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<document ID-DOI="http://doi.org/10.5281/zenodo.6870843" ID-ISBN="978-84-16728-08-4" ID-Zenodo-Dep="6870843" approvalRequired="304" approvalRequired_for_illustrations="45" approvalRequired_for_matCits="75" approvalRequired_for_taxonomicNames="40" approvalRequired_for_textStreams="75" approvalRequired_for_treatments="69" checkinTime="1658335596803" checkinUser="felipe" docAuthor="Russell A. Mittermeier &amp; Don E. Wilson" docDate="2018" docId="3D474A54A05787C4FA1BADED138DFCD9" docLanguage="en" docName="hbmw_8_Soricidae_0332.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos, Barcelona: Lynx Edicions" docTitle="Crocidura mariquensis" docType="treatment" docVersion="3" lastPageNumber="525" masterDocId="C17E322CA0288744FF8DAB47125EFFF9" masterDocTitle="Soricidae" masterLastPageNumber="551" masterPageNumber="332" pageNumber="524" updateTime="1658350298165" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Soricidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2018</mods:dateIssued>
<mods:dateOther type="pubDate">2018-07-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>332</mods:start>
<mods:end>551</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6870843</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-08-4</mods:identifier>
<mods:identifier type="Zenodo-Dep">6870843</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6870401" ID-Zenodo-Dep="6870401" LSID="urn:lsid:plazi:treatment:3D474A54A05787C4FA1BADED138DFCD9" httpUri="http://treatment.plazi.org/id/3D474A54A05787C4FA1BADED138DFCD9" lastPageId="128" lastPageNumber="525" pageId="127" pageNumber="524">
<heading pageId="127" pageNumber="524">
<subSubSection box="[1430,1512,1706,1752]" pageId="127" pageNumber="524" type="multiple">
<paragraph blockId="127.[1427,2611,1706,1832]" box="[1430,1512,1706,1752]" pageId="127" pageNumber="524">
<figureCitation box="[1430,1512,1706,1752]" captionStart="Plate 23: Soricidae" captionStartId="122.[130,160,3095,3120]" captionTargetBox="[13,2739,15,3635]" captionTargetPageId="121" captionText="346. Mdumas White-toothed Shrew (Crocidura mdumaz), 347. Montane White-toothed Shrew (Crocidura montis), 348. Newmarks White-toothed Shrew (Crocidura newmarki), 349. Munissis White-toothed Shrew (Crocidura munissi), 350. Tanzanian White-toothed Shrew (Crocidura tansaniana), 351. Usambara White-toothed Shrew (Crocidura usambarae), 352. Kivu White-toothed Shrew (Crocidura kivuana), 353. Niobes White-toothed Shrew (Crocidura niobe), 354. Desert White-toothed Shrew (Crocidura smith), 355. Sahelian Tiny White-toothed Shrew (Crocidura pasha), 356. Roosevelt's White-toothed Shrew (Crocidura roosevelti), 357. Long-footed White-toothed Shrew (Crocidura crenata), 358. Grasses White-toothed Shrew (Crocidura grasset), 359. Bicolored African White-toothed Shrew (Crocidura fuscomurina), 360. Flat-headed White-toothed Shrew (Crocidura planiceps), 361. Crosses White-toothed Shrew (Crocidura crosser), 362. Jouvenets White-toothed Shrew (Crocidura jouvenetae), 363. Mauritanian White-toothed Shrew (Crocidura lusitania), 364. Naked-tailed White-toothed Shrew (Crocidura lttoralis), 365. Gracile White-toothed Shrew (Crocidura maurisca), 366. Reddish-gray White-toothed Shrew (Crocidura cyanea), 367. Swamp White-toothed Shrew (Crocidura mariquensis), 368. Lesser Gray-brown White-toothed Shrew (Crocidura silacea), 369. Dent's White-toothed Shrew (Crocidura denti), 370. Hildegardes White-toothed Shrew (Crocidura hildegardeae), 371. Mamfe White-toothed Shrew (Crocidura virgata), 372. Batess White-toothed Shrew (Crocidura batesi), 373. Sao Tome White-toothed Shrew (Crocidura thomensis), 374. Fingui White-toothed Shrew (Crocidura fingui), 375. Frasers White-toothed Shrew (Crocidura poensis), 376. Nigerian White-toothed Shrew (Crocidura nigeriae), 377. Hun White-toothed Shrew (Crocidura attila), 378. Foxs White-toothed Shrew (Crocidura foxi), 379. Bittikofers White-toothed Shrew (Crocidura buettikoferi), 380. Thereses White-toothed Shrew (Crocidura theresae), 381. Large-headed White-toothed Shrew (Crocidura grandiceps), 382. Wimmers White-toothed Shrew (Crocidura wimmenr), 383. African Black White-toothed Shrew (Crocidura nigrofusca), 384. Savanna Dwarf White-toothed Shrew (Crocidura nanilla), 385. Nimba White-toothed Shrew (Crocidura nimbae)" figureDoi="http://doi.org/10.5281/zenodo.6871925" httpUri="https://zenodo.org/record/6871925/files/figure.png" pageId="127" pageNumber="524">367.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[1530,2134,1706,1752]" pageId="127" pageNumber="524" type="vernacular_names">
<paragraph blockId="127.[1427,2611,1706,1832]" box="[1530,2134,1706,1752]" pageId="127" pageNumber="524">
<vernacularName box="[1530,2134,1706,1752]" pageId="127" pageNumber="524">Swamp White-toothed Shrew</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[2207,2610,1706,1752]" pageId="127" pageNumber="524" type="nomenclature">
<paragraph blockId="127.[1427,2611,1706,1832]" box="[2207,2610,1706,1752]" pageId="127" pageNumber="524">
<taxonomicName baseAuthorityName="A. Smith" baseAuthorityYear="1844" box="[2207,2610,1706,1752]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="mariquensis">
<emphasis box="[2207,2610,1706,1752]" italics="true" pageId="127" pageNumber="524">Crocidura mariquensis</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="524" type="vernacular_names">
<paragraph blockId="127.[1427,2611,1706,1832]" box="[1429,2454,1769,1790]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1429,1504,1769,1790]" pageId="127" pageNumber="524">French:</emphasis>
<vernacularName box="[1513,1725,1769,1790]" pageId="127" pageNumber="524">Crocidure des marais</vernacularName>
/
<emphasis bold="true" box="[1746,1837,1769,1790]" pageId="127" pageNumber="524">German:</emphasis>
<vernacularName box="[1846,2120,1769,1790]" pageId="127" pageNumber="524">Sumpf-WeiRzahnspitzmaus</vernacularName>
/
<emphasis bold="true" box="[2141,2233,1769,1790]" pageId="127" pageNumber="524">Spanish:</emphasis>
<vernacularName box="[2242,2454,1769,1790]" pageId="127" pageNumber="524">Musarana de ciénaga</vernacularName>
</paragraph>
<paragraph blockId="127.[1427,2611,1706,1832]" box="[1428,1894,1808,1829]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1428,1675,1808,1829]" pageId="127" pageNumber="524">Other common names:</emphasis>
<vernacularName box="[1685,1894,1808,1829]" pageId="127" pageNumber="524">Swamp Musk Shrew</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection pageId="127" pageNumber="524" type="reference_group">
<paragraph blockId="127.[2039,2635,1876,2303]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[2040,2195,1876,1909]" pageId="127" pageNumber="524">Taxonomy.</emphasis>
<taxonomicName authority="A. Smith, 1844" authorityName="A. Smith" authorityYear="1844" class="Mammalia" family="Soricidae" genus="Sorex" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="mariqguensis">Sorex mariquensis A. Smith, 1844</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="524" type="materials_examined">
<paragraph blockId="127.[2039,2635,1876,2303]" pageId="127" pageNumber="524">
<materialsCitation country="South Africa" location="A wooded ravine near the Tropic of Capricorn" pageId="127" pageNumber="524" specimenCount="1" stateProvince="North West Province">
<location LSID="urn:lsid:plazi:treatment:3D474A54A05787C4FA1BADED138DFCD9:B031AD99A057873BF7D5AC3B1AD7F83D" country="South Africa" name="A wooded ravine near the Tropic of Capricorn" pageId="127" pageNumber="524" stateProvince="North West Province">A wooded ravine near the Tropic of Capricorn</location>
.” Clarified by N. J. Dippenaar in 1979 as
<location LSID="urn:lsid:plazi:treatment:3D474A54A05787C4FA1BADED138DFCD9:B031AD99A057873BF71AAC8C1B10F815" box="[2199,2382,1995,2028]" country="South Africa" name="Marico River" pageId="127" pageNumber="524" stateProvince="North West Province">Marico River</location>
<locationDeviation pageId="127" pageNumber="524">near its junction with Limpopo River</locationDeviation>
,
<collectingRegion box="[2333,2625,2034,2067]" country="South Africa" name="North West" pageId="127" pageNumber="524">North West Province</collectingRegion>
,
<collectingCountry box="[2040,2216,2081,2106]" name="South Africa" pageId="127" pageNumber="524">South Africa</collectingCountry>
, based on lectotype selection
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="524" type="discussion">
<paragraph blockId="127.[2039,2635,1876,2303]" pageId="127" pageNumber="524">
<taxonomicName baseAuthorityName="A. Smith" baseAuthorityYear="1844" box="[2042,2327,2113,2146]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="mariquensis">Crocidura mariquensis</taxonomicName>
seems to be basal to a large clade including
<taxonomicName baseAuthorityName="Duvernoy" baseAuthorityYear="1838" box="[2357,2481,2152,2185]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="cyanea">C. cyanea</taxonomicName>
,
<taxonomicName authorityName="Thomas" authorityYear="1895" box="[2502,2625,2152,2185]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="silacea">C. silacea</taxonomicName>
,
<taxonomicName authorityName="Thomas" authorityYear="1904" box="[2042,2242,2192,2225]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="528" phylum="Chordata" rank="species" species="hildegardeae">C. hildegardeae</taxonomicName>
,
<taxonomicName authorityName="Dollman" authorityYear="1915" box="[2270,2383,2192,2225]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="batesi">C. batesi</taxonomicName>
,
<taxonomicName authorityName="Dollman" authorityYear="1915" box="[2411,2501,2192,2225]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="538" phylum="Chordata" rank="species" species="foxi">C. foxi</taxonomicName>
,
<taxonomicName authorityName="Jentink" authorityYear="1888" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="buettikoferi">C. buettikoferi</taxonomicName>
,
<taxonomicName box="[2143,2273,2231,2264]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="0" phylum="Chordata" rank="species" species="theresae">C. theresae</taxonomicName>
,
<taxonomicName authorityName="Hutterer" authorityYear="1983" box="[2290,2458,2231,2264]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="538" phylum="Chordata" rank="species" species="grandiceps">C. grandiceps</taxonomicName>
, and
<taxonomicName class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="0" phylum="Chordata" rank="species" species="wimmeri">C. wimmeri</taxonomicName>
. This large clade is sister to a clade
</paragraph>
<paragraph blockId="127.[1421,2631,2309,3482]" pageId="127" pageNumber="524">
including
<taxonomicName authorityName="Brosset, Dubost &amp; Heim de Balsac" authorityYear="1965" box="[1574,1702,2309,2342]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="crenata">C. crenata</taxonomicName>
,
<taxonomicName baseAuthorityName="Heuglin" baseAuthorityYear="1865" box="[1720,1915,2309,2342]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="542" phylum="Chordata" rank="species" species="fuscomurina">C. fuscomurina</taxonomicName>
,
<taxonomicName authorityName="Thomas" authorityYear="1895" box="[1934,2047,2309,2342]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="crossei">C. crossei</taxonomicName>
,
<taxonomicName box="[2065,2233,2309,2342]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="525" phylum="Chordata" rank="species" species="jouvenetae">C. jouvenetae</taxonomicName>
, and
<taxonomicName authorityName="Dollman" authorityYear="1915" box="[2314,2465,2309,2342]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="lusitania">C. lusitania</taxonomicName>
. Three subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="524" type="synonymic_list">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6871723" ID-Zenodo-Dep="6871723" httpUri="https://zenodo.org/record/6871723/files/figure.png" inLine="true" pageId="127" pageNumber="524" targetBox="[1425,2016,1884,2297]" targetPageId="127">
<paragraph blockId="127.[1421,2631,2309,3482]" box="[1426,1826,2386,2419]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1426,1826,2386,2419]" pageId="127" pageNumber="524">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="127.[1421,2631,2309,3482]" box="[1429,2560,2427,2460]" pageId="127" pageNumber="524">
<taxonomicName authority="A. Smith, 1844" authorityName="A. Smith" authorityYear="1844" baseAuthorityName="A. Smith" baseAuthorityYear="1844" box="[1429,1870,2427,2460]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="subSpecies" species="mariquensis" subSpecies="mariquensis">C.m.mariquensisA.Smith,1844—ESouthAfrica,Swaziland,andSMozambique.</taxonomicName>
</paragraph>
<paragraph blockId="127.[1421,2631,2309,3482]" box="[1429,2416,2466,2499]" pageId="127" pageNumber="524">
<taxonomicName authority="Wroughton, 1907" authorityName="Wroughton" authorityYear="1907" box="[1429,1840,2466,2499]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="subSpecies" species="mariquensis" subSpecies="neavet">C.m.neavetWroughton,1907—WCAngola,SDRCongo,andZambia.</taxonomicName>
</paragraph>
<paragraph blockId="127.[1421,2631,2309,3482]" box="[1429,2628,2510,2539]" pageId="127" pageNumber="524">
<taxonomicName authority="St. Leger, 1932" authorityName="St. Leger" authorityYear="1932" box="[1429,1838,2510,2539]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="subSpecies" species="mariquensis" subSpecies="shortridgei">C. m. shortridgei St. Leger, 1932</taxonomicName>
— NE Namibia, N Botswana, and extreme NW Zimbabwe.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="127" pageNumber="524" type="description">
<paragraph blockId="127.[1421,2631,2309,3482]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1426,1681,2545,2578]" pageId="127" pageNumber="524">Descriptive notes.</emphasis>
Head-body 63-105 mm (males) and 62-90 mm (females), tail 45-72 mm (males) and 45-71 mm (females), ear 7-11 mm (males) and 6-10 mm (females), hindfoot 15-20 mm (males) and 14-19 mm (females); weight 9-8-16-5 g (males) and 6-1-13-9 g (females). The Swamp White-toothed Shrew is medium-sized to relatively large, with long pelage. Males, on average, are larger than females. Dorsal pelage is blackish brown, grizzled lightly with reddish fawn (hairs are slate-gray-based with reddish fawn to reddish brown subterminal bands and blackish brown tips). Venter is paler dark brown, grizzled with fawn (hairs are slate-gray basally with fawn to pale brown tips). Feet are yellowish brown, brown, or blackish brown. Tail is ¢.70% of head-body length, relatively hairy, and dark. Females have six inguinal nipples. When walking, tail is uniquely curved slightly upward. Nominate
<taxonomicName baseAuthorityName="A. Smith" baseAuthorityYear="1844" box="[2371,2522,2939,2972]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="mariquensis">mariquensis</taxonomicName>
is dark and larger than neavei, which is also dark, and shortridgei is paler than the other two subspecies. There are three unicuspids.
</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="524" type="biology_ecology">
<paragraph blockId="127.[1421,2631,2309,3482]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1424,1535,3060,3089]" pageId="127" pageNumber="524">Habitat.</emphasis>
Highly associated with swampy riverine regions and semi-aquatic vegetation and reed beds, generally only close to open water such as marshes.
</paragraph>
</subSubSection>
<subSubSection box="[1424,2556,3135,3168]" pageId="127" pageNumber="524" type="food_feeding">
<paragraph blockId="127.[1421,2631,2309,3482]" box="[1424,2556,3135,3168]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1424,1685,3135,3168]" pageId="127" pageNumber="524">Food and Feeding.</emphasis>
Captive Swamp White-toothed Shrews eat snails and termites.
</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="524" type="breeding">
<paragraph blockId="127.[1421,2631,2309,3482]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1423,1558,3174,3207]" pageId="127" pageNumber="524">Breeding.</emphasis>
Reproductive activity of the Swamp White-toothed Shrew has been recorded in the wet season (September—February) in South Africa (subspecies
<taxonomicName baseAuthorityName="A. Smith" baseAuthorityYear="1844" box="[2389,2545,3218,3247]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="127" pageNumber="524" phylum="Chordata" rank="species" species="mariquensis">mariquensis</taxonomicName>
), with litters of 2-7 young (average 4-1). In Botswana (subspecies shortridgei), they were also recorded in wetter months (August—April), with embryo counts of 2-5/female (average 3-3). Three pregnant females were recorded in August-October, February, and April in Zambia (subspecies neaver), with embryo counts of 2-5/female (average 3-8).
</paragraph>
</subSubSection>
<subSubSection lastPageId="128" lastPageNumber="525" pageId="127" pageNumber="524" type="activity">
<paragraph blockId="127.[1421,2631,2309,3482]" pageId="127" pageNumber="524">
<emphasis bold="true" box="[1421,1653,3410,3443]" pageId="127" pageNumber="524">Activity patterns.</emphasis>
Swamp White-toothed Shrews are primarily nocturnal, but they have been recorded during the day. Nests have been found in clumps of tussock grass and
</paragraph>
<paragraph blockId="128.[125,1332,293,804]" pageId="128" pageNumber="525">debris 30 cm above the ground. Nests have not been observed in captivity, although they seem to creep into nesting materials in captivity and creating a hollow in the material.</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="525" type="biology_ecology">
<paragraph blockId="128.[125,1332,293,804]" pageId="128" pageNumber="525">
<emphasis bold="true" box="[126,822,372,405]" pageId="128" pageNumber="525">Movements, Home range and Social organization.</emphasis>
The Swamp White-toothed Shrew is not very aggressive toward conspecifics, but aggressive encounters do occur. Vocalizations accompanying these encounters include short, sharp squeaks and short “chirrs.” Although scent marking has not been observed, their bodies and fecal piles have very strong scents.
</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="525" type="conservation">
<paragraph blockId="128.[125,1332,293,804]" pageId="128" pageNumber="525">
<emphasis bold="true" box="[127,490,569,602]" pageId="128" pageNumber="525">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Swamp White-toothed Shrew has a wide distribution, and there are no major threats affecting populations.
</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="525" type="bibRefCitation_list">
<paragraph blockId="128.[125,1332,293,804]" pageId="128" pageNumber="525">
<emphasis bold="true" box="[128,281,695,720]" pageId="128" pageNumber="525">Bibliography.</emphasis>
Baxter &amp; Dippenaar (2013d), Baxter &amp; Meester (1980), Baxter et al. (1979a), Cassola (2016an), Dippenaar (1977, 1978, 1979), Goulden &amp; Meester (1978), Meester (1963), Rathbun (2005), Taylor (1998), Taylor et al. (1994), Vogelet al. (2013).
</paragraph>
</subSubSection>
</treatment>
</document>