401 lines
67 KiB
XML
401 lines
67 KiB
XML
<document id="D7C29E3F0A67A24F2FB45B17D34C080D" ID-CLB-Dataset="53164" ID-DOI="10.11646/zootaxa.5124.1.1" ID-GBIF-Dataset="9666d599-2c7d-4a52-bce2-a8794f03265b" ID-ISSN="1175-5326" ID-Zenodo-Dep="6404674" ID-ZooBank="C015F8CB-799B-4A92-90AE-02B4C576089E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648795871367" checkinUser="plazi" docAuthor="O’Hara, Timothy D. & Thuy, Ben" docDate="2022" docId="C72D87A1537CFFB5FEA38290FCC1FA6F" docLanguage="en" docName="zootaxa.5124.1.1.pdf" docOrigin="Zootaxa 5124 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphiura remota O’Hara & Thuy 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="38" masterDocId="3B14FFD9535EFF90FE348503FFA8FFAE" masterDocTitle="Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean" masterLastPageNumber="49" masterPageNumber="1" pageNumber="35" updateTime="1699310728982" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9A663B44F9B10A6F00DABB7CE90023D7">Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean</mods:title>
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<mods:name id="A8721A43B0AE4171CAC59FAA2A39761C" type="personal">
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<mods:namePart id="0D77986CA17DC2330B967372E5766485">O’Hara, Timothy D.</mods:namePart>
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<mods:affiliation id="115E937B833E9460F526B54C0359E6A3">Museums Victoria, GPO Box 666 E, Melbourne, 3001, AUSTRALIA,</mods:affiliation>
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<mods:namePart id="2CB75847DB5CF69127C6E6527760A156">Thuy, Ben</mods:namePart>
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<mods:affiliation id="14676AD24D369A5D5737B19972A4FDA6">Natural History Museum of Luxembourg, 24 Rue Münster, 2160 Luxembourg</mods:affiliation>
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<mods:title id="85C299D6DF46BC1D3635FA527359EAF7">Zootaxa</mods:title>
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<mods:date id="22C445D7C16EBA63E78C9286D3A44BB4">2022</mods:date>
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<treatment id="C72D87A1537CFFB5FEA38290FCC1FA6F" ID-DOI="http://doi.org/10.5281/zenodo.6404743" ID-GBIF-Taxon="194442299" ID-Zenodo-Dep="6404743" LSID="urn:lsid:plazi:treatment:C72D87A1537CFFB5FEA38290FCC1FA6F" httpUri="http://treatment.plazi.org/id/C72D87A1537CFFB5FEA38290FCC1FA6F" lastPageId="37" lastPageNumber="38" pageId="34" pageNumber="35">
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<subSubSection id="079E653C537CFFB2FEA38290FE7CF800" box="[151,468,1939,1966]" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="4F3B36B7537CFFB2FEA38290FE7CF800" blockId="34.[151,1154,1939,2039]" box="[151,468,1939,1966]" pageId="34" pageNumber="35">
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<heading id="147381DB537CFFB2FEA38290FE7CF800" bold="true" box="[151,468,1939,1966]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
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<emphasis id="7DF0EAA5537CFFB2FEA38290FE7CF800" bold="true" box="[151,468,1939,1966]" pageId="34" pageNumber="35">
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<taxonomicName id="88844D34537CFFB2FEA38290FEC7F800" ID-CoL="8ZDNJ" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[151,367,1939,1966]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="34" pageNumber="35" phylum="Echinodermata" rank="species" species="remota" status="sp. nov.">
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<emphasis id="7DF0EAA5537CFFB2FEA38290FEC7F800" bold="true" box="[151,367,1939,1966]" italics="true" pageId="34" pageNumber="35">Amphiura remota</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="66C357DE537CFFB2FF428297FE7CF800" box="[374,468,1940,1966]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="079E653C537CFFB2FEA382BBFB2AF859" pageId="34" pageNumber="35" type="description">
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<paragraph id="4F3B36B7537CFFB2FEA382BBFED9F87C" blockId="34.[151,1154,1939,2039]" box="[151,369,1976,2002]" pageId="34" pageNumber="35">
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<figureCitation id="D7BF2A32537CFFB2FEA382BBFEBCF87C" box="[151,276,1976,2002]" captionStart="FIGURE 13" captionStartId="33.[151,250,1879,1903]" captionTargetBox="[166,1421,181,1855]" captionTargetId="figure-17@33.[166,1421,181,1855]" captionTargetPageId="33" captionText="FIGURE 13. (A–B) Ophiomyxa vivipara, MNHN IE.2009.1606, A: dorsal and B: ventral views. (C–D) Ophiactis abyssicola, MNHN IE.2009.1582, C: dorsal and D: ventral views. (E–I) Amphiura remota sp. nov., holotype, MNHN IE.2019.4748, E: dorsal outline, F: dorsal, G: ventral views, with H: an inset showing the denticulate disc scales. (I-J) MNHN IE.2009.1537 (non type), I: ventral view of four-rayed specimen, J: inset showing the lateral thorn at the arm spine tip." figureDoi="http://doi.org/10.5281/zenodo.6404714" httpUri="https://zenodo.org/record/6404714/files/figure.png" pageId="34" pageNumber="35">Fig. 13E–J</figureCitation>
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,
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<figureCitation id="D7BF2A32537CFFB2FF2B82BBFED9F87C" box="[287,369,1976,2002]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="34" pageNumber="35">14A–Q</figureCitation>
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</paragraph>
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<paragraph id="4F3B36B7537CFFB2FEA382DFFB2AF859" blockId="34.[151,1154,1939,2039]" box="[151,1154,2012,2039]" pageId="34" pageNumber="35">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="079E653C537DFFB5FEA38594FE23FD2B" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="materials_examined">
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<paragraph id="4F3B36B7537DFFB3FEA38594FB92FF7B" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">
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<emphasis id="7DF0EAA5537DFFB3FEA38594FEE8FF1E" bold="true" box="[151,320,150,177]" pageId="35" pageNumber="36">Type material</emphasis>
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.
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<materialsCitation id="FFEC3CEA537DFFB3FF648595FEA4FF7B" ID-GBIF-Occurrence="3746276318" collectingDate="1986-07-24" collectionCode="MD" elevation="1515" latitude="-38.407166" location="Ile Saint-Paul" longLatPrecision="11" longitude="77.47733" pageId="35" pageNumber="36" specimenCode="MD50" specimenCount="1">
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<specimenCode id="1F229ECC537DFFB3FF648595FE31FF1F" box="[336,409,150,177]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="35" pageNumber="36">MD50</specimenCode>
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DC167, north of
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<location id="4A5B606C537DFFB3FC53859BFCAEFF1F" LSID="urn:lsid:plazi:treatment:C72D87A1537CFFB5FEA38290FCC1FA6F:4A5B606C537DFFB3FC53859BFCAEFF1F" box="[615,774,151,179]" latitude="-38.407166" longLatPrecision="11" longitude="77.47733" name="Ile Saint-Paul" pageId="35" pageNumber="36">Île Saint-Paul</location>
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,
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<geoCoordinate id="2AB05070537DFFB3FD268594FC34FF1F" box="[786,924,151,177]" degrees="38" direction="south" minutes="24.43" orientation="latitude" pageId="35" pageNumber="36" precision="9" value="-38.407166">38° 24.43´S</geoCoordinate>
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,
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<geoCoordinate id="2AB05070537DFFB3FD9E8594FB9FFF1F" box="[938,1079,151,177]" degrees="77" direction="east" minutes="28.64" orientation="longitude" pageId="35" pageNumber="36" precision="9" value="77.47733">77° 28.64´E</geoCoordinate>
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,
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<quantity id="887C9B52537DFFB3FA708594FB72FF1F" box="[1092,1242,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.515" metricValueMax="1.6" metricValueMin="1.43" pageId="35" pageNumber="36" unit="m" value="1515.0" valueMax="1600.0" valueMin="1430.0">
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<elevation id="C4A9D184537DFFB3FA708594FB72FF1F" box="[1092,1242,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.515" metricValueMax="1.6" metricValueMin="1.43" pageId="35" pageNumber="36" unit="m" value="1515.0" valueMax="1600.0" valueMin="1430.0">1430–1600m</elevation>
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, Charcot dredge,
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<date id="3B3A1077537DFFB3FEA385B8FEA4FF7B" box="[151,268,187,213]" pageId="35" pageNumber="36" value="1986-07-24">
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<collectingDate id="2B7EE99F537DFFB3FEA385B8FEA4FF7B" box="[151,268,187,213]" pageId="35" pageNumber="36" value="1986-07-24">24/7/1986</collectingDate>
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</date>
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,
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<materialsCitation id="FFEC3CEA537DFFB3FF2385B8FD22FF78" ID-GBIF-Occurrence="3746276326" box="[279,650,187,214]" collectionCode="MNHN, IE" pageId="35" pageNumber="36" specimenCount="1" typeStatus="holotype">
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<collectionCode id="2995AE72537DFFB3FF2385B8FEC5FF7B" box="[279,365,187,213]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
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<collectionCode id="2995AE72537DFFB3FF4785B8FE26FF7B" box="[371,398,187,213]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="35" pageNumber="36">IE</collectionCode>
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.2019.4748 (
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<typeStatus id="903F8815537DFFB3FC2F85B8FD2BFF78" box="[539,643,187,214]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
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)
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</materialsCitation>
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,
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<materialsCitation id="FFEC3CEA537DFFB3FCA185B8FB9EFF7B" ID-GBIF-Occurrence="3746276305" box="[661,1078,186,213]" collectionCode="MNHN, IE" pageId="35" pageNumber="36" specimenCount="16" typeStatus="paratype">
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<collectionCode id="2995AE72537DFFB3FCA185B8FD43FF7B" box="[661,747,187,213]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
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<collectionCode id="2995AE72537DFFB3FCC685B9FCA5FF7B" box="[754,781,186,213]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="35" pageNumber="36">IE</collectionCode>
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.2009.1557 (
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<specimenCount id="5982FD3E537DFFB3FDAE85B8FB87FF7B" box="[922,1071,187,213]" pageId="35" pageNumber="36" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="4F3B36B7537DFFB3FEF385DDFC3CFEB3" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">
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<emphasis id="7DF0EAA5537DFFB3FEF385DDFE55FF56" bold="true" box="[199,509,222,248]" pageId="35" pageNumber="36">Other material examined.</emphasis>
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<collectionCode id="2995AE72537DFFB3FC3185DDFD9AFF57" box="[517,562,222,249]" country="Germany" name="Museum Donaueschingen" pageId="35" pageNumber="36">MD</collectionCode>
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50 DC34,
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<collectionCode id="2995AE72537DFFB3FC9385DCFD55FF57" box="[679,765,223,249]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
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<collectionCode id="2995AE72537DFFB3FD3185DDFC88FF57" box="[773,800,222,249]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="35" pageNumber="36">IE</collectionCode>
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.2009.1535 (2).
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<collectionCode id="2995AE72537DFFB3FDE085DDFBA9FF57" box="[980,1025,222,249]" country="Germany" name="Museum Donaueschingen" pageId="35" pageNumber="36">MD</collectionCode>
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50 DC55,
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<collectionCode id="2995AE72537DFFB3FA4385DCFB65FF57" box="[1143,1229,223,249]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
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<collectionCode id="2995AE72537DFFB3FAE085DDFB47FF57" box="[1236,1263,222,249]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="35" pageNumber="36">IE</collectionCode>
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.2009.1537 (1).
|
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<collectionCode id="2995AE72537DFFB3FEA38401FF6CFEB3" box="[151,196,258,285]" country="Germany" name="Museum Donaueschingen" pageId="35" pageNumber="36">MD</collectionCode>
|
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50 DC137,
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<collectionCode id="2995AE72537DFFB3FF728400FE34FEB3" box="[326,412,259,285]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
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<collectionCode id="2995AE72537DFFB3FF978401FE16FEB3" box="[419,446,258,285]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="35" pageNumber="36">IE</collectionCode>
|
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.2009.1552 (1),
|
||
<collectionCode id="2995AE72537DFFB3FC458400FD6FFEB3" box="[625,711,259,285]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
|
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<collectionCode id="2995AE72537DFFB3FCF98401FD40FEB3" box="[717,744,258,285]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="35" pageNumber="36">IE</collectionCode>
|
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.2009.1551 (3).
|
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</paragraph>
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<paragraph id="4F3B36B7537DFFB3FEF38424FCD4FD07" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">
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<emphasis id="7DF0EAA5537DFFB3FEF38424FE68FEEE" bold="true" box="[199,448,294,321]" pageId="35" pageNumber="36">
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<typeStatus id="903F8815537DFFB3FEF38424FE9CFEEF" box="[199,308,295,321]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
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description
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</emphasis>
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. Disc is
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||
<quantity id="887C9B52537DFFB3FC2A8424FDDBFEEF" box="[542,627,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="35" pageNumber="36" unit="mm" value="4.4">4.4 mm</quantity>
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dd, petaloid, indented interradially, covered in overlapping scales, 15–16 scales from disc centre to margin, some primary scales are evident (0.3 long,
|
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<quantity id="887C9B52537DFFB3FDC28448FBE4FECB" box="[1014,1100,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="35" pageNumber="36" unit="mm" value="0.4">0.4 mm</quantity>
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wide), separate, scales in disc centre
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<quantity id="887C9B52537DFFB3FED0846CFE3AFE27" box="[228,402,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="3.0" metricValueMin="1.7999999999999998" pageId="35" pageNumber="36" unit="mm" value="0.24" valueMax="0.3" valueMin="0.18">0.18 to 0.3 mm</quantity>
|
||
wide, becoming abruptly smaller and denser near the disc margin and outside of the radial shields (
|
||
<quantity id="887C9B52537DFFB3FEC08491FE1EFE03" box="[244,438,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="1.75" metricValueMin="1.25" pageId="35" pageNumber="36" unit="mm" value="0.15" valueMax="0.175" valueMin="0.125">
|
||
<geoCoordinate id="2AB05070537DFFB3FEC08491FE9CFE03" box="[244,308,402,429]" degrees="0.125" direction="north" orientation="latitude" pageId="35" pageNumber="36" precision="55" value="0.125">0.125</geoCoordinate>
|
||
<geoCoordinate id="2AB05070537DFFB3FF008491FE2BFE03" box="[308,387,402,429]" degrees="0.175" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="55" value="-0.175">–0.175</geoCoordinate>
|
||
mm
|
||
</quantity>
|
||
wide). A slender triangular scale (
|
||
<quantity id="887C9B52537DFFB3FD0F8490FC0AFE03" box="[827,930,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="35" pageNumber="36" unit="mm" value="0.86">0.86 mm</quantity>
|
||
long) separates the radial shields (
|
||
<figureCitation id="D7BF2A32537DFFB3FB1F8490FA38FE03" box="[1323,1424,403,429]" captionStart="FIGURE 13" captionStartId="33.[151,250,1879,1903]" captionTargetBox="[166,1421,181,1855]" captionTargetId="figure-17@33.[166,1421,181,1855]" captionTargetPageId="33" captionText="FIGURE 13. (A–B) Ophiomyxa vivipara, MNHN IE.2009.1606, A: dorsal and B: ventral views. (C–D) Ophiactis abyssicola, MNHN IE.2009.1582, C: dorsal and D: ventral views. (E–I) Amphiura remota sp. nov., holotype, MNHN IE.2019.4748, E: dorsal outline, F: dorsal, G: ventral views, with H: an inset showing the denticulate disc scales. (I-J) MNHN IE.2009.1537 (non type), I: ventral view of four-rayed specimen, J: inset showing the lateral thorn at the arm spine tip." figureDoi="http://doi.org/10.5281/zenodo.6404714" httpUri="https://zenodo.org/record/6404714/files/figure.png" pageId="35" pageNumber="36">Fig. 13F</figureCitation>
|
||
). Scales have microscopic glassy beads, marginal scales have finely denticulate margins (
|
||
<figureCitation id="D7BF2A32537DFFB3FA4A84B4FB40FE7F" box="[1150,1256,439,465]" captionStart="FIGURE 13" captionStartId="33.[151,250,1879,1903]" captionTargetBox="[166,1421,181,1855]" captionTargetId="figure-17@33.[166,1421,181,1855]" captionTargetPageId="33" captionText="FIGURE 13. (A–B) Ophiomyxa vivipara, MNHN IE.2009.1606, A: dorsal and B: ventral views. (C–D) Ophiactis abyssicola, MNHN IE.2009.1582, C: dorsal and D: ventral views. (E–I) Amphiura remota sp. nov., holotype, MNHN IE.2019.4748, E: dorsal outline, F: dorsal, G: ventral views, with H: an inset showing the denticulate disc scales. (I-J) MNHN IE.2009.1537 (non type), I: ventral view of four-rayed specimen, J: inset showing the lateral thorn at the arm spine tip." figureDoi="http://doi.org/10.5281/zenodo.6404714" httpUri="https://zenodo.org/record/6404714/files/figure.png" pageId="35" pageNumber="36">Fig. 13H</figureCitation>
|
||
). Radial shields are long (
|
||
<quantity id="887C9B52537DFFB3FF3784D8FEF3FE5B" box="[259,347,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="mm" value="0.3">0.3 mm</quantity>
|
||
wide and
|
||
<quantity id="887C9B52537DFFB3FFE584D8FD80FE5B" box="[465,552,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="35" pageNumber="36" unit="mm" value="1.1">1.1 mm</quantity>
|
||
long), contiguous distally but separated proximally by the long triangular and a few smaller rounded scales, rounded distally, tapered for the proximal 2/3 of their length and terminating in a sharp angle. Ventral disc surfaces (
|
||
<figureCitation id="D7BF2A32537DFFB3FC168720FD20FD93" box="[546,648,547,573]" captionStart="FIGURE 13" captionStartId="33.[151,250,1879,1903]" captionTargetBox="[166,1421,181,1855]" captionTargetId="figure-17@33.[166,1421,181,1855]" captionTargetPageId="33" captionText="FIGURE 13. (A–B) Ophiomyxa vivipara, MNHN IE.2009.1606, A: dorsal and B: ventral views. (C–D) Ophiactis abyssicola, MNHN IE.2009.1582, C: dorsal and D: ventral views. (E–I) Amphiura remota sp. nov., holotype, MNHN IE.2019.4748, E: dorsal outline, F: dorsal, G: ventral views, with H: an inset showing the denticulate disc scales. (I-J) MNHN IE.2009.1537 (non type), I: ventral view of four-rayed specimen, J: inset showing the lateral thorn at the arm spine tip." figureDoi="http://doi.org/10.5281/zenodo.6404714" httpUri="https://zenodo.org/record/6404714/files/figure.png" pageId="35" pageNumber="36">Fig. 13G</figureCitation>
|
||
) have similar scales to dorsal margin, becoming less overlapped near oral shield. Genital slits are open along the first 3 arm segments, bordered by a prominent adradial genital scale that persists proximally until the 2nd set of arm spines, where it is followed along the slit by a series of stout plates until the distal projection of the oral plates to the rear of the oral shield.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7537DFFB3FEF387B0FBD0FC67" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">
