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<document id="3C3E253E18F7709AD3B3A178B7B1FF87" ID-CLB-Dataset="53164" ID-DOI="10.11646/zootaxa.5124.1.1" ID-GBIF-Dataset="9666d599-2c7d-4a52-bce2-a8794f03265b" ID-ISSN="1175-5326" ID-Zenodo-Dep="6404674" ID-ZooBank="C015F8CB-799B-4A92-90AE-02B4C576089E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648795871367" checkinUser="plazi" docAuthor="OHara, Timothy D. &amp; Thuy, Ben" docDate="2022" docId="C72D87A15354FF9AFEA3846DFC7DF955" docLanguage="en" docName="zootaxa.5124.1.1.pdf" docOrigin="Zootaxa 5124 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthophiura ingolfi Fasmer 1930" docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="3B14FFD9535EFF90FE348503FFA8FFAE" masterDocTitle="Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean" masterLastPageNumber="49" masterPageNumber="1" pageNumber="11" updateTime="1699310728982" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8F85EB1656665CC6945AB776E6962D9C">Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean</mods:title>
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<treatment id="C72D87A15354FF9AFEA3846DFC7DF955" ID-DOI="http://doi.org/10.5281/zenodo.6411766" ID-GBIF-Taxon="194442318" ID-Zenodo-Dep="6411766" LSID="urn:lsid:plazi:treatment:C72D87A15354FF9AFEA3846DFC7DF955" httpUri="http://treatment.plazi.org/id/C72D87A15354FF9AFEA3846DFC7DF955" lastPageNumber="11" pageId="10" pageNumber="11">
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<paragraph id="4F3B36B75354FF9AFEA3846DFD90FE26" blockId="10.[151,568,366,429]" box="[151,568,366,393]" pageId="10" pageNumber="11">
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<taxonomicName id="88844D345354FF9AFEA3846DFD90FE26" ID-CoL="5VFXD" authority="Fasmer, 1930" authorityName="Fasmer" authorityYear="1930" box="[151,568,366,393]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="ingolfi">
<emphasis id="7DF0EAA55354FF9AFEA3846DFD90FE26" bold="true" box="[151,568,366,393]" pageId="10" pageNumber="11">
<emphasis id="7DF0EAA55354FF9AFEA3846DFE23FE27" bold="true" box="[151,395,366,393]" italics="true" pageId="10" pageNumber="11">Anthophiura ingolfi</emphasis>
<bibRefCitation id="2B154B465354FF9AFFA6846CFD90FE26" author="Fasmer, H." box="[402,568,366,393]" pageId="10" pageNumber="11" pagination="1 - 7" refId="ref30977" refString="Fasmer, H. (1930) On the genus Anthophiura. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 90, 1 - 7." type="journal article" year="1930">Fasmer, 1930</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="079E653C5354FF9AFEA38490FEA5FE03" box="[151,269,403,429]" pageId="10" pageNumber="11" type="description">
<paragraph id="4F3B36B75354FF9AFEA38490FEA5FE03" blockId="10.[151,568,366,429]" box="[151,269,403,429]" pageId="10" pageNumber="11">
<figureCitation id="D7BF2A325354FF9AFEA38490FEA5FE03" box="[151,269,403,429]" captionStart="FIGURE 4" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[151,1435,561,1830]" captionTargetId="figure-183@11.[151,1436,561,1830]" captionTargetPageId="11" captionText="FIGURE 4. (AB) Anthophiura ingolfi, MNHN IE.2009.1609, A: dorsal and B: ventral views. (CD) Amphiophiura sp., juvenile, MNHN IE.2009.1615, C: dorsal and D: ventral views. (E) Ophiomyces sp., juvenile, MNHN IE.2009.1577, lateral view. (FH) Ophiophrura tripapillata, MNHN IE.2009.1573, F: dorsal view, G: ventral view and H: lateral arm view." figureDoi="http://doi.org/10.5281/zenodo.6404684" httpUri="https://zenodo.org/record/6404684/files/figure.png" pageId="10" pageNumber="11">Fig. 4AB</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="079E653C5354FF9AFEA384D4FC28FDA0" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="4F3B36B75354FF9AFEA384D4FC28FDA0" blockId="10.[151,1437,471,526]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="6F9411995354FF9AFEA384D4FC28FDA0" pageId="10" pageNumber="11">
