317 lines
33 KiB
XML
317 lines
33 KiB
XML
<document id="26AC6E2EA1C21039901202CCB499B6CC" ENCODING="UTF-8" ID-GBIF-Dataset="21d21894-7003-4d6d-882b-5322ea07968d" ModsDocAuthor="Longino, J. T." ModsDocID="21311" checkinTime="1244213549108" checkinUser="thomas" docAuthor="Longino, J. T." docDate="2007" docId="8514CF76288578BDFA9BE3279E8B281E" docLanguage="en" docName="21311_gg2.xml" docOrigin="Zootaxa 1491" docSource="http://www.antbase.org/ants/publications/21311/21311.pdf" docTitle="Azteca flavigaster Longino, 2007, new species" docType="treatment" docVersion="12" lastPageNumber="31" masterDocId="A75589B0B733EFAA7DECE020EC76B43D" masterDocTitle="A taxonomic review of the genus Azteca (Hymenoptera: Formicidae) in Costa Rica and a global revision of the aurita group." masterLastPageNumber="63" masterPageNumber="1" pageNumber="30" updateTime="1701311503130" updateUser="plazi">
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<mods:titleInfo id="735BE17FAB9D20535F6C24D46F6D1021">
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<mods:title id="4BE0D0394D5B51480253A2CF361F2553">A taxonomic review of the genus Azteca (Hymenoptera: Formicidae) in Costa Rica and a global revision of the aurita group.</mods:title>
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<mods:name id="BD0A7B562043F55AFB4005222F84AD59" type="personal">
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<mods:namePart id="2CCD7DFFD5A8E955D16F21484CA5E950">Longino, J. T.</mods:namePart>
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<mods:typeOfResource id="5961619932DB4BD2B61A158D20567B5E">text</mods:typeOfResource>
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<mods:title id="6376605A63FD7A48CCE5A6EE14EB65F8">Zootaxa</mods:title>
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<mods:part id="B124A03D1925D1B43FDD2ED02ABB56A2">
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<mods:date id="7B6CB548B02F7C0AE03955C1E47ACA84">2007</mods:date>
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<mods:detail id="2D33EECB9C8203ECCABBD185DEE3F9D9" type="volume">
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<mods:number id="2FB0D506FB4A37AEFE67C56FA8E05B75">1491</mods:number>
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<mods:start id="8A99BD7C5FF07F0468CB334BCABDB4DF">1</mods:start>
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<mods:location id="B36507D693D9326619A1A3EE28900F7C">
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<mods:url id="29B472B0E20A4EB0A10056125F35E05F">http://www.antbase.org/ants/publications/21311/21311.pdf</mods:url>
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<mods:classification id="8A6A4A72A951A5C30D52D592780C77B6">journal article</mods:classification>
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<mods:identifier id="62B7F458612E289C0FF9E421115D2EAE" type="HNS-Pub">21311</mods:identifier>
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<mods:identifier id="E7410E7C6C53A280CA0FCD71D9451AC6" type="ZooBank">C31A1226-724D-4D1A-8471-E6BB441EE3EF</mods:identifier>
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<treatment id="8514CF76288578BDFA9BE3279E8B281E" ID-DOI="http://doi.org/10.5281/zenodo.6246510" ID-GBIF-Taxon="100125914" ID-Zenodo-Dep="6246510" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E" httpUri="http://treatment.plazi.org/id/8514CF76288578BDFA9BE3279E8B281E" lastPageNumber="31" pageNumber="30">
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<subSubSection id="2512747358F7F745D46BD6E70C2CAAA0" type="nomenclature">
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<paragraph id="45938E10E8E4C4456FFF743994D3FBEA" pageNumber="30">
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<taxonomicName id="812CD9D7E4DD3346ADEB8031FADB2094" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239261" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca flavigaster" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="flavigaster">Azteca flavigaster</taxonomicName>
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<taxonomicNameLabel id="F779521802EBAD2D809185CB6ED1125E" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="9519634437F812F8950D429262EF4C4B" type="description">
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<paragraph id="89C57611B99F7D764AC2BC7F7379D043" pageNumber="30">Figures 3,4A,5,6A,6E,6F,8.</paragraph>
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</subSubSection>
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<subSubSection id="BD9587604D5BA307EA91DA2265000F2C" type="nomenclature">
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<paragraph id="5660C6E9061C6C614EA79610798E5C9A" pageNumber="30">
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<materialsCitation id="9E4775A9D14181586BFD26969DA9D6EE" ID-GBIF-Occurrence="923890898" collectingDate="1989-06-04" collectionCode="INBC" collectorName="J. Longino" country="Costa Rica" latitude="9.4" location="Punta Quepos, 5 m" longitude="-84.1666667" specimenCode="JTLC 000005689" stateProvince="Prov. Puntarenas" typeStatus="Holotype">
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<typeStatus id="DC5E7D1173F2D6A76E75B2A72E4AED0D">Holotype</typeStatus>
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alate queen:
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<collectingCountry id="F0807804D9E785C0589A6D1C58ABEA89">Costa Rica</collectingCountry>
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,
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<collectingRegion id="C340268AD68777A258092F2B54FF335F">Prov. Puntarenas</collectingRegion>
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,
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<location id="DB028553B975ABAF3F78EB29FD101829" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:DB028553B975ABAF3F78EB29FD101829" country="Costa Rica" latitude="9.4" longitude="-84.166664" name="Punta Quepos" stateProvince="Prov. Puntarenas">Punta Quepos</location>
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,
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<geoCoordinate id="A73501F8577A2387D76790917FBA6BE0" direction="north" orientation="latitude" value="9.4">9°24'N</geoCoordinate>
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,
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<geoCoordinate id="52BA850268DBBCE9F66F3A49324E441D" direction="west" orientation="longitude" value="-84.166664">84°10'W</geoCoordinate>
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,
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<location id="2E5F4AB55E088294923BC031C0DF8CDD" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:2E5F4AB55E088294923BC031C0DF8CDD" country="Costa Rica" latitude="9.4" longitude="-84.166664" name="5 m" stateProvince="Prov. Puntarenas">5m</location>
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,
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<date id="048CC39E0BB83D323A2104F33E6758CD" day="04" month="06" value="1989-06-04" year="1989">
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<collectingDate id="E70FF5E3CF684D42AC5F9918D5D61716" day="04" month="06" value="1989-06-04" year="1989">4 Jun 1989</collectingDate>
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</date>
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(
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<collectorName id="480C18610D37170676FE8C5E1D0A17AC">J. Longino</collectorName>
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) [
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<collectionCode id="6F0F607D28318978D55886584AA1072A" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
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, specimen (pin) code
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<specimenCode id="9B473BCA55F389443B427F4DCFB84B25">JTLC000005689</specimenCode>
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].
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</materialsCitation>
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</paragraph>
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<paragraph id="63DE6E69EEBC48BE3EDBB25B1688680E" country="Costa Rica" pageNumber="30">
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<materialsCitation id="277FCF2E1ADAD5E6D1FCA1E566E6D992" ID-GBIF-Occurrence="923890884" collectingDate="1989-06-04" collectionCode="INBC, USNM, MCZC, LACM" collectorName="J. Longino" country="Costa Rica" latitude="9.4" location="Punta Quepos, 5 m" longitude="-84.1666667" specimenCode="JTLC000005689, JTLC000005687, JTLC000005688, INBIOCRI001281548" stateProvince="Prov. Puntarenas" typeStatus="Paratypes">
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<typeStatus id="9BF49A8CFE84676BFC4F473631B2067E">Paratypes</typeStatus>
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: same data as holotype; 4 workers, 3 alate queens [
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<specimenCode id="CEF29544C50E31B1870AC67E441C392C">JTLC000005689</specimenCode>
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,
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<collectionCode id="290377D82C8ACCDE4B4E3FF9A3D1C56C" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
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;
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<specimenCode id="43985BC072C774E7142397751517C4FC">JTLC000005687</specimenCode>
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,
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<collectionCode id="A33C2364C4B6ACA745264B028BBB1377" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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;
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<specimenCode id="A6EA15DE91350CF6B6D0B1FE07F172D2">JTLC000005688</specimenCode>
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,
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<collectionCode id="3D8D2778603A48C279603D50A0100158" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
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;
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<specimenCode id="329EA7F70F63AF306F7491F7B83A6A2E">INBIOCRI001281548</specimenCode>
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,
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<collectionCode id="22E64FA294F0B4C1F13B96D3A8DD7957" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
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].
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6A6CE413AA164C78E94ECEB75522FC3C" type="description">
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<paragraph id="9C7AEDC7FDC21A061C3397BE723B17BD" pageNumber="30">Measurements of holotype. HLA 1.62, HLB 1.59, HW 1.53, HW 1.53, SL 1.03, EL 0.44, OC 0.10, MTSC 30.</paragraph>
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</subSubSection>
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<subSubSection id="56D23779D0B4C5F78B7828E480967CBF" type="diagnosis">
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<paragraph id="810010371C09FCEC258066DEB7F52444" pageNumber="30">
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Diagnosis. Among the species with 6,4 palpal formula,
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<taxonomicName id="48D28945A322E2CC32E9E94167657C04" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239261" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca flavigaster" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="flavigaster">A. flavigaster</taxonomicName>
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has the smallest queen head size. The most similar species are
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<taxonomicName id="6CF528BD91EF0302329BC8E38A242C8B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239264" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca quadraticeps" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="quadraticeps">A. quadraticeps</taxonomicName>
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,
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<taxonomicName id="A6B6C9F788A506BD157A1084925FE1D9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26035" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca velox Forel" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="velox">A. velox</taxonomicName>
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, and
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<taxonomicName id="43E3A4B135CFBA350ABEE8BBAAD9BF88" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239262" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca nigra" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="nigra">A. nigra</taxonomicName>
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, all of which have larger queen head size. The workers have a unique coloration, with bright yellow gastral dorsum and contrastingly dark brown mesosoma.
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</paragraph>
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</subSubSection>
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<subSubSection id="41D4B402DFCA29EF4DE8D826F0796977" type="description">
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<paragraph id="7A99F60A7E9DE10AA765B0DE64B7C9F7" pageNumber="30">Queen characters. Measurements (n=5): HLA 1.55 (1.54-1.60), HW 1.52 (1.51-1.55), SL 1.04 (1.01- 1.04), CI 98 (96-100), SI 67 (65-67).</paragraph>
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<paragraph id="B54A3D674740918F90D0043299EFFE7B" pageNumber="30">Palpal formula 6,4; ocelli small; middle and hind tibia with prominent pectinate apical spur; dorsal surface of mandible with small piligerous puncta, setae in puncta short, subdecumbent, interspaces between puncta smooth and shiny on apical half of mandible, gradually becoming faintly microareolate at base; medial and lateral clypeal lobes at about same level; head subquadrate, posterior margin not strongly cordate, very shallowly excavate; petiolar node tall, strongly compressed into thin scale at apex; posteroventral petiolar lobe evenly convex from front to back; scape with abundant erect setae, about as long as one half maximum width of scape; middle and hind tibia with abundant erect setae, longest of these about as long as one half maximum width of tibia (MTSC 20-35); sides of head without erect setae; posterior margin of head with erect setae; pronotum with erect setae on posterior margin; mesoscutum, scutellum, and propodeum with abundant erect setae; petiolar node with irregular pubescence and sparse short erect setae, 0-2 pairs of erect setae extending above apex in profile, posterior border of sternal lobe of petiole with uniform layer of short erect setae; gastral terga with sparse long erect setae; most of head dark brown, with lighter orange coloration restricted to anterior malar area near mandibular insertions.</paragraph>
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<paragraph id="AA788D72C940AFC72DD435A25A9A13D1" pageNumber="30">Worker characters. Measurements (n=4): HLA 1.19 (1.16-1.23), HW 1.20 (1.15-1.26), SL 0.96 (0.94- 0.98), CI 101 (99-102), SI 81 (80-82).</paragraph>
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<paragraph id="4DC83A3DFFC2C98D8ABA861ECC677AD6" lastPageNumber="31" pageNumber="30">Palpal formula 6,4; middle and hind tibia with prominent pectinate apical spur; dorsal surface of mandible smooth and shining, with moderately abundant small piligerous puncta, setae in puncta short, erect, larger puncta with long setae near masticatory margin; medial and lateral clypeal lobes at about same level; head with convex sides, strongly cordate posterior margin; in lateral profile promesonotum forming single convexity; scape with abundant erect setae, length of setae about one half maximum width of scape; mid and hind tibia with abundant erect setae, longest setae about one half maximum width of tibia; side of head with 1-5 erect setae on malar area, otherwise lacking setae; posterior margin of head with abundant erect setae; pronotum, mesonotum, and dorsal face of propodeum with abundant long erect setae; anterior and anterolateral portions of head light yellow brown, variable extent of darker brown on medial vertex and posteriorly, mesosoma variably light to dark brown, gastral dorsum bright yellow, contrasting with darker mesosoma.</paragraph>
