageNumber="460">acutus</emphasis>
|
||
, meaning sharp or pointed, and the Latin noun,
|
||
<emphasis id="DE908E12FF86FFB1D14AD91BFD1861B3" box="[638,722,380,404]" italics="true" pageId="9" pageNumber="460">angulus</emphasis>
|
||
, meaning corner or angle, here used as a noun in apposition. It refers to the apical angles of the pronotum, which are exceptionally narrow and pointed in members of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4FE018BFF86FFB1D39FD9BAFDDF62D1" pageId="9" pageNumber="460" type="diagnosis">
|
||
<paragraph id="EC5B5200FF86FFB1D39FD9BAFDDF62D1" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
|
||
<emphasis id="DE908E12FF86FFB1D39FD9BAFED561D3" bold="true" box="[171,287,477,500]" pageId="9" pageNumber="460">Diagnosis.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="2BE42983FF86FFB1D2A5D9BAFDF861D3" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[401,562,476,500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="DE908E12FF86FFB1D2A5D9BAFDF861D3" box="[401,562,476,500]" italics="true" pageId="9" pageNumber="460">C. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from those of all other
|
||
<taxonomicName id="2BE42983FF86FFB1D720D9BBFF026232" pageId="9" pageNumber="460">
|
||
<emphasis id="DE908E12FF86FFB1D720D9BBFB5F61D3" box="[1044,1173,476,500]" italics="true" pageId="9" pageNumber="460">Cyclosomus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in Asia by the following combination of character states: Dorsal habitus as in
|
||
<figureCitation id="74DF4E85FF86FFB1D70DD99AFB5B6232" box="[1081,1169,509,533]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="9" pageNumber="460">Fig. 7A</figureCitation>
|
||
; pronotum (
|
||
<figureCitation id="74DF4E85FF86FFB1D3C6DA7AFE986212" box="[242,338,541,565]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="9" pageNumber="460">Fig. 13A</figureCitation>
|
||
) with anterior angles acute, narrow, lateral margins slightly to distinctly sinuate near anterior angles, lateral pale bands very narrow. Members of this species can be distinguished from those of other cyclosomines in the study area by the following combination of character states: Size larger, BL = more than
|
||
<quantity id="2B1CFFE5FF86FFB1D175DA19FD5362B2" box="[577,665,638,662]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="460" unit="mm" value="6.8">6.8 mm</quantity>
|
||
; pronotum (
|
||
<figureCitation id="74DF4E85FF86FFB1D02BDA19FC4B62B1" box="[799,897,638,662]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="9" pageNumber="460">Fig. 13A</figureCitation>
|
||
) very broad, trapezoidal, almost as wide as elytra at humeral angles, distinctly widest at base; elytra broadly rounded apically, elytral color pattern as in
|
||
<figureCitation id="74DF4E85FF86FFB1D2FFDAD9FDD562F1" box="[459,543,702,726]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="9" pageNumber="460">Fig. 7A</figureCitation>
|
||
; front tarsomeres 1 to 3 markedly expanded laterally (
|
||
<figureCitation id="74DF4E85FF86FFB1D754DAD9FF6062D1" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="9" pageNumber="460">Figs. 14A</figureCitation>
|
||
,
|
||
<figureCitation id="74DF4E85FF86FFB1D383DAB9FF2F62D1" box="[183,229,734,758]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="9" pageNumber="460">15A</figureCitation>
|
||
) in both males and females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4FE018BFF86FFB4D39FDA98FDF06134" lastPageId="12" lastPageNumber="463" pageId="9" pageNumber="460" type="description">
|
||
<paragraph id="EC5B5200FF86FFB1D39FDA98FF1E6311" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
|
||
<emphasis id="DE908E12FF86FFB1D39FDA98FEFC6331" bold="true" box="[171,310,767,790]" pageId="9" pageNumber="460">Description.</emphasis>
|
||
(
|
||
<figureCitation id="74DF4E85FF86FFB1D270DA99FE516331" box="[324,411,766,790]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="9" pageNumber="460">Fig. 7A</figureCitation>
|
||
). Size. Medium for genus, BL males =
|
||
<quantity id="2B1CFFE5FF86FFB1D077DA99FC166331" box="[835,988,766,790]" metricMagnitude="-3" metricUnit="m" metricValue="7.6499999999999995" metricValueMax="8.2" metricValueMin="7.1" pageId="9" pageNumber="460" unit="mm" value="7.6499999999999995" valueMax="8.2" valueMin="7.1">7.1 to 8.2 mm</quantity>
|
||
, females =
|
||
<quantity id="2B1CFFE5FF86FFB1D76CDA99FF1A6311" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.2" metricValueMin="6.8" pageId="9" pageNumber="460" unit="mm" value="7.5" valueMax="8.2" valueMin="6.8">6.8 to 8.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF86FFB1D39FDB58FF69649E" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
|
||
Color. Head piceous except clypeus, labrum, and venter rufous or rufopiceous; antennae, mandibles, maxillae, and maxillary and labial palpi pale yellow-tan. Pronotum piceous with very narrow pale (yellow-tan or rufous) lateral band). Elytra mainly yellow-tan, with black to piceous markings; basal dark band extended laterally to humerus on base anterior to basal margination but only to interval 5 posterior to basal margination; interval 1 piceous to apex or nearly so, middle transverse dark band moderately thick, irregular, extended from intervals 2 to 6, 7, or slightly onto interval
|
||
<quantity id="2B1CFFE5FF86FFB1D3E1DC67FF35643F" box="[213,255,1024,1048]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="9" pageNumber="460" unit="in" value="8.0">8 in</quantity>
|
||
some specimens, portion on intervals 2 to 4 located at about three-fourths of elytral length and with spot on interval 3 offset anteriorly relative to those on intervals 2 and 4, portion on interval 5 to 7 or 8 near midlength of elytra and formed more or less in a transverse row; preapical dark spot present or absent, if present then small. Venter piceous laterally, rufo-piceous medially, elytral epipleurae and apical portion of last abdominal sternite pale yellow-tan. Legs pale yellow-tan.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF86FFB1D39FDCA6FCEF651D" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">Reflection, luster, and microsculpture. Dorsum and venter without metallic reflection. Head and pronotum dull, with microsculpture moderately impressed and comprised of isodiametric meshes; elytra slightly shiny with isodiametric meshes slightly less deeply impressed; venter with isodiametric to slightly transverse meshes shallowly impressed.</paragraph>
|
||
<paragraph id="EC5B5200FF86FFB1D39FDD25FECC659C" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">Head. Eyes large, hemispheric; antennae slightly short, extended only to basal one-fifth of elytra; clypeus bisetose; labrum with three pairs of setae in most individuals, four pairs in a few; mentum asetose, with a broad, apically emarginate medial tooth; submentum anteriorly with a single pair of setae.</paragraph>
|
||
<paragraph id="EC5B5200FF86FFB1D39FDDA4FBA2669A" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
|
||
Prothorax. Pronotum (
|
||
<figureCitation id="74DF4E85FF86FFB1D2ADDDA4FE3365FC" box="[409,505,1475,1499]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="9" pageNumber="460">Fig. 13A</figureCitation>
|
||
) broad but slightly narrow for genus (ratio PWM/PL = 2.12 to 2.43), trapezoidal, almost as wide as elytra at humeral angles, with anterior angles acute, narrow; apical margin markedly concave between anterior angles; lateral margins slightly to distinctly sinuate near anterior angles, otherwise smoothly curved or nearly straight; basal margin markedly bisinuate, lobate medially; anterior and lateral margination absent or extremely faintly impressed, basal margination distinct laterally, obsolete medially; midlateral setae inserted slightly anterior to mid-length, basolateral setae inserted on lateral edge of pronotum and just anterior to posterior angles. Prosternal intercoxal process moderately long, lanceolate, with complete margination.
|
||
</paragraph>
|
||
<caption id="B89B0288FF85FFB2D3AFDE54FEA766B5" ID-DOI="http://doi.org/10.5281/zenodo.11067301" ID-Zenodo-Dep="11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="10" pageNumber="461" startId="10.[155,167,1587,1606]" targetBox="[126,1180,216,1575]" targetPageId="10" targetType="figure">
|
||
<paragraph id="EC5B5200FF85FFB2D3AFDE54FEA766B5" blockId="10.[123,1174,1587,1683]" pageId="10" pageNumber="461">
|
||
<smallCapsWord id="EABDC4DCFF85FFB2D3AFDE54FF126662" baselines="1601,1601" box="[155,216,1587,1606]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="10" pageNumber="461">FIGURE</smallCapsWord>
|
||
7.