|
||
Jaws are wider than long, slit open and teeth form a buccal funnel (sensu
|
||
<bibRefCitation id="2B154B46537DFFB3FDD887B0FB21FD63" author="Hendler, G." box="[1004,1161,691,717]" pageId="35" pageNumber="36" pagination="189 - 311" refId="ref31642" refString="Hendler, G. (2018) Armed to the teeth: a new paradigm for the buccal skeleton of brittle stars (Echinodermata: Ophiuroidea). Contributions in Science, 526, 189 - 311. https: // doi. org / 10.5962 / p. 324539" type="journal article" year="2018">Hendler 2018</bibRefCitation>
|
||
). Oral shield spear-headshaped, as wide as long, with a small distal lobe. Madreporite is larger than other shields, longer than wide with a prominent wide distal lobe. Adoral shields are slightly separated interradially, with a concave proximal margin, rounded lateral angles and extending distally to border the genital slit; exposed surface of the oral plate rectangular, is separated distally. Four teeth are visible, upper ones rounded, lower ones truncate and longer, all bordered by imperforate stereom. Infradental papillae are pentagonal, square at their base and pointed proximally, a small conical buccal scale adjacent to each infradental papilla on the oral plate, tall flask-shaped to conical adoral shield spine, 2 times as long as wide, sitting proximoventral to the large exposed 2nd oral tentacle pore.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7537DFFB3FEF386D1FA81FB6C" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">DAPs are fan-shaped then transversely ovoid, 1.3 times wider than long, with the glassy distal margin broadly convex and the proximal side produced into a very obtuse angle, separate. First VAPs are hourglass-shaped, with concave sides formed by the rounded edge of the adoral shields, a little longer than wide, with an obtuse proximal angle and truncate distal edge, contiguous with the 2nd VAP which is 1.25 times longer than wide with a straight proximal margin, divergent proximolateral sides, concave lateral margins around the large tentacle pores, and a rounded distal edge. Succeeding plates are similar but becoming progressively narrower and more tapered proximally, so that they are 2 times as long as wide by the 10th segment and only narrowly contiguous with neighbouring plates.</paragraph>
|
||
<paragraph id="4F3B36B7537DFFB3FEF381CEFB6CFADB" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">
|
||
Two arm spines are on the first segments, thereafter 3 arise from the lateral side of the arm. Spines are long and rounded, gradually tapering to a fine rounded tip, composed of sculptured stereom and fine thorns, the 2 ventral ones are longest, to
|
||
<quantity id="887C9B52537DFFB3FF088010FE0AFA80" box="[316,418,1299,1326]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="35" pageNumber="36" unit="mm" value="0.55">0.55 mm</quantity>
|
||
, the dorsal one is more slender and 0.6 times shorter. One or two middle arm spines have a diminutive thorn at the tip. Two small tentacle scales occur at right angles to each other around each tentacle pore, one on the LAP and a slightly smaller one on the VAP, which do not completely cover the pore.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7537DFFB3FEF3807CFE01FA37" blockId="35.[151,1437,150,2005]" box="[199,425,1407,1434]" pageId="35" pageNumber="36">Colour (dry): white.</paragraph>
|
||
<paragraph id="4F3B36B7537DFFB3FEF380A1FC80F9E5" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">
|
||
<emphasis id="7DF0EAA5537DFFB3FEF380A1FE13FA12" bold="true" box="[199,443,1442,1468]" pageId="35" pageNumber="36">
|
||
<typeStatus id="903F8815537DFFB3FEF380A1FE9CFA12" box="[199,308,1442,1468]" pageId="35" pageNumber="36" type="paratype">Paratype</typeStatus>
|
||
variations.
|
||
</emphasis>
|
||
Other specimens from stn DC167 measure
|
||
<quantity id="887C9B52537DFFB3FD9F80A1FBE2FA13" box="[939,1098,1442,1469]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.5" metricValueMin="3.5" pageId="35" pageNumber="36" unit="mm" value="5.5" valueMax="7.5" valueMin="3.5">3.5 to 7.5 mm</quantity>
|
||
dd. One has two oral tentacle scales on one jaw. On some specimens, the adoral shield spine is very tall and slender after the swollen base, up to 4 times as tall as wide. On smaller specimens the primary plates are relatively large. All specimens have a white or pale-grey disc and white arms. A non-type specimen from stn DC55 is four-rayed (
|
||
<figureCitation id="D7BF2A32537DFFB3FA13830EFB28F986" box="[1063,1152,1549,1576]" captionStart="FIGURE 13" captionStartId="33.[151,250,1879,1903]" captionTargetBox="[166,1421,181,1855]" captionTargetId="figure-17@33.[166,1421,181,1855]" captionTargetPageId="33" captionText="FIGURE 13. (A–B) Ophiomyxa vivipara, MNHN IE.2009.1606, A: dorsal and B: ventral views. (C–D) Ophiactis abyssicola, MNHN IE.2009.1582, C: dorsal and D: ventral views. (E–I) Amphiura remota sp. nov., holotype, MNHN IE.2019.4748, E: dorsal outline, F: dorsal, G: ventral views, with H: an inset showing the denticulate disc scales. (I-J) MNHN IE.2009.1537 (non type), I: ventral view of four-rayed specimen, J: inset showing the lateral thorn at the arm spine tip." figureDoi="http://doi.org/10.5281/zenodo.6404714" httpUri="https://zenodo.org/record/6404714/files/figure.png" pageId="35" pageNumber="36">Fig. 13I</figureCitation>
|
||
) and has notable distallydirected thorns on the tips of middle arm spines (
|
||
<figureCitation id="D7BF2A32537DFFB3FC888332FCB3F9E5" box="[700,795,1585,1611]" captionStart="FIGURE 13" captionStartId="33.[151,250,1879,1903]" captionTargetBox="[166,1421,181,1855]" captionTargetId="figure-17@33.[166,1421,181,1855]" captionTargetPageId="33" captionText="FIGURE 13. (A–B) Ophiomyxa vivipara, MNHN IE.2009.1606, A: dorsal and B: ventral views. (C–D) Ophiactis abyssicola, MNHN IE.2009.1582, C: dorsal and D: ventral views. (E–I) Amphiura remota sp. nov., holotype, MNHN IE.2019.4748, E: dorsal outline, F: dorsal, G: ventral views, with H: an inset showing the denticulate disc scales. (I-J) MNHN IE.2009.1537 (non type), I: ventral view of four-rayed specimen, J: inset showing the lateral thorn at the arm spine tip." figureDoi="http://doi.org/10.5281/zenodo.6404714" httpUri="https://zenodo.org/record/6404714/files/figure.png" pageId="35" pageNumber="36">Fig. 13J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7537DFFB3FEF38356FB88F87B" blockId="35.[151,1437,150,2005]" pageId="35" pageNumber="36">
|
||
<emphasis id="7DF0EAA5537DFFB3FEF38356FD8CF9C1" bold="true" box="[199,548,1621,1647]" pageId="35" pageNumber="36">
|
||
<typeStatus id="903F8815537DFFB3FEF38356FE9CF9C1" box="[199,308,1621,1647]" pageId="35" pageNumber="36" type="paratype">Paratype</typeStatus>
|
||
ossicle description.
|
||
</emphasis>
|
||
Proximal (
|
||
<figureCitation id="D7BF2A32537DFFB3FC9E8356FCBCF9C1" box="[682,788,1621,1647]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="35" pageNumber="36">Fig. 14A</figureCitation>
|
||
) and median (
|
||
<figureCitation id="D7BF2A32537DFFB3FDF58356FBE3F9C1" box="[961,1099,1621,1647]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="35" pageNumber="36">Fig. 14B–C</figureCitation>
|
||
) LAPs are two times higher than long, strongly arched, relatively thin, with strongly protruding ventral portion; dorsal edge conspicuously angular; outer surface with moderately finely meshed stereom with trabecular intersections transformed into tiny tubercles; outer proximal edge with a central area of more finely meshed stereom. Spine articulations (
|
||
<figureCitation id="D7BF2A32537DFFB3FB1983C3FA3CF974" box="[1325,1428,1728,1754]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="35" pageNumber="36">Fig. 14D</figureCitation>
|
||
) are freestanding in the centre of the distal edge, on the same level as the outer surface stereom, moderately large, composed of parallel and completely separated lobes, the dorsal one relatively slender and slightly arched and the ventral one shorter and thicker, encompassing a moderately large muscle opening and a slightly smaller nerve opening. Inner side of LAP has three round, moderately well-defined knobs, a large one close to the centre of the inner proximal edge, a second smaller knob in the dorso-proximal corner of the inner side of the LAP, and a third smaller knob between the large one and the distal edge of the LAP; single small perforation on the ventro-distal edge of the third small knob; tentacle notch large but very shallow and inconspicuous.
|
||
</paragraph>
|
||
<caption id="1BFB663F537AFFB4FEA3821CFD4CF845" ID-DOI="http://doi.org/10.5281/zenodo.6404716" ID-Zenodo-Dep="6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1823,1847]" targetBox="[177,1410,181,1799]" targetPageId="36">
|
||
<paragraph id="4F3B36B7537AFFB4FEA3821CFD4CF845" blockId="36.[151,1437,1823,2027]" pageId="36" pageNumber="37">
|
||
<emphasis id="7DF0EAA5537AFFB4FEA3821CFEB7F899" bold="true" box="[151,287,1823,1847]" pageId="36" pageNumber="37">FIGURE 14.</emphasis>
|
||
(A–Q)
|
||
<taxonomicName id="88844D34537AFFB4FF59821CFDB7F899" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[365,543,1823,1847]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="36" pageNumber="37" phylum="Echinodermata" rank="species" species="remota" status="sp. nov.">
|
||
<emphasis id="7DF0EAA5537AFFB4FF59821CFDB7F899" box="[365,543,1823,1847]" italics="true" pageId="36" pageNumber="37">Amphiura remota</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7DF0EAA5537AFFB4FC10821CFDDCF899" bold="true" box="[548,628,1823,1847]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="66C357DE537AFFB4FC10821CFDDCF899" box="[548,628,1823,1847]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, MNHN IE.2009.1557. A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4F3B36B7537BFFB5FEF38594FDE3FEEF" blockId="37.[151,1437,151,1473]" pageId="37" pageNumber="38">
|
||
Vertebrae (
|
||
<figureCitation id="D7BF2A32537BFFB5FF728594FE6DFF1F" box="[326,453,151,177]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14F–J</figureCitation>
|
||
) are small, delicate, with relatively small, rounded muscle fossae on the distal and proximal faces; articulations very small, typically zygospondylous, with a large zygosphene on the distal face; dorso-distal muscle fossae deeply carved into the lateral side of the vertebra, with converging dorso-distal tips strongly elongated and pointed, almost projecting beyond zygocondyles; very large, deep ventral furrow, with large, moderately well-defined podial basins.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7537BFFB5FEF38448FE23FD2B" blockId="37.[151,1437,151,1473]" pageId="37" pageNumber="38">
|
||
Oral plates (
|
||
<figureCitation id="D7BF2A32537BFFB5FF648448FE12FECB" box="[336,442,331,357]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14M</figureCitation>
|
||
) are nearly as high as long, with adradial muscle fossa transformed into a vertical spoonshaped depression lining the entire lateral edge of articulation area. Dental plates (
|
||
<figureCitation id="D7BF2A32537BFFB5FA16846CFB2EFE27" box="[1058,1158,367,393]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14N</figureCitation>
|
||
) are undivided, rounded, with single row of large, tooth sockets surrounded by separate, strongly protruding knobs; dorsalmost tooth socket very large, perforating dental plate; second dorsalmost tooth socket smaller, penetrating dental plate but with a vertical septum. Abradial genital plates (
|
||
<figureCitation id="D7BF2A32537BFFB5FC6C84D8FD15FE5B" box="[600,701,475,501]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14O</figureCitation>
|
||
) are slightly shorter than the adradial one (
|
||
<figureCitation id="D7BF2A32537BFFB5FAAC84D8FB52FE5B" box="[1176,1274,475,501]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14P</figureCitation>
|
||
), very slender, bar-like, bent, without ridges or groves and with a small, concave adradio-distal tip. Radial shields (
|
||
<figureCitation id="D7BF2A32537BFFB5FAC784FCFAF1FDB7" box="[1267,1369,511,537]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14Q</figureCitation>
|
||
) have an irregular, rounded triangular outline, slightly oblique, with a more finely-meshed stereom in the middle part of the outer surface. A proximal dorsal arm spine (
|
||
<figureCitation id="D7BF2A32537BFFB5FC868744FCB0FDCF" box="[690,792,583,609]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14E</figureCitation>
|
||
), dorsal arm plate (
|
||
<figureCitation id="D7BF2A32537BFFB5FDC68744FBF3FDCF" box="[1010,1115,583,609]" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14K</figureCitation>
|
||
), and ventral arm plate (
|
||
<figureCitation id="D7BF2A32537BFFB5FB5B8744FF6EFD2B" captionStart="FIGURE 14" captionStartId="36.[151,250,1823,1847]" captionTargetBox="[177,1410,181,1799]" captionTargetId="figure-26@36.[177,1410,181,1799]" captionTargetPageId="36" captionText="FIGURE 14. (A–Q) Amphiura remota sp. nov., paratype, MNHN IE.2009.1557.A: proximal lateral arm plate in external view. B–C: median lateral arm plate in external (B) and internal (C) views. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N: dental plate. O: abradial genital plate. P: adradial genital plate. Q: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; K = knob; MO = muscle opening; NO = nerve opening; P = perforation; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404716" httpUri="https://zenodo.org/record/6404716/files/figure.png" pageId="37" pageNumber="38">Fig. 14L</figureCitation>
|
||
) are also figured.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="079E653C537BFFB5FEF3878DFDFAFD07" box="[199,594,654,681]" pageId="37" pageNumber="38" type="distribution">
|
||
<paragraph id="4F3B36B7537BFFB5FEF3878DFDFAFD07" blockId="37.[151,1437,151,1473]" box="[199,594,654,681]" pageId="37" pageNumber="38">
|
||
<emphasis id="7DF0EAA5537BFFB5FEF3878DFEF1FD06" bold="true" box="[199,345,654,680]" pageId="37" pageNumber="38">Distribution</emphasis>
|
||
. SPA (
|
||
<quantity id="887C9B52537BFFB5FF91878DFDEEFD07" box="[421,582,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.6075" metricValueMax="2.2" metricValueMin="1.015" pageId="37" pageNumber="38" unit="m" value="1607.5" valueMax="2200.0" valueMin="1015.0">1015–2200 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="079E653C537BFFB5FEF387B0FE38FAD7" pageId="37" pageNumber="38" type="discussion">
|
||
<paragraph id="4F3B36B7537BFFB5FEF387B0FB67FC0B" blockId="37.[151,1437,151,1473]" pageId="37" pageNumber="38">
|
||
<emphasis id="7DF0EAA5537BFFB5FEF387B0FE9CFD63" bold="true" box="[199,308,691,717]" pageId="37" pageNumber="38">Remarks</emphasis>
|
||
. This species shares many morphological features with the
|
||
<taxonomicName id="88844D34537BFFB5FDED87B0FB08FD63" authorityName="Koehler" authorityYear="1912" box="[985,1184,691,717]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="joubini">
|
||
<emphasis id="7DF0EAA5537BFFB5FDED87B0FB08FD63" box="[985,1184,691,717]" italics="true" pageId="37" pageNumber="38">Amphiura joubini</emphasis>
|
||
</taxonomicName>
|
||
-group of species, that includes
|
||
<taxonomicName id="88844D34537BFFB5FECF87D4FDBAFD5F" authority="Koehler, 1912" authorityName="Koehler" authorityYear="1912" box="[251,530,727,753]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="joubini">
|
||
<emphasis id="7DF0EAA5537BFFB5FECF87D4FEC1FD5F" box="[251,361,727,753]" italics="true" pageId="37" pageNumber="38">A. joubini</emphasis>
|
||
<bibRefCitation id="2B154B46537BFFB5FF4487D4FDBAFD5F" author="Koehler, R." box="[368,530,727,753]" pageId="37" pageNumber="38" refId="ref32381" refString="Koehler, R. (1912) Echinodermes (Asteries, Ophiures et Echinides). In: Deuxieme Expedition Antarctique Francaise (1908 - 10), Paris, pp. 1 - 272, 16 pls. https: // doi. org / 10.5962 / bhl. title. 85347" type="book" year="1912">Koehler, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D34537BFFB5FC2887D4FC3AFD5F" authority="Hertz, 1927 a" authorityName="Hertz" authorityYear="1927" box="[540,914,727,753]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="subSpecies" species="joubini" subSpecies="abyssalis">
|
||
<emphasis id="7DF0EAA5537BFFB5FC2887D4FD50FD5F" box="[540,760,727,753]" italics="true" pageId="37" pageNumber="38">A. joubini abyssalis</emphasis>
|
||
<bibRefCitation id="2B154B46537BFFB5FCCB87D4FC3AFD5F" author="Hertz, M." box="[767,914,727,753]" pageId="37" pageNumber="38" pagination="1 - 56" refId="ref31692" refString="Hertz, M. (1927 a) Die Ophiuroiden der Deutschen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition, 19, 1 - 56, pls. 1 - 9." type="journal article" year="1927">Hertz, 1927a</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D34537BFFB5FDA987D4FA9EFD5F" authority="Mortensen, 1936" authorityName="Mortensen" authorityYear="1936" box="[925,1334,727,753]" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="acutus">
|
||
<emphasis id="7DF0EAA5537BFFB5FDA987D4FBDAFD5E" box="[925,1138,727,753]" italics="true" pageId="37" pageNumber="38">Amphioplus acutus</emphasis>
|
||
<bibRefCitation id="2B154B46537BFFB5FA4D87D4FA9EFD5F" author="Mortensen, T." box="[1145,1334,727,753]" pageId="37" pageNumber="38" pagination="199 - 348" refId="ref34461" refString="Mortensen, T. (1936) Echinoidea and Ophiuroidea. Discovery Reports, 12, 199 - 348, 9 pls. https: // doi. org / 10.5962 / bhl. part. 8051" type="journal article" year="1936">Mortensen, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Southern Ocean;
|
||
<taxonomicName id="88844D34537BFFB5FF6387F8FD77FCBB" authority="Lyman, 1879" authorityName="Lyman" authorityYear="1879" box="[343,735,763,789]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="lanceolata">
|
||
<emphasis id="7DF0EAA5537BFFB5FF6387F8FDEBFCBB" box="[343,579,763,789]" italics="true" pageId="37" pageNumber="38">Amphiura lanceolata</emphasis>
|
||
<bibRefCitation id="2B154B46537BFFB5FC7D87F8FD77FCBB" author="Lyman, T." box="[585,735,763,789]" pageId="37" pageNumber="38" pagination="17 - 83" refId="ref33210" refString="Lyman, T. (1879) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger under Prof. Sir Wyville Thomson, F. R. S. Part 2. Bulletin of the Museum of Comparative Zoology, Harvard University, 6, 17 - 83, pls. 11 - 19." type="journal article" year="1879">Lyman, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D34537BFFB5FCDD87F8FB25FCBB" authority="Baker, 1977" authorityName="Baker" authorityYear="1977" box="[745,1165,763,789]" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="ctenacantha">
|
||
<emphasis id="7DF0EAA5537BFFB5FCDD87F8FC56FCBB" box="[745,1022,763,789]" italics="true" pageId="37" pageNumber="38">Amphioplus ctenacantha</emphasis>
|
||
<bibRefCitation id="2B154B46537BFFB5FA3087F8FB25FCBB" author="Baker, A. N." box="[1028,1165,763,789]" pageId="37" pageNumber="38" pagination="149 - 160" refId="ref29158" refString="Baker, A. N. (1977) Some deep-sea Ophiuroidea from New Zealand. Records of the National Museum of New Zealand, 1, 149 - 160." type="journal article" year="1977">Baker, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D34537BFFB5FAA387F8FEC8FC97" authority="Baker, 1977" authorityName="Baker" authorityYear="1977" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="cipus" subGenus="Unioplus">
|
||
<emphasis id="7DF0EAA5537BFFB5FAA387F8FF7AFC96" italics="true" pageId="37" pageNumber="38">Amphioplus (Unioplus) cipus</emphasis>
|
||
<bibRefCitation id="2B154B46537BFFB5FEED861CFEC8FC97" author="Baker, A. N." box="[217,352,799,825]" pageId="37" pageNumber="38" pagination="149 - 160" refId="ref29158" refString="Baker, A. N. (1977) Some deep-sea Ophiuroidea from New Zealand. Records of the National Museum of New Zealand, 1, 149 - 160." type="journal article" year="1977">Baker, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="37937627537BFFB5FF90861CFD95FC97" box="[420,573,799,825]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
|
||
; and
|
||
<taxonomicName id="88844D34537BFFB5FC43861CFC41FC97" authority="Ljungman, 1872" authorityName="Ljungman" authorityYear="1872" box="[631,1001,799,825]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="otteri">
|
||
<emphasis id="7DF0EAA5537BFFB5FC43861CFC82FC96" box="[631,810,799,825]" italics="true" pageId="37" pageNumber="38">Amphiura otteri</emphasis>
|
||
<bibRefCitation id="2B154B46537BFFB5FD05861CFC41FC97" author="Ljungman, A. V." box="[817,1001,799,825]" pageId="37" pageNumber="38" pagination="615 - 657" refId="ref32907" refString="Ljungman, A. V. (1872) Forteckning ofver uti Vestindien af Dr. A. Goes samt under korvetten Josefinas expedition i Atlantiska Oceanen samlade Ophiurider. Ofversigt af Kongl. Vetenskaps-Akademiens Forhandlingar. Stockholm, 28, 615 - 657." type="journal article" year="1872">Ljungman, 1872</bibRefCitation>
|
||
</taxonomicName>
|
||
from the North Atlantic. They all form a monophyletic clade on the phylogenies in
|
||
<bibRefCitation id="2B154B46537BFFB5FCB28640FCC4FCF3" author="O'Hara, T. D. & Hugall, A. F. & Thuy, B. & Stohr, S. & Martynov, A. V." box="[646,876,835,861]" pageId="37" pageNumber="38" pagination="415 - 430" refId="ref34885" refString="O'Hara, T. D., Hugall, A. F., Thuy, B., Stohr, S. & Martynov, A. V. (2017) Restructuring higher taxonomy using broad-scale phylogenomics: the living Ophiuroidea. Molecular Phylogenetics and Evolution, 107, 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
|
||
O’Hara
|
||
<emphasis id="7DF0EAA5537BFFB5FCD58640FCB2FCF3" box="[737,794,835,861]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="2B154B46537BFFB5FD968640FB7FFCF3" author="Christodoulou, M. & O'Hara, T. D. & Hugall, A. F. & Arbizu, P. M." box="[930,1239,835,861]" pageId="37" pageNumber="38" pagination="3909 - 3912" refId="ref29723" refString="Christodoulou, M., O'Hara, T. D., Hugall, A. F. & Arbizu, P. M. (2019) Dark Ophiuroid Biodiversity in a Prospective Abyssal Mine Field. Current Biology, 29 (e 3), 3909 - 3912." type="journal article" year="2019">
|
||
Christodoulou
|
||
<emphasis id="7DF0EAA5537BFFB5FA7E8640FB2AFCF3" box="[1098,1154,835,861]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
. They share discs with long divergent radial shields, fine disc scales, spear-head-shaped oral shields, one to two long spiniform adoral shield spines, long pointed arm spines often with a terminal thorn, and two small tentacle scales.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7537BFFB5FEF386ACFC2BFBD3" blockId="37.[151,1437,151,1473]" pageId="37" pageNumber="38">
|
||
Here, all these species are reassigned to
|
||
<taxonomicName id="88844D34537BFFB5FCA186ACFCADFC67" authorityName="Forbes" authorityYear="1843" box="[661,773,943,969]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA5537BFFB5FCA186ACFCADFC67" box="[661,773,943,969]" italics="true" pageId="37" pageNumber="38">Amphiura</emphasis>
|
||
</taxonomicName>
|
||
pending a revision of this large genus. The fact that these species are spread over
|
||
<taxonomicName id="88844D34537BFFB5FF8286D0FD8EFC43" authorityName="Forbes" authorityYear="1843" box="[438,550,979,1005]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA5537BFFB5FF8286D0FD8EFC43" box="[438,550,979,1005]" italics="true" pageId="37" pageNumber="38">Amphiura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D34537BFFB5FC0E86D0FD68FC43" authorityName="Verrill" authorityYear="1899" box="[570,704,979,1005]" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA5537BFFB5FC0E86D0FD68FC43" box="[570,704,979,1005]" italics="true" pageId="37" pageNumber="38">Amphioplus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D34537BFFB5FCE786D0FC76FC43" authority="" box="[723,990,979,1005]" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="subGenus" subGenus="Unioplus">
|
||
<emphasis id="7DF0EAA5537BFFB5FCE786D0FCF1FC43" box="[723,857,979,1005]" italics="true" pageId="37" pageNumber="38">Amphioplus</emphasis>
|
||
(
|
||
<emphasis id="7DF0EAA5537BFFB5FD5A86D0FC7DFC43" box="[878,981,979,1005]" italics="true" pageId="37" pageNumber="38">Unioplus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
and (formerly)
|
||
<taxonomicName id="88844D34537BFFB5FA9486D0FAB3FC43" authorityName="Verrill" authorityYear="1899" box="[1184,1307,979,1005]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiodia" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA5537BFFB5FA9486D0FAB3FC43" box="[1184,1307,979,1005]" italics="true" pageId="37" pageNumber="38">Amphiodia</emphasis>
|
||
</taxonomicName>
|
||
shows the artificiality of these genera (
|
||
<bibRefCitation id="2B154B46537BFFB5FFE786F4FD04FBBF" author="O'Hara, T. D. & Hugall, A. F. & Thuy, B. & Stohr, S. & Martynov, A. V." box="[467,684,1015,1041]" pageId="37" pageNumber="38" pagination="415 - 430" refId="ref34885" refString="O'Hara, T. D., Hugall, A. F., Thuy, B., Stohr, S. & Martynov, A. V. (2017) Restructuring higher taxonomy using broad-scale phylogenomics: the living Ophiuroidea. Molecular Phylogenetics and Evolution, 107, 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
|
||
O’Hara
|
||
<emphasis id="7DF0EAA5537BFFB5FC0486F4FDC1FBBF" box="[560,617,1015,1041]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). Species in the
|
||
<taxonomicName id="88844D34537BFFB5FD5686F4FC19FBBF" authorityName="Koehler" authorityYear="1912" box="[866,945,1015,1041]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="joubini">
|
||
<emphasis id="7DF0EAA5537BFFB5FD5686F4FC19FBBF" box="[866,945,1015,1041]" italics="true" pageId="37" pageNumber="38">joubini</emphasis>
|
||
</taxonomicName>
|
||
-group have been referred to
|
||
<taxonomicName id="88844D34537BFFB5FACC86F4FAD6FBBF" authorityName="Verrill" authorityYear="1899" box="[1272,1406,1015,1041]" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA5537BFFB5FACC86F4FAD6FBBF" box="[1272,1406,1015,1041]" italics="true" pageId="37" pageNumber="38">Amphioplus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="88844D34537BFFB5FEA38118FEBAFB9B" authorityName="Verrill" authorityYear="1899" box="[151,274,1051,1077]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiodia" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA5537BFFB5FEA38118FEBAFB9B" box="[151,274,1051,1077]" italics="true" pageId="37" pageNumber="38">Amphiodia</emphasis>
|
||
</taxonomicName>
|
||
on the basis that some specimens can have two distal oral papillae. However, the presence of two distal oral papillae is homoplasic, and in the case of joubini-group can be interpreted as paired adoral shield spines rather than one being a superficial oral tentacle spine (sensu
|
||
<bibRefCitation id="2B154B46537BFFB5FCC18160FCDEFBD3" author="Clark, A. M." box="[757,886,1123,1149]" pageId="37" pageNumber="38" pagination="1 - 81" refId="ref29819" refString="Clark, A. M. (1970) Notes on the family Amphiuridae (Ophiuroidea). Bulletin of the British Museum (Natural History), Zoology, 19, 1 - 81. https: // doi. org / 10.5962 / bhl. part. 24085" type="journal article" year="1970">Clark 1970</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4F3B36B7537BFFB5FEF38184FE38FAD7" blockId="37.[151,1437,151,1473]" pageId="37" pageNumber="38">
|
||
The new species differs from the other species in having only three arm spines, whereas all the other species have four to eight. Other differences include the bihamulate (two lateral terminal thorns) arm spines on
|
||
<taxonomicName id="88844D34537BFFB5FAFC81A8FAC6FB6B" authorityName="Baker" authorityYear="1977" box="[1224,1390,1195,1221]" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="ctenacantha">
|
||
<emphasis id="7DF0EAA5537BFFB5FAFC81A8FAC6FB6B" box="[1224,1390,1195,1221]" italics="true" pageId="37" pageNumber="38">A. ctenacantha</emphasis>
|
||
</taxonomicName>
|
||
, the notably denticulate disc scales on
|
||
<taxonomicName id="88844D34537BFFB5FC2581CCFD2AFB47" authorityName="Koehler" authorityYear="1912" box="[529,642,1231,1257]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="joubini">
|
||
<emphasis id="7DF0EAA5537BFFB5FC2581CCFD2AFB47" box="[529,642,1231,1257]" italics="true" pageId="37" pageNumber="38">O. joubini</emphasis>
|
||
</taxonomicName>
|
||
, and the naked ventral side of the disc on
|
||
<taxonomicName id="88844D34537BFFB5FA6781CCFB4FFB47" authorityName="Lyman" authorityYear="1879" box="[1107,1255,1231,1257]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="lanceolata">
|
||
<emphasis id="7DF0EAA5537BFFB5FA6781CCFB4FFB47" box="[1107,1255,1231,1257]" italics="true" pageId="37" pageNumber="38">A. lanceolata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D34537BFFB5FAC181CCFA34FB47" authorityName="Baker" authorityYear="1977" box="[1269,1436,1231,1257]" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="ctenacantha">
|
||
<emphasis id="7DF0EAA5537BFFB5FAC181CCFA34FB47" box="[1269,1436,1231,1257]" italics="true" pageId="37" pageNumber="38">A. ctenacantha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="88844D34537BFFB5FEF281F0FE0CFAA3" authorityName="Hertz" authorityYear="1927" box="[198,420,1267,1293]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="subSpecies" species="joubini" subSpecies="abyssalis">
|
||
<emphasis id="7DF0EAA5537BFFB5FEF281F0FE0CFAA3" box="[198,420,1267,1293]" italics="true" pageId="37" pageNumber="38">O. joubini abyssalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="88844D34537BFFB5FF8481F0FD89FAA3" authorityName="Koehler" authorityYear="1912" box="[432,545,1267,1293]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="joubini">
|
||
<emphasis id="7DF0EAA5537BFFB5FF8481F0FD89FAA3" box="[432,545,1267,1293]" italics="true" pageId="37" pageNumber="38">O. joubini</emphasis>
|
||
</taxonomicName>
|
||
is partially scaled ventrally, with few scales next to the oral shields.
|
||
<taxonomicName id="88844D34537BFFB5FB2281F0FF48FA9E" authorityName="Mortensen" authorityYear="1936" class="Ophiuroidea" family="Amphiuridae" genus="Amphioplus" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="acutus">
|
||
<emphasis id="7DF0EAA5537BFFB5FB2281F0FF48FA9E" italics="true" pageId="37" pageNumber="38">Amphioplus acutus</emphasis>
|
||
</taxonomicName>
|
||
is a variant of
|
||
<taxonomicName id="88844D34537BFFB5FFB98014FE57FA9F" authorityName="Koehler" authorityYear="1912" box="[397,511,1303,1329]" class="Ophiuroidea" family="Amphiuridae" genus="Amphiura" kingdom="Animalia" order="Amphilepidida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="joubini">
|
||
<emphasis id="7DF0EAA5537BFFB5FFB98014FE57FA9F" box="[397,511,1303,1329]" italics="true" pageId="37" pageNumber="38">O. joubini</emphasis>
|
||
</taxonomicName>
|
||
that has a third tiny oral papilla near the end of the jaw slit. The species limits of these species need to be determined. However, the morphology of the MD
|
||
<specimenCount id="5982FD3E537BFFB5FDD58039FBD1FAFB" box="[993,1145,1338,1365]" pageId="37" pageNumber="38" type="generic">50 specimens</specimenCount>
|
||
do not conform to any of these existing species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="079E653C537BFFB5FEF38080FCC1FA6F" pageId="37" pageNumber="38" type="etymology">
|
||
<paragraph id="4F3B36B7537BFFB5FEF38080FCC1FA6F" blockId="37.[151,1437,151,1473]" pageId="37" pageNumber="38">
|
||
<emphasis id="7DF0EAA5537BFFB5FEF38080FEE0FA33" bold="true" box="[199,328,1411,1437]" pageId="37" pageNumber="38">Etymology</emphasis>
|
||
. Named from “remota”, a female Latin adjective that means distant or far away, in reference to the presence of this species in the middle of the south Indian Ocean.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |