165 lines
17 KiB
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165 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="8BDDA5A9CED55F05944C8ABB749B5AB4" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina hungerfordiana Nevill 1878" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="199839034" LSID="urn:lsid:plazi:treatment:8BDDA5A9CED55F05944C8ABB749B5AB4" httpUri="http://treatment.plazi.org/id/8BDDA5A9CED55F05944C8ABB749B5AB4" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="8BDDA5A9-CED5-5F05-944C-8ABB749B5AB4" authority="Nevill, 1878" authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina hungerfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hungerfordiana">Pupina hungerfordiana Nevill, 1878</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">Figs 21U, V</figureCitation>
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D)." figureDoi="10.3897/zookeys.1119.85400.figure26" httpUri="https://binary.pensoft.net/fig/735387" pageId="0" pageNumber="1">, 26B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina hungerfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hungerfordiana">Pupina hungerfordiana</taxonomicName>
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Nevill, 1878: 300, 301. Type locality: Hsaddan Koo, Salween Valley [Hasaddan Koo, the cave on the limestone hill south of Hpa-An in Ein Du Village, Hpa-An Township, Hpa-An District, Kayin State, Myanmar].
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<bibRefCitation author="Nevill, G" journalOrPublisher="Journal of the Asiatic Society of Bengal, Part II" pageId="0" pageNumber="1" pagination="125 - 167" refId="B105" refString="Nevill, G, 1881. New or little-known Mollusca of the Indo-Malayan fauna. Journal of the Asiatic Society of Bengal, Part II 50 (3): 125 - 167" title="New or little-known Mollusca of the Indo-Malayan fauna." volume="50" year="1881">Nevill 1881</bibRefCitation>
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: 148, pl. 6, fig. 6.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="G.Nevill" authorityYear="1878" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina hungerfordi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hungerfordi">Pupina hungerfordi</taxonomicName>
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[sic]-
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<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen 1897</bibRefCitation>
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: 41, 42, pl. 69, fig. 7, 7a.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) hungerfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hungerfordiana" subGenus="Tylotoechus">Pupina (Tylotoechus) hungerfordiana</taxonomicName>
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-
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 314.
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<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
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: 194, 195.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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of
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<taxonomicName authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina hungerfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hungerfordiana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina hungerfordiana</emphasis>
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</taxonomicName>
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figured in
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<bibRefCitation author="Nevill, G" journalOrPublisher="Journal of the Asiatic Society of Bengal, Part II" pageId="0" pageNumber="1" pagination="125 - 167" refId="B105" refString="Nevill, G, 1881. New or little-known Mollusca of the Indo-Malayan fauna. Journal of the Asiatic Society of Bengal, Part II 50 (3): 125 - 167" title="New or little-known Mollusca of the Indo-Malayan fauna." volume="50" year="1881">Nevill (1881</bibRefCitation>
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: pl. 6, fig. 6).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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NHMUK 91.3.14.686-7 (2 shells; Figs
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21U, V</figureCitation>
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,
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D)." figureDoi="10.3897/zookeys.1119.85400.figure26" httpUri="https://binary.pensoft.net/fig/735387" pageId="0" pageNumber="1">26B</figureCitation>
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) from Hsaddan Koo.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate; last whorl ca. two thirds of shell height. Apertural lip thickened. Parietal tooth thickened, long fin-shaped, reaching beyond the middle of last whorl, outer border curved, covering posterior canal; columellar tooth somewhat thickened, curvedly triangular shaped, located next to slit-like anterior canal.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina hungerfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hungerfordiana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina hungerfordiana</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
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<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. bensoni" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="bensoni">
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<emphasis italics="true" pageId="0" pageNumber="1">P. bensoni</emphasis>
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</taxonomicName>
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sp. nov. in shell shape, but different from
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
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<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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by the long, thickened, fin-shaped parietal tooth, reaching beyond the middle of last whorl, and different from
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. bensoni" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="bensoni">
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<emphasis italics="true" pageId="0" pageNumber="1">P. bensoni</emphasis>
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</taxonomicName>
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sp. nov. by the lack of furrow between the inner and outer peristomes.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Known only from the type locality (
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<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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As
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. hungerfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="hungerfordiana">
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<emphasis italics="true" pageId="0" pageNumber="1">P. hungerfordiana</emphasis>
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</taxonomicName>
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was described based on a single specimen as explicitly stated in the original description, that specimen is the holotype fixed by monotypy (
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<bibRefCitation author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
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: Art. 73.1.2).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |