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<mods:title id="64B7215A676B2A5C108917485A306346">The systematic status of two enigmatic ocypodoid crabs, &quot; Paracleistostoma &quot; dentatum Tesch, 1918, and &quot; Paracleistostoma &quot; fossulum Barnard, 1955 (Crustacea: Decapoda: Brachyura)</mods:title>
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<treatment id="F224110FED1AD036FF2F6CFFFEECF82A" ID-DOI="http://doi.org/10.5281/zenodo.6170625" ID-GBIF-Taxon="119509384" ID-Zenodo-Dep="6170625" LSID="urn:lsid:plazi:treatment:F224110FED1AD036FF2F6CFFFEECF82A" httpUri="http://treatment.plazi.org/id/F224110FED1AD036FF2F6CFFFEECF82A" lastPageId="2" lastPageNumber="60" pageId="1" pageNumber="59">
<subSubSection id="3297F392ED1AD035FF2F6CFFFDE6FB95" pageId="1" pageNumber="59" type="nomenclature">
<paragraph id="7A32A019ED1AD035FF2F6CFFFDB6FC0C" blockId="1.[151,630,976,1002]" box="[151,630,976,1002]" pageId="1" pageNumber="59">
<heading id="217A1775ED1AD035FF2F6CFFFDB6FC0C" bold="true" box="[151,630,976,1002]" fontSize="11" level="1" pageId="1" pageNumber="59" reason="1">
<emphasis id="48F97C0BED1AD035FF2F6CFFFDB6FC0C" bold="true" box="[151,630,976,1002]" pageId="1" pageNumber="59">
Family
<taxonomicName id="BD8DDB9AED1AD035FF4E6CFFFDB6FC0C" ID-CoL="7NJ9F" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[246,630,976,1002]" class="Malacostraca" family="Dotillidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="family">
DOTILLIDAE
<bibRefCitation id="1E1CDDE8ED1AD035FE016CFFFDB6FC0C" author="Stimpson" box="[441,630,976,1002]" pageId="1" pageNumber="59" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="7A32A019ED1AD035FF2F6B39FDE6FBD6" blockId="1.[151,550,1045,1072]" box="[151,550,1045,1072]" pageId="1" pageNumber="59">
<emphasis id="48F97C0BED1AD035FF2F6B39FE7CFBD6" bold="true" box="[151,444,1046,1072]" pageId="1" pageNumber="59">
<emphasis id="48F97C0BED1AD035FF2F6B39FE40FBD6" bold="true" box="[151,384,1046,1072]" italics="true" pageId="1" pageNumber="59">Lazarocleistostoma</emphasis>
Štev
</emphasis>
ě
<emphasis id="48F97C0BED1AD035FE716B39FE11FBD6" bold="true" box="[457,465,1046,1072]" pageId="1" pageNumber="59">i</emphasis>
ć,
<emphasis id="48F97C0BED1AD035FE556B39FDE6FBD6" bold="true" box="[493,550,1046,1072]" pageId="1" pageNumber="59">2011</emphasis>
</paragraph>
<paragraph id="7A32A019ED1AD035FF2F6B72FDE6FB95" blockId="1.[151,1107,1117,1170]" box="[151,550,1117,1139]" pageId="1" pageNumber="59">
<emphasis id="48F97C0BED1AD035FF2F6B72FE9CFB95" box="[151,348,1117,1139]" italics="true" pageId="1" pageNumber="59">Lazarocleistostoma</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FEDB6B72FE2FFB95" author="Stevcic" box="[355,495,1117,1139]" pageId="1" pageNumber="59" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčić, 2011</bibRefCitation>
: 136.
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED1AD035FF2F6B53FB93FB74" box="[151,1107,1147,1170]" pageId="1" pageNumber="59" type="materials_examined">
<paragraph id="7A32A019ED1AD035FF2F6B53FB93FB74" blockId="1.[151,1107,1117,1170]" box="[151,1107,1147,1170]" pageId="1" pageNumber="59">
<emphasis id="48F97C0BED1AD035FF2F6B53FEDBFB74" bold="true" box="[151,283,1147,1170]" pageId="1" pageNumber="59">
<typeStatus id="A5361EBBED1AD035FF2F6B53FF0AFB77" box="[151,202,1148,1169]" pageId="1" pageNumber="59">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="BD8DDB9AED1AD035FE916B53FD02FB77" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[297,706,1147,1170]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED1AD035FE916B53FDFFFB74" box="[297,575,1148,1170]" italics="true" pageId="1" pageNumber="59">Paracleistostoma dentatum</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FDFD6B54FD02FB77" author="Tesch" box="[581,706,1147,1169]" pageId="1" pageNumber="59" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, by original designation; gender neuter.
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED1AD035FF2F6B91FC0FFA10" pageId="1" pageNumber="59" type="diagnosis">
<paragraph id="7A32A019ED1AD035FF2F6B91FC0FFA10" blockId="1.[151,1436,1214,2031]" pageId="1" pageNumber="59">
<emphasis id="48F97C0BED1AD035FF2F6B91FED1FB31" bold="true" box="[151,273,1214,1239]" pageId="1" pageNumber="59">Diagnosis.</emphasis>
Carapace transversely hexagonal, regions just discernible (
<figureCitation id="E2B6BC9CED1AD035FC1F6B91FC31FB30" box="[935,1009,1214,1238]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Figs. 1</figureCitation>
A, D). Front narrow, entire, with shallow concave indentation medially; lateral margins angular, with low epigastric lobes; external orbital tooth prominent; anterolateral margin deeply concave with distinct lateral tooth; posterolateral margin short, convex (
<figureCitation id="E2B6BC9CED1AD035FA866A29FABEFAF8" box="[1342,1406,1286,1310]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
A, B, D). Eyes prominent, peduncle long, slender (
<figureCitation id="E2B6BC9CED1AD035FD0E6A05FD36FAA4" box="[694,758,1322,1346]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
A, D). Posterior margin of epistome with triangular median lobe, lateral margins deeply concave (
<figureCitation id="E2B6BC9CED1AD035FDF16A61FD4AFA80" box="[585,650,1358,1382]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
E). Third maxillipeds operculiform, almost completely filling buccal cavity; exopod slender, partially hidden under ischium, reaching just beyond distal part of merus (
<figureCitation id="E2B6BC9CED1AD035FB646A5DFADBFA6C" box="[1244,1307,1394,1418]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
C, F). Cheliped very slender, short, symmetrical; merus subcylindrical; chela slender, cutting edges of fingers unarmed setae (
<figureCitation id="E2B6BC9CED1AD035FF276A95FF1FFA34" box="[159,223,1466,1490]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
A, G). Ambulatory legs relatively long, third leg longest; merus unarmed; dactyli of first to third legs falciform; that of last leg spatuliform, lined with numerous setae (
<figureCitation id="E2B6BC9CED1AD035FCFE6AF1FC45FA10" box="[838,901,1502,1526]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
A, H).
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED1AD036FF7F692CFD65FDB9" lastPageId="2" lastPageNumber="60" pageId="1" pageNumber="59" type="discussion">
<paragraph id="7A32A019ED1AD035FF7F692CFC77F94C" blockId="1.[151,1436,1214,2031]" pageId="1" pageNumber="59">
<emphasis id="48F97C0BED1AD035FF7F692CFEFBF9FD" bold="true" box="[199,315,1539,1563]" pageId="1" pageNumber="59">Remarks.</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FEFC692DFD00F9FC" author="Manning" box="[324,704,1538,1562]" pageId="1" pageNumber="59" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis (1981: 209)</bibRefCitation>
made it clear that
<taxonomicName id="BD8DDB9AED1AD035FC21692CFAA3F9FC" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[921,1379,1538,1563]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED1AD035FC21692CFB0EF9FD" box="[921,1230,1539,1563]" italics="true" pageId="1" pageNumber="59">Paracleistostoma dentatum</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FB6F692DFAA3F9FC" author="Tesch" box="[1239,1379,1538,1562]" pageId="1" pageNumber="59" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, was not a species of
<emphasis id="48F97C0BED1AD035FEEE6909FDD2F9D8" box="[342,530,1574,1598]" italics="true" pageId="1" pageNumber="59">Paracleistosoma</emphasis>
or of a camptandriid but did not know for certain where to place it. Ng
<emphasis id="48F97C0BED1AD035FAD96907FA5CF9D8" box="[1377,1436,1574,1599]" italics="true" pageId="1" pageNumber="59">et al.</emphasis>
(2008: 236) agreed and commented that the only known
<typeStatus id="A5361EBBED1AD035FC9F6964FC97F985" box="[807,855,1611,1635]" pageId="1" pageNumber="59">type</typeStatus>
female of this species had been examined and the structure of its frontal carapace margin and third maxillipeds indicated it was likely to be a dotillid, probably close to the genus
<taxonomicName id="BD8DDB9AED1AD035FE9D69BCFDFAF94C" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[293,570,1682,1707]" class="Malacostraca" family="Dotillidae" genus="Ilyoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="48F97C0BED1AD035FE9D69BCFE41F94D" box="[293,385,1683,1707]" italics="true" pageId="1" pageNumber="59">Ilyoplax</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FE3F69BDFDFAF94C" author="Stimpson" box="[391,570,1682,1706]" pageId="1" pageNumber="59" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
, and certainly not a camptandriid.
</paragraph>
<paragraph id="7A32A019ED1AD035FF716999FCCDF8DC" blockId="1.[151,1436,1214,2031]" pageId="1" pageNumber="59">
<bibRefCitation id="1E1CDDE8ED1AD035FF716999FEB1F928" author="Stevcic" box="[201,369,1718,1743]" pageId="1" pageNumber="59" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčiċ (2011)</bibRefCitation>
established a new genus,
<emphasis id="48F97C0BED1AD035FD146999FC46F928" box="[684,902,1718,1742]" italics="true" pageId="1" pageNumber="59">Lazarocleistostoma</emphasis>
, for
<taxonomicName id="BD8DDB9AED1AD035FC7D6999FA52F928" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[965,1426,1718,1742]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED1AD035FC7D6999FB3DF928" box="[965,1277,1718,1742]" italics="true" pageId="1" pageNumber="59">Paracleistostoma dentatum</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FAB16999FA52F928" author="Tesch" box="[1289,1426,1718,1742]" pageId="1" pageNumber="59" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, without providing any explanation for why a new taxon was necessary. He also decided to recognise a new family, Lazarocleistostomidae
<bibRefCitation id="1E1CDDE8ED1AD035FE2369D1FDF4F8F0" author="Stevcic" box="[411,564,1790,1815]" pageId="1" pageNumber="59" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčiċ, 2011</bibRefCitation>
(
<bibRefCitation id="1E1CDDE8ED1AD035FDFC69D1FCD5F8F0" author="Stevcic" box="[580,789,1790,1815]" pageId="1" pageNumber="59" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčiċ 2011: 127</bibRefCitation>
), again without comment or regard to earlier discussions by
<bibRefCitation id="1E1CDDE8ED1AD035FF02680DFE34F8DC" author="Manning" box="[186,500,1826,1850]" pageId="1" pageNumber="59" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis (1981)</bibRefCitation>
or Ng
<emphasis id="48F97C0BED1AD035FDFB680BFDBCF8DD" box="[579,636,1827,1851]" italics="true" pageId="1" pageNumber="59">et al.</emphasis>
(2008: 236).
</paragraph>
<paragraph id="7A32A019ED1AD036FF716869FAC0FEFA" blockId="1.[151,1436,1214,2031]" lastBlockId="2.[151,1436,151,607]" lastPageId="2" lastPageNumber="60" pageId="1" pageNumber="59">
The
<typeStatus id="A5361EBBED1AD035FF446868FEECF8B9" box="[252,300,1863,1887]" pageId="1" pageNumber="59">type</typeStatus>
specimen is broken and in poor condition, but the narrow front, relatively long eyes and orbits, form of the female chelae as well as operculiform third maxillipeds (
<figureCitation id="E2B6BC9CED1AD035FCE36845FC5BF864" box="[859,923,1898,1922]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="1" pageNumber="59">Fig. 1</figureCitation>
) strongly suggest it is a member of the
<taxonomicName id="BD8DDB9AED1AD035FAE16845FF11F840" class="Malacostraca" family="Dotillidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="family">Dotillidae</taxonomicName>
. Overall, this species is similar to
<taxonomicName id="BD8DDB9AED1AD035FDF468A1FCA1F840" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[588,865,1934,1958]" class="Malacostraca" family="Dotillidae" genus="Ilyoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="48F97C0BED1AD035FDF468A1FD68F840" box="[588,680,1934,1958]" italics="true" pageId="1" pageNumber="59">Ilyoplax</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FD1768A1FCA1F840" author="Stimpson" box="[687,865,1934,1958]" pageId="1" pageNumber="59" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
, or
<taxonomicName id="BD8DDB9AED1AD035FC3168A1FB76F840" authority="Tweedie, 1950" authorityName="Tweedie" authorityYear="1950" box="[905,1206,1934,1958]" class="Malacostraca" family="Dotillidae" genus="Dotilloplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="48F97C0BED1AD035FC3168A1FBC8F840" box="[905,1032,1934,1958]" italics="true" pageId="1" pageNumber="59">Dotilloplax</emphasis>
<bibRefCitation id="1E1CDDE8ED1AD035FBB768A1FB76F840" author="Tweedie" box="[1039,1206,1934,1958]" pageId="1" pageNumber="59" refString="Tweedie, M. W. F. (1950) Grapsoid crabs from Labuan and Sarawak. The Sarawak Museum Journal, 5 (2), 338 - 369." type="journal article" year="1950">Tweedie, 1950</bibRefCitation>
</taxonomicName>
. It is known that
<taxonomicName id="BD8DDB9AED1AD035FACD68A1FF15F82D" class="Malacostraca" family="Dotillidae" genus="Ilyoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="48F97C0BED1AD035FACD68A1FF15F82D" italics="true" pageId="1" pageNumber="59">Ilyoplax</emphasis>
</taxonomicName>
is polyphyletic (Davie &amp; Naruse 2011;
<bibRefCitation id="1E1CDDE8ED1AD035FD2C689DFCAAF82C" author="Kitaura" box="[660,874,1970,1995]" pageId="1" pageNumber="59" refString="Kitaura, J., Wada, K. &amp; Nishida, M. (1998) Molecular phylogeny and evolution of unique mud-using territorial behavior in ocypodid crabs (Crustacea: Brachyura: Ocypodidae). Molecular Biology and Evolution, 15, 626 - 637." type="journal article" year="1998">
Kitaura
<emphasis id="48F97C0BED1AD035FD48689BFCE8F82D" box="[752,808,1971,1995]" italics="true" pageId="1" pageNumber="59">et al.</emphasis>
1998
</bibRefCitation>
;
<bibRefCitation id="1E1CDDE8ED1AD035FCCD689DFBB5F82C" author="Kitaura" box="[885,1141,1970,1994]" pageId="1" pageNumber="59" refString="Kitaura, J. &amp; Wada, K. (2006) New species of Ilyoplax (Brachyura: Ocypodidae: Dotillinae) from the Philippines and Indonesia: behavioral, molecular, and morphological evidence. Raffles Bulletin of Zoology, 54 (2), 373 - 379." type="journal article" year="2006">Kitaura &amp; Wada, 2006</bibRefCitation>
; P.J.F. Davie, unpublished data) and the carapace of
<taxonomicName id="BD8DDB9AED1AD035FE0468F7FD81F808" box="[444,577,2006,2031]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="59" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED1AD035FE0468F7FD81F808" box="[444,577,2006,2031]" italics="true" pageId="1" pageNumber="59">P. dentatum</emphasis>
</taxonomicName>
is unusual for a dotillid or even an ocypodoid. The carapace is atypical for ocypodoids in that there is a wide space between the external orbital tooth and first lateral tooth, resulting in a relatively short posterolateral margin. The anterolateral margin is not as long even in species like
<taxonomicName id="BD8DDB9AED19D036FB686F93FEFBFF11" authority="Tweedie, 1950" authorityName="Tweedie" authorityYear="1950" class="Malacostraca" family="Dotillidae" genus="Dotilloplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="kempi">
<emphasis id="48F97C0BED19D036FB686F93FA5CFF32" box="[1232,1436,188,212]" italics="true" pageId="2" pageNumber="60">Dotilloplax kempi</emphasis>
<bibRefCitation id="1E1CDDE8ED19D036FF2F6FF0FEFBFF11" author="Tweedie" box="[151,315,223,247]" pageId="2" pageNumber="60" refString="Tweedie, M. W. F. (1950) Grapsoid crabs from Labuan and Sarawak. The Sarawak Museum Journal, 5 (2), 338 - 369." type="journal article" year="1950">Tweedie, 1950</bibRefCitation>
</taxonomicName>
(
<typeStatus id="A5361EBBED19D036FEF16FF0FEBBFF11" box="[329,379,223,247]" pageId="2" pageNumber="60">type</typeStatus>
locality Labuan, Borneo), which has well separated, sharp external orbital and lateral carapace teeth. A separate genus for
<taxonomicName id="BD8DDB9AED19D036FE716E2BFC4DFEFA" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[457,909,260,284]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED19D036FE716E2BFD3CFEFA" box="[457,764,260,284]" italics="true" pageId="2" pageNumber="60">Paracleistostoma dentatum</emphasis>
<bibRefCitation id="1E1CDDE8ED19D036FCBB6E2BFC4DFEFA" author="Tesch" box="[771,909,260,284]" pageId="2" pageNumber="60" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, is therefore probably warranted.
</paragraph>
<paragraph id="7A32A019ED19D036FF716E07FB17FE82" blockId="2.[151,1436,151,607]" pageId="2" pageNumber="60">
By referring
<taxonomicName id="BD8DDB9AED19D036FEE16E07FCDCFED9" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[345,796,295,320]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED19D036FEE16E07FD4CFEA6" box="[345,652,296,320]" italics="true" pageId="2" pageNumber="60">Paracleistostoma dentatum</emphasis>
<bibRefCitation id="1E1CDDE8ED19D036FD2B6E08FCDCFED9" author="Tesch" box="[659,796,295,319]" pageId="2" pageNumber="60" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, to
<emphasis id="48F97C0BED19D036FCFC6E07FBDEFEA6" box="[836,1054,296,320]" italics="true" pageId="2" pageNumber="60">Lazarocleistostoma</emphasis>
<bibRefCitation id="1E1CDDE8ED19D036FB9D6E08FB7FFED9" author="Stevcic" box="[1061,1215,295,320]" pageId="2" pageNumber="60" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčiċ, 2011</bibRefCitation>
, and not to
<taxonomicName id="BD8DDB9AED19D036FAF86E07FA5CFEA6" box="[1344,1436,296,320]" class="Malacostraca" family="Dotillidae" genus="Ilyoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="48F97C0BED19D036FAF86E07FA5CFEA6" box="[1344,1436,296,320]" italics="true" pageId="2" pageNumber="60">Ilyoplax</emphasis>
</taxonomicName>
also solves the homonymy problem with
<taxonomicName id="BD8DDB9AED19D036FDDD6E63FC68FE82" authority="Ward, 1933" authorityName="Ward" authorityYear="1933" box="[613,936,332,356]" class="Malacostraca" family="Dotillidae" genus="Ilyoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="48F97C0BED19D036FDDD6E63FCDBFE82" box="[613,795,332,356]" italics="true" pageId="2" pageNumber="60">Ilyoplax dentata</emphasis>
<bibRefCitation id="1E1CDDE8ED19D036FC9B6E63FC68FE82" author="Ward" box="[803,936,332,356]" pageId="2" pageNumber="60" refString="Ward, M. (1933) New genera and species of marine Decapod Brachyura. From the coasts of New South Wales and Queensland. Australian Zoologist, 7, 377 - 394, pls. 21 - 23." type="journal article" year="1933">Ward, 1933</bibRefCitation>
</taxonomicName>
, at least for the time being.
</paragraph>
<paragraph id="7A32A019ED19D036FF716E40FD65FDB9" blockId="2.[151,1436,151,607]" pageId="2" pageNumber="60">
Members of
<taxonomicName id="BD8DDB9AED19D036FEE46E5FFE78FE6E" box="[348,440,368,392]" class="Malacostraca" family="Dotillidae" genus="Ilyoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="48F97C0BED19D036FEE46E5FFE78FE6E" box="[348,440,368,392]" italics="true" pageId="2" pageNumber="60">Ilyoplax</emphasis>
</taxonomicName>
(and clearly all dotillids), however, are intertidal, and the
<typeStatus id="A5361EBBED19D036FBED6E40FB77FE61" box="[1109,1207,367,391]" pageId="2" pageNumber="60" type="holotype">holotype</typeStatus>
of
<emphasis id="48F97C0BED19D036FB586E5EFAA7FE6E" box="[1248,1383,368,392]" italics="true" pageId="2" pageNumber="60">
L.
<taxonomicName id="BD8DDB9AED19D036FB476E5FFAA7FE6E" box="[1279,1383,368,392]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="dentatum">dentatum</taxonomicName>
</emphasis>
was collected from depths of
<quantity id="BD750DFCED19D036FE0D6EBBFDE8FE4A" box="[437,552,404,428]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="3.4" metricValueMin="0.9" pageId="2" pageNumber="60" unit="m" value="21.5" valueMax="34.0" valueMin="9.0">
9 to
<specimenCount id="6C8B6B90ED19D036FE536EBBFDE8FE4A" box="[491,552,404,428]" pageId="2" pageNumber="60" type="generic">34 m</specimenCount>
</quantity>
. An allied family which lives in subtidal waters,
<taxonomicName id="BD8DDB9AED19D036FBE46EBBFF12FE29" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" class="Malacostraca" family="Xenophthalmidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="family">
Xenophthalmidae
<bibRefCitation id="1E1CDDE8ED19D036FA946EBBFF12FE29" author="Stimpson" pageId="2" pageNumber="60" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
, was recognised by Ng
<emphasis id="48F97C0BED19D036FE646E96FDD4FE36" box="[476,532,440,464]" italics="true" pageId="2" pageNumber="60">et al.</emphasis>
(2008: 245), and while it shared similar mouthparts and chelipeds with dotillids, differed from them markedly by the form of their epistome, orbits, eyes, ambulatory legs and carapace. In summary, with regard to the Lazarocleistostomidae
<bibRefCitation id="1E1CDDE8ED19D036FD156ED0FC88FDF1" author="Stevcic" box="[685,840,511,536]" pageId="2" pageNumber="60" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčiċ, 2011</bibRefCitation>
, the structure of the mouthparts, ambulatory legs and chelae as discussed above leave little doubt that it is a dotillid. As such, Lazarocleistostomidae
<bibRefCitation id="1E1CDDE8ED19D036FB586D0BFABBFDDA" author="Stevcic" box="[1248,1403,548,573]" pageId="2" pageNumber="60" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčiċ, 2011</bibRefCitation>
, is synonymised in the
<taxonomicName id="BD8DDB9AED19D036FEC06D68FD61FDB9" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[376,673,583,607]" class="Malacostraca" family="Dotillidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="family">
Dotillidae
<bibRefCitation id="1E1CDDE8ED19D036FE566D68FD61FDB9" author="Stimpson" box="[494,673,583,607]" pageId="2" pageNumber="60" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED19D036FF2F6D9CFD57FC95" pageId="2" pageNumber="60" type="reference_group">
<paragraph id="7A32A019ED19D036FF2F6D9CFF2BFD09" blockId="2.[151,651,691,751]" pageId="2" pageNumber="60">
Lazarocleistostoma
<taxonomicName id="BD8DDB9AED19D036FEC46D9CFD4BFD2A" authority="(Tesch, 1918)" authorityName="Tesch" authorityYear="1918" baseAuthorityName="Tesch" baseAuthorityYear="1918" box="[380,651,691,716]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED19D036FEC46D9CFD43FD2A" bold="true" box="[380,643,691,716]" pageId="2" pageNumber="60">
<emphasis id="48F97C0BED19D036FEC46D9CFE28FD2A" bold="true" box="[380,488,691,716]" italics="true" pageId="2" pageNumber="60">dentatum</emphasis>
(
<bibRefCitation id="1E1CDDE8ED19D036FE4F6D9CFD43FD2A" author="Tesch" box="[503,643,691,716]" pageId="2" pageNumber="60" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</emphasis>
)
</taxonomicName>
(
<figureCitation id="E2B6BC9CED19D036FF276DF8FF20FD09" box="[159,224,727,751]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="7A32A019ED19D036FF2F6C0FFEE7FCB2" blockId="2.[151,1437,800,883]" pageId="2" pageNumber="60">
<treatmentCitationGroup id="5A9D8737ED19D036FF2F6C0FFEE7FCB2" pageId="2" pageNumber="60">
<taxonomicName id="BD8DDB9AED19D036FF2F6C0FFD9CFCD0" authority="Tesch, 1918: 63" authorityName="Tesch" authorityPageNumber="63" authorityYear="1918" box="[151,604,800,822]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="dentatum">
<emphasis id="48F97C0BED19D036FF2F6C0FFE6EFCD0" box="[151,430,800,822]" italics="true" pageId="2" pageNumber="60">Paracleistostoma dentatum</emphasis>
<treatmentCitation id="FB2C8608ED19D036FE0E6C0FFD9CFCD0" author="Tesch" box="[438,604,800,822]" page="63" pageId="2" pageNumber="60" year="1918">
<bibRefCitation id="1E1CDDE8ED19D036FE0E6C0FFDF5FCD3" author="Tesch" box="[438,565,800,822]" pageId="2" pageNumber="60" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
: 63
</treatmentCitation>
</taxonomicName>
, pl. 3 fig. 2;
<treatmentCitation id="FB2C8608ED19D036FD5E6C0FFC5EFCD3" author="Serene" box="[742,926,800,822]" page="101" pageId="2" pageNumber="60" year="1968">
<bibRefCitation id="1E1CDDE8ED19D036FD5E6C0FFCA9FCD0" author="Serene" box="[742,873,800,822]" pageId="2" pageNumber="60" refString="Serene, R. (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, No. 1, 33 - 120." type="book chapter" year="1968">Serène 1968</bibRefCitation>
: 101
</treatmentCitation>
;
<treatmentCitation id="FB2C8608ED19D036FC126C0FFB33FCD3" author="Manning" box="[938,1267,800,822]" page="209" pageId="2" pageNumber="60" year="1981">
<bibRefCitation id="1E1CDDE8ED19D036FC126C0FFB7EFCD3" author="Manning" box="[938,1214,800,822]" pageId="2" pageNumber="60" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis 1981</bibRefCitation>
: 209
</treatmentCitation>
; Ng
<emphasis id="48F97C0BED19D036FA9E6C0EFA9BFCD0" box="[1318,1371,800,822]" italics="true" pageId="2" pageNumber="60">et al.</emphasis>
2008: 235, 236.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7A32A019ED19D036FF2F6C72FD57FC95" blockId="2.[151,1437,800,883]" box="[151,663,861,883]" pageId="2" pageNumber="60">
<treatmentCitationGroup id="5A9D8737ED19D036FF2F6C72FD57FC95" box="[151,663,861,883]" pageId="2" pageNumber="60">
<emphasis id="48F97C0BED19D036FF2F6C72FE01FC95" box="[151,449,861,883]" italics="true" pageId="2" pageNumber="60">
Lazarocleistostoma
<taxonomicName id="BD8DDB9AED19D036FEDB6C72FE01FC95" box="[355,449,861,883]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="60" phylum="Arthropoda" rank="species" species="dentatum">dentatum</taxonomicName>
</emphasis>
<treatmentCitation id="FB2C8608ED19D036FE636C72FD53FC95" author="Stevcic" box="[475,659,861,883]" page="136" pageId="2" pageNumber="60" year="2011">
<bibRefCitation id="1E1CDDE8ED19D036FE636C72FDA0FC94" author="Stevcic" box="[475,608,861,883]" pageId="2" pageNumber="60" refString="Stevcic, Z. (2011) Addition to the reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Part I. New taxa. Natura Croatica (Fauna Croatica), 20 (1), 125 - 139." type="journal article" year="2011">Števčić 2011</bibRefCitation>
: 136
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED19D036FF2F6CB0FB79FC3A" pageId="2" pageNumber="60" type="materials_examined">
<paragraph id="7A32A019ED19D036FF2F6CB0FB79FC3A" blockId="2.[151,1437,927,1996]" pageId="2" pageNumber="60">
<emphasis id="48F97C0BED19D036FF2F6CB0FE40FC5E" bold="true" box="[151,384,927,952]" pageId="2" pageNumber="60">Material examined.</emphasis>
<typeStatus id="A5361EBBED19D036FE3F6CB0FE2FFC51" box="[391,495,927,951]" pageId="2" pageNumber="60" type="holotype">Holotype</typeStatus>
ovigerous female (6.2 × 3.3 mm) (
<collectionCode id="1C9C38DCED19D036FC346CB0FC0CFC51" box="[908,972,927,951]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="2" pageNumber="60">ZMA</collectionCode>
De.102.997), Saleyer Island, south Celebes (Sulawesi),
<collectingCountry id="029AE089ED19D036FEF16CEBFE79FC3A" box="[329,441,964,988]" name="Indonesia" pageId="2" pageNumber="60">Indonesia</collectingCountry>
,
<quantity id="BD750DFCED19D036FE7C6CEBFDDBFC3A" box="[452,539,964,988]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="3.4" metricValueMin="0.9" pageId="2" pageNumber="60" unit="m" value="21.5" valueMax="34.0" valueMin="9.0">934 m</quantity>
depth, coll. M. Webber, Siboga Expedition,
<date id="0E3386D9ED19D036FBB56CEBFB75FC3A" box="[1037,1205,964,989]" pageId="2" pageNumber="60" value="1899-05-07" valueMax="1899-05-08" valueMin="1899-05-07">78 May 1899</date>
.
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED19D036FF716CC8FC85F8DA" pageId="2" pageNumber="60" type="description">
<paragraph id="7A32A019ED19D036FF716CC8FB07FAA2" blockId="2.[151,1437,927,1996]" pageId="2" pageNumber="60">
<emphasis id="48F97C0BED19D036FF716CC8FEB6FBE6" bold="true" box="[201,374,999,1024]" pageId="2" pageNumber="60">Redescription.</emphasis>
Carapace transversely hexagonal, dorsal surface almost smooth, regions discernible but not prominent; cardiac region convex, prominent (
<figureCitation id="E2B6BC9CED19D036FD276B23FD29FBC2" box="[671,745,1036,1060]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Figs. 1</figureCitation>
A, D). Front narrow, entire, with shallow concave indentation medially; lateral margins angular, sharply separated from sinuous supraorbital margin; low epigastric lobes discernible (
<figureCitation id="E2B6BC9CED19D036FEB86B7BFEFEFB8A" box="[256,318,1108,1132]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, B, D). External orbital tooth prominent, directly obliquely anteriorly; anterolateral margin deeply concave with distinct tooth demarcating antero-, posterolateral junctions; posterolateral margin short, convex (
<figureCitation id="E2B6BC9CED19D036FAD76B58FF61FB52" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, D). Posterior carapace margin almost straight (
<figureCitation id="E2B6BC9CED19D036FD636BB3FCDBFB52" box="[731,795,1180,1204]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, D). Eyes prominent, entirely filling orbit; cornea distinctly pigmented; peduncle long, slender (
<figureCitation id="E2B6BC9CED19D036FD3F6B90FD09FB31" box="[647,713,1215,1239]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, D). Antennules folding obliquely (
<figureCitation id="E2B6BC9CED19D036FB3A6B90FB05FB31" box="[1154,1221,1215,1239]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
E). Antenna short, basal articles not fused to carapace; flagellum very short (
<figureCitation id="E2B6BC9CED19D036FC906BCBFCA8FB1A" box="[808,872,1252,1276]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
E). Suborbital, subhepatic, sub-branchial surfaces relatively smooth; suborbital margin crenulated to dentate on outer two-thirds (
<figureCitation id="E2B6BC9CED19D036FBA66A27FB9FFAC6" box="[1054,1119,1288,1312]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
E). Epistome longitudinally narrow; posterior margin with triangular median lobe, lateral margins deeply concave (
<figureCitation id="E2B6BC9CED19D036FBDC6A03FB63FAA2" box="[1124,1187,1324,1348]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
E).
</paragraph>
<paragraph id="7A32A019ED19D036FF716A7FFC05FA32" blockId="2.[151,1437,927,1996]" pageId="2" pageNumber="60">
Third maxillipeds operculiform, almost completely filling buccal cavity (
<figureCitation id="E2B6BC9CED19D036FBB36A7FFB8CFA8E" box="[1035,1100,1360,1384]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
C, F). Palp short, carpus longest, partially hidden under merus (
<figureCitation id="E2B6BC9CED19D036FD9C6A5BFDA4FA6A" box="[548,612,1396,1420]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
C, F). Merus subtriangular with margins rounded, without prominent sulcus (
<figureCitation id="E2B6BC9CED19D036FF756AB7FECFFA56" box="[205,271,1432,1456]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
C, F). Ischium rectangular, symmetrical (
<figureCitation id="E2B6BC9CED19D036FD516AB7FCEAFA56" box="[745,810,1432,1456]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
C, F). Exopod slender, partially hidden under ischium, reaching just beyond distal part of merus, flagellum distinct (
<figureCitation id="E2B6BC9CED19D036FCFA6A93FC41FA32" box="[834,897,1468,1492]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
C, F).
</paragraph>
<paragraph id="7A32A019ED19D036FF716ACFFF2CF982" blockId="2.[151,1437,927,1996]" pageId="2" pageNumber="60">
Cheliped very slender, short, symmetrical (
<figureCitation id="E2B6BC9CED19D036FD7E6ACFFCCAFA1E" box="[710,778,1504,1528]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, G). Merus subcylindrical, surface minutely granular, unarmed (
<figureCitation id="E2B6BC9CED19D036FEB0692BFE88F9FA" box="[264,328,1540,1564]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A). Carpus elongated, surface minutely granular, unarmed (
<figureCitation id="E2B6BC9CED19D036FC5D692BFBE4F9FA" box="[997,1060,1540,1564]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A). Chela slender, scattered short setae on inner subventral margin; fingers almost as long as palm, cutting edges unarmed with tufts of setae (
<figureCitation id="E2B6BC9CED19D036FAD76907FF61F982" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, G).
</paragraph>
<paragraph id="7A32A019ED19D036FF71695FFC4AF936" blockId="2.[151,1437,927,1996]" pageId="2" pageNumber="60">
Ambulatory legs relatively long, third leg longest (
<figureCitation id="E2B6BC9CED19D036FCAB695FFC96F96E" box="[787,854,1648,1672]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
). Merus laterally flattened; unarmed (
<figureCitation id="E2B6BC9CED19D036FAB6695FFA8FF96E" box="[1294,1359,1648,1672]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, H). Propodus laterally flattened with long setae on both margins (
<figureCitation id="E2B6BC9CED19D036FCEE69BBFC56F94A" box="[854,918,1684,1708]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, H). Dactyli of first to third legs falciform; that of last leg spatuliform, lined with numerous setae (
<figureCitation id="E2B6BC9CED19D036FCB86997FC80F936" box="[768,832,1720,1744]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
A, H).
</paragraph>
<paragraph id="7A32A019ED19D036FF7169F3FD26F8FE" blockId="2.[151,1437,927,1996]" pageId="2" pageNumber="60">
Thoracic sternum badly damaged, detailed structures not discernible. Abdomen rounded, covering most of thoracic sternum; all somites freely articulating (
<figureCitation id="E2B6BC9CED19D036FD2C682FFD15F8FE" box="[660,725,1792,1816]" captionStart="FIGURE 1" captionStartId="3.[151,250,1836,1859]" captionTargetBox="[168,1421,218,1794]" captionTargetId="figure@3.[151,1436,195,1815]" captionTargetPageId="3" captionText="FIGURE 1. Lazarocleistostoma dentatum (Tesch, 1918). Holotype female (6.2 × 3.3 mm) (ZMA De. 102.997), Saleyer Island. A, overall dorsal view; B, frontal view of frontal margin; C, buccal cavity showing third maxillipeds; D, dorsal view of carapace; E, frontal view of carapace; F, right third maxilliped; G, outer view of right chela; H, right last ambulatory leg. (A-C, after Tesch 1918: 63, pl. 3 fig. 2; other drawings by C. G. S. Tan). Scales = 1.0 mm." httpUri="https://zenodo.org/record/208600/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="7A32A019ED19D036FF71680CFC85F8DA" blockId="2.[151,1437,927,1996]" box="[201,837,1827,1852]" pageId="2" pageNumber="60">
<emphasis id="48F97C0BED19D036FF71680CFEE1F8DA" bold="true" box="[201,289,1827,1852]" pageId="2" pageNumber="60">Colour.</emphasis>
“Uniform ivory-white colour” (
<bibRefCitation id="1E1CDDE8ED19D036FD30680BFCF8F8DA" author="Tesch" box="[648,824,1828,1852]" pageId="2" pageNumber="60" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga Expedition. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls. 1 - 6." type="journal article" year="1918">Tesch 1918: 65</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED19D036FF716867FC79F886" box="[201,953,1863,1888]" pageId="2" pageNumber="60" type="materials_examined">
<paragraph id="7A32A019ED19D036FF716867FC79F886" blockId="2.[151,1437,927,1996]" box="[201,953,1863,1888]" pageId="2" pageNumber="60">
<emphasis id="48F97C0BED19D036FF716867FEA5F886" bold="true" box="[201,357,1863,1888]" pageId="2" pageNumber="60">
<typeStatus id="A5361EBBED19D036FF716867FEC2F886" box="[201,258,1864,1888]" pageId="2" pageNumber="60">Type</typeStatus>
locality.
</emphasis>
Saleyer Island, south Celebes (Sulawesi),
<collectingCountry id="029AE089ED19D036FCFC6867FC75F886" box="[836,949,1864,1888]" name="Indonesia" pageId="2" pageNumber="60">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3297F392ED19D036FF716843FEECF82A" pageId="2" pageNumber="60" type="discussion">
<paragraph id="7A32A019ED19D036FF716843FEECF82A" blockId="2.[151,1437,927,1996]" pageId="2" pageNumber="60">
<emphasis id="48F97C0BED19D036FF716843FEFDF862" bold="true" box="[201,317,1900,1924]" pageId="2" pageNumber="60">Remarks.</emphasis>
The
<typeStatus id="A5361EBBED19D036FEC26842FE6AF863" box="[378,426,1901,1925]" pageId="2" pageNumber="60">type</typeStatus>
specimen is in poor condition and most of its ventral structures are not discernible. The legs are also detached. Fortunately, Teschs (1918) description is detailed and the available structures confirm his observations.
</paragraph>
</subSubSection>
</treatment>
</document>