|
||
<taxonomicName id="2BE42983FF85FFB2D3C6DE54FD106661" authority="Kavanaugh & Cueva-Dabkoski" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[242,730,1587,1606]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
|
||
<emphasis id="DE908E12FF85FFB2D3C6DE54FE0D6661" box="[242,455,1587,1606]" italics="true" pageId="10" pageNumber="461">Cyclosomus acutangulus</emphasis>
|
||
Kavanaugh & Cueva-Dabkoski
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C5A33369FF85FFB2D1D6DE54FCEF6661" box="[738,805,1587,1606]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC5B5200FF84FFB3D39FD8BCFE2461F5" pageId="11" pageNumber="462">Elytra. Broadly ovate, not or only slightly longer than wide (ratio EL/EW = 1.00 to 1.02), broadly rounded apically, humeri obtusely angulate; elytral striae deeply impressed throughout, elytral intervals slightly convex, smooth; parascutellar setiferous pore present at base of interval 1 near junction of striae 1 and 2; two discal setiferous pores present on interval three adjacent to stria 2, one near elytral mid-length and the other near apical one-third; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 11 to 13 setae.</paragraph>
|
||
<paragraph id="EC5B5200FF84FFB3D39FD9BCFE966334" pageId="11" pageNumber="462">
|
||
Legs. Tarsal claws, smooth, edentate. Males (
|
||
<figureCitation id="74DF4E85FF84FFB3D3F9D99CFEFB6234" box="[205,305,507,531]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="11" pageNumber="462">Fig. 14A</figureCitation>
|
||
) and females with front tarsi similar in shape and width, but males with tarsomeres 1 to 3 with two rows of adhesive squamosetae (
|
||
<bibRefCitation id="88752FF1FF84FFB3D224DA3CFE436254" author="STORK, N. E." box="[272,393,603,627]" pageId="11" pageNumber="462" pagination="173 - 306" refId="ref17965" refString="STORK, N. E. 1980. A scanning electron microscope study of tarsal adhesive setae in Coleoptera). Zoological Journal of the Linnaean Society 68: 173 - 306." type="journal article" year="1980">Stork 1980</bibRefCitation>
|
||
) ventrally (
|
||
<figureCitation id="74DF4E85FF84FFB3D137DA3CFDAE6254" box="[515,612,603,627]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="11" pageNumber="462">Fig. 15A</figureCitation>
|
||
), females without such setae; middle tarsi with tarsomeres 1 to 3 distinctly broader in males (
|
||
<figureCitation id="74DF4E85FF84FFB3D3B7DADCFF2F62F4" box="[131,229,699,723]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="11" pageNumber="462">Fig. 16A</figureCitation>
|
||
) than in females and with two rows of adhesive squamosetae ventrally (
|
||
<figureCitation id="74DF4E85FF84FFB3D134DABCFDAE62D4" box="[512,612,731,755]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="11" pageNumber="462">Fig. 17A</figureCitation>
|
||
), absent from females.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF84FFB3D39FDB7CFEB963F4" pageId="11" pageNumber="462">
|
||
Male genitalia. Median lobe (
|
||
<figureCitation id="74DF4E85FF84FFB3D2D3DB7CFDEC6314" box="[487,550,795,819]" captionStart="FIGURE" captionStartId="11.[687,699,975,994]" captionTargetBox="[657,1156,383,944]" captionTargetId="figure-502@11.[652,1176,357,962]" captionText="FIGURE 8. Male genitalia, Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012642, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Dorsal aspect; B. Left lateral aspect; scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12628241" httpUri="https://zenodo.org/record/12628241/files/figure.png" pageId="11" pageNumber="462">Fig. 8</figureCitation>
|
||
) stout, smoothly arcuate, and with apical lamella tapered to a rounded point apically in lateral view, slightly sinusoidal and with apical lamella broadly rounded in dorsal view. Internal sac without evident spines.
