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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1144.98644" ID-GBIF-Dataset="3b01e63b-186d-4a0a-ac41-8e2a8a4435c1" ID-Pensoft-Pub="1313-2970-1144-171" ID-Pensoft-UUID="D148129EDAED5CD0ACA97302FBE35254" ID-ZooBank="C11074779CE243F9998521D42836A8F4" ModsDocID="1313-2970-1144-171" checkinTime="1675346916091" checkinUser="pensoft" docAuthor="Tran, Ngat Thi, Engel, Michael S., Nguyen, Cuong Quang, Tran, Duong Dinh &amp; Nguyen, Lien Thi Phuong" docDate="2023" docId="8B8108AE254D57BB8C115FE6F05F0C94" docLanguage="en" docName="ZooKeys 1144: 171-196" docOrigin="ZooKeys 1144" docPubDate="2023-02-02" docSource="http://dx.doi.org/10.3897/zookeys.1144.98644" docTitle="Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen 2023, sp. nov." docType="treatment" docUuid="4CF71E2F-C15B-457A-B383-5C3BF9AC34B1" docUuidSource="ZooBank" docVersion="2" id="D148129EDAED5CD0ACA97302FBE35254" lastPageNumber="171" masterDocId="D148129EDAED5CD0ACA97302FBE35254" masterDocTitle="The bee genus Anthidiellum in Vietnam: descriptions of five new species and the first male of Anthidiellum coronum (Hymenoptera, Megachilidae)" masterLastPageNumber="196" masterPageNumber="171" pageNumber="171" updateTime="1675347537077" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The bee genus Anthidiellum in Vietnam: descriptions of five new species and the first male of Anthidiellum coronum (Hymenoptera, Megachilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tran, Ngat Thi</mods:namePart>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier type="email">tranthingat1012@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Engel, Michael S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3067-077X</mods:nameIdentifier>
<mods:affiliation>Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 &amp; th</mods:affiliation>
<mods:nameIdentifier type="email">msengel@ku.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Cuong Quang</mods:namePart>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tran, Duong Dinh</mods:namePart>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Lien Thi Phuong</mods:namePart>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1144</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>171</mods:start>
<mods:end>196</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1144.98644</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1144.98644</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1144-171</mods:identifier>
<mods:identifier type="ZooBank">C11074779CE243F9998521D42836A8F4</mods:identifier>
<mods:identifier type="Pensoft-UUID">D148129EDAED5CD0ACA97302FBE35254</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:4CF71E2F-C15B-457A-B383-5C3BF9AC34B1" httpUri="http://treatment.plazi.org/id/8B8108AE254D57BB8C115FE6F05F0C94" lastPageNumber="171" pageId="0" pageNumber="171">
<subSubSection pageId="0" pageNumber="171" type="nomenclature">
<paragraph pageId="0" pageNumber="171">
<taxonomicName LSID="https://zoobank.org/4CF71E2F-C15B-457A-B383-5C3BF9AC34B1" authority="Tran, Engel &amp; Nguyen" authorityName="Tran, Engel &amp; Nguyen" authorityYear="2023" class="Insecta" family="Megachilidae" genus="Anthidiellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthidiellum (Pycnanthidium) ayun" order="Hymenoptera" pageId="0" pageNumber="171" phylum="Arthropoda" rank="species" species="ayun" status="sp. nov." subGenus="Pycnanthidium">Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="171">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="description">
<paragraph pageId="0" pageNumber="171">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">Fig. 2A-H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="type material">
<paragraph pageId="0" pageNumber="171">Type material.</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Holotype</emphasis>
.
</emphasis>
Vietnam: ♀, Gia Lai, Mang Yang, Ayun, Kon Ka Kinh NP,
<geoCoordinate degrees="14" direction="north" minutes="13" orientation="latitude" precision="15" seconds="18" value="14.221666">14°13'18&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="19" orientation="longitude" precision="15" seconds="02" value="108.31722">108°19'02&quot;E</geoCoordinate>
, alt. 907 m, 26.iii.2022 [26 March 2022], Lien Thi Phuong Nguyen, Ngat Thi Tran, Cuong Quang Nguyen leg. [IEBR].
</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Paratypes</emphasis>
.
</emphasis>
Vietnam: 51♀♀, same data as holotype [41♀♀ in IEBR; 10♀♀ in AMNH]; 30♀♀, Gia Lai, Mang Yang, Ayun, Kon Ka Kinh NP,
<geoCoordinate degrees="14" direction="north" minutes="12" orientation="latitude" precision="15" seconds="11" value="14.203055">14°12'11&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="18" orientation="longitude" precision="15" seconds="58" value="108.31612">108°18'58&quot;E</geoCoordinate>
, alt. 834 m, 25.iii.2022 [25 March 2022], Lien Thi Phuong Nguyen, Ngat Thi Tran, Cuong Quang Nguyen leg.; 4♀♀, Gia Lai, Mang Yang, Ayun, Kon Ka Kinh NP,
<geoCoordinate degrees="14" direction="north" minutes="12" orientation="latitude" precision="15" seconds="11" value="14.203055">14°12'11&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="18" orientation="longitude" precision="15" seconds="58" value="108.31612">108°18'58&quot;E</geoCoordinate>
, alt. 834 m, 26.iv.2022 [25 April 2022], Lien Thi Phuong Nguyen, Ngat Thi Tran leg. [IEBR].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="diagnosis">
<paragraph pageId="0" pageNumber="171">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="171">
This female of this species is most similar to
<taxonomicName lsidName="A. (P.) chumomray" pageId="0" pageNumber="171" rank="species" species="chumomray" subGenus="P.">A. (P.) chumomray</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="171">vide infra</emphasis>
), sharing with it the following characters: metatibia and metabasitarsus with a longitudinal carina on prolateral surfaces, and metasomal T2, supraclypeal area, pronotal lobe, and axillae black. It can be separated from that species easily by the following characters: mandible tridentate, teeth gradually longer and teeth sharp apically; clypeus approximately 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as broad as long and with trapezoidal yellow marking medially; mesoscutellum convex and bigibbous; paraocular area black.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="description">
<paragraph pageId="0" pageNumber="171">Description.</paragraph>
<paragraph pageId="0" pageNumber="171">♀: body length 7.0-7.5 mm (holotype = 7.5 mm). Forewing length 6.5-7.0 mm (holotype = 7.0 mm).</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Structure</emphasis>
.
</emphasis>
Head broader than long, approximately 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as broad as long (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2C</figureCitation>
). Compound eyes about 2.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, about 1.4
<normalizedToken originalValue="×">x</normalizedToken>
genal width. Mandible tridentate, teeth gradually longer and sharp apically, interspace between first two teeth broad, second tooth nearly contiguous with third tooth (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2C</figureCitation>
). Clypeus slightly convex medially, approximately 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as broad as long. Supraclypeal area slightly convex medially. Scape slender, about 3.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, pedicel approximately 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad and about 1.3
<normalizedToken originalValue="×">x</normalizedToken>
length of F1; F1 as long as broad and 1.5
<normalizedToken originalValue="×">x</normalizedToken>
length of F2; F3-F5, F6-F7, F8-F9 subequal in length; F10 longest flagellomere, about 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad. Mesosoma approximately 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as broad as long; mesoscutum with medioapical margin slightly sharp; mesoscutellum convex, with mediolongitudinal depression apically giving bigibbous surface (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2D</figureCitation>
). Prolateral surface of metatibia and metabasitarsus each with a longitudinal carina (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2G</figureCitation>
). Metapretarsal arolium present (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2H</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">Figure 2.</emphasis>
<taxonomicName authorityName="Tran, Engel &amp; Nguyen" authorityYear="2023" class="Insecta" family="Megachilidae" genus="Anthidiellum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthidiellum (Pycnanthidium) ayun" order="Hymenoptera" pageId="0" pageNumber="171" phylum="Arthropoda" rank="species" species="ayun" subGenus="Pycnanthidium">Anthidiellum (Pycnanthidium) ayun</taxonomicName>
Tran, Engel &amp; Nguyen, sp. nov., holotype, female
<emphasis bold="true" pageId="0" pageNumber="171">A</emphasis>
dorsolateral oblique habitus
<emphasis bold="true" pageId="0" pageNumber="171">B</emphasis>
dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="171">C</emphasis>
facial view
<emphasis bold="true" pageId="0" pageNumber="171">D</emphasis>
mesosoma in dorsal view
<emphasis bold="true" pageId="0" pageNumber="171">E</emphasis>
head and anterior mesosoma in profile, red arrow indicating omaular carina
<emphasis bold="true" pageId="0" pageNumber="171">F</emphasis>
dorsal oblique view of metasoma
<emphasis bold="true" pageId="0" pageNumber="171">G</emphasis>
prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae
<emphasis bold="true" pageId="0" pageNumber="171">H</emphasis>
metapretarsal arolium (red arrow). Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="171">A-G</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="171">H</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Sculpturing and texture</emphasis>
.
</emphasis>
Outer surface of mandible with abundant, small, dense punctures except larger, wrinkled punctures on proximal half. Clypeus with dense, shallow punctures, puncture much smaller on apical margin than on disc. Paraocular area, supraclypeal area, and frons with large, coarse, contiguous punctures. Vertex with round, dense punctures of unequal sizes, puncture sizes smaller than those on frons. Mesoscutum with large, coarse, contiguous punctures; mesoscutellum with contiguous punctures of unequal sizes, puncture sizes smaller than those of mesoscutum (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2D</figureCitation>
). Metasomal T1-T6 with small, round, dense punctures except punctures larger laterally.
</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Color</emphasis>
.
</emphasis>
Body black except as follows: clypeus with trapezoid yellow marking except lateral and apical margins black (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2C</figureCitation>
). Mesoscutum with short, thin yellow markings latero-anteriorly (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2D</figureCitation>
). Metasomal T1 with yellow markings laterally; T3 basally with yellow band and large interrupted medially; T4 basally with yellow band and narrow interrupted medially (near as continuously in some paratypes); T5 basally with yellow band (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Anthidiellum (Pycnanthidium) ayun Tran, Engel &amp; Nguyen, sp. nov., holotype, female A dorsolateral oblique habitus B dorsal habitus C facial view D mesosoma in dorsal view E head and anterior mesosoma in profile, red arrow indicating omaular carina F dorsal oblique view of metasoma G prolateral surfaces of metatibia and metatarsus, with red arrows indicating longitudinal carinae H metapretarsal arolium (red arrow). Scale bars: 1 mm (A-G); 0.5 mm (H)." figureDoi="10.3897/zookeys.1144.98644.figure2" httpUri="https://binary.pensoft.net/fig/803872" pageId="0" pageNumber="171">2A, B, F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">
<emphasis italics="true" pageId="0" pageNumber="171">Pubescence</emphasis>
.
</emphasis>
Ventral margin of mandible with abundant, short, yellowish setae intermixed with long setae. Clypeus with short yellowish setae except sparse, erect, yellow setae on apical margin. Paraocular area, supraclypeal area, frons, and vertex with short yellowish setae. Diffuse tufts of long, dense, plumose, yellowish setae above antennal toruli. Mesoscutum and mesoscutellum with abundant short yellowish setae; mesoscutellum with long, erect, yellowish setae ventro-apically; propodeum with long, plumose, dense, yellowish setae. Prolateral surfaces of basitarsi and tarsi with dense, erect, yellowish setae; retrolateral surfaces of these same podites with tawny yellow setae. Metasomal T1-T6 with short, sparse, yellowish setae except short, yellowish setae on T6 apically; S2-S5 with long, dense, tawny yellow scopal setae.
</paragraph>
<paragraph pageId="0" pageNumber="171">♂: Latet.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="etymology">
<paragraph pageId="0" pageNumber="171">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="171">The specific epithet is a toponym for type locality, the Ayun commune in Gia Lai Province. The name is treated as a noun in apposition.</paragraph>
</subSubSection>
</treatment>
</document>