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<mods:title id="14AD24112398A3280FA6FAB8C206C732">Euscorpius thracicus sp n (Scorpiones: Euscorpiidae) from Bulgaria</mods:title>
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<taxonomicName id="A6B54D36DD3BE3111F2746740512FCFB" ID-CoL="4H4X" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[303,579,846,870]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD3BE3111F27467406FAFCFB" bold="true" box="[303,427,846,870]" italics="true" pageId="4" pageNumber="3">Euscorpius</emphasis>
Thorell, 1876
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(
<figureCitation id="F98E2A30DD3BE3111F3946540690FC1B" box="[305,449,878,902]" captionStart-0="Figures 12" captionStart-1="Figures 36" captionStart-10="Figure 61" captionStart-2="Figures 716" captionStart-3="Figures 1722" captionStart-4="Figures 2331" captionStart-5="Figures 3239" captionStart-6="Figures 4043" captionStart-7="Figures 4449" captionStart-8="Figures5058" captionStart-9="Figures 5960" captionStartId-0="3.[113,190,2016,2038]" captionStartId-1="6.[115,192,1981,2003]" captionStartId-10="15.[116,184,886,908]" captionStartId-2="7.[115,192,1958,1980]" captionStartId-3="8.[116,193,1988,2010]" captionStartId-4="9.[115,192,1979,2001]" captionStartId-5="10.[113,190,1958,1980]" captionStartId-6="11.[116,193,1198,1220]" captionStartId-7="12.[115,192,1991,2013]" captionStartId-8="13.[115,192,1982,2004]" captionStartId-9="14.[116,193,2015,2037]" captionTargetBox-0="[113,1474,234,2005]" captionTargetBox-1="[138,1430,274,1952]" captionTargetBox-10="[113,1474,234,877]" captionTargetBox-2="[113,1474,248,1887]" captionTargetBox-3="[127,1464,262,1953]" captionTargetBox-4="[129,1472,285,1928]" captionTargetBox-5="[125,1441,250,1915]" captionTargetBox-6="[113,1474,274,1183]" captionTargetBox-7="[151,1429,222,1983]" captionTargetBox-8="[115,1472,214,1975]" captionTargetBox-9="[113,1474,268,2003]" captionTargetId-0="figure-27@3.[113,1474,234,2007]" captionTargetId-1="figure-59@6.[116,1477,257,1971]" captionTargetId-10="figure-360@15.[113,1474,234,880]" captionTargetId-2="figure-74@7.[113,1474,248,1911]" captionTargetId-3="figure-69@8.[113,1474,234,1973]" captionTargetId-4="figure-54@9.[115,1476,272,1960]" captionTargetId-5="figure-66@10.[113,1474,244,1951]" captionTargetId-6="figure-530@11.[113,1474,234,1184]" captionTargetId-7="figure-80@12.[113,1474,214,1991]" captionTargetId-8="figure-136@13.[113,1474,211,1977]" captionTargetId-9="figure-35@14.[113,1474,232,2006]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-10="15" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="14" captionText-0="Figures 12. E. thracicus sp. n., male (1) and female (2) paratypes in vivo habitus." captionText-1="Figures 36: E. thracicus sp. n. Figures 34. Male holotype in dorsal (3) and ventral (4) views. Figures 56. Female paratype in dorsal (5) and ventral (6) views. Scale bar: 10 mm." captionText-10="Figure 61. Map of distribution of Euscorpius populations analysed by DNA (see Table 2)." captionText-2="Figures 716: E. thracicus sp. n. Figures 7, 9, 1115. Male holotype, carapace and tergites IIV (7), coxosternal area (9), left legs IIV, retrolateral aspect (1114 respectively), and telson lateral (15). Figures 8, 10, 16. Female paratype, carapace and tergites IIII (8), coxosternal area and sternites IIIIV (10), and telson lateral (16)." captionText-3="Figures 1722: E. thracicus sp. n. Figures 1719. Female paratype, metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Figures 2022. Male holotype, metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Scale bars: 10 mm (1719, 2022)." captionText-4="Figures 2331. E. thracicus sp. n., male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable finger dentition (31)." captionText-5="Figures 3239. E. thracicus sp. n., female paratype, pedipalp segments. Chela dorsal (32), external (33) and ventral (34) views. Patella dorsal (35), external (36) and ventral (37) views. Trochanter and femur dorsal (38) and ventral (39) views. Trichobothrial pattern is indicated by white circles (3238)." captionText-6="Figures 4043. Chromosomes of E. thracicus sp. n., karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm." captionText-7="Figures 4449. Euscorpius thracicus sp. n., left hemispermatophore. Figures 4445. Whole hemispermatophore in convex (44) and concave (45) views. Figures 4649. Capsule of left hemispermatophore, 3-dimensional form displayed in cross stereoscopic pairs, in posterior-convex (46), anterior (47), convex (48) and concave (49) views. Paratype No. 1864. Scale bar: 1 mm (Figs. 4449)." captionText-8="Figures5058. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 5058." captionText-9="Figures 5960. E. thracicus sp. n., type locality, Bulgaria, Krumovgrad, Bryagovets, 41.6485862°N 25.8132185°E." figureDoi-0="http://doi.org/10.5281/zenodo.4648926" figureDoi-1="http://doi.org/10.5281/zenodo.4648928" figureDoi-10="http://doi.org/10.5281/zenodo.4648946" figureDoi-2="http://doi.org/10.5281/zenodo.4648930" figureDoi-3="http://doi.org/10.5281/zenodo.4648932" figureDoi-4="http://doi.org/10.5281/zenodo.4648934" figureDoi-5="http://doi.org/10.5281/zenodo.4648936" figureDoi-6="http://doi.org/10.5281/zenodo.4648938" figureDoi-7="http://doi.org/10.5281/zenodo.4648940" figureDoi-8="http://doi.org/10.5281/zenodo.4648942" figureDoi-9="http://doi.org/10.5281/zenodo.4648944" httpUri-0="https://zenodo.org/record/4648926/files/figure.png" httpUri-1="https://zenodo.org/record/4648928/files/figure.png" httpUri-10="https://zenodo.org/record/4648946/files/figure.png" httpUri-2="https://zenodo.org/record/4648930/files/figure.png" httpUri-3="https://zenodo.org/record/4648932/files/figure.png" httpUri-4="https://zenodo.org/record/4648934/files/figure.png" httpUri-5="https://zenodo.org/record/4648936/files/figure.png" httpUri-6="https://zenodo.org/record/4648938/files/figure.png" httpUri-7="https://zenodo.org/record/4648940/files/figure.png" httpUri-8="https://zenodo.org/record/4648942/files/figure.png" httpUri-9="https://zenodo.org/record/4648944/files/figure.png" pageId="4" pageNumber="3">Figures 161</figureCitation>
,
<tableCitation id="2C37030EDD3BE3111FC346540511FC1B" box="[459,576,878,902]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="5.[115,171,993,1014]" captionStartId-1="15.[112,168,1430,1451]" captionTargetBox-1="[117,1471,936,1402]" captionTargetId-0="graphics-542@5.[113,1483,237,975]" captionTargetId-1="graphics-361@15.[111,1476,929,1407]" captionTargetPageId-0="5" captionTargetPageId-1="15" captionText-0="Table 1. Comparative measurements of adults of Euscorpius thracicus sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 2. A matrix of genetic distances among the COI genes of the Euscorpius species from Bulgaria, eastern Serbia, and northern Greece. p-distance (below the diagonal); Kimura 2-parameter distance (above the diagonal). In the brackets are mentioned specimen codes/accession number from GenBank." httpUri-0="http://table.plazi.org/id/35CA663DDD3AE3101E7B46DB055EFB8E" httpUri-1="http://table.plazi.org/id/35CA663DDD30E31A1E7840AC0607FA7B" pageId="4" pageNumber="3" tableUuid-0="35CA663DDD3AE3101E7B46DB055EFB8E" tableUuid-1="35CA663DDD30E31A1E7840AC0607FA7B">Tables 12</tableCitation>
)
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<emphasis id="53C1EAA7DD3BE3111E7F46B4052AFC5B" bold="true" pageId="4" pageNumber="3">http://zoobank.org/urn:lsid:zoobank.org:act:CEC71123- 0F84-45DD-857E-2D4702C2FD80</emphasis>
</paragraph>
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<paragraph id="610A36B5DD3BE3111E7846D105C9FBDE" blockId="4.[112,770,1002,1093]" pageId="4" pageNumber="3">
<taxonomicName id="A6B54D36DD3BE3111E7846D106FEFB9F" ID-CoL="4H4X" authority="Thorell, 1876: 15" authorityName="Thorell" authorityPageNumber="15" authorityYear="1876" box="[112,431,1002,1027]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD3BE3111E7846D107B9FB9F" box="[112,232,1003,1026]" italics="true" pageId="4" pageNumber="3">Euscorpius</emphasis>
Thorell, 1876: 15
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; Fet &amp; Sissom, 2000: 357377 (in part, complete reference list until 1998); KovařÍk &amp; Šťáhlavský, 2020: 137, figs. 1186, tabs. 14.
</paragraph>
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<paragraph id="610A36B5DD3BE3111E78415205D8FBE2" blockId="4.[112,649,1127,1152]" box="[112,649,1127,1152]" pageId="4" pageNumber="3">
<materialsCitation id="D1DD3CE8DD3BE3111E78415205D8FBE2" ID-GBIF-Occurrence="3114128307" box="[112,649,1127,1152]" collectionCode="SPECIES" pageId="4" pageNumber="3" specimenCount="1" typeStatus="holotype">
<typeStatus id="BE0E8817DD3BE3111E78415207F7FBE2" box="[112,166,1128,1151]" pageId="4" pageNumber="3">TYPE</typeStatus>
<collectionCode id="07A4AE70DD3BE3111EA5415207AFFB1D" box="[173,254,1128,1152]" pageId="4" pageNumber="3">SPECIES</collectionCode>
.
<taxonomicName id="A6B54D36DD3BE3111F0F415205D8FBE2" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[263,649,1127,1151]" class="Arachnida" family="Scorpionidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="carpathicus">
<emphasis id="53C1EAA7DD3BE3111F0F4152068EFBE2" box="[263,479,1127,1151]" italics="true" pageId="4" pageNumber="3">Scorpio carpathicus</emphasis>
Linnaeus, 1767
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<subSubSection id="29AF653EDD3BE3111E78419E06A2F981" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="610A36B5DD3BE3111E78419E06A2F981" blockId="4.[112,771,1188,1564]" pageId="4" pageNumber="3">
DIAGNOSIS. Total length
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. Movable fingers of pedipalp chela with MD aligned in a single straight row along entire length of chela fingers; ID, IAD (
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number) and OD present. OD are displaced to outer aspect of fingers. Pedipalp chela flat in appearance. Trichobothrial pattern
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C. Two subdistal denticles present on cheliceral movable finger dorsal edge. Ventral edge of cheliceral movable finger smooth; serrula absent. Two pedal spurs present on legs. Tarsal spurs on legs absent. Sternum pentagonal in shape. Hemispermatophore lamelliform in shape with broad trunk and distal lamina. Telson without subaculear tubercle. Metasoma IIV with single median carina usually absent.
</paragraph>
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</treatment>
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