|
||
</paragraph>
|
||
<caption id="B89B0288FF84FFB3D19BDBA8FBF5646F" ID-DOI="http://doi.org/10.5281/zenodo.12628241" ID-Zenodo-Dep="12628241" httpUri="https://zenodo.org/record/12628241/files/figure.png" pageId="11" pageNumber="462" startId="11.[687,699,975,994]" targetBox="[657,1156,383,944]" targetPageId="11">
|
||
<paragraph id="EC5B5200FF84FFB3D19BDBA8FBF5646F" pageId="11" pageNumber="462">
|
||
<subScript id="70605045FF84FFB3D19BDBA8FB5F6432" attach="right" fontSize="8" pageId="11" pageNumber="462">
|
||
<smallCapsWord id="EABDC4DCFF84FFB3D19BDBA8FD2763C6" baselines="989,989" box="[687,749,975,994]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="11" pageNumber="462">FIGURE</smallCapsWord>
|
||
8. Male genitalia,
|
||
<taxonomicName id="2BE42983FF84FFB3D087DBA9FB5C63C6" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[947,1174,974,993]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="DE908E12FF84FFB3D087DBA9FB5C63C6" box="[947,1174,974,993]" italics="true" pageId="11" pageNumber="462">Cyclosomus acutangulus</emphasis>
|
||
</taxonomicName>
|
||
Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012642, Longchuan Jiang at Longjiang Bridge, Wuhe
|
||
</subScript>
|
||
Township, Tengchong County,
|
||
<collectingRegion id="2E209CE2FF84FFB3D091DC7BFC226408" box="[933,1000,1052,1071]" country="China" name="Yunnan" pageId="11" pageNumber="462">Yunnan</collectingRegion>
|
||
,
|
||
<collectingCountry id="94F31290FF84FFB3D0C2DC7CFBE76409" box="[1014,1069,1051,1070]" name="China" pageId="11" pageNumber="462">China</collectingCountry>
|
||
). A. Dorsal
|
||
<superScript id="1B91FF48FF84FFB3D1BBDC52FBF5646F" attach="right" box="[655,1087,1077,1096]" fontSize="8" pageId="11" pageNumber="462">
|
||
aspect; B. Left lateral aspect; scale line =
|
||
<quantity id="2B1CFFE5FF84FFB3D0C3DC52FBF6646F" box="[1015,1084,1077,1096]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="462" unit="mm" value="0.5">0.5 mm</quantity>
|
||
.
|
||
</superScript>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC5B5200FF84FFB3D39FDBBCFEA86474" blockId="11.[123,626,987,1107]" pageId="11" pageNumber="462">
|
||
Geographical variation. The two specimens from
|
||
<collectingCountry id="94F31290FF84FFB3D236DB9CFEF36434" box="[258,313,1019,1043]" name="India" pageId="11" pageNumber="462">India</collectingCountry>
|
||
lack a preapical dark spot found in specimens from the study area, but otherwise are similar.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF84FFB3D39FDC3BFB5C64D4" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">
|
||
<emphasis id="DE908E12FF84FFB3D39FDC3BFE526454" bold="true" box="[171,408,1116,1139]" pageId="11" pageNumber="462">Habitat distribution.</emphasis>
|
||
Specimens of this species were collected mainly at night, with the aid of headlamps, on the upper edges of open sandy beaches along the Longchuan River (
|
||
<figureCitation id="74DF4E85FF84FFB3D717DC1CFBBB64B4" box="[1059,1137,1147,1171]" captionStart="FIGURE" captionStartId="38.[155,167,1623,1642]" captionTargetBox="[136,1161,216,1607]" captionTargetId="figure-10@38.[136,1161,216,1607]" captionTargetPageId="38" captionText="FIGURE 21. Photographs of habitat for cyclosomine species, Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov., Tetragonoderus elegans Andrewes, T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., and T. punctatus Wiedemann at Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China. A. lower river bank; B. sandy upper bank with sparse vegetation. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067337" httpUri="https://zenodo.org/record/11067337/files/figure.png" pageId="11" pageNumber="462">Fig. 21</figureCitation>
|
||
) at an elevation of
|
||
<quantity id="2B1CFFE5FF84FFB3D216DCFCFEBC6495" box="[290,374,1179,1203]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="11" pageNumber="462" unit="m" value="1215.0">1215 m</quantity>
|
||
, where they were found active on the surface of dry or only slightly damp sandy substrate with no or only widely scattered and low vegetation. Members of this genus are known to burrow quickly into dry sand to hide when disturbed during the daytime (
|
||
<bibRefCitation id="88752FF1FF84FFB3D0CCDCBCFB4364D4" author="NIETNER, J." box="[1016,1161,1243,1267]" pageId="11" pageNumber="462" pagination="132 - 153" refId="ref17620" refString="NIETNER, J. 1857. Entomological papers, being chiefly descriptions of new Ceylon Coleoptera with such observations on their habits etc., as appear in any way interesting. Journal of the Asiatic Society of Bengal 26: 132 - 153." type="journal article" year="1857">Nietner 1857</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF84FFB3D39FDC9BFBAA6514" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">
|
||
<emphasis id="DE908E12FF84FFB3D39FDC9BFCD96534" bold="true" box="[171,787,1276,1299]" pageId="11" pageNumber="462">Geographical distribution within the Gaoligong Shan.</emphasis>
|
||
<figureCitation id="74DF4E85FF84FFB3D02EDC9CFCA56535" box="[794,879,1275,1299]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="11" pageNumber="462">Fig. 7B</figureCitation>
|
||
. We examined a total of
|
||
<specimenCount id="FAE29989FF84FFB3D74FDC9CFF206514" count="29" pageId="11" pageNumber="462" type="generic">29 specimens</specimenCount>
|
||
(
|
||
<specimenCount id="FAE29989FF84FFB3D3CEDD7CFE906514" box="[250,346,1307,1331]" count="13" pageId="11" pageNumber="462" type="male">13 males</specimenCount>
|
||
and
|
||
<specimenCount id="FAE29989FF84FFB3D2BBDD7CFDCC6514" box="[399,518,1307,1331]" count="16" pageId="11" pageNumber="462" type="female">16 females</specimenCount>
|
||
) from the study area (see list of
|
||
<typeStatus id="335FECA2FF84FFB3D050DD7CFC586514" box="[868,914,1307,1331]" pageId="11" pageNumber="462">type</typeStatus>
|
||
specimens above).
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF84FFB3D39FDD5CFF196554" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">Members of this species were collected only in the southwestern part of the study area (Core Area 6).</paragraph>
|
||
<paragraph id="EC5B5200FF84FFB3D39FDD1BFE276614" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">
|
||
<emphasis id="DE908E12FF84FFB3D39FDD1BFDFA65B4" bold="true" box="[171,560,1404,1427]" pageId="11" pageNumber="462">Overall geographical distribution.</emphasis>
|
||
<figureCitation id="74DF4E85FF84FFB3D103DD1CFD4E65B4" box="[567,644,1403,1427]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="11" pageNumber="462">Fig. 23</figureCitation>
|
||
. At present, this species is known only from single areas in western
|
||
<collectingRegion id="2E209CE2FF84FFB3D26EDDFCFDDC6594" box="[346,534,1435,1459]" country="China" name="Yunnan" pageId="11" pageNumber="462">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="94F31290FF84FFB3D116DDFCFDA86594" box="[546,610,1435,1459]" name="China" pageId="11" pageNumber="462">China</collectingCountry>
|
||
and northcentral
|
||
<collectingRegion id="2E209CE2FF84FFB3D014DDFCFCA16594" box="[800,875,1435,1459]" country="India" name="Assam" pageId="11" pageNumber="462">Assam</collectingRegion>
|
||
,
|
||
<collectingCountry id="94F31290FF84FFB3D04CDDFCFC786594" box="[888,946,1435,1459]" name="India" pageId="11" pageNumber="462">India</collectingCountry>
|
||
. These two areas are about
|
||
<quantity id="2B1CFFE5FF84FFB3D38BDDDCFED865F4" box="[191,274,1467,1491]" metricMagnitude="5" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="462" unit="km" value="650.0">650 km</quantity>
|
||
apart, and it is likely that this species occurs also in suitable habitats in the intervening region, including northern
|
||
<collectingCountry id="94F31290FF84FFB3D2D1DDBCFD8765D4" box="[485,589,1499,1523]" name="Myanmar" pageId="11" pageNumber="462">Myanmar</collectingCountry>
|
||
(
|
||
<collectingRegion id="2E209CE2FF84FFB3D16EDDBCFD2D65D4" box="[602,743,1499,1523]" country="Myanmar" name="Kachin" pageId="11" pageNumber="462">Kachin State</collectingRegion>
|
||
), and northeasternmost
|
||
<collectingCountry id="94F31290FF84FFB3D0D7DDBCFBD665D4" box="[995,1052,1499,1523]" name="India" pageId="11" pageNumber="462">India</collectingCountry>
|
||
, at low elevations (below
|
||
<quantity id="2B1CFFE5FF84FFB3D217DD9CFEB26634" box="[291,376,1531,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="462" unit="m" value="1500.0">1500 m</quantity>
|
||
) along rivers draining the western and southern slopes of the Himalayan ranges in these areas, respectively.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF84FFB4D39FDE5BFDF06134" blockId="11.[123,1174,1115,1715]" lastBlockId="12.[123,1174,219,275]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
|
||
<emphasis id="DE908E12FF84FFB3D39FDE5BFCF76674" bold="true" box="[171,829,1595,1619]" pageId="11" pageNumber="462">
|
||
Geographical relationships with other
|
||
<taxonomicName id="2BE42983FF84FFB3D169DE5CFCF76674" box="[605,829,1595,1619]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="DE908E12FF84FFB3D169DE5CFD2B6674" bold="true" box="[605,737,1595,1619]" italics="true" pageId="11" pageNumber="462">Cyclosomus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</emphasis>
|
||
No other species of
|
||
<taxonomicName id="2BE42983FF84FFB3D720DE5CFB5F6674" authorityName="Latreille" authorityYear="1829" box="[1044,1173,1595,1619]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DE908E12FF84FFB3D720DE5CFB5F6674" box="[1044,1173,1595,1619]" italics="true" pageId="11" pageNumber="462">Cyclosomus</emphasis>
|
||
</taxonomicName>
|
||
has been recorded from the Gaoligong Shan region, but members of
|
||
<taxonomicName id="2BE42983FF84FFB3D05ADE3CFBDB6654" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[878,1041,1627,1651]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="DE908E12FF84FFB3D05ADE3CFBDB6654" box="[878,1041,1627,1651]" italics="true" pageId="11" pageNumber="462">C. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
were found syntopic with
|
||
<taxonomicName id="2BE42983FF84FFB3D227DE1CFE5F66B4" authorityName="Fabricius" authorityYear="1775" box="[275,405,1659,1683]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="flexuosus">
|
||
<emphasis id="DE908E12FF84FFB3D227DE1CFE5F66B4" box="[275,405,1659,1683]" italics="true" pageId="11" pageNumber="462">C. flexuosus</emphasis>
|
||
</taxonomicName>
|
||
members at the locality in
|
||
<collectingRegion id="2E209CE2FF84FFB3D18DDE1CFCC866B4" box="[697,770,1659,1683]" country="India" name="Assam" pageId="11" pageNumber="462">Assam</collectingRegion>
|
||
[NHMUK], which is the easternmost known locality for the latter species and the westernmost for
|
||
<taxonomicName id="2BE42983FF84FFB3D023DEFCFC706694" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[791,954,1691,1715]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="DE908E12FF84FFB3D023DEFCFC706694" box="[791,954,1691,1715]" italics="true" pageId="11" pageNumber="462">C. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
. Other cyclosomine species found syntopic with
|
||
<taxonomicName id="2BE42983FF83FFB4D29ED8BCFD8160D4" authorityName="Kavanaugh & Cueva-Dabkoski & Liang" authorityYear="2023" box="[426,587,219,243]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="DE908E12FF83FFB4D29ED8BCFD8160D4" box="[426,587,219,243]" italics="true" pageId="12" pageNumber="463">C. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
at the
|
||
<typeStatus id="335FECA2FF83FFB4D1A7D8BBFD0B60D3" box="[659,705,220,244]" pageId="12" pageNumber="463">type</typeStatus>
|
||
locality along the Longchuan River include
|
||
<taxonomicName id="2BE42983FF83FFB4D34FD89CFEB16134" box="[123,379,251,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="DE908E12FF83FFB4D34FD89CFEB16134" box="[123,379,251,275]" italics="true" pageId="12" pageNumber="463">Tetragonoderus elegans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2BE42983FF83FFB4D285D89BFDF96134" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[433,563,252,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="DE908E12FF83FFB4D285D89BFDF96134" box="[433,563,252,275]" italics="true" pageId="12" pageNumber="463">T. punctatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |