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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.545.6470" ID-GBIF-Dataset="52de69a6-afe1-447f-9ace-cbc6ab610abf" ID-PMC="PMC4714369" ID-Pensoft-Pub="1313-2970-545-89" ID-PubMed="26798296" ID-ZBK="61C2313F341741698E3559D134F884E0" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-545-89" ModsDocOrigin="ZooKeys 545" ModsDocTitle="Two new fossil genera and species of Cerocephalinae (Hymenoptera, Chalcidoidea, Pteromalidae), including the first record from the Eocene" checkinTime="1451243628961" checkinUser="pensoft" docAuthor="Blaeser, Marcel, Krogmann, Lars &amp; Peters, Ralph S." docDate="2015" docId="BF06359B59BF50D47F5769B41694165C" docLanguage="en" docName="ZooKeys 545: 89-100" docOrigin="ZooKeys 545" docSource="http://dx.doi.org/10.3897/zookeys.545.6470" docTitle="Tenuicornus dominicus Blaeser, Krogmann &amp; Peters, 2015, sp. n." docType="treatment" docUuid="8968FC81-D138-43FB-AA79-CD15E283D82C" docUuidSource="ZooBank" docVersion="6" lastPageNumber="96" masterDocId="B67CFFF5FFF0B032DD50FFE4FFF76B6C" masterDocTitle="Two new fossil genera and species of Cerocephalinae (Hymenoptera, Chalcidoidea, Pteromalidae), including the first record from the Eocene" masterLastPageNumber="100" masterPageNumber="89" pageNumber="95" updateTime="1668162325483" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new fossil genera and species of Cerocephalinae (Hymenoptera, Chalcidoidea, Pteromalidae), including the first record from the Eocene</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Blaeser, Marcel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krogmann, Lars</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peters, Ralph S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>545</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>89</mods:start>
<mods:end>100</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.545.6470</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.545.6470</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-545-89</mods:identifier>
<mods:identifier type="ZBK">61C2313F341741698E3559D134F884E0</mods:identifier>
<mods:identifier type="ZooBank">61C2313F341741698E3559D134F884E0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127897705" LSID="urn:lsid:zoobank.org:act:8968FC81-D138-43FB-AA79-CD15E283D82C" httpUri="http://treatment.plazi.org/id/BF06359B59BF50D47F5769B41694165C" lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95">
<subSubSection pageId="6" pageNumber="95" type="multiple">
<paragraph pageId="6" pageNumber="95">Taxon classification Animalia Hymenoptera Pteromalidae</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="nomenclature">
<paragraph pageId="6" pageNumber="95">
<taxonomicName LSID="http://zoobank.org/8968FC81-D138-43FB-AA79-CD15E283D82C" class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus dominicus" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="dominicus">Tenuicornus dominicus</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="95">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="diagnosis">
<paragraph pageId="6" pageNumber="95">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="95">See genus.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="description">
<paragraph pageId="6" pageNumber="95">Description.</paragraph>
<paragraph pageId="6" pageNumber="95">
Female: total body length 2.07 mm; length of mesosoma 0.92 mm, of metasoma 1.02 mm. Body without metallic luster. Head: height 0.39 mm, width 0.38 mm. Face deeply impressed. Shape of head cuboid; mandibles not visible, hidden in facial depression. Margin of facial depression with rough surface and with a thickened seta on each side (length of setae 85
<normalizedToken originalValue="µm">µm</normalizedToken>
). Inner antennal process (iap) positioned distinctly above level of toruli; nail-like, 0.11 mm long, with basis wider than apex, orientated anteroventrally. Eyes large and egg-shaped, 0.25 mm high and 0.2 mm wide. Antennae: inserted at ventral margin of eyes; scape long and slightly curved, length 0.25 mm; pedicel length 75
<normalizedToken originalValue="µm">µm</normalizedToken>
, rather short and stout; funicle 6-segmented with first two funicular segments longer than wide (F1: length: 71
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
width: 52
<normalizedToken originalValue="µm">µm</normalizedToken>
; F2: 68
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
60
<normalizedToken originalValue="µm">µm</normalizedToken>
) and F3-F6 wider than long, distinctly broadening distally, club-like (F3: 59
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
72
<normalizedToken originalValue="µm">µm</normalizedToken>
; F4: 60
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
74
<normalizedToken originalValue="µm">µm</normalizedToken>
; F5: 58
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
80
<normalizedToken originalValue="µm">µm</normalizedToken>
; F6: 52
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
86
<normalizedToken originalValue="µm">µm</normalizedToken>
). Clava egg-shaped: 97
<normalizedToken originalValue="µm">µm</normalizedToken>
long and 77
<normalizedToken originalValue="µm">µm</normalizedToken>
wide. Wings: fore wing long and slender (length 1.53 mm and width 0.55 mm); submarginal vein 0.57 mm long, marginal vein 0.38 mm long, postmarginal vein 0.11 mm long, stigmal vein 99
<normalizedToken originalValue="µm">µm</normalizedToken>
long; stigma slightly thickened and uncus visible. Wing disc bare, setal bases absent. Elongate admarginal setae present. Hind wing length 1.27 mm and width 0.27 mm, three hamuli present. Mesosoma: pronotum and anterior part of mesoscutum with slight traces of strigulate surface sculpture; mesonotum otherwise polished. Notauli foveolate. Length of pronotum 0.37 mm, length of mesoscutum 0.22 mm, length of mesoscutellum 0.22 mm; no frenal line. Axillae medially connected. Mesopleuron (in lateral view) height 0.29 mm, width 0.36 mm. Prepectus enlarged and triangular in shape (0.14 mm
<normalizedToken originalValue="×">x</normalizedToken>
0.17 mm). Length of metanotum 39
<normalizedToken originalValue="µm">µm</normalizedToken>
; length of propodeum 72
<normalizedToken originalValue="µm">µm</normalizedToken>
. Propodeum without a median carina or plicae. Legs: coxae strong and stout. Femora and tibiae long and slender (Fe1: 0.45 mm; Fe2: 0.32 mm; Fe3: 0.41 mm; Ti1: 0.39 mm; Ti2: 0.49 mm; Ti3: 0.42 mm). Metasoma: petiole (Mt1) short and transverse, hardly visible. Metasomal tergites Mt2 to Mt9 smooth, Mt2: 0.28 mm, Mt3: 0.17 mm, Mt4: 0.15 mm, Mt5: 0.16 mm, Mt6: 0.11 mm, Mt7 60
<normalizedToken originalValue="µm">µm</normalizedToken>
, Mt8/9: 71
<normalizedToken originalValue="µm">µm</normalizedToken>
. Ovipositor 0.23 mm (top view) exerted beyond end of gaster.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95" type="taxonomic remarks">
<paragraph pageId="6" pageNumber="95">Taxonomic remarks.</paragraph>
<paragraph lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95">
The two genera that are most similar to
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
are
<taxonomicName class="Insecta" family="Pteromalidae" genus="Neocalosoter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocalosoter" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Neocalosoter</taxonomicName>
and
<taxonomicName class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
.
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
runs to
<taxonomicName class="Insecta" family="Pteromalidae" genus="Neocalosoter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocalosoter" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Neocalosoter</taxonomicName>
in the key of
<bibRefCitation author="Krogmann, L" journalOrPublisher="Historical Biology" pageId="10" pageNumber="99" pagination="43 - 49" title="First fossil record of cerocephaline wasps with a description of new genus and species from Dominican amber (Hymenoptera: Chalcidoidea: Pteromalidae: Cerocephalinae)." url="10.1080/08912963.2012.685729" volume="25" year="2013">Krogmann (2013)</bibRefCitation>
but differs from this genus in various features:
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
possesses a head that is deeply impressed (convex in
<taxonomicName class="Insecta" family="Pteromalidae" genus="Neocalosoter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocalosoter" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Neocalosoter</taxonomicName>
), the dark transverse bands on
<pageBreakToken pageId="7" pageNumber="96" start="start">its</pageBreakToken>
fore wing are absent, and the propodeum has no traces of a median carina or plicae (Fig. 2B). The tentative similarity between
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
and
<taxonomicName class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
is based on the deeply impressed face. However,
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
differs from
<taxonomicName class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
by the absence of a tuft of setae on the parastigma on the fore wings (Fig. 2E), one of the most distinct characters for the generic classification of
<taxonomicName lsidName="" pageId="7" pageNumber="96" rank="subFamily" subFamily="Cerocephalinae">Cerocephalinae</taxonomicName>
.
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
further differs from
<taxonomicName class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
by the shape and position of the inner antennal process. In
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
, the process is shaped like a nail and positioned above the level of the toruli at the upper margin of the facial depression (Fig. 2C). In
<taxonomicName class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
, the process is broader and stouter and positioned at or slightly below the level of the toruli. Furthermore,
<taxonomicName class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
is described to have all funicular segments longer than wide (
<bibRefCitation author="Westwood, JO" journalOrPublisher="Philosophical Magazine" pageId="10" pageNumber="99" title="Descriptions of several new British forms amongst the parasitic hymenopterous insects." volume="1" year="1832">Westwood 1832</bibRefCitation>
), while in
<taxonomicName class="Insecta" family="Pteromalidae" genus="Tenuicornus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tenuicornus" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Tenuicornus</taxonomicName>
only the first two funicular segments are longer than wide. The funicular segments three to six are wider than long (Fig. 2D).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="96" type="etymology">
<paragraph pageId="7" pageNumber="96">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="96">The name dominicus is derived from the amber deposit in which the fossil was found.</paragraph>
</subSubSection>
</treatment>
</document>