<treatmentCitation id="CE2510A65354FF9AFEA384D4FD84FE41" author="Fasmer, H." box="[151,556,471,495]" page="7" pageId="10" pageNumber="11" year="1930">
<taxonomicName id="88844D345354FF9AFEA384D4FD84FE41" ID-CoL="5VFXD" authority="Fasmer, 1930: 4 - 7" authorityName="Fasmer" authorityPageNumber="4 - 7" authorityYear="1930" box="[151,556,471,495]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="ingolfi">
<emphasis id="7DF0EAA55354FF9AFEA384D4FECAFE41" box="[151,354,471,495]" italics="true" pageId="10" pageNumber="11">Anthophiura ingolfi</emphasis>
<bibRefCitation id="2B154B465354FF9AFF5E84DBFD84FE41" author="Fasmer, H." box="[362,556,471,495]" pageId="10" pageNumber="11" pagination="1 - 7" refId="ref30977" refString="Fasmer, H. (1930) On the genus Anthophiura. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 90, 1 - 7." type="journal article" year="1930">Fasmer, 1930: 47</bibRefCitation>
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, fig. 35.—
<treatmentCitation id="CE2510A65354FF9AFC9184D4FC13FE41" author="Mortensen, T." box="[677,955,471,495]" page="99" pageId="10" pageNumber="11" year="1933">
<bibRefCitation id="2B154B465354FF9AFC9184D4FC13FE41" author="Mortensen, T." box="[677,955,471,495]" pageId="10" pageNumber="11" pagination="401 - 472" refId="ref34407" refString="Mortensen, T. (1933 b) The echinoderms of St. Helena. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KObenhavn, 93, 401 - 472." type="journal article" year="1933">Mortensen 1933b: 99100</bibRefCitation>
</treatmentCitation>
, fig.52.—
<treatmentCitation id="CE2510A65354FF9AFA2B84D4FAC6FE41" author="Vadon, C. &amp; Guille, A." box="[1055,1390,471,495]" page="593" pageId="10" pageNumber="11" year="1984">
<bibRefCitation id="2B154B465354FF9AFA2B84D4FAC6FE41" author="Vadon, C. &amp; Guille, A." box="[1055,1390,471,495]" pageId="10" pageNumber="11" pagination="583 - 615" refId="ref36538" refString="Vadon, C. &amp; Guille, A. (1984) Les Ophiuridae (Ophiuroidea, Echinodermata) de la campagne MD 32 du Marion-Dufresne autour de l'ile de La Reunion. Bulletin du Museum National d'Histoire Naturelle. Paris, 6, 583 - 615." type="journal article" year="1984">Vadon &amp; Guille 1984: 593595</bibRefCitation>
</treatmentCitation>
, fig. 1(ab), pl. 4(46).—
<treatmentCitation id="CE2510A65354FF9AFFA184F5FDF7FDA0" author="Paterson, G. L. J." box="[405,607,502,526]" page="140" pageId="10" pageNumber="11" year="1985">
<bibRefCitation id="2B154B465354FF9AFFA184F5FDF7FDA0" author="Paterson, G. L. J." box="[405,607,502,526]" pageId="10" pageNumber="11" pagination="1 - 162" refId="ref35334" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History) Zoology, 49, 1 - 162." type="journal article" year="1985">Paterson 1985: 140</bibRefCitation>
</treatmentCitation>
, fig. 53.—
<treatmentCitation id="CE2510A65354FF9AFCFC84F5FCD4FDA0" author="Vadon, C." box="[712,892,502,526]" page="348" pageId="10" pageNumber="11" year="1991">
<bibRefCitation id="2B154B465354FF9AFCFC84F5FCD4FDA0" author="Vadon, C." box="[712,892,502,526]" pageId="10" pageNumber="11" pagination="335 - 356" refId="ref36500" refString="Vadon, C. (1991) Echinodermata: Ophiuridae profunds de Novelle-Caledonie: Formes paedomorphes. Resultats des Campagnes MUSORSTOM Vol. 8. Memoires du Museum National d'Histoire Naturelle, 151, 335 - 356." type="journal article" year="1991">Vadon 1991: 348</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="079E653C5354FF9AFEA3873FFAB0FDF8" box="[151,1304,572,599]" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="4F3B36B75354FF9AFEA3873FFAB0FDF8" blockId="10.[151,1437,572,1787]" box="[151,1304,572,599]" pageId="10" pageNumber="11">
<emphasis id="7DF0EAA55354FF9AFEA3873FFED1FDF8" bold="true" box="[151,377,572,598]" pageId="10" pageNumber="11">Material examined</emphasis>
.
