479 lines
71 KiB
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479 lines
71 KiB
XML
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<mods:title id="61055D3500BC7C591CB917D5FEF80A4B">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:namePart id="E299DC83424B87854D5931A3F432CDDA">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="5065AC71F77FDC2A84463DCC9A6E5696">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:namePart id="F576E004261BD4CD6FAA8D13C2ECD90B">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="B4EAF16612037B96EC72191F215ECAE2">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:namePart id="DB88CA1F2873310127FF593B2C6A8D1B">Liang, Hongbin</mods:namePart>
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<mods:affiliation id="E659CAB03BA269E670FC03C5CA08E7AE">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
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<mods:date id="DC727E2CBE3B1325333721E64408B192">2023</mods:date>
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<mods:number id="E3AA0B00A3A46B659AFB69A923651DDE">2023-02-15</mods:number>
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<treatment id="644DE316FF83FFB7D34FDB25FC2A61DF" ID-DOI="http://doi.org/10.5281/zenodo.11067315" ID-GBIF-Taxon="234412272" ID-Zenodo-Dep="11067315" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF" httpUri="http://treatment.plazi.org/id/644DE316FF83FFB7D34FDB25FC2A61DF" lastPageId="15" lastPageNumber="466" pageId="12" pageNumber="463">
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<paragraph id="EC5B5200FF83FFB4D34FDB25FE2C637E" blockId="12.[123,675,834,888]" box="[123,486,834,858]" pageId="12" pageNumber="463">
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<heading id="B713E56CFF83FFB4D34FDB25FE2C637E" box="[123,486,834,858]" fontSize="10" level="2" pageId="12" pageNumber="463" reason="4">
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<taxonomicName id="2BE42983FF83FFB4D34FDB25FE2C637E" ID-CoL="55MYV" authority="Dejean" authorityName="Dejean" authorityYear="1829" box="[123,486,834,858]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">
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<emphasis id="DE908E12FF83FFB4D34FDB25FE2C637E" bold="true" box="[123,486,834,858]" pageId="12" pageNumber="463">
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<emphasis id="DE908E12FF83FFB4D34FDB25FE44637D" bold="true" box="[123,398,834,858]" italics="true" pageId="12" pageNumber="463">Tetragonoderus arcuatus</emphasis>
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Dejean
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</emphasis>
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</taxonomicName>
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</paragraph>
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<subSubSection id="A4FE018BFF83FFB4D34FDB05FD69635F" box="[123,675,866,888]" pageId="12" pageNumber="463" type="description">
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<paragraph id="EC5B5200FF83FFB4D34FDB05FD69635F" blockId="12.[123,675,834,888]" box="[123,675,866,888]" pageId="12" pageNumber="463">
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<figureCitation id="74DF4E85FF83FFB4D34FDB05FF1D635F" box="[123,215,866,888]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="12" pageNumber="463">Figures 9</figureCitation>
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,
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<figureCitation id="74DF4E85FF83FFB4D3D6DB05FEC1635F" box="[226,267,866,888]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="12" pageNumber="463">13B</figureCitation>
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,
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<figureCitation id="74DF4E85FF83FFB4D222DB05FE8B635F" box="[278,321,866,888]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="12" pageNumber="463">14B</figureCitation>
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,
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<figureCitation id="74DF4E85FF83FFB4D27FDB05FEBF635F" box="[331,373,866,888]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="12" pageNumber="463">15B</figureCitation>
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,
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<figureCitation id="74DF4E85FF83FFB4D2B4DB05FE60635F" box="[384,426,866,888]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="12" pageNumber="463">16B</figureCitation>
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,
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<figureCitation id="74DF4E85FF83FFB4D280DB05FE14635F" box="[436,478,866,888]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="12" pageNumber="463">17B</figureCitation>
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,
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<figureCitation id="74DF4E85FF83FFB4D2DDDB05FDCA635F" box="[489,512,866,888]" captionStart="FIGURE" captionStartId="35.[155,167,1036,1055]" captionTargetBox="[130,1144,252,995]" captionTargetId="figure-68@35.[120,1176,216,1033]" captionTargetPageId="35" captionText="FIGURE 18. Male genitalia, Tetragonoderus spp. A, B. T. arcuatus Dejean (Nepalgunj, Banke District, Nepal); C, D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). A, C. Dorsal aspect; B, D. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067329" httpUri="https://zenodo.org/record/11067329/files/figure.png" pageId="12" pageNumber="463">18</figureCitation>
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A-B, 22, and 23.
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</paragraph>
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</subSubSection>
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<subSubSection id="A4FE018BFF83FFB4D34FDBEBFC92644A" pageId="12" pageNumber="463" type="reference_group">
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<paragraph id="EC5B5200FF83FFB4D34FDBEBFEC76414" blockId="12.[123,1174,908,1133]" pageId="12" pageNumber="463">
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<treatmentCitation id="6D457411FF83FFB4D34FDBEBFDEE6385" author="DEJEAN, P. F. M. A." box="[123,548,908,930]" page="495" pageId="12" pageNumber="463" year="1829">
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<taxonomicName id="2BE42983FF83FFB4D34FDBEBFDEE6385" ID-CoL="55MYV" authority="Dejean, 1829: 495" authorityName="Dejean" authorityPageNumber="495" authorityYear="1829" box="[123,548,908,930]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">
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<emphasis id="DE908E12FF83FFB4D34FDBEBFEA76385" box="[123,365,908,930]" italics="true" pageId="12" pageNumber="463">Tetragonoderus arcuatus</emphasis>
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<bibRefCitation id="88752FF1FF83FFB4D241DBEBFDEE6385" author="DEJEAN, P. F. M. A." box="[373,548,908,930]" pageId="12" pageNumber="463" refId="ref16379" refString="DEJEAN, P. F. M. A. 1829. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829:495</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="335FECA2FF83FFB4D11BDBEBFD706385" box="[559,698,908,930]" pageId="12" pageNumber="463" type="lectotype">LECTOTYPE</typeStatus>
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, here designated, a female, deposited in MNHN and labeled: “
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<collectingCountry id="94F31290FF83FFB4D22ADBCEFEAF6398" box="[286,357,937,959]" name="Egypt" pageId="12" pageNumber="463">Egypte</collectingCountry>
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” [handwritten]/ “TYPE arcuatus” [red label]/
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<taxonomicName id="2BE42983FF83FFB4D02BDBCEFC096398" ID-CoL="55MYV" authority="Dejean" authorityName="Dejean" authorityYear="1829" box="[799,963,937,959]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">“arcuatus Dejean</taxonomicName>
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<collectingCountry id="94F31290FF83FFB4D0FCDBCEFBC66399" box="[968,1036,937,958]" name="Egypt" pageId="12" pageNumber="463">Egypte</collectingCountry>
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Coll. Dejean” [handwritten, white label with black border]/ “
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<typeStatus id="335FECA2FF83FFB4D149DBA1FCC263FB" box="[637,776,966,988]" pageId="12" pageNumber="463" type="lectotype">LECTOTYPE</typeStatus>
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<taxonomicName id="2BE42983FF83FFB4D027DBA1FB5F63FB" ID-CoL="55MYV" authority="Dejean 1829" authorityName="Dejean" authorityYear="1829" box="[787,1173,966,988]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">
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<emphasis id="DE908E12FF83FFB4D027DBA1FBC363FB" box="[787,1033,966,988]" italics="true" pageId="12" pageNumber="463">Tetragonoderus arcuatus</emphasis>
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<bibRefCitation id="88752FF1FF83FFB4D721DBA1FB5F63FB" author="DEJEAN, P. F. M. A." box="[1045,1173,966,988]" pageId="12" pageNumber="463" refId="ref16379" refString="DEJEAN, P. F. M. A. 1829. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean 1829</bibRefCitation>
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</taxonomicName>
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design. by D.H. Kavanaugh & M. Cueva-Dabkoski 2022” [red label]. Type locality:
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<collectingCountry id="94F31290FF83FFB4D0D5DB83FBD063DE" box="[993,1050,996,1017]" name="Egypt" pageId="12" pageNumber="463">Egypt</collectingCountry>
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[but Dejean viewed this record as uncertain].