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</subSubSection>
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<subSubSection id="EDAE4555F6FAA2AED194B08ED0395EB9" type="etymology">
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<paragraph id="FA4BB04AE6F45F476713F096A00E45BB" pageNumber="31">Etymology. The name refers to the bright yellow gaster of the worker.</paragraph>
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</subSubSection>
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<subSubSection id="8478BDF7E4CA45291FF98CAAEBC4A572" type="distribution">
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<paragraph id="C5E45B928B44DD3F00343C54F5C25397" pageNumber="31">Range. Costa Rica, Panama, Ecuador.</paragraph>
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</subSubSection>
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<subSubSection id="43692A7F3BCA7DFD34A003D2708BFC4F" type="biology_ecology">
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<paragraph id="5A76E3E8D8D087CB38635DEB1E2D06FF" pageNumber="31">
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Biology. Most observations of
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<taxonomicName id="4AA70F131A93DC7E56F7CD0A729C92A3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239261" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca flavigaster" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="flavigaster">A. flavigaster</taxonomicName>
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have been made in the southern Pacific lowlands of Costa Rica, in moderately seasonal evergreen forest.
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<taxonomicName id="0C7B4385DF465AC8FBD32CA1B9EA72A4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239261" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca flavigaster" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="flavigaster">Azteca flavigaster</taxonomicName>
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is a generalist cavity nester with conspicuous surface-foraging workers. In a degraded patch of forest near Quepos, at a pasture edge, I observed abundant workers on the trunk of a live, 40cm diameter tree, emerging from a long narrow fissure at the base. Abundant workers were also on the base of another small tree in the same forest patch. This tree had a dead branch at ground level with workers, brood, and scattered alate females in two parallel cavities, each with horizontal carton partitions. Large columns of workers went up the trunk to a larger dead branch further up. In a patch of mature forest near Ciudad Neily an aggregation of workers were in cavities in the main stem of a sapling of
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<taxonomicName id="86306FE1179F66BE80B95BCC83675538" class="Magnoliopsida" family="Lecythidaceae" genus="Grias" higherTaxonomySource="CoL" kingdom="Plantae" order="Ericales" pageNumber="31" phylum="Tracheophyta" rank="genus">Grias</taxonomicName>
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(Lecythidaceae). At Sirena in Corcovado National Park I made several observations of this species. I observed a column of foraging workers on the ground surface, flowing in and out of a 4cm diameter hole extending horizontally into the ground. I observed workers 18m high in the canopy of a
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<taxonomicName id="EFC4DE9230C6C84D6BA81A3A52A67C44" class="Magnoliopsida" family="Moraceae" genus="Perebea" higherTaxonomySource="CoL" kingdom="Plantae" order="Rosales" pageNumber="31" phylum="Tracheophyta" rank="species" species="trophophylla">Perebea trophophylla</taxonomicName>
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(Moraceae) tree, with an aggregation of workers in a small knot. I observed an incipient colony in an
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<taxonomicName id="771AF1341CAD68C44D72305CD761B1BE" class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageNumber="31" phylum="Tracheophyta" rank="species" species="alleni">Acacia alleni</taxonomicName>
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tree, the common myrmecophytic acacia in Corcovado. The tree was in poor shape, without a dominant
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<taxonomicName id="753ECBC359C5B859B1F73BD02D45609B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24858" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pseudomyrmex Lund" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="genus">Pseudomymex</taxonomicName>
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colony. The
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<taxonomicName id="C5D6688A9B1FC4B7D303B91BE4A2FB3B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2370" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca Forel" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="genus">Azteca</taxonomicName>
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occupied several of the thorns, one of which contained a physogastric queen and small brood. Finally, I observed two founding queens in a dead and tattered
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<taxonomicName id="11AFCDE0F1EC498BFECD239ED4FF1859" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" order="Zingiberales" pageNumber="31" phylum="Tracheophyta" rank="genus">Costus</taxonomicName>
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(Costaceae) inflorescence. The inflorescence was filled with many old chambers containing dead
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<taxonomicName id="1D725E9F99604417E32144D7FA8DEF83" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2370" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca Forel" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="genus">Azteca</taxonomicName>
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queen remains. The two living queens were in adjacent chambers, but not in contact. Each had brood, and one had nanitic workers. When I put the queens together in the same vial, they immediately locked mandibles in combat. Other researchers at Sirena have collected founding queens in chambers of
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<taxonomicName id="6D7D7E60C5FED2CE2E290BDADF871E10" class="Magnoliopsida" family="Salicaceae" genus="Tetrathylacium" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageNumber="31" phylum="Tracheophyta" rank="species" species="costaricensis">Tetrathylacium costaricensis</taxonomicName>
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and
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<taxonomicName id="CFDC0284194369EA35DA861F526CDDA2" class="Magnoliopsida" family="Salicaceae" genus="Tetrathylacium" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageNumber="31" phylum="Tracheophyta" rank="species" species="macrophyllum">T. macrophyllum</taxonomicName>
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.
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</paragraph>
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<paragraph id="9D6C28F49AB8A6109706D2816A281525" pageNumber="31">
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Workers with the same coloration and habitus of
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<taxonomicName id="C3200DAF576645F0FD1427931915ED55" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239261" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca flavigaster" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="flavigaster">Azteca flavigaster</taxonomicName>
|
|
occur in the Atlantic lowlands of Costa Rica, but are rare. At La Selva Biological Station these workers have been collected twice. A worker was collected in a Berlese sample of canopy soil and epiphytes, and a nest or nest fragment was found in a live branch of
|
|
<taxonomicName id="E465D81B08B213F08357462CCFEBD1C8" class="Magnoliopsida" family="Urticaceae" genus="Coussapoa" higherTaxonomySource="CoL" kingdom="Plantae" order="Rosales" pageNumber="31" phylum="Tracheophyta" rank="genus">Coussapoa</taxonomicName>
|
|
(Cecropiaceae). The nest also contained workers of
|
|
<taxonomicName id="27B119106395C8D46110F139F18D0779" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26217" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus atriceps (Smith)" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="atriceps">Camponotus atriceps</taxonomicName>
|
|
, in a parabiotic association.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7E3212B6F1423072D10C43DE18B941DF" type="discussion">
|
|
<paragraph id="77A3E31C922A196165EEFECB70A7DF7D" pageNumber="31">
|
|
Comments. Although workers matching
|
|
<taxonomicName id="F343CA970C61D8B33C1425DEA17AB6FE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239261" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca flavigaster" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="flavigaster">A. flavigaster</taxonomicName>
|
|
have been found on the Atlantic side of Costa Rica and in other countries, no queens have been collected to confirm these identifications. However, the
|
|
<taxonomicNameLabel id="3EF306E746716AB0BA2C870B11A89DD4" rank="species">new species</taxonomicNameLabel>
|
|
<taxonomicName id="7573FC180F877996D15F03807D381155" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239264" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca quadraticeps" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="quadraticeps">A. quadraticeps</taxonomicName>
|
|
is known only from queens from the Atlantic lowlands. It is possible that
|
|
<taxonomicName id="2CC7A431960AD35CEE2BB41623EBAB67" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239264" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca quadraticeps" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="quadraticeps">A. quadraticeps</taxonomicName>
|
|
and
|
|
<taxonomicName id="C5C27EA4ED9217790ADD41FD0F444BA3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239261" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca flavigaster" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="flavigaster">A. flavigaster</taxonomicName>
|
|
are allopatric or parapatric sister taxa with similar workers, and that the Atlantic slope workers with yellow gasters are actually the workers of
|
|
<taxonomicName id="EDAE787B4548A5B394AE0CB5317C1E08" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239264" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca quadraticeps" order="Hymenoptera" pageNumber="31" phylum="Arthropoda" rank="species" species="quadraticeps">A. quadraticeps</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="23208654AFA416EA1484F6435E53E829" type="materials_examined">
|
|
<paragraph id="4EE7E9169C96C8ECCC5DEEAF5EC445E3" pageNumber="31">
|
|
Additional material examined.
|
|
<materialsCitation id="C2295677670A4650BA5197C5A0A4BE99" ID-GBIF-Occurrence="923890910" collectingDate="1993-11-08" collectionCode="INBC" collectorName="ALAS" country="COSTA RICA" latitude="10.433333" location="La Selva Biological Station, 50 m" longitude="-84.01667" stateProvince="Heredia">
|
|
<collectingCountry id="2A9D7E3796D99351B2499A194321DBF0">COSTA RICA</collectingCountry>
|
|
:
|
|
<collectingRegion id="4DEAA52BB819A8A97F4884518122AB0F">Heredia</collectingRegion>
|
|
:
|
|
<location id="9D2D4F08E8D1A57A3D6B4DAEBA905166" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:9D2D4F08E8D1A57A3D6B4DAEBA905166" country="COSTA RICA" latitude="10.433333" longitude="-84.01667" name="La Selva Biological Station" stateProvince="Heredia">La Selva Biological Station</location>
|
|
,
|
|
<geoCoordinate id="F1A02FED7E1381325D78D4E09D819DF9" direction="north" orientation="latitude" value="10.433333">10°26'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="0367DEBB617946780328F4193049C02B" direction="west" orientation="longitude" value="-84.01667">84°01'W</geoCoordinate>
|
|
,
|
|
<location id="6B7E01290807201DAEFD0366BD688D12" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:6B7E01290807201DAEFD0366BD688D12" country="COSTA RICA" latitude="10.433333" longitude="-84.01667" name="50 m" stateProvince="Heredia">50m</location>
|
|
,
|
|
<date id="41D0EA9D236EC10B64D2E24DE265FC49" day="08" month="11" value="1993-11-08" year="1993">
|
|
<collectingDate id="A86CE6CE6EB7F044C000E1AA428AC54E" day="08" month="11" value="1993-11-08" year="1993">8 Nov 1993</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="D4E4A923374D41D3A62C26D69E80465D">ALAS</collectorName>
|
|
) - worker [
|
|
<collectionCode id="68C999DF837345D889BB611051D11DD0" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="2A56FA04F715B6816CE8EA2877561F09" ID-GBIF-Occurrence="923890932" collectingDate="2004-04-20" collectorName="A. Fitch" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station, 50 m" longitude="-84.01667" stateProvince="Heredia">
|
|
same locality,
|
|
<date id="6E301707D8036E18167BAB70357A8BD9" day="20" month="04" value="2004-04-20" year="2004">
|
|
<collectingDate id="C3570742D28D2E0A292C5FF3F36DBB6B" day="20" month="04" value="2004-04-20" year="2004">20 Apr 2004</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="B236A8285F847FFC0EA10CE9783797BC">A. Fitch</collectorName>
|
|
) - worker
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="9FD4D743ECA039D9E716C97734AE9A88" ID-GBIF-Occurrence="923890902" collectingDate="2000-04" collectorName="M. Schmidt" country="Costa Rica" latitude="8.483334" location="Sirena, Corcovado National Park" longitude="-83.6" stateProvince="Puntarenas">
|
|
<collectingRegion id="262E1EF028496DEDEEA52EAF395432CE">Puntarenas</collectingRegion>
|
|
:
|
|
<location id="72B64FE211C3D000093708D29911BBC3" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:72B64FE211C3D000093708D29911BBC3" country="Costa Rica" latitude="8.483334" longitude="-83.6" name="Sirena, Corcovado National Park" stateProvince="Puntarenas">Sirena, Corcovado National Park</location>
|
|
,
|
|
<geoCoordinate id="7BEDC068AEC1D875661522DF39D96E5B" direction="north" orientation="latitude" value="8.483334">8°29'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="1F2A2A7B87582E6C6C1D45565434F64C" direction="west" orientation="longitude" value="-83.6">83°36'W</geoCoordinate>
|
|
,
|
|
<collectingDate id="77866188495921EF5F9CC15222A89396">Feb-Apr 2000</collectingDate>
|
|
(
|
|
<collectorName id="FEE05906340D3F1572CF20D68207F194">M. Schmidt</collectorName>
|
|
) - dealate queen
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="4A1434C92D58433B3201D56D83A41C8A" ID-GBIF-Occurrence="923890925" collectorName="J. Longino" country="Costa Rica" latitude="8.483334" location="Sirena, Corcovado National Park" longitude="-83.6" stateProvince="Puntarenas">
|
|
same locality, multiple collections (
|
|
<collectorName id="714B5C22A13DFD317C54FDFF32683EBE">J. Longino</collectorName>
|
|
) - workers, queens
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="7576C8B68CC7E1737A289CF60BCE0E0E" ID-GBIF-Occurrence="923890911" collectingDate="1987-07-22" collectorName="L. Tennant" country="Costa Rica" latitude="8.483334" location="Sirena, Corcovado National Park" longitude="-83.6" stateProvince="Puntarenas">
|
|
same locality,
|
|
<date id="33E60A2A55575B65422FD7A3EDC51311" day="22" month="07" value="1987-07-22" year="1987">
|
|
<collectingDate id="B912B77F4D5324951E1717E27A6D32B7" day="22" month="07" value="1987-07-22" year="1987">22 Jul 1987</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="03754747073D2AF99FE7290E32C3236C">L. Tennant</collectorName>
|
|
) - dealate queen
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="879A878AD82BCF3DF8F6A92AC92BB6C7" ID-GBIF-Occurrence="923890933" collectingDate="1990-03-26" collectorName="J. Longino" country="Costa Rica" latitude="8.483334" location="Ciudad Neily, 20 m" locationDeviation="19 km S" longitude="-82.96667" stateProvince="Puntarenas">
|
|
<locationDeviation id="06462E812AA484094387675A6CDB20B4">19km S</locationDeviation>
|
|
<location id="85CA3D0682E41753B00F889B0F36A525" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:85CA3D0682E41753B00F889B0F36A525" country="Costa Rica" latitude="8.483334" longitude="-82.96667" name="Ciudad Neily" stateProvince="Puntarenas">Ciudad Neily</location>
|
|
,
|
|
<geoCoordinate id="7F752BD3557B2EB4C71B8BBDE3FD554A" direction="north" orientation="latitude" value="8.483334">8°29'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="922D1C7C115CBFBD30D785A99D6AC548" direction="west" orientation="longitude" value="-82.96667">82°58'W</geoCoordinate>
|
|
,
|
|
<location id="4B2C8738FE478D54B33C2EE02533E379" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:4B2C8738FE478D54B33C2EE02533E379" country="Costa Rica" latitude="8.483334" longitude="-82.96667" name="20 m" stateProvince="Puntarenas">20m</location>
|
|
,
|
|
<date id="D5DE1E62D09BF31357E79B44D3695A2D" day="26" month="03" value="1990-03-26" year="1990">
|
|
<collectingDate id="A3E6E0C5138B87ABEA89B529644224ED" day="26" month="03" value="1990-03-26" year="1990">26 Mar 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="5E4893ACCAB65F4E4106D4A32DC44249">J. Longino</collectorName>
|
|
) - workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="B0B21DB3EC25F718FE2D0C185650AB21" ID-GBIF-Occurrence="923890916" collectingDate="1990-03-25" collectorName="P. S. Ward" country="Costa Rica" latitude="8.483334" location="Ciudad Neily, 20 m" locationDeviation="19 km S" longitude="-82.96667" stateProvince="Puntarenas">
|
|
same locality,
|
|
<date id="BFED7B5F93AE127FC023031C0D1AF87A" day="25" month="03" value="1990-03-25" year="1990">
|
|
<collectingDate id="5C72467610B413C6EB61788E057E60D1" day="25" month="03" value="1990-03-25" year="1990">25 Mar 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="422209222F5698798361C4F4B63BF5F8">P. S. Ward</collectorName>
|
|
) - worker
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="F621D5CE5DF662E2C1ECF6261D914C70" ID-GBIF-Occurrence="923890918" collectingDate="1991-08-11" collectorName="P. S. Ward" country="ECUADOR" latitude="-1.2333333" location="Jauneche, 19 km WSW Mocache, 60 m" longitude="-79.666664" stateProvince="Los Rios">
|
|
<collectingCountry id="13D6DCD19032D98D5E0E5D310B49F54C">ECUADOR</collectingCountry>
|
|
:
|
|
<collectingRegion id="1A11F8D7002FA5C9B3A9A78631937395">Los Rios</collectingRegion>
|
|
:
|
|
<location id="09516B0E2F5B3A323CAFDC6C7DDA9639" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:09516B0E2F5B3A323CAFDC6C7DDA9639" country="ECUADOR" latitude="-1.2333333" longitude="-79.666664" name="Jauneche, 19 km WSW Mocache" stateProvince="Los Rios">Jauneche, 19km WSW Mocache</location>
|
|
,
|
|
<geoCoordinate id="6430401DC7844A3B0D03D7AF7A705F71" direction="south" orientation="latitude" value="-1.2333333">1°14'S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="62F616D53D5F7BBF41F91AA1BA1EE2B8" direction="west" orientation="longitude" value="-79.666664">79°40'W</geoCoordinate>
|
|
,
|
|
<location id="6B4841AF5566DB91967D0C508E9FC23D" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:6B4841AF5566DB91967D0C508E9FC23D" country="ECUADOR" latitude="-1.2333333" longitude="-79.666664" name="60 m" stateProvince="Los Rios">60m</location>
|
|
,
|
|
<date id="1D077C0AEC26DE188CDB02078AFA34E1" day="11" month="08" value="1991-08-11" year="1991">
|
|
<collectingDate id="8544AC2EA2EBF4E9DF30A91792BFBDB1" day="11" month="08" value="1991-08-11" year="1991">11 Aug 1991</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="48666B1151D10264277AAADB772EFDE6">P. S. Ward</collectorName>
|
|
) - workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="D46D4AE16037345CF04FA406633B4A70" ID-GBIF-Occurrence="923890921" collectingDate="1990-09-10" collectorName="D. M. Olson" country="PANAMA" latitude="9.116667" location="Pipeline Road, 50 m" longitude="-79.73333" stateProvince="Canal Zone">
|
|
<collectingCountry id="EC571558649D0EBA9F37C8369779EDFC">PANAMA</collectingCountry>
|
|
:
|
|
<collectingRegion id="1DB17AC6ACDAAF2DE9A9CAC236EB4081">Canal Zone</collectingRegion>
|
|
:
|
|
<location id="F141EAF08FC492D7C5AFE5DD25B6FD53" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:F141EAF08FC492D7C5AFE5DD25B6FD53" country="PANAMA" latitude="9.116667" longitude="-79.73333" name="Pipeline Road" stateProvince="Canal Zone">Pipeline Road</location>
|
|
,
|
|
<geoCoordinate id="E7E190A6A9326700072A18E73AB880BA" direction="north" orientation="latitude" value="9.116667">9°07'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="9472A91F6164ADD01ABDF7164503D608" direction="west" orientation="longitude" value="-79.73333">79°44'W</geoCoordinate>
|
|
,
|
|
<location id="63F8B02260D140068F5C9F57A63CBF38" LSID="urn:lsid:plazi:treatment:8514CF76288578BDFA9BE3279E8B281E:63F8B02260D140068F5C9F57A63CBF38" country="PANAMA" latitude="9.116667" longitude="-79.73333" name="50 m" stateProvince="Canal Zone">50m</location>
|
|
,
|
|
<date id="E4DE19EAD159F4BF5F7CB67682DF1F12" day="10" month="09" value="1990-09-10" year="1990">
|
|
<collectingDate id="9E5693CE8DD9F8332AEB7C8FC9646ECC" day="10" month="09" value="1990-09-10" year="1990">10 Sep 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="C8E504B2A1249A7ED24A56B7914D2838">D. M. Olson</collectorName>
|
|
) - worker
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |