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<document ID-DOI="http://doi.org/10.5281/zenodo.6870843" ID-ISBN="978-84-16728-08-4" ID-Zenodo-Dep="6870843" approvalRequired="304" approvalRequired_for_illustrations="45" approvalRequired_for_matCits="75" approvalRequired_for_taxonomicNames="40" approvalRequired_for_textStreams="75" approvalRequired_for_treatments="69" checkinTime="1658335596803" checkinUser="felipe" docAuthor="Russell A. Mittermeier &amp; Don E. Wilson" docDate="2018" docId="3D474A54A0788715FAFAAC511337FA12" docLanguage="en" docName="hbmw_8_Soricidae_0332.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos, Barcelona: Lynx Edicions" docTitle="Crocidura nigripes G. S. Miller &amp; Hollister 1921" docType="treatment" docVersion="3" lastPageNumber="476" masterDocId="C17E322CA0288744FF8DAB47125EFFF9" masterDocTitle="Soricidae" masterLastPageNumber="551" masterPageNumber="332" pageNumber="475" updateTime="1658350298165" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Soricidae</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Don E. Wilson</mods:namePart>
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<mods:dateIssued>2018</mods:dateIssued>
<mods:dateOther type="pubDate">2018-07-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
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<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos</mods:title>
</mods:titleInfo>
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<mods:start>332</mods:start>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6870843</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-08-4</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6870082" ID-Zenodo-Dep="6870082" LSID="urn:lsid:plazi:treatment:3D474A54A0788715FAFAAC511337FA12" httpUri="http://treatment.plazi.org/id/3D474A54A0788715FAFAAC511337FA12" lastPageId="81" lastPageNumber="476" pageId="80" pageNumber="475">
<heading pageId="80" pageNumber="475">
<subSubSection box="[1399,1481,1814,1864]" pageId="80" pageNumber="475" type="multiple">
<paragraph blockId="80.[1396,2600,1814,1943]" box="[1399,1481,1814,1864]" pageId="80" pageNumber="475">
<figureCitation box="[1399,1481,1814,1864]" captionStart="Plate 20: Soricidae" captionStartId="80.[125,155,3104,3125]" captionTargetBox="[12,2740,14,3636]" captionTargetPageId="79" captionText="226. Malayan White-toothed Shrew (Crocidura malayana), 227. Peninsular White-toothed Shrew (Crocidura negligens), 228. Black-footed White-toothed Shrew (Crocidura nigripes), 229. Bornean White-toothed Shrew (Crocidura foetida), 230. Kinabalu White-toothed Shrew (Crocidura baluensis), 231. Thick-tailed White-toothed Shrew (Crocidura brunnea), 232. Oriental White-toothed Shrew (Crocidura orientalis), 233. Bangka White-toothed Shrew (Crocidura vosmaeri), 234. Hutan White-toothed Shrew (Crocidura hutanis), 235. Beccaris White-toothed Shrew (Crocidura beccarii), 236. Sumatran Giant White-toothed Shrew (Crocidura lepidura), 237. Mount Malindang White-toothed Shrew (Crocidura grandis), 238. Mindanao White-toothed Shrew (Crocidura beata), 239. Mindoro White-toothed Shrew (Crocidura mindorus), 240. Panay White-toothed Shrew (Crocidura panayensis), 241. Negros White-toothed Shrew (Crocidura negrina), 242. Luzon White-toothed Shrew (Crocidura grayi), 243. Sibuyan White-toothed Shrew (Crocidura ninoyi), 244. Palawan White-toothed Shrew (Crocidura palawanensis), 245. Javan Long-tailed White-toothed Shrew (Crocidura abscondita), 246. Gathornes White-toothed Shrew (Crocidura gathornei), 247. Horsfields White-toothed Shrew (Crocidura horsfieldii), 248. Sinharaja White-toothed Shrew (Crocidura hikmiya), 249. Sri Lankan White-toothed Shrew (Crocidura miya), 250. Lesser Ryukyu White-toothed Shrew (Crocidura wataser), 251. Southeast Asian White-toothed Shrew (Crocidura fuliginosa), 252. Large White-toothed Shrew (Crocidura dracula), 253. Sumatran Long-tailed White-toothed Shrew (Crocidura paradoxura), 254. Dsinezumi White-toothed Shrew (Crocidura dsinezuma), 255. Chinese White-toothed Shrew (Crocidura rapax), 256. Ussuri White-toothed Shrew (Crocidura lasiura), 257. Asian Gray White-toothed Shrew (Crocidura attenuata), 258. Zaitsevs White-toothed Shrew (Crocidura zaitsevi), 259. Indochinese White-toothed Shrew (Crocidura indochinensis), 260. Hainan White-toothed Shrew (Crocidura wuchihensis), 261. Timor White-toothed Shrew (Crocidura tenuis), 262. Javan White-toothed Shrew (Crocidura maxi), 263. Javan Ghost White-toothed Shrew (Crocidura umbra), 264. Sunda White-toothed Shrew (Crocidura monticola), 265. Neglected White-toothed Shrew (Crocidura neglecta)" figureDoi="http://doi.org/10.5281/zenodo.6871919" httpUri="https://zenodo.org/record/6871919/files/figure.png" pageId="80" pageNumber="475">228.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[1497,2208,1814,1864]" pageId="80" pageNumber="475" type="vernacular_names">
<paragraph blockId="80.[1396,2600,1814,1943]" box="[1497,2208,1814,1864]" pageId="80" pageNumber="475">
<vernacularName box="[1497,2208,1814,1864]" pageId="80" pageNumber="475">Black-footed White-toothed Shrew</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[2279,2599,1814,1864]" pageId="80" pageNumber="475" type="nomenclature">
<paragraph blockId="80.[1396,2600,1814,1943]" box="[2279,2599,1814,1864]" pageId="80" pageNumber="475">
<taxonomicName authorityName="G. S. Miller &amp; Hollister" authorityYear="1921" box="[2279,2599,1814,1864]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="nigripes">
<emphasis box="[2279,2599,1814,1864]" italics="true" pageId="80" pageNumber="475">Crocidura nigripes</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="475" type="vernacular_names">
<paragraph blockId="80.[1396,2600,1814,1943]" box="[1398,2555,1882,1903]" pageId="80" pageNumber="475">
<emphasis bold="true" box="[1398,1474,1882,1903]" pageId="80" pageNumber="475">French:</emphasis>
<vernacularName box="[1483,1733,1882,1903]" pageId="80" pageNumber="475">Crocidure a pattes noires</vernacularName>
/
<emphasis bold="true" box="[1754,1844,1882,1903]" pageId="80" pageNumber="475">German:</emphasis>
<vernacularName box="[1854,2179,1882,1903]" pageId="80" pageNumber="475">SchwarzfulR-Weil3zahnspitzmaus</vernacularName>
/
<emphasis bold="true" box="[2201,2292,1882,1903]" pageId="80" pageNumber="475">Spanish:</emphasis>
<vernacularName box="[2302,2555,1882,1903]" pageId="80" pageNumber="475">Musarana de pies negros</vernacularName>
</paragraph>
<paragraph blockId="80.[1396,2600,1814,1943]" box="[1398,1850,1922,1943]" pageId="80" pageNumber="475">
<emphasis bold="true" box="[1398,1645,1922,1943]" pageId="80" pageNumber="475">Other common names:</emphasis>
<vernacularName box="[1655,1850,1922,1943]" pageId="80" pageNumber="475">Black-footed Shrew</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection pageId="80" pageNumber="475" type="reference_group">
<paragraph blockId="80.[2010,2603,1987,2414]" pageId="80" pageNumber="475">
<emphasis bold="true" box="[2010,2165,1987,2020]" pageId="80" pageNumber="475">Taxonomy.</emphasis>
<taxonomicName authority="G. S. Miller &amp; Hollister, 1921" authorityName="G. S. Miller &amp; Hollister" authorityYear="1921" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="nigripes">Crocidura nigripes G. S. Miller &amp; Hollister, 1921</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="475" type="materials_examined">
<paragraph blockId="80.[2010,2603,1987,2414]" pageId="80" pageNumber="475">
<materialsCitation country="Indonesia" location="south-west from Tondano Lake" pageId="80" pageNumber="475" specimenCount="1" stateProvince="north-eastern Sulawesi">
<location LSID="urn:lsid:plazi:treatment:3D474A54A0788715FAFAAC511337FA12:B031AD99A0788714F730ACB41A7FF7CD" country="Indonesia" name="south-west from Tondano Lake" pageId="80" pageNumber="475" stateProvince="north-eastern Sulawesi">south-west from Tondano Lake</location>
,
<location LSID="urn:lsid:plazi:treatment:3D474A54A0788715FAFAAC511337FA12:B031AD99A0788714F7B7A35C1A9FF7CD" box="[2106,2241,2075,2100]" country="Indonesia" name="Temboan" pageId="80" pageNumber="475" stateProvince="north-eastern Sulawesi">Temboan</location>
,
<collectingRegion box="[2268,2595,2075,2100]" pageId="80" pageNumber="475">north-eastern Sulawesi</collectingRegion>
,
<collectingCountry box="[2011,2151,2106,2139]" name="Indonesia" pageId="80" pageNumber="475">Indonesia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="475" type="discussion">
<paragraph blockId="80.[2010,2603,1987,2414]" pageId="80" pageNumber="475">Unlike the other shrews endemic to Sulawesi, C. mnigripes is phylogenetically unrelated to the “Old Sulawesian” clade and is more closely related to species from the Sunda Shelf, suggesting a relatively recent (perhaps late Pliocene) colonization across the Makassar Strait. Its karyotype,</paragraph>
<paragraph blockId="80.[1399,2603,2421,3005]" pageId="80" pageNumber="475">
consisting of 38 oy supports such a recent biogeographic origin, as this formula is shared with most other species from the Sunda Shelf, while other Sulawesian
<taxonomicName box="[1467,1591,2504,2533]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
have a decreased chromosome count (2n = 30-34). Races differ in size and in a single fixed diagnostic locus over the 32 assayed allozyme loci; such a difference might reflect the large geographic distance separating the samples rather than having true taxonomic implications; there is a possible zone of overlap in east-central Sulawesi, supporting animals of intermediate size. Two subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="475" type="synonymic_list">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6871392" ID-Zenodo-Dep="6871392" httpUri="https://zenodo.org/record/6871392/files/figure.png" inLine="true" pageId="80" pageNumber="475" targetBox="[1395,1986,1998,2412]" targetPageId="80">
<paragraph blockId="80.[1399,2603,2421,3005]" box="[1399,1799,2697,2730]" pageId="80" pageNumber="475">
<emphasis bold="true" box="[1399,1799,2697,2730]" pageId="80" pageNumber="475">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="80.[1399,2603,2421,3005]" box="[1402,2586,2736,2769]" pageId="80" pageNumber="475">
<taxonomicName authority="G. S. Miller &amp; Hollister, 1921" authorityName="Miller &amp; Hollister" authorityYear="1921" box="[1402,1979,2736,2769]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="subSpecies" species="nigripes" subSpecies="nigripes">C.n.nigripesG.S.Miller&amp;Hollister,1921—NEpeninsulaofSulawesiandLembehI.</taxonomicName>
</paragraph>
<paragraph blockId="80.[1399,2603,2421,3005]" pageId="80" pageNumber="475">
<taxonomicName authority="G. S. Miller &amp; Hollister, 1921" authorityName="G. S. Miller &amp; Hollister" authorityYear="1921" box="[1402,1956,2784,2809]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="subSpecies" species="nigripes" subSpecies="ipara">C. n. ipara G. S. Miller &amp; Hollister, 1921</taxonomicName>
— C Sulawesi. Its absence from the S peninsulas should be ascertained by adequate surveys.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="81" lastPageNumber="476" pageId="80" pageNumber="475" type="description">
<paragraph blockId="80.[1399,2603,2421,3005]" pageId="80" pageNumber="475">
<emphasis bold="true" box="[1399,1650,2854,2887]" pageId="80" pageNumber="475">Descriptive notes.</emphasis>
Head—body 72-86 mm, tail 50-65 mm, ear 6 mm, hindfoot 13-1-15-5 mm; weight 8-12 g. The Black-footed White-toothed Shrew is a medium-sized shrew with dark brown to black pelage, but molting individuals show a browner tinge to the fur, particularly on the throat and upper venter, which can be chestnut in color.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6871919" ID-Zenodo-Dep="6871919" httpUri="https://zenodo.org/record/6871919/files/figure.png" pageId="80" pageNumber="475" startId="80.[125,155,3104,3125]" targetBox="[12,2740,14,3636]" targetPageId="79">
<paragraph blockId="80.[123,2580,3103,3483]" pageId="80" pageNumber="475">
On following pages: 229. Bornean White-toothed Shrew (
<taxonomicName authorityName="Peters" authorityYear="1870" box="[755,943,3104,3125]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="foetida">Crocidura foetida</taxonomicName>
); 230. Kinabalu White-toothed Shrew Shrew (
<taxonomicName box="[210,422,3139,3164]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="orientalis">Crocidura orientalis</taxonomicName>
); 233. Bangka White-toothed Shrew (
<taxonomicName authorityName="Jentink" authorityYear="1888" box="[835,1047,3139,3164]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="477" phylum="Chordata" rank="species" species="vosmaeri">Crocidura vosmaeri</taxonomicName>
); 234. Hutan White-toothed White-toothed Shrew (
<taxonomicName authorityName="Lyon" authorityYear="1908" box="[376,575,3178,3203]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="lepidura">Crocidura lepidura</taxonomicName>
); 237. Mount Malindang White-toothed Shrew (
<taxonomicName box="[1097,1288,3178,3203]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="grandis">Crocidura grandis</taxonomicName>
); 238. 240. Panay White-toothed Shrew (
<taxonomicName authorityName="Hutterer" authorityYear="2007" box="[502,734,3217,3242]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="panayensis">Crocidura panayensis</taxonomicName>
); 241. Negros White-toothed Shrew (
<taxonomicName authorityName="Rabor" authorityYear="1952" box="[1144,1337,3217,3242]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="negrina">Crocidura negrina</taxonomicName>
); 242 Palawan White-toothed Shrew (
<taxonomicName authorityName="E. H. Taylor" authorityYear="1934" box="[472,732,3257,3282]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="palawanensis">Crocidura palawanensis</taxonomicName>
); 245. Javan Long-tailed White-toothed Shrew (
<taxonomicName box="[1255,1358,3257,3282]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
(
<taxonomicName baseAuthorityName="Tomes" baseAuthorityYear="1856" box="[132,350,3296,3321]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="horsfieldii">Crocidura horsfieldii</taxonomicName>
); 248. Sinharaja White-toothed Shrew (
<taxonomicName box="[781,884,3296,3321]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
hikmiya); 249. Sri Lankan White-toothed Shrew White-toothed Shrew (
<taxonomicName baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[377,593,3335,3360]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="fuliginosa">Crocidura fuliginosa</taxonomicName>
); 252. Large White-toothed Shrew (
<taxonomicName authorityName="Thomas" authorityYear="1912" box="[988,1178,3335,3360]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="dracula">Crocidura dracula</taxonomicName>
); 253. Sumatran
<taxonomicName authorityName="Temminck" authorityYear="1842" box="[126,237,3375,3400]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="dsinezumi">dsinezumi</taxonomicName>
); 255. Chinese White-toothed Shrew (
<taxonomicName box="[659,830,3375,3400]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="rapax">Crocidura rapax</taxonomicName>
); 256. Ussuri White-toothed Shrew (
<taxonomicName authorityName="Dobson" authorityYear="1890" box="[1233,1415,3375,3400]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="lasiura">Crocidura lasiura</taxonomicName>
(
<taxonomicName box="[132,236,3414,3439]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
zaitsevi); 259. Indochinese White-toothed Shrew (
<taxonomicName box="[790,1051,3414,3439]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="indochinensis">Crocidura indochinensis</taxonomicName>
); 260. Hainan White-toothed Shrew (
<taxonomicName authorityName="Sody" authorityYear="1936" box="[211,375,3453,3478]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="maxi">Crocidura maxi</taxonomicName>
); 263. Javan Ghost White-toothed Shrew (
<taxonomicName box="[843,946,3453,3478]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
umbra); 264. Sunda White-toothed Shrew (
<taxonomicName authorityName="Thomas" authorityYear="1898" box="[1368,1582,3104,3125]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="476" phylum="Chordata" rank="species" species="baluensis">Crocidura baluensis</taxonomicName>
); 231. Thick-tailed White-toothed Shrew (
<taxonomicName authorityName="Jentink" authorityYear="1888" box="[2032,2233,3104,3125]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="brunnea">Crocidura brunnea</taxonomicName>
); 232. Oriental White-toothed Shrew (
<taxonomicName box="[1447,1638,3139,3164]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="478" phylum="Chordata" rank="species" species="hutanis">Crocidura hutanis</taxonomicName>
); 235. Beccari's White-toothed Shrew (
<taxonomicName authorityName="Dobson" authorityYear="1887" box="[2070,2266,3139,3164]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="beccarii">Crocidura beccarii</taxonomicName>
); 236. Sumatran Giant Mindanao White-toothed Shrew (
<taxonomicName box="[1731,1834,3178,3203]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
beata); 239. Mindoro White-toothed Shrew (
<taxonomicName box="[2329,2543,3178,3203]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="mindorus">Crocidura mindorus</taxonomicName>
);. Luzon White-toothed Shrew (
<taxonomicName authorityName="Dobson" authorityYear="1890" box="[1736,1900,3217,3242]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="grayi">Crocidura grayi</taxonomicName>
); 243. Sibuyan White-toothed Shrew (
<taxonomicName box="[2318,2421,3217,3242]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
ninoyi); 244. abscondita); 246. Gathorne's White-toothed Shrew (
<taxonomicName box="[1937,2040,3257,3282]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
gathornei); 247. Horsfield's White-toothed Shrew (
<taxonomicName authorityName="Phillips" authorityYear="1929" box="[1424,1588,3296,3321]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="483" phylum="Chordata" rank="species" species="miya">Crocidura miya</taxonomicName>
); 250. Lesser Ryukyu White-toothed Shrew (
<taxonomicName box="[2083,2187,3296,3321]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
watasel); 251. Southeast Asian Long-tailed White-toothed Shrew (
<taxonomicName authorityName="Dobson" authorityYear="1886" box="[1746,1978,3335,3360]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="paradoxura">Crocidura paradoxura</taxonomicName>
); 254. Dsinezumi White-toothed Shrew (
<taxonomicName box="[2427,2531,3335,3360]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
); 257. Asian Gray White-toothed Shrew (
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[1860,2073,3375,3400]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="attenuata">Crocidura attenuata</taxonomicName>
); 258. Zaitsevs White-toothed Shrew
</paragraph>
<paragraph blockId="80.[123,2580,3103,3483]" pageId="80" pageNumber="475">
Shrew (
<taxonomicName authorityName="Wang" authorityYear="1966" box="[1460,1708,3414,3439]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="wuchihensis">Crocidura wuchihensis</taxonomicName>
); 261. Timor White-toothed Shrew (
<taxonomicName baseAuthorityName="Muller" baseAuthorityYear="1840" box="[2098,2277,3414,3439]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="486" phylum="Chordata" rank="species" species="tenuis">Crocidura tenuis</taxonomicName>
); 262. Javan White-toothed (
<taxonomicName authorityName="Peters" authorityYear="1870" box="[1426,1645,3453,3478]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="487" phylum="Chordata" rank="species" species="monticola">Crocidura monticola</taxonomicName>
); 265. Neglected White-toothed Shrew (
<taxonomicName authorityName="Jentink" authorityYear="1888" box="[2091,2297,3453,3478]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="80" pageNumber="475" phylum="Chordata" rank="species" species="neglecta">Crocidura neglecta</taxonomicName>
).
</paragraph>
</caption>
<paragraph blockId="81.[155,1365,305,1516]" pageId="81" pageNumber="476">
Face, ears, feet, and tail also blackish. Tail cylindrical, relatively short (60-80% headbody length) and essentially naked; a few long bristles present only along its basal third. Skull slender with narrow rostral parts and typically parallel tooth rows. Subspecies lipara is distinctly larger, particularly in skull dimensions (mean greatest length of skull 22:9 mm vs. 21-7 mm). Chromosomal complement has 2n = 38 and FN = 56, with submetacentric X and subtelocentric Y chromosomes. Chromosomal complement very similar to those of most other Sundaland
<taxonomicName box="[824,948,535,568]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="81" pageNumber="476" phylum="Chordata" rank="genus">Crocidura</taxonomicName>
but differs significantly from those of all other tested species from Sulawesi, stressing a different biogeographic origin of the Black-footed White-toothed Shrew.
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="476" type="biology_ecology">
<paragraph blockId="81.[155,1365,305,1516]" pageId="81" pageNumber="476">
<emphasis bold="true" box="[157,268,654,687]" pageId="81" pageNumber="476">Habitat.</emphasis>
Lowland evergreen to montane rainforests. Known altitudinal range 30-2200 m. The nominate race apparently occurs exclusively in lowland rainforests; race
<taxonomicName box="[159,234,732,765]" pageId="81" pageNumber="476" rank="subSpecies" subSpecies="lipara">lipara</taxonomicName>
occupies a much broader elevational range of habitats, up to 2200 m, being most abundant at moderate altitudes (800-900 m).
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="476" type="food_feeding">
<paragraph blockId="81.[155,1365,305,1516]" pageId="81" pageNumber="476">
<emphasis bold="true" box="[157,424,811,844]" pageId="81" pageNumber="476">Food and Feeding.</emphasis>
Black-footed White-toothed Shrews feed on invertebrates but no precise data are available.
</paragraph>
</subSubSection>
<subSubSection box="[157,1254,890,923]" pageId="81" pageNumber="476" type="breeding">
<paragraph blockId="81.[155,1365,305,1516]" box="[157,1254,890,923]" pageId="81" pageNumber="476">
<emphasis bold="true" box="[157,291,890,923]" pageId="81" pageNumber="476">Breeding.</emphasis>
In August, several pregnant females carried 1-2 near-term embryos.
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="476" type="activity">
<paragraph blockId="81.[155,1365,305,1516]" pageId="81" pageNumber="476">
<emphasis bold="true" box="[155,400,929,962]" pageId="81" pageNumber="476">Activity patterns.</emphasis>
Black-footed White-toothed Shrews are terrestrial, being found among leaflitter on the forest floor, near logs or dead trees. They are essentially nocturnal or crepuscular.
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="476" type="biology_ecology">
<paragraph blockId="81.[155,1365,305,1516]" pageId="81" pageNumber="476">
<emphasis bold="true" box="[157,856,1048,1081]" pageId="81" pageNumber="476">Movements, Home range and Social organization.</emphasis>
Black-footed White-toothed Shrews may be sympatric with all other shrew species present in Sulawesi, but no precise data on their natural history are available.
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="476" type="conservation">
<paragraph blockId="81.[155,1365,305,1516]" pageId="81" pageNumber="476">
<emphasis bold="true" box="[158,515,1166,1199]" pageId="81" pageNumber="476">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List in view of its widespread occurrence in Sulawesi and expected large population size, but its relative tolerance of anthropic changes to its habitat is unknown. The Black-footed White-toothed Shrew is locally common as demonstrated by pitfall trapping. It occurs in protected areas such as Lore Lindu National Park and Bogani Nani Wartabone National Park.
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="476" type="bibRefCitation_list">
<paragraph blockId="81.[155,1365,305,1516]" pageId="81" pageNumber="476">
<emphasis bold="true" box="[158,311,1411,1436]" pageId="81" pageNumber="476">Bibliography.</emphasis>
Dubey, Salamin et al. (2008), Esselstyn &amp; Brown (2009), Esselstyn et al. (2009), Hutterer (2005b), Lunde (2008), Miller &amp; Hollister (1921), Musser (1987), Ruedi (1995, 1996), Ruedi &amp; Vogel (1995), Ruedi et al. (1998).
</paragraph>
</subSubSection>
</treatment>
</document>