<collectionCode id="2995AE725354FF9AFFB3873FFE1CFDF9" box="[391,436,572,599]" country="Germany" name="Museum Donaueschingen" pageId="10" pageNumber="11">MD</collectionCode>
50 DC55,
<collectionCode id="2995AE725354FF9AFC1C873EFDD6FDF9" box="[552,638,573,599]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
<collectionCode id="2995AE725354FF9AFCB1873FFD08FDF8" box="[645,672,572,598]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="10" pageNumber="11">IE</collectionCode>
.2009.1608 (2).
<collectionCode id="2995AE725354FF9AFD67873FFC28FDF9" box="[851,896,572,599]" country="Germany" name="Museum Donaueschingen" pageId="10" pageNumber="11">MD</collectionCode>
50 DC64,
<collectionCode id="2995AE725354FF9AFDC0873EFBE2FDF9" box="[1012,1098,573,599]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
<collectionCode id="2995AE725354FF9AFA65873FFBC4FDF8" box="[1105,1132,572,598]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="10" pageNumber="11">IE</collectionCode>
.2009.1609 (1).
</paragraph>
</subSubSection>
<subSubSection id="079E653C5354FF9AFEF38763FB7AFD6D" pageId="10" pageNumber="11" type="distribution">
<paragraph id="4F3B36B75354FF9AFEF38763FB7AFD6D" blockId="10.[151,1437,572,1787]" pageId="10" pageNumber="11">
<emphasis id="7DF0EAA55354FF9AFEF38763FEF1FDD4" bold="true" box="[199,345,608,634]" pageId="10" pageNumber="11">Distribution</emphasis>
. Arctic (2448
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), NW Atlantic (
<quantity id="887C9B525354FF9AFD288763FC17FDD5" box="[796,959,608,635]" metricMagnitude="3" metricUnit="m" metricValue="3.466" metricValueMax="3.796" metricValueMin="3.136" pageId="10" pageNumber="11" unit="m" value="3466.0" valueMax="3796.0" valueMin="3136.0">31363796 m</quantity>
), NE Atlantic (
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), E Atlantic (
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), W Indian (
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), E Indo-W Pacific (
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), E Pacific (
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), S Africa (
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), S
<collectingCountry id="379376275354FF9AFF5C87ABFE67FD6C" box="[360,463,680,706]" name="Australia" pageId="10" pageNumber="11">Australia</collectingCountry>
(
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),
<collectingCountry id="379376275354FF9AFCA487AAFC8EFD6C" box="[656,806,680,707]" name="New Zealand" pageId="10" pageNumber="11">New Zealand</collectingCountry>
(
<quantity id="887C9B525354FF9AFD0087ABFC7DFD6D" box="[820,981,680,707]" metricMagnitude="3" metricUnit="m" metricValue="1.219" metricValueMax="1.222" metricValueMin="1.216" pageId="10" pageNumber="11" unit="m" value="1219.0" valueMax="1222.0" valueMin="1216.0">12161222 m</quantity>
). SPA (
<quantity id="887C9B525354FF9AFA1287ABFB6EFD6D" box="[1062,1222,680,707]" metricMagnitude="3" metricUnit="m" metricValue="1.0625" metricValueMax="1.125" metricValueMin="1.0" pageId="10" pageNumber="11" unit="m" value="1062.5" valueMax="1125.0" valueMin="1000.0">10001125 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="079E653C5354FF9AFEF387CFFC7DF955" pageId="10" pageNumber="11" type="discussion">
<paragraph id="4F3B36B75354FF9AFEF387CFFC81FBE0" blockId="10.[151,1437,572,1787]" pageId="10" pageNumber="11">
<emphasis id="7DF0EAA55354FF9AFEF387CFFE9CFD48" bold="true" box="[199,308,716,742]" pageId="10" pageNumber="11">Remarks</emphasis>
. The MD50 material, to
<quantity id="887C9B525354FF9AFC7B87CFFD0EFD49" box="[591,678,716,743]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="11" unit="mm" value="3.4">3.4 mm</quantity>
dd, has been identified as
<taxonomicName id="88844D345354FF9AFDE187CFFB96FD48" authorityName="Fasmer" authorityYear="1930" box="[981,1086,716,742]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="ingolfi">
<emphasis id="7DF0EAA55354FF9AFDE187CFFB96FD48" box="[981,1086,716,742]" italics="true" pageId="10" pageNumber="11">A. ingolfi</emphasis>
</taxonomicName>
on the basis of the disc scales, which are few in number and thickened around the distal edges, broadly contiguous radial shields, rudimentary genital papillae, no arm comb, second oral tentacle pores slit like and positioned well back out of the jaw, pentagonal oral shields, ventral disc dominated by a large pentagonal scale adjacent to the oral shield and two separated tuberculated marginal scales, no DAPs, very reduced VAPs, two arm spines and tentacle pores present down the arm with one spine like tentacle scale (
<figureCitation id="D7BF2A325354FF9AFF988683FD8FFC34" box="[428,551,896,922]" captionStart="FIGURE 4" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[151,1435,561,1830]" captionTargetId="figure-183@11.[151,1436,561,1830]" captionTargetPageId="11" captionText="FIGURE 4. (AB) Anthophiura ingolfi, MNHN IE.2009.1609, A: dorsal and B: ventral views. (CD) Amphiophiura sp., juvenile, MNHN IE.2009.1615, C: dorsal and D: ventral views. (E) Ophiomyces sp., juvenile, MNHN IE.2009.1577, lateral view. (FH) Ophiophrura tripapillata, MNHN IE.2009.1573, F: dorsal view, G: ventral view and H: lateral arm view." figureDoi="http://doi.org/10.5281/zenodo.6404684" httpUri="https://zenodo.org/record/6404684/files/figure.png" pageId="10" pageNumber="11">Fig. 4AB</figureCitation>
). Previous authors have emphasised the elevated star-shaped centrodorsal plate, but this plate is not always shaped like this, and can sometimes be pentagonal (
<bibRefCitation id="2B154B465354FF9AFA2086A7FB04FC10" author="Fasmer, H." box="[1044,1196,932,958]" pageId="10" pageNumber="11" pagination="1 - 7" refId="ref30977" refString="Fasmer, H. (1930) On the genus Anthophiura. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 90, 1 - 7." type="journal article" year="1930">Fasmer 1930</bibRefCitation>
). Fasmer emphasised the split nature of the genital plates on the
<typeStatus id="903F88155354FF9AFC5A86CAFD36FC4D" box="[622,670,969,995]" pageId="10" pageNumber="11">type</typeStatus>
but this hasnt been reported by others (e.g.,
<bibRefCitation id="2B154B465354FF9AFAA786CAFA9DFC4D" author="Paterson, G. L. J." box="[1171,1333,968,995]" pageId="10" pageNumber="11" pagination="1 - 162" refId="ref35334" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History) Zoology, 49, 1 - 162." type="journal article" year="1985">Paterson 1985</bibRefCitation>
). He also described three arm spines (for animals
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dd) while
<bibRefCitation id="2B154B465354FF9AFD1586EEFC7CFBA9" author="Paterson, G. L. J." box="[801,980,1004,1031]" pageId="10" pageNumber="11" pagination="1 - 162" refId="ref35334" refString="Paterson, G. L. J. (1985) The deep-sea Ophiuroidea of the North Atlantic Ocean. Bulletin of the British Museum (Natural History) Zoology, 49, 1 - 162." type="journal article" year="1985">Paterson (1985)</bibRefCitation>
re-examining the
<typeStatus id="903F88155354FF9AFA9286EEFB49FBA9" box="[1190,1249,1005,1031]" pageId="10" pageNumber="11">types</typeStatus>
noted only two. The SPA material and that of
<bibRefCitation id="2B154B465354FF9AFFD58113FD5FFB84" author="Vadon, C. &amp; Guille, A." box="[481,759,1040,1066]" pageId="10" pageNumber="11" pagination="583 - 615" refId="ref36538" refString="Vadon, C. &amp; Guille, A. (1984) Les Ophiuridae (Ophiuroidea, Echinodermata) de la campagne MD 32 du Marion-Dufresne autour de l'ile de La Reunion. Bulletin du Museum National d'Histoire Naturelle. Paris, 6, 583 - 615." type="journal article" year="1984">Vadon and Guille (1984)</bibRefCitation>
from off
<collectingCountry id="379376275354FF9AFD568113FBA4FB84" box="[866,1036,1040,1066]" name="Reunion Island" pageId="10" pageNumber="11">Reunion Island</collectingCountry>
differ from those reported from the North Atlantic in having tuberculated marginal disc scales.
</paragraph>
<paragraph id="4F3B36B75354FF9AFEF3815BFD81F98C" blockId="10.[151,1437,572,1787]" pageId="10" pageNumber="11">
There are several other species in this little known genus. The
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species
<taxonomicName id="88844D345354FF9AFDCF815BFA91FBDC" authority="H. L. Clark, 1911" authorityName="H. L. Clark" authorityYear="1911" box="[1019,1337,1112,1139]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="axiologa">
<emphasis id="7DF0EAA55354FF9AFDCF815BFBD2FBDC" box="[1019,1146,1112,1138]" italics="true" pageId="10" pageNumber="11">A. axiologa</emphasis>
H.L.
<bibRefCitation id="2B154B465354FF9AFA8C815BFA91FBDC" author="Clark, H. L." box="[1208,1337,1112,1138]" pageId="10" pageNumber="11" pagination="1 - 302" refId="ref29926" refString="Clark, H. L. (1911) North Pacific ophiurans in the collection of the United States National Museum. Bulletin of the United States National Museum, 75, 1 - 302, figs. 1 - 144. https: // doi. org / 10.5962 / bhl. title. 32556" type="journal article" year="1911">Clark, 1911</bibRefCitation>
</taxonomicName>
from the North Pacific (
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,
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dd) is distinguished by having disc scales that are neither tuberculated nor thickened, a wide marginal disc scale ventrally and arm spines that become hooked only from the 10th segment.
<taxonomicName id="88844D345354FF9AFB3B81A3FE49FB70" authority="(H. L. Clark, 1939)" baseAuthorityName="H. L. Clark" baseAuthorityYear="1939" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="granulata">
<emphasis id="7DF0EAA55354FF9AFB3B81A3FEADFB70" italics="true" pageId="10" pageNumber="11">Anthophiura granulata</emphasis>
(H.L. Clark, 1939)
</taxonomicName>
from the North Indian Ocean (
<quantity id="887C9B525354FF9AFD7581C7FC33FB71" box="[833,923,1220,1247]" metricMagnitude="3" metricUnit="m" metricValue="3.84" pageId="10" pageNumber="11" unit="m" value="3840.0">3840 m</quantity>
,
<quantity id="887C9B525354FF9AFD9381C7FC41FB71" box="[935,1001,1220,1247]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="11" unit="mm" value="4.0">4 mm</quantity>
dd) has disc scales that are sculptured and pitted, a wide marginal disc scale, and prominent genital papillae.
<taxonomicName id="88844D345354FF9AFD9881EBFAF7FAAC" authority="Fasmer, 1930" authorityName="Fasmer" authorityYear="1930" box="[940,1375,1256,1283]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="challengeri">
<emphasis id="7DF0EAA55354FF9AFD9881EBFB69FAAC" box="[940,1217,1256,1282]" italics="true" pageId="10" pageNumber="11">Anthophiura challengeri</emphasis>
<bibRefCitation id="2B154B465354FF9AFAFC81EAFAF7FAAC" author="Fasmer, H." box="[1224,1375,1256,1283]" pageId="10" pageNumber="11" pagination="1 - 7" refId="ref30977" refString="Fasmer, H. (1930) On the genus Anthophiura. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 90, 1 - 7." type="journal article" year="1930">Fasmer, 1930</bibRefCitation>
</taxonomicName>
from the central South Pacific (
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,
<quantity id="887C9B525354FF9AFC13800FFDC1FA89" box="[551,617,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="mm" value="6.0">6 mm</quantity>
dd) has small disc scales that surround the primary plates and the large ventral scale, and separate the radial shields proximally. It also has a few prominent genital papillae.
<taxonomicName id="88844D345354FF9AFB3B8033FE06FAC0" authority="Tommasi, 1976" authorityName="Tommasi" authorityYear="1976" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="dilatata">
<emphasis id="7DF0EAA55354FF9AFB3B8033FF46FAC0" italics="true" pageId="10" pageNumber="11">Anthophiura dilatata</emphasis>
<bibRefCitation id="2B154B465354FF9AFEC28057FE06FAC0" author="Tommasi, L. R." box="[246,430,1364,1391]" pageId="10" pageNumber="11" pagination="281 - 318" refId="ref36406" refString="Tommasi, L. R. (1976) Ophiuroidea collected in the Peru-Chile Trench by the USNS &quot; Eltanin &quot; during cruise III. Papeis Avulsos de Zoologia, Sao Paulo, 29, 281 - 318." type="journal article" year="1976">Tommasi, 1976</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="379376275354FF9AFC2B8056FDFAFAC1" box="[543,594,1365,1391]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
Trench (
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,
<quantity id="887C9B525354FF9AFD528057FC15FAC1" box="[870,957,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="mm" value="2.5">2.5 mm</quantity>
dd) has only one ventral disc scale that is flanked by the genital plates (
<bibRefCitation id="2B154B465354FF9AFFD1807BFD76FA3C" author="Vadon, C. &amp; Guille, A." box="[485,734,1400,1426]" pageId="10" pageNumber="11" pagination="583 - 615" refId="ref36538" refString="Vadon, C. &amp; Guille, A. (1984) Les Ophiuridae (Ophiuroidea, Echinodermata) de la campagne MD 32 du Marion-Dufresne autour de l'ile de La Reunion. Bulletin du Museum National d'Histoire Naturelle. Paris, 6, 583 - 615." type="journal article" year="1984">Vadon &amp; Guille 1984</bibRefCitation>
).
<bibRefCitation id="2B154B465354FF9AFCC5807BFC23FA3C" author="Vadon, C." box="[753,907,1400,1426]" pageId="10" pageNumber="11" pagination="335 - 356" refId="ref36500" refString="Vadon, C. (1991) Echinodermata: Ophiuridae profunds de Novelle-Caledonie: Formes paedomorphes. Resultats des Campagnes MUSORSTOM Vol. 8. Memoires du Museum National d'Histoire Naturelle, 151, 335 - 356." type="journal article" year="1991">Vadon (1991)</bibRefCitation>
transferred the species
<taxonomicName id="88844D345354FF9AFAA3807BFEE6FA18" authority="Litvinova, 1981" authorityName="Litvinova" authorityYear="1981" class="Ophiuroidea" family="Astrophiuridae" genus="Ophiophycis" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="nixastrum">
<emphasis id="7DF0EAA55354FF9AFAA3807BFA34FA3C" box="[1175,1436,1400,1426]" italics="true" pageId="10" pageNumber="11">Ophiophycis nixastrum</emphasis>
<bibRefCitation id="2B154B465354FF9AFEA3809FFEE6FA18" author="Litvinova, N. M." box="[151,334,1436,1462]" pageId="10" pageNumber="11" pagination="113 - 131" refId="ref32739" refString="Litvinova, N. M. (1981) Brittle-stars (Ophiuroidea). In: Benthos of the submarine mountains Marcus-Necker and adjacent Pacific regions. Academy of Science of the USSR. P. P. Shirshov Institute of Oceanology, pp. 113 - 131." type="book chapter" year="1981">Litvinova, 1981</bibRefCitation>
</taxonomicName>
from the central North Pacific (
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,
<quantity id="887C9B525354FF9AFD49809FFBA0FA19" box="[893,1032,1436,1463]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="10" pageNumber="11" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.82.1 mm</quantity>
dd) to
<taxonomicName id="88844D345354FF9AFA57809FFB59FA18" authorityName="H. L. Clark" authorityYear="1911" box="[1123,1265,1436,1462]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA55354FF9AFA57809FFB59FA18" box="[1123,1265,1436,1462]" italics="true" pageId="10" pageNumber="11">Anthophiura</emphasis>
</taxonomicName>
. However, the attenuated arm, large enlarged basal LAPs, prominent arm spines, and disc plating are more similar to
<taxonomicName id="88844D345354FF9AFB3B80C3FA35FA74" authorityName="Koehler" authorityYear="1901" box="[1295,1437,1472,1498]" class="Ophiuroidea" family="Astrophiuridae" genus="Ophiophycis" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA55354FF9AFB3B80C3FA35FA74" box="[1295,1437,1472,1498]" italics="true" pageId="10" pageNumber="11">Ophiophycis</emphasis>
</taxonomicName>
and in many ways the figures of the
<typeStatus id="903F88155354FF9AFC7C80E6FDD0FA51" box="[584,632,1509,1535]" pageId="10" pageNumber="11">type</typeStatus>
are reminiscent of
<taxonomicName id="88844D345354FF9AFD6980E7FB50FA50" authority="McKnight, 2003" authorityName="McKnight" authorityYear="2003" box="[861,1272,1508,1534]" class="Ophiuroidea" family="Astrophiuridae" genus="Ophiophycis" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="johni">
<emphasis id="7DF0EAA55354FF9AFD6980E7FB87FA50" box="[861,1071,1508,1534]" italics="true" pageId="10" pageNumber="11">Ophiophycis johni</emphasis>
<bibRefCitation id="2B154B465354FF9AFA0D80E7FB50FA50" author="McKnight, D. G." box="[1081,1272,1508,1534]" pageId="10" pageNumber="11" pagination="1 - 36" refId="ref34183" refString="McKnight, D. G. (2003) New brittle-stars (Echinodermata: Ophiuroidea) from New Zealand waters. Zootaxa, 352, 1 - 36. https: // doi. org / 10.11646 / zootaxa. 352.1.1" type="journal article" year="2003">McKnight, 2003</bibRefCitation>
</taxonomicName>
, including the prominent star-shaped centrodorsal.
</paragraph>
<paragraph id="4F3B36B75354FF9AFEF3832EFC7DF955" blockId="10.[151,1437,572,1787]" pageId="10" pageNumber="11">
No genetic sequences are known from
<taxonomicName id="88844D345354FF9AFCBC832FFCBEF9E8" authorityName="H. L. Clark" authorityYear="1911" box="[648,790,1580,1606]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Anthophiura" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="7DF0EAA55354FF9AFCBC832FFCBEF9E8" box="[648,790,1580,1606]" italics="true" pageId="10" pageNumber="11">Anthophiura</emphasis>
</taxonomicName>
and its family level placement is uncertain. However, it is likely to be a member of the
<taxonomicName id="88844D345354FF9AFC3F8353FD1DF9C4" authorityName="Perrier" authorityYear="1893" box="[523,693,1616,1642]" class="Ophiuroidea" family="Ophiopyrgidae" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="family">Ophiopyrgidae</taxonomicName>
, given the position, shape and scales of the second oral tentacle pore, and the presence of pores all along the arm. However, the tube feet of the arm emerging though a hole in the lateral arm plate, the reduced DAPs and VAPs and the arm spines are also characteristic of the
<taxonomicName id="88844D345354FF9AFA80839BFAC0F91C" box="[1204,1384,1688,1714]" class="Ophiuroidea" family="Ophiomusaidae" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="family">Ophiomusaidae</taxonomicName>
. The presence in the SPA material of two tuberculated scales along the interradial disc margin is similar to
<taxonomicName id="88844D345354FF9AFB2F83BFFF40F954" authorityName="Lyman" authorityYear="1878" class="Ophiuroidea" family="Ophiomusaidae" genus="Ophiomusa" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="scalare">
<emphasis id="7DF0EAA55354FF9AFB2F83BFFF40F954" italics="true" pageId="10" pageNumber="11">Ophiomusa scalare</emphasis>
</taxonomicName>
from which it differs by having open tentacle pores along the arm.
</paragraph>
</subSubSection>
</treatment>
</document>