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<treatmentCitation id="6D457411FF83FFB4D2E9DC67FD5E6431" author="CHAUDOIR, M." box="[477,660,1024,1046]" page="38" pageId="12" pageNumber="463" year="1876">
|
||
<bibRefCitation id="88752FF1FF83FFB4D2E9DC67FD5E6431" author="CHAUDOIR, M." box="[477,660,1024,1046]" pageId="12" pageNumber="463" pagination="1 - 84" refId="ref16317" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir, 1876:38</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="6D457411FF83FFB4D1ABDC67FCF36431" author="FELIX, R. F. F. L." box="[671,825,1024,1046]" page="116" pageId="12" pageNumber="463" year="2009">
|
||
<bibRefCitation id="88752FF1FF83FFB4D1ABDC67FCF36431" author="FELIX, R. F. F. L." box="[671,825,1024,1046]" pageId="12" pageNumber="463" pagination="66 - 141" refId="ref16680" refString="FELIX, R. F. F. L. 2009. Order Coleoptera, family Carabidae. Arthropod Fauna of the UAE 2: 66 - 141." type="journal article" year="2009">Felix, 2009:116</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="6D457411FF83FFB4D077DC67FBFA6431" author="ASSMANN, T. & E. BOUTAUD & J. BUSE & V. CHIKATUNOV & C. DREES & A. - L. - L. FRIEDMAN & W. HARDTLE & K. HOMBURG & T. MARCUS & I. RENAN & D. W. WRASE" box="[835,1072,1024,1046]" page="57" pageId="12" pageNumber="463" year="2015">
|
||
<bibRefCitation id="88752FF1FF83FFB4D077DC67FBFA6431" author="ASSMANN, T. & E. BOUTAUD & J. BUSE & V. CHIKATUNOV & C. DREES & A. - L. - L. FRIEDMAN & W. HARDTLE & K. HOMBURG & T. MARCUS & I. RENAN & D. W. WRASE" box="[835,1072,1024,1046]" pageId="12" pageNumber="463" pagination="49 - 69" refId="ref15907" refString="ASSMANN, T., E. BOUTAUD, J. BUSE, V. CHIKATUNOV, C. DREES, A. - L. - L. FRIEDMAN, W. HARDTLE, K. HOMBURG, T. MARCUS, I. RENAN, AND D. W. WRASE. 2015. The ground beetle tribe Cyclosomini s. l. in Israel (Coleoptera, Carabidae). Spixiana 38: 49 - 69." type="journal article" year="2015">Assmann et al., 2015:57</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="6D457411FF83FFB4D70FDC66FEC36414" author="BOUSQUET, Y." page="498" pageId="12" pageNumber="463" year="2017">
|
||
<bibRefCitation id="88752FF1FF83FFB4D70FDC66FEC36414" author="BOUSQUET, Y." pageId="12" pageNumber="463" refId="ref16205" refString="BOUSQUET, Y. 2017. Tribe Cyclosomini. Pages 498 - 499 in Lobl, I., and D. Lobl (editors) Catalogue of Palaearctic Coleoptera. Volume 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden, xxxiv + 1143 pp." type="book" year="2017">Bousquet (2017:498)</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF83FFB4D34FDC5DFC92644A" blockId="12.[123,1174,908,1133]" pageId="12" pageNumber="463">
|
||
<treatmentCitationGroup id="CCF4752EFF83FFB4D34FDC5DFC92644A" pageId="12" pageNumber="463">
|
||
<treatmentCitation id="6D457411FF83FFB4D34FDC5DFDC06477" author="BATES, H. W." box="[123,522,1082,1104]" page="338" pageId="12" pageNumber="463" year="1891">
|
||
<taxonomicName id="2BE42983FF83FFB4D34FDC5DFDC06477" ID-CoL="55N34" authority="Bates, 1891: 338" authorityName="Bates" authorityPageNumber="338" authorityYear="1891" box="[123,522,1082,1104]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cardoni">
|
||
<emphasis id="DE908E12FF83FFB4D34FDC5DFEA96477" box="[123,355,1082,1104]" italics="true" pageId="12" pageNumber="463">Tetragonoderus cardoni</emphasis>
|
||
<bibRefCitation id="88752FF1FF83FFB4D25EDC5CFDC06477" author="BATES, H. W." box="[362,522,1082,1104]" pageId="12" pageNumber="463" pagination="324 - 340" refId="ref16102" refString="BATES, H. W. 1891. List of the Carabidae (ord. Coleoptera) obtained by Pere Cardon in Chota-Nagpore. Annales de la Societe entomologique de Belgique 35: 324 - 340." type="journal article" year="1891">Bates, 1891:338</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
<typeStatus id="335FECA2FF83FFB4D120DC5DFD5F6477" box="[532,661,1082,1104]" pageId="12" pageNumber="463" type="holotype">HOLOTYPE</typeStatus>
|
||
, a female, deposited in MNHN. Type locality:
|
||
<collectingCountry id="94F31290FF83FFB4D76ADC5DFB586477" box="[1118,1170,1082,1104]" name="India" pageId="12" pageNumber="463">India</collectingCountry>
|
||
,
|
||
<collectingRegion id="2E209CE2FF83FFB4D3AFDC30FECA644A" box="[155,256,1111,1133]" country="India" name="Jharkhand" pageId="12" pageNumber="463">Jharkhand</collectingRegion>
|
||
, Konbir.
|
||
<treatmentCitation id="6D457411FF83FFB4D26EDC3FFE3E644A" author="BATES, H. W." box="[346,500,1111,1133]" page="416" pageId="12" pageNumber="463" year="1892">
|
||
<bibRefCitation id="88752FF1FF83FFB4D26EDC3FFE3E644A" author="BATES, H. W." box="[346,500,1111,1133]" pageId="12" pageNumber="463" pagination="267 - 428" refId="ref16139" refString="BATES, H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. Annali del Museuo Civico di Storia Naturale di Genova (Serie 2 a) 12: 267 - 428." type="journal article" year="1892">Bates 1892:416</bibRefCitation>
|
||
</treatmentCitation>
|
||
. Synonymy by
|
||
<treatmentCitation id="6D457411FF83FFB4D1BDDC30FC9E644A" author="ANDREWES, H. E." box="[649,852,1111,1133]" page="150" pageId="12" pageNumber="463" year="1921">
|
||
<bibRefCitation id="88752FF1FF83FFB4D1BDDC30FC9E644A" author="ANDREWES, H. E." box="[649,852,1111,1133]" pageId="12" pageNumber="463" pagination="145 - 195" refId="ref15718" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes, 1921:150</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4FE018BFF83FFB6D39FDCE6FD156252" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463" type="materials_examined">
|
||
<paragraph id="EC5B5200FF83FFB4D39FDCE6FC47655D" blockId="12.[123,1174,1152,1726]" pageId="12" pageNumber="463">
|
||
<emphasis id="DE908E12FF83FFB4D39FDCE6FDFD64BF" bold="true" box="[171,567,1153,1176]" pageId="12" pageNumber="463">Notes on nomenclature and types.</emphasis>
|
||
<materialsCitation id="5C8C585DFF83FFB4D175DCE6FC43655D" ID-GBIF-Occurrence="4902827334" collectionCode="MNHN, ZMHB" country="Egypt" county="Recommendation" location="Bernd Jaeger" municipality="International Code of Zoological Nomenclature" pageId="12" pageNumber="463" specimenCount="5" specimenCount-male="1" typeStatus="lectotype">
|
||
In his original description,
|
||
<bibRefCitation id="88752FF1FF83FFB4D059DCE7FBC064BF" author="DEJEAN, P. F. M. A." box="[877,1034,1152,1176]" pageId="12" pageNumber="463" refId="ref16379" refString="DEJEAN, P. F. M. A. 1829. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean (1829)</bibRefCitation>
|
||
was unclear with regard to how many specimens he had examined (Thierry Deuve, personal communication). He noted that he had received material from Klug and that he had not seen a male specimen. In addition to a single female specimen in
|
||
<collectionCode id="8AF5CAC5FF83FFB4D11ADC85FD4864DE" box="[558,642,1250,1273]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="463">MNHN</collectionCode>
|
||
, there are
|
||
<specimenCount id="FAE29989FF83FFB4D1C1DC86FC5264DE" box="[757,920,1249,1273]" count="4" pageId="12" pageNumber="463" type="generic">four specimens</specimenCount>
|
||
from
|
||
<collectingCountry id="94F31290FF83FFB4D0E9DC85FBD764DD" box="[989,1053,1250,1274]" name="Egypt" pageId="12" pageNumber="463">Egypt</collectingCountry>
|
||
in
|
||
<collectionCode id="8AF5CAC5FF83FFB4D776DC85FB5864DE" box="[1090,1170,1250,1273]" pageId="12" pageNumber="463">ZMHB</collectionCode>
|
||
, which were probably part of Klug’s original series and include
|
||
<specimenCount id="FAE29989FF83FFB4D018DD65FC45653E" box="[812,911,1281,1305]" count="1" pageId="12" pageNumber="463" type="male">one male</specimenCount>
|
||
(
|
||
<location id="E93B04DBFF83FFB4D0AADD66FBE5653D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D0AADD66FBE5653D" box="[926,1071,1281,1306]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="Bernd Jaeger" pageId="12" pageNumber="463">Bernd Jaeger</location>
|
||
, personal communication).
|
||
<location id="E93B04DBFF83FFB4D208DD44FE86651D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D208DD44FE86651D" box="[316,332,1315,1338]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="It" pageId="12" pageNumber="463">It</location>
|
||
seems most likely that
|
||
<location id="E93B04DBFF83FFB4D156DD45FD64651D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D156DD45FD64651D" box="[610,686,1314,1338]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="Dejean" pageId="12" pageNumber="463">Dejean</location>
|
||
saw only the single female specimen now deposited in
|
||
<collectionCode id="8AF5CAC5FF83FFB4D232DD24FE90657D" box="[262,346,1347,1370]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="463">MNHN</collectionCode>
|
||
, but we cannot be sure.
|
||
<location id="E93B04DBFF83FFB4D156DD25FD3C657D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D156DD25FD3C657D" box="[610,758,1346,1370]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="Consequently" pageId="12" pageNumber="463">Consequently</location>
|
||
, we designate that female as
|
||
<typeStatus id="335FECA2FF83FFB4D700DD25FB5C657D" box="[1076,1174,1346,1370]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<collectingMunicipality id="0C3FC87AFF83FFB4D3B6DD05FD4F655D" box="[130,645,1378,1402]" pageId="12" pageNumber="463">International Code of Zoological Nomenclature</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="053A2A8CFF83FFB4D1A4DD05FC86655D" box="[656,844,1378,1402]" pageId="12" pageNumber="463">Recommendation</collectingCounty>
|
||
73F)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF83FFB6D39FDDE4FDFD61B3" blockId="12.[123,1174,1152,1726]" lastBlockId="14.[123,1174,219,1725]" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463">
|
||
When he proposed the synonymy of
|
||
<taxonomicName id="2BE42983FF83FFB4D10DDDE3FD2265BD" authority="Bates" authorityName="Bates" authorityYear="1891" box="[569,744,1411,1435]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cardoni">
|
||
<emphasis id="DE908E12FF83FFB4D10DDDE3FD6C65BC" box="[569,678,1411,1435]" italics="true" pageId="12" pageNumber="463">T. cardoni</emphasis>
|
||
Bates
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="2BE42983FF83FFB4D013DDE3FB5F65BC" authority=", Andrewes (1921: 150)" authorityName="Andrewes" authorityPageNumber="150" authorityYear="1921" box="[807,1173,1411,1435]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF83FFB4D013DDE3FC5465BC" box="[807,926,1412,1435]" italics="true" pageId="12" pageNumber="463">T. arcuatus</emphasis>
|
||
,
|
||
<bibRefCitation id="88752FF1FF83FFB4D09FDDE4FB5F65BC" author="ANDREWES, H. E." box="[939,1173,1411,1435]" pageId="12" pageNumber="463" pagination="145 - 195" refId="ref15718" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:150)</bibRefCitation>
|
||
</taxonomicName>
|
||
also noted the pronounced variation in the degree of development of the sericeous sheen seen among specimens of this species. Specimens from the study area and adjacent regions appear to have the most markedly sericeous elytra while those from areas to the west, including the
|
||
<typeStatus id="335FECA2FF83FFB4D75CDD83FB5C65DB" box="[1128,1174,1508,1532]" pageId="12" pageNumber="463">type</typeStatus>
|
||
locality (
|
||
<collectingCountry id="94F31290FF83FFB4D3EDDE63FED4663B" box="[217,286,1540,1564]" name="Egypt" pageId="12" pageNumber="463">Egypt</collectingCountry>
|
||
), have the least sericeous elytra. There is also conspicuous variation in the distinctness and extent of the subapical pale band of spots. In the single specimen from the study area, and also in those from adjacent parts of the eastern end of the range of the species (e.g., Assam,
|
||
<collectingCountry id="94F31290FF83FFB4D77ADE22FF6E665B" name="Myanmar" pageId="12" pageNumber="463">Myanmar</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="94F31290FF83FFB4D20DDE03FE56665B" box="[313,412,1636,1660]" name="Thailand" pageId="12" pageNumber="463">Thailand</collectingCountry>
|
||
), the subapical pale band of spots is faint to very faint, almost invisible in some specimens, and apparent only on intervals 7 and 8 or extremely faintly visible also on more mediad intervals in some specimens. In contrast, specimens from the western part of the range of the species (i.e., North Africa, the Middle East and Central Asia) have the subapical band much more distinct and extended from interval 2 to interval 8. The female
|
||
<typeStatus id="335FECA2FF81FFB6D068D89BFC406133" box="[860,906,252,276]" pageId="14" pageNumber="465">type</typeStatus>
|
||
of
|
||
<taxonomicName id="2BE42983FF81FFB6D09AD89BFBD16134" authorityName="Bates" authorityYear="1891" box="[942,1051,251,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="cardoni">
|
||
<emphasis id="DE908E12FF81FFB6D09AD89BFBD16134" box="[942,1051,251,275]" italics="true" pageId="14" pageNumber="465">T. cardoni</emphasis>
|
||
</taxonomicName>
|
||
is intermediate between specimens from the western part of the species range and those from the study area and adjacent regions in both of these features. Because of this pronounced variation, features noted in the key above and in the diagnosis presented below are applicable specifically for specimens from the study area and adjacent regions.
|
||
</paragraph>
|
||
<caption id="B89B0288FF82FFB5D3AFDE5FFE376659" ID-DOI="http://doi.org/10.5281/zenodo.11067303" ID-Zenodo-Dep="11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="13" pageNumber="464" startId="13.[155,167,1592,1611]" targetBox="[128,1175,226,1577]" targetPageId="13" targetType="figure">
|
||
<paragraph id="EC5B5200FF82FFB5D3AFDE5FFE376659" blockId="13.[123,1174,1592,1663]" pageId="13" pageNumber="464">
|
||
<smallCapsWord id="EABDC4DCFF82FFB5D3AFDE5FFF12666D" baselines="1606,1606" box="[155,216,1592,1611]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="13" pageNumber="464">FIGURE</smallCapsWord>
|
||
9.
|
||
<taxonomicName id="2BE42983FF82FFB5D3C7DE5FFDD1666C" authority="(Dejean)" baseAuthorityName="Dejean" box="[243,539,1592,1611]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF82FFB5D3C7DE5FFE03666C" box="[243,457,1592,1611]" italics="true" pageId="13" pageNumber="464">Tetragonoderus arcuatus</emphasis>
|
||
(Dejean)
|
||
</taxonomicName>
|
||
. A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC5B5200FF81FFB6D39FD9FBFD156252" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
|
||
Adults of several additional Asian taxa presently treated as distinct species are very similar to those of
|
||
<taxonomicName id="2BE42983FF81FFB6D3EAD9DAFE9D61F3" box="[222,343,445,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF81FFB6D3EAD9DAFE9D61F3" box="[222,343,445,468]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
|
||
</taxonomicName>
|
||
. These include
|
||
<taxonomicName id="2BE42983FF81FFB6D13FD9DBFC1561F3" authority="Jedlicka, 1964" authorityName="Jedlicka" authorityYear="1964" box="[523,991,444,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="assamensis">
|
||
<emphasis id="DE908E12FF81FFB6D13FD9DBFCFF61F3" box="[523,821,444,468]" italics="true" pageId="14" pageNumber="465">Tetragonoderus assamensis</emphasis>
|
||
<bibRefCitation id="88752FF1FF81FFB6D00BD9DBFC1561F3" author="JEDLICKA, A." box="[831,991,444,468]" pageId="14" pageNumber="465" pagination="305 - 318" refId="ref16866" refString="JEDLICKA, A. 1964. Neue Carabiden aus Indien (Coleoptera-Carabidae). Entomologische Arbeiten aus dem Museum G. Frey 15: 305 - 318." type="journal article" year="1964">Jedlička, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BE42983FF81FFB6D0D9D9DBFE4C61D3" authority="Jedlicka, 1964" authorityName="Jedlicka" authorityYear="1964" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="cinchona">
|
||
<emphasis id="DE908E12FF81FFB6D0D9D9DBFF1761D3" italics="true" pageId="14" pageNumber="465">Tetragonoderus cinchona</emphasis>
|
||
<bibRefCitation id="88752FF1FF81FFB6D3D2D9BBFE4C61D3" author="JEDLICKA, A." box="[230,390,476,500]" pageId="14" pageNumber="465" pagination="305 - 318" refId="ref16866" refString="JEDLICKA, A. 1964. Neue Carabiden aus Indien (Coleoptera-Carabidae). Entomologische Arbeiten aus dem Museum G. Frey 15: 305 - 318." type="journal article" year="1964">Jedlička, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BE42983FF81FFB6D2A0D9BBFC9761D3" authority="Solsky, 1874" authorityName="Solsky" authorityYear="1874" box="[404,861,476,500]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="DE908E12FF81FFB6D2A0D9BBFD0E61D3" box="[404,708,476,500]" italics="true" pageId="14" pageNumber="465">Tetragonoderus intermedius</emphasis>
|
||
<bibRefCitation id="88752FF1FF81FFB6D1F9D9BBFC9761D3" author="SOLSKY, S. M." box="[717,861,476,500]" pageId="14" pageNumber="465" refId="ref17856" refString="SOLSKY, S. M. 1874. Zhestkokrylye (Coleoptera). In: Fedchenko, A. P. Puteshestvie v Turkestan. Izvestiya Imperatorskogo Obshchestva Lyubitelei Estestvoznaniya, Anthropologii I Etnografii (5) 11: iv + 222 pp. + 1 pl." type="book" year="1874">Solsky, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2BE42983FF81FFB6D0AFD9BBFED06232" authority="Jedlicka, 1966" authorityName="Jedlicka" authorityYear="1966" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="nakaoi">
|
||
<emphasis id="DE908E12FF81FFB6D0AFD9BBFB5C61D3" box="[923,1174,476,500]" italics="true" pageId="14" pageNumber="465">Tetragonoderus nakaoi</emphasis>
|
||
<bibRefCitation id="88752FF1FF81FFB6D34FD99AFED06232" author="JEDLICKA, A." box="[123,282,509,533]" pageId="14" pageNumber="465" pagination="11 - 16" refId="ref16894" refString="JEDLICKA, A. 1966. 2. Family Carabidae. Neue Carabiden aus Vietnam und Thailand (Coleoptera-Carabidae). Memoirs of the Faculty of Education. Kagawa University (Part II) 140: 11 - 16" type="journal article" year="1966">Jedlička, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
. The taxonomic status of these names is currently under review, but none of them have been recorded from the study area and all are more recently described than
|
||
<taxonomicName id="2BE42983FF81FFB6D0EFDA79FB986212" box="[987,1106,542,565]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF81FFB6D0EFDA79FB986212" box="[987,1106,542,565]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
|
||
</taxonomicName>
|
||
. Consequently, none of them would challenge
|
||
<taxonomicName id="2BE42983FF81FFB6D171DA59FD776272" box="[581,701,574,597]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF81FFB6D171DA59FD776272" box="[581,701,574,597]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
|
||
</taxonomicName>
|
||
as the valid name of this species if one or more are synonymized with the latter name in the future.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A4FE018BFF81FFB7D39FDA19FC2A61DF" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465" type="diagnosis">
|
||
<paragraph id="EC5B5200FF81FFB6D39FDA19FC29641F" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
|
||
<emphasis id="DE908E12FF81FFB6D39FDA19FED562B2" bold="true" box="[171,287,638,661]" pageId="14" pageNumber="465">Diagnosis.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="2BE42983FF81FFB6D2BADA18FDC962B1" box="[398,515,639,662]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF81FFB6D2BADA18FDC962B1" box="[398,515,639,662]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from those of other cyclosomine species in the study area by the following combination of character states: Body size medium for genus, BL males =
|
||
<quantity id="2B1CFFE5FF81FFB6D3CADAD9FE5C62F1" box="[254,406,702,726]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.1" metricValueMin="4.9" pageId="14" pageNumber="465" unit="mm" value="5.0" valueMax="5.1" valueMin="4.9">4.9 to 5.1 mm</quantity>
|
||
, females = 5.0 to
|
||
<quantity id="2B1CFFE5FF81FFB6D160DAD9FD6062F1" box="[596,682,702,726]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="465" unit="mm" value="5.7">5.7 mm</quantity>
|
||
; forebody uniformly dark, elytra dark without pale markings in basal half, in the study area with faint pale band of spots in apical half (
|
||
<figureCitation id="74DF4E85FF81FFB6D70FDAB9FB5B62D1" box="[1083,1169,734,758]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="14" pageNumber="465">Fig. 9A</figureCitation>
|
||
, mainly visible on intervals 7 and 8, femora dark black or piceous, tibiae paler; dorsal surfaces dull, with slight to distinct bronze/copper metallic reflection, elytra sericeous (silky) and mottled, especially in posterior half, with elytral microsculpture comprised of a complex pattern of transverse, longitudinal, and oblique patches of elongate meshes, most distorted near discal and umbilicate setal pore punctures; pronotum widest at or anterior to middle; elytra slightly to distinctly and obliquely truncate apically, posterior discal setiferous puncture on interval 3 inserted distinctly anterior to level of pale subapical markings; front tarsomeres 1 to 3 without lateral expansions (
|
||
<figureCitation id="74DF4E85FF81FFB6D3B7DB87FF2563DF" box="[131,239,992,1016]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="14" pageNumber="465">Figs. 14B</figureCitation>
|
||
,
|
||
<figureCitation id="74DF4E85FF81FFB6D3CEDB87FEE363DF" box="[250,297,992,1016]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="14" pageNumber="465">15B</figureCitation>
|
||
), middle tarsi of males (
|
||
<figureCitation id="74DF4E85FF81FFB6D104DB87FD5A63DF" box="[560,656,992,1016]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="14" pageNumber="465">Fig. 16B</figureCitation>
|
||
) with tarsomeres 1 to 4 broader than in females and with pads of adhesive setae ventrally (
|
||
<figureCitation id="74DF4E85FF81FFB6D170DC67FD6C643F" box="[580,678,1024,1048]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="14" pageNumber="465">Fig. 17B</figureCitation>
|
||
); median lobe of male genitalia (
|
||
<figureCitation id="74DF4E85FF81FFB6D73DDC67FB47643F" box="[1033,1165,1024,1048]" captionStart="FIGURE" captionStartId="35.[155,167,1036,1055]" captionTargetBox="[130,1144,252,995]" captionTargetId="figure-68@35.[120,1176,216,1033]" captionTargetPageId="35" captionText="FIGURE 18. Male genitalia, Tetragonoderus spp. A, B. T. arcuatus Dejean (Nepalgunj, Banke District, Nepal); C, D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). A, C. Dorsal aspect; B, D. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067329" httpUri="https://zenodo.org/record/11067329/files/figure.png" pageId="14" pageNumber="465">Figs. 18A,B</figureCitation>
|
||
) long and slender, with apical lamella also elongate, slender and rounded apically.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF81FFB6D39FDC26FB5C649E" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
|
||
In the study area, specimens of
|
||
<taxonomicName id="2BE42983FF81FFB6D2C9DC25FDB9647E" box="[509,627,1090,1113]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF81FFB6D2C9DC25FDB9647E" box="[509,627,1090,1113]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="74DF4E85FF81FFB6D1B6DC26FD1C647E" box="[642,726,1089,1113]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="14" pageNumber="465">Fig. 9A</figureCitation>
|
||
) are similar only to those of
|
||
<taxonomicName id="2BE42983FF81FFB6D73EDC25FB5C647E" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[1034,1174,1089,1113]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="parviculus">
|
||
<emphasis id="DE908E12FF81FFB6D73EDC25FB5C647E" box="[1034,1174,1089,1113]" italics="true" pageId="14" pageNumber="465">T. parviculus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="74DF4E85FF81FFB6D3B7DC06FF2C645E" box="[131,230,1121,1145]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="14" pageNumber="465">Fig. 11A</figureCitation>
|
||
) but differ from them in having larger body size, elytra with less distinct and more restricted subapical band and distinctly sericeous sheen, and male genitalia with the median lobe more slender and with a longer and narrower apical lamella (refer to key for additional differences).
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF81FFB6D39FDCA5FD94651D" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
|
||
<emphasis id="DE908E12FF81FFB6D39FDCA5FE5064FE" bold="true" box="[171,410,1218,1241]" pageId="14" pageNumber="465">Habitat distribution.</emphasis>
|
||
The lone specimen collected in the study area was found in daytime under a stone on an open beach of the Longchuan River on a mix of sandy and rocky substrate and at an elevation of
|
||
<quantity id="2B1CFFE5FF81FFB6D275DD65FE5C653D" box="[321,406,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" pageId="14" pageNumber="465" unit="m" value="1445.0">1445 m</quantity>
|
||
.
|
||
<bibRefCitation id="88752FF1FF81FFB6D295DD65FD58653D" author="ASSMANN, T. & E. BOUTAUD & J. BUSE & V. CHIKATUNOV & C. DREES & A. - L. - L. FRIEDMAN & W. HARDTLE & K. HOMBURG & T. MARCUS & I. RENAN & D. W. WRASE" box="[417,658,1282,1306]" pageId="14" pageNumber="465" pagination="49 - 69" refId="ref15907" refString="ASSMANN, T., E. BOUTAUD, J. BUSE, V. CHIKATUNOV, C. DREES, A. - L. - L. FRIEDMAN, W. HARDTLE, K. HOMBURG, T. MARCUS, I. RENAN, AND D. W. WRASE. 2015. The ground beetle tribe Cyclosomini s. l. in Israel (Coleoptera, Carabidae). Spixiana 38: 49 - 69." type="journal article" year="2015">Assmann et al. (2015)</bibRefCitation>
|
||
found members of this species “on heavy soils close to water in semi-desert areas” in
|
||
<collectingCountry id="94F31290FF81FFB6D128DD45FD91651D" box="[540,603,1314,1338]" name="Israel" pageId="14" pageNumber="465">Israel</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF81FFB6D39FDD24FF79659C" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
|
||
<emphasis id="DE908E12FF81FFB6D39FDD24FD16657D" bold="true" box="[171,732,1347,1370]" pageId="14" pageNumber="465">Geographical distribution within the Gaoligong</emphasis>
|
||
<materialsCitation id="5C8C585DFF81FFB6D1DCDD24FF65659C" ID-GBIF-Occurrence="4902827340" collectingDate="2003-10-21" collectorName="Liang H. - B. & Shi X. - C." county="Tengchong County" elevation="1445" latitude="25.23944" location="Qushi Township" longLatPrecision="1" longitude="98.61667" municipality="We" pageId="14" pageNumber="465" specimenCount="1" stateProvince="Shan">
|
||
<emphasis id="DE908E12FF81FFB6D1DCDD24FCE3657D" bold="true" box="[744,809,1347,1370]" pageId="14" pageNumber="465">
|
||
<collectingRegion id="2E209CE2FF81FFB6D1DCDD24FCEE657D" box="[744,804,1347,1370]" country="Myanmar" name="Shan" pageId="14" pageNumber="465">Shan</collectingRegion>
|
||
.
|
||
</emphasis>
|
||
<figureCitation id="74DF4E85FF81FFB6D000DD25FC46657D" box="[820,908,1346,1370]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="14" pageNumber="465">Fig. 9B</figureCitation>
|
||
.
|
||
<collectingMunicipality id="0C3FC87AFF81FFB6D0A8DD24FC75657D" box="[924,959,1347,1370]" pageId="14" pageNumber="465">We</collectingMunicipality>
|
||
examined a single female specimen from the following locality:
|
||
<emphasis id="DE908E12FF81FFB6D1A3DD04FCB0655C" bold="true" box="[663,890,1379,1403]" italics="true" pageId="14" pageNumber="465">
|
||
<collectingCounty id="053A2A8CFF81FFB6D1A3DD04FCB9655C" box="[663,883,1379,1403]" pageId="14" pageNumber="465">Tengchong County</collectingCounty>
|
||
:
|
||
</emphasis>
|
||
<location id="E93B04DBFF81FFB6D0BDDD05FB8B655D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D0BDDD05FB8B655D" box="[905,1089,1378,1402]" county="Tengchong County" latitude="25.23944" longLatPrecision="1" longitude="98.61667" municipality="We" name="Qushi Township" pageId="14" pageNumber="465" stateProvince="Shan">Qushi Township</location>
|
||
(
|
||
<location id="E93B04DBFF81FFB6D763DD05FF2B65BC" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D763DD05FF2B65BC" county="Tengchong County" latitude="25.23944" longLatPrecision="1" longitude="98.61667" municipality="We" name="Xiaojiangqiao" pageId="14" pageNumber="465" stateProvince="Shan">Xiaojiangqiao</location>
|
||
,
|
||
<geoCoordinate id="89D034C7FF81FFB6D3D9DDE4FEBA65BC" box="[237,368,1411,1435]" degrees="25.23944" direction="north" orientation="latitude" pageId="14" pageNumber="465" precision="1" value="25.23944">N25.23944°</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="89D034C7FF81FFB6D24CDDE4FDCF65BC" box="[376,517,1411,1435]" degrees="098.61667" direction="east" orientation="longitude" pageId="14" pageNumber="465" precision="1" value="98.61667">E098.61667°</geoCoordinate>
|
||
,
|
||
<quantity id="2B1CFFE5FF81FFB6D125DDE4FDAC65BC" box="[529,614,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" pageId="14" pageNumber="465" unit="m" value="1445.0">
|
||
<elevation id="67C9B533FF81FFB6D125DDE4FDAC65BC" box="[529,614,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" pageId="14" pageNumber="465" unit="m" value="1445.0">1445 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="985A74C0FF81FFB6D146DDE4FCE165BC" box="[626,811,1411,1435]" pageId="14" pageNumber="465" value="2003-10-21">
|
||
<collectingDate id="881E8D28FF81FFB6D146DDE4FCE165BC" box="[626,811,1411,1435]" pageId="14" pageNumber="465" value="2003-10-21">21 October 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="411137D6FF81FFB6D003DDE4FC7365BC" box="[823,953,1411,1435]" pageId="14" pageNumber="465">Liang H.-B.</collectorName>
|
||
&
|
||
<collectorName id="411137D6FF81FFB6D0EADDE4FB8F65BC" box="[990,1093,1411,1435]" pageId="14" pageNumber="465">Shi X.-C.</collectorName>
|
||
collectors)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF81FFB6D39FDDA4FEDF65DC" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">At present, this species has been recorded only from the southwestern part of the study area (Core Area 6).</paragraph>
|
||
<paragraph id="EC5B5200FF81FFB7D39FDE63FDEF6134" blockId="14.[123,1174,219,1725]" lastBlockId="15.[123,1174,219,505]" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465">
|
||
<emphasis id="DE908E12FF81FFB6D39FDE63FDE7663C" bold="true" box="[171,557,1540,1563]" pageId="14" pageNumber="465">Overall geographical distribution.</emphasis>
|
||
<figureCitation id="74DF4E85FF81FFB6D106DE63FD4A663B" box="[562,640,1540,1564]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="14" pageNumber="465">Fig. 23</figureCitation>
|
||
.
|
||
<collectingMunicipality id="0C3FC87AFF81FFB6D1BDDE63FD78663B" box="[649,690,1540,1564]" pageId="14" pageNumber="465">The</collectingMunicipality>
|
||
known range of this species as presently conceived extends from
|
||
<location id="E93B04DBFF81FFB6D26ADE43FE2D661B" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D26ADE43FE2D661B" box="[350,487,1572,1596]" country="Niger, Chad, Sudan" county="Lancang County" latitude="22.57925" longLatPrecision="1" longitude="99.99849" municipality="The" name="North Africa" pageId="14" pageNumber="465" stateProvince="Bihar">North Africa</location>
|
||
(
|
||
<collectingCountry id="94F31290FF81FFB6D2C2DE43FD04661B" box="[502,718,1572,1596]" isEnumeration="true" pageId="14" pageNumber="465">Niger, Chad, Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="94F31290FF81FFB6D1EFDE42FCD7661A" box="[731,797,1573,1597]" name="Egypt" pageId="14" pageNumber="465">Egypt</collectingCountry>
|
||
, and
|
||
<collectingCountry id="94F31290FF81FFB6D06CDE43FC7E661B" box="[856,948,1572,1596]" name="Ethiopia" pageId="14" pageNumber="465">Ethiopia</collectingCountry>
|
||
) eastward across the
|
||
<location id="E93B04DBFF81FFB6D34FDE23FF34667B" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D34FDE23FF34667B" box="[123,254,1604,1628]" country="Niger, Chad, Sudan" county="Lancang County" latitude="22.57925" longLatPrecision="1" longitude="99.99849" municipality="The" name="Middle East" pageId="14" pageNumber="465" stateProvince="Bihar">Middle East</location>
|
||
(
|
||
<collectingCountry id="94F31290FF81FFB6D239DE23FD4B667B" box="[269,641,1604,1628]" isEnumeration="true" pageId="14" pageNumber="465">Israel, Iraq, United Arab Emirates</collectingCountry>
|
||
,
|
||
<collectingCountry id="94F31290FF81FFB6D1B8DE23FD1A667B" box="[652,720,1604,1628]" name="Oman" pageId="14" pageNumber="465">Oman</collectingCountry>
|
||
, and
|
||
<collectingCountry id="94F31290FF81FFB6D03EDE23FC93667B" box="[778,857,1604,1628]" name="Yemen" pageId="14" pageNumber="465">Yemen</collectingCountry>
|
||
),
|
||
<location id="E93B04DBFF81FFB6D058DE23FC38667B" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D058DE23FC38667B" box="[876,1010,1604,1628]" country="Niger, Chad, Sudan" county="Lancang County" latitude="22.57925" longLatPrecision="1" longitude="99.99849" municipality="The" name="Central Asia" pageId="14" pageNumber="465" stateProvince="Bihar">Central Asia</location>
|
||
(
|
||
<collectingCountry id="94F31290FF81FFB6D735DE23FBE4667B" box="[1025,1070,1604,1628]" name="Iran" pageId="14" pageNumber="465">Iran</collectingCountry>
|
||
and
|
||
<collectingCountry id="94F31290FF81FFB6D751DE23FF7B665B" name="Pakistan" pageId="14" pageNumber="465">Pakistan</collectingCountry>
|
||
),
|
||
<collectingCountry id="94F31290FF81FFB6D3F3DE03FEC3665B" box="[199,265,1636,1660]" name="Nepal" pageId="14" pageNumber="465">Nepal</collectingCountry>
|
||
,
|
||
<collectingCountry id="94F31290FF81FFB6D22CDE03FE85665B" box="[280,335,1636,1660]" name="India" pageId="14" pageNumber="465">India</collectingCountry>
|
||
(
|
||
<collectingRegion id="2E209CE2FF81FFB6D255DE03FE55665B" box="[353,415,1636,1660]" country="India" name="Bihar" pageId="14" pageNumber="465">Bihar</collectingRegion>
|
||
,
|
||
<location id="E93B04DBFF81FFB6D29ADE03FDDB665B" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D29ADE03FDDB665B" box="[430,529,1636,1660]" country="Niger, Chad, Sudan" county="Lancang County" latitude="22.57925" longLatPrecision="1" longitude="99.99849" municipality="The" name="Jharkand" pageId="14" pageNumber="465" stateProvince="Bihar">Jharkand</location>
|
||
,
|
||
<collectingRegion id="2E209CE2FF81FFB6D114DE03FDBA665B" box="[544,624,1636,1660]" country="India" name="Sikkim" pageId="14" pageNumber="465">Sikkim</collectingRegion>
|
||
, and
|
||
<collectingRegion id="2E209CE2FF81FFB6D19ADE03FD32665B" box="[686,760,1636,1660]" country="India" name="Assam" pageId="14" pageNumber="465">Assam</collectingRegion>
|
||
)
|
||
<collectingCountry id="94F31290FF81FFB6D03FDE03FC22665B" box="[779,1000,1636,1660]" isEnumeration="true" pageId="14" pageNumber="465">Bhutan, Bangladesh</collectingCountry>
|
||
,
|
||
<collectingCountry id="94F31290FF81FFB6D0C3DE02FBAA665A" box="[1015,1120,1637,1661]" name="Myanmar" pageId="14" pageNumber="465">Myanmar</collectingCountry>
|
||
, and western
|
||
<collectingCountry id="94F31290FF81FFB6D3E3DEE3FEDD66BB" box="[215,279,1668,1692]" name="China" pageId="14" pageNumber="465">China</collectingCountry>
|
||
(western
|
||
<collectingRegion id="2E209CE2FF81FFB6D2B0DEE2FDF566BB" box="[388,575,1668,1692]" country="China" name="Yunnan" pageId="14" pageNumber="465">Yunnan Province</collectingRegion>
|
||
) and south to northwestern
|
||
<collectingCountry id="94F31290FF81FFB6D047DEE3FC1F66BB" box="[883,981,1668,1692]" name="Thailand" pageId="14" pageNumber="465">Thailand</collectingCountry>
|
||
. A large series of specimens (a total of
|
||
<specimenCount id="FAE29989FF81FFB6D246DEC2FE1C669A" box="[370,470,1701,1725]" count="95" pageId="14" pageNumber="465" type="male">95 males</specimenCount>
|
||
and females, in
|
||
<collectionCode id="8AF5CAC5FF81FFB6D1ADDEC2FD0D669A" box="[665,711,1701,1725]" pageId="14" pageNumber="465">IOZ</collectionCode>
|
||
) from
|
||
<location id="E93B04DBFF81FFB6D02DDEC2FCA5669A" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D02DDEC2FCA5669A" box="[793,879,1701,1725]" country="Niger, Chad, Sudan" county="Lancang County" latitude="22.57925" longLatPrecision="1" longitude="99.99849" municipality="The" name="Mafang" pageId="14" pageNumber="465" stateProvince="Bihar">Mafang</location>
|
||
,
|
||
<collectingCounty id="053A2A8CFF81FFB6D04BDEC2FBFC669A" box="[895,1078,1701,1725]" pageId="14" pageNumber="465">Lancang County</collectingCounty>
|
||
,
|
||
<collectorName id="411137D6FF81FFB6D770DEC2FB5F669B" box="[1092,1173,1701,1724]" pageId="14" pageNumber="465">Yunnan</collectorName>
|
||
(
|
||
<geoCoordinate id="89D034C7FF80FFB7D3B6D8BCFECE60D4" box="[130,260,219,243]" degrees="22.57925" direction="north" orientation="latitude" pageId="15" pageNumber="466" precision="1" value="22.57925">N22.57925°</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="89D034C7FF80FFB7D220D8BCFE6960D4" box="[276,419,219,243]" degrees="099.99849" direction="east" orientation="longitude" pageId="15" pageNumber="466" precision="1" value="99.99849">E099.99849°</geoCoordinate>
|
||
,
|
||
<quantity id="2B1CFFE5FF80FFB7D29BD8BCFE3660D4" box="[431,508,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="15" pageNumber="466" unit="m" value="1723.0">
|
||
<elevation id="67C9B533FF80FFB7D29BD8BCFE3660D4" box="[431,508,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="15" pageNumber="466" unit="m" value="1723.0">1723m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="411137D6FF80FFB7D132D8BCFDAF60D4" box="[518,613,219,243]" pageId="15" pageNumber="466">W.B. Gu</collectorName>
|
||
collector,
|
||
<date id="985A74C0FF80FFB7D1E2D8BCFC4A60D4" box="[726,896,219,243]" pageId="15" pageNumber="466" value="2003-12">
|
||
<collectingDate id="881E8D28FF80FFB7D1E2D8BCFC4A60D4" box="[726,896,219,243]" pageId="15" pageNumber="466" value="2003-12">December 2003</collectingDate>
|
||
</date>
|
||
-
|
||
<date id="985A74C0FF80FFB7D0B3D8BCFBD160D4" box="[903,1051,219,243]" pageId="15" pageNumber="466" value="2005-01">
|
||
<collectingDate id="881E8D28FF80FFB7D0B3D8BCFBD160D4" box="[903,1051,219,243]" pageId="15" pageNumber="466" value="2005-01">January 2005</collectingDate>
|
||
</date>
|
||
) represents the eastern most record for this species.
|
||
</paragraph>
|
||
<paragraph id="EC5B5200FF80FFB7D39FD97AFC2A61DF" blockId="15.[123,1174,219,505]" pageId="15" pageNumber="466">
|
||
<emphasis id="DE908E12FF80FFB7D39FD97AFCBA6113" bold="true" box="[171,880,284,308]" pageId="15" pageNumber="466">
|
||
Geographical relationships with other
|
||
<taxonomicName id="2BE42983FF80FFB7D152D97BFCBA6113" box="[614,880,284,308]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="DE908E12FF80FFB7D152D97BFCD86113" bold="true" box="[614,786,284,308]" italics="true" pageId="15" pageNumber="466">Tetragonoderus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</emphasis>
|
||
The geographical range of
|
||
<taxonomicName id="2BE42983FF80FFB7D34FD959FF396172" box="[123,243,318,341]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF80FFB7D34FD959FF396172" box="[123,243,318,341]" italics="true" pageId="15" pageNumber="466">T. arcuatus</emphasis>
|
||
</taxonomicName>
|
||
overlaps with several other species in different parts of its range, and these relationships will be addressed in a forthcoming review of the Asian
|
||
<taxonomicName id="2BE42983FF80FFB7D015D93AFC036152" authorityName="Dejean" authorityYear="1829" box="[801,969,349,373]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DE908E12FF80FFB7D015D93AFC036152" box="[801,969,349,373]" italics="true" pageId="15" pageNumber="466">Tetragonoderus</emphasis>
|
||
</taxonomicName>
|
||
fauna (in preparation). In the study area, its range overlaps with those of
|
||
<taxonomicName id="2BE42983FF80FFB7D1FAD918FCF361B1" box="[718,825,382,406]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="DE908E12FF80FFB7D1FAD918FCF361B1" box="[718,825,382,406]" italics="true" pageId="15" pageNumber="466">T. elegans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2BE42983FF80FFB7D073D918FC1861B1" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[839,978,382,406]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="parviculus">
|
||
<emphasis id="DE908E12FF80FFB7D073D918FC1861B1" box="[839,978,382,406]" italics="true" pageId="15" pageNumber="466">T. parviculus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2BE42983FF80FFB7D739D918FB4561B1" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[1037,1167,383,406]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="DE908E12FF80FFB7D739D918FB4561B1" box="[1037,1167,383,406]" italics="true" pageId="15" pageNumber="466">T. punctatus</emphasis>
|
||
</taxonomicName>
|
||
, although it has not yet been found syntopic with any of these species. As noted below,
|
||
<taxonomicName id="2BE42983FF80FFB7D727D9C7FF5561F0" authorityName="Jian & Tian" authorityYear="2009" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="microthorax">
|
||
<emphasis id="DE908E12FF80FFB7D727D9C7FF5561F0" italics="true" pageId="15" pageNumber="466">T. microthorax</emphasis>
|
||
</taxonomicName>
|
||
(see treatment for
|
||
<taxonomicName id="2BE42983FF80FFB7D244D9A7FE3F61F0" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[368,501,448,471]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="DE908E12FF80FFB7D244D9A7FE3F61F0" box="[368,501,448,471]" italics="true" pageId="15" pageNumber="466">T. punctatus</emphasis>
|
||
</taxonomicName>
|
||
) has not yet been recorded from the study area, but its range overlaps with that of
|
||
<taxonomicName id="2BE42983FF80FFB7D254D986FE1D61DF" box="[352,471,481,504]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="DE908E12FF80FFB7D254D986FE1D61DF" box="[352,471,481,504]" italics="true" pageId="15" pageNumber="466">T. arcuatus</emphasis>
|
||
</taxonomicName>
|
||
elsewhere and may also do so in the study area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |