treatments-xml/data/65/75/0D/65750D68FFE6FF93FF2B3082FD097690.xml

<document id="B6E894F6E2EF0DC3ED7BD56AAAF1EDA8" ID-CLB-Dataset="4054" ID-DOI="10.11646/zootaxa.5048.3.5" ID-GBIF-Dataset="8a115b28-99ef-4210-8729-c7cc6afad14d" ID-ISSN="1175-5326" ID-Zenodo-Dep="5556467" ID-ZooBank="E468FA7C-0A49-4BA2-A98C-9A9FE6C5D873" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633692456878" checkinUser="plazi" docAuthor="Joharchi, Omid &amp; Halliday, Bruce" docDate="2021" docId="65750D68FFE6FF93FF2B3082FD097690" docLanguage="en" docName="zootaxa.5048.3.5.pdf" docOrigin="Zootaxa 5048 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hyposternus ceylonicus Joharchi &amp; Halliday 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="402" masterDocId="994C7510FFE4FF98FFBC3015FF837439" masterDocTitle="A new genus and species of Laelapidae Canestrini from Sri Lanka (Acari: Mesostigmata)" masterLastPageNumber="406" masterPageNumber="391" pageNumber="393" updateTime="1699459111764" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9C08A8D9A5E469AD3A2637922513B6F6">A new genus and species of Laelapidae Canestrini from Sri Lanka (Acari: Mesostigmata)</mods:title>
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<mods:namePart id="7E5556BB4C48282A5F6456511B86B82D">Joharchi, Omid</mods:namePart>
<mods:affiliation id="E2E5BC83A24D80EBEF4A7B913E2DC546">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="D0E4892CF9BD157D28BD684E4699A85C">Halliday, Bruce</mods:namePart>
<mods:affiliation id="696C92F65AE54EA95E91BD3EB924DBC2">Australian National Insect Collection, CSIRO, GPO Box 1700, Canberra ACT 2601, Australia.</mods:affiliation>
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<mods:date id="DA29EC5A3B505BB468ACABFF38E019F3">2021</mods:date>
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<treatment id="65750D68FFE6FF93FF2B3082FD097690" ID-DOI="http://doi.org/10.5281/zenodo.5556465" ID-GBIF-Taxon="188638646" ID-Zenodo-Dep="5556465" LSID="urn:lsid:plazi:treatment:65750D68FFE6FF93FF2B3082FD097690" httpUri="http://treatment.plazi.org/id/65750D68FFE6FF93FF2B3082FD097690" lastPageId="11" lastPageNumber="402" pageId="2" pageNumber="393">
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<heading id="B62B0B12FFE6FF9AFF2B3082FD9F7488" bold="true" box="[151,540,151,178]" fontSize="11" level="1" pageId="2" pageNumber="393" reason="1">
<emphasis id="DFA8606CFFE6FF9AFF2B3082FD9F7488" bold="true" box="[151,540,151,178]" pageId="2" pageNumber="393">
<taxonomicName id="2ADCC7FDFFE6FF9AFF2B3082FE34748B" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[151,439,151,178]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="ceylonicus" status="sp. nov.">
<emphasis id="DFA8606CFFE6FF9AFF2B3082FE34748B" bold="true" box="[151,439,151,178]" italics="true" pageId="2" pageNumber="393">Hyposternus ceylonicus</emphasis>
</taxonomicName>
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</emphasis>
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<subSubSection id="A5C6EFF5FFE6FF9AFF2B30AEFEBF74EC" box="[151,316,187,213]" pageId="2" pageNumber="393" type="description">
<paragraph id="ED63BC7EFFE6FF9AFF2B30AEFEBF74EC" blockId="2.[151,540,151,213]" box="[151,316,187,213]" pageId="2" pageNumber="393">
(
<figureCitation id="75E7A0FBFFE6FF9AFF2330AEFEB674EC" box="[159,309,187,213]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–11" captionStart-2="FIGURES 12–17" captionStart-3="FIGURES 18–21" captionStart-4="FIGURES 22–23" captionStart-5="FIGURES 24–29" captionStart-6="FIGURES 30–34" captionStartId-0="3.[151,264,1962,1986]" captionStartId-1="4.[151,264,1943,1967]" captionStartId-2="5.[151,264,1943,1967]" captionStartId-3="6.[151,264,1943,1967]" captionStartId-4="7.[151,264,1671,1695]" captionStartId-5="9.[151,264,1943,1967]" captionStartId-6="10.[151,264,1908,1932]" captionTargetBox-0="[211,1381,245,1927]" captionTargetBox-1="[179,1406,181,1920]" captionTargetBox-2="[184,1403,182,1918]" captionTargetBox-3="[174,1424,186,1915]" captionTargetBox-4="[169,1420,199,1632]" captionTargetBox-5="[192,1407,185,1914]" captionTargetBox-6="[164,1426,182,1885]" captionTargetId-0="figure-46@3.[192,1395,237,1939]" captionTargetId-1="figure-25@4.[179,1408,181,1920]" captionTargetId-2="figure-17@5.[184,1403,181,1919]" captionTargetId-3="figure-25@6.[151,1436,181,1919]" captionTargetId-4="figure-25@7.[151,1436,181,1648]" captionTargetId-5="figure-17@9.[164,1424,181,1919]" captionTargetId-6="figure-25@10.[161,1426,181,1885]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." captionText-1="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." captionText-2="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." captionText-3="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." captionText-4="FIGURES 22–23. Hyposternus ceylonicus gen. nov, sp. nov., male. 22, ventral idiosoma; 23, chelicera." captionText-5="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." captionText-6="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi-0="http://doi.org/10.5281/zenodo.5556471" figureDoi-1="http://doi.org/10.5281/zenodo.5556473" figureDoi-2="http://doi.org/10.5281/zenodo.5556475" figureDoi-3="http://doi.org/10.5281/zenodo.5556477" figureDoi-4="http://doi.org/10.5281/zenodo.5556479" figureDoi-5="http://doi.org/10.5281/zenodo.5556481" figureDoi-6="http://doi.org/10.5281/zenodo.5556483" httpUri-0="https://zenodo.org/record/5556471/files/figure.png" httpUri-1="https://zenodo.org/record/5556473/files/figure.png" httpUri-2="https://zenodo.org/record/5556475/files/figure.png" httpUri-3="https://zenodo.org/record/5556477/files/figure.png" httpUri-4="https://zenodo.org/record/5556479/files/figure.png" httpUri-5="https://zenodo.org/record/5556481/files/figure.png" httpUri-6="https://zenodo.org/record/5556483/files/figure.png" pageId="2" pageNumber="393">Figures 1–34</figureCitation>
)
</paragraph>
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<paragraph id="ED63BC7EFFE6FF9AFF2B3116FA92755F" blockId="2.[151,1437,259,2014]" pageId="2" pageNumber="393">
<emphasis id="DFA8606CFFE6FF9AFF2B3116FEC47524" bold="true" box="[151,327,259,285]" pageId="2" pageNumber="393">Type material:</emphasis>
<materialsCitation id="5DB4B623FFE6FF9AFEF33116FF7D755C" ID-GBIF-Occurrence="3383508301" collectingDate="2019-01-24" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="2" pageNumber="393" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
<typeStatus id="326702DCFFE6FF9AFEF33116FE397524" box="[335,442,259,285]" pageId="2" pageNumber="393" type="holotype">Holotype</typeStatus>
, female, 
<collectingCountry id="95CBFCEEFFE6FF9AFD9C3116FD127524" box="[544,657,259,285]" name="Sri Lanka" pageId="2" pageNumber="393">Sri Lanka</collectingCountry>
, 
<collectingRegion id="2F18729CFFE6FF9AFD203116FC377524" box="[668,948,259,285]" country="Sri Lanka" name="Sabaragamuwa" pageId="2" pageNumber="393">Sabaragamuwa Province</collectingRegion>
, 
<location id="E803EAA5FFE6FF9AFC7C3116FBCA7524" LSID="urn:lsid:plazi:treatment:65750D68FFE6FF93FF2B3082FD097690:E803EAA5FFE6FF9AFC7C3116FBCA7524" box="[960,1097,259,285]" country="Sri Lanka" latitude="6.45" longLatPrecision="1304" longitude="80.2" name="Polgampola" pageId="2" pageNumber="393" stateProvince="Sabaragamuwa">Polgampola</location>
, 
<geoCoordinate id="88E8DAB9FFE6FF9AFBE83116FB3E7527" box="[1108,1213,259,286]" degrees="06" direction="north" minutes="27" orientation="latitude" pageId="2" pageNumber="393" precision="925" value="6.45">06°27’ N</geoCoordinate>
, 
<geoCoordinate id="88E8DAB9FFE6FF9AFB743116FAAE7527" box="[1224,1325,259,286]" degrees="80" direction="east" minutes="12" orientation="longitude" pageId="2" pageNumber="393" precision="925" value="80.2">80°12’ E</geoCoordinate>
, alt. 
<quantity id="2A24119BFFE6FF9AFADF3116FA1B7524" box="[1379,1432,259,285]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="393" unit="m" value="42.0">
<elevation id="66F15B4DFFE6FF9AFADF3116FA1B7524" box="[1379,1432,259,285]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="393" unit="m" value="42.0">42m</elevation>
</quantity>
, 
<date id="99629ABEFFE6FF9AFF2B3132FED17578" box="[151,338,295,322]" pageId="2" pageNumber="393" value="2019-01-24">
<collectingDate id="89266356FFE6FF9AFF2B3132FED17578" box="[151,338,295,322]" pageId="2" pageNumber="393" value="2019-01-24">24 January 2019</collectingDate>
</date>
, 
<collectorName id="4029D9A8FFE6FF9AFEE13132FE637578" box="[349,480,295,321]" pageId="2" pageNumber="393">O. Joharchi</collectorName>
, 
<collectorName id="4029D9A8FFE6FF9AFE573132FDFC7578" box="[491,639,295,321]" pageId="2" pageNumber="393">S.G. Ermilov</collectorName>
&amp; 
<collectorName id="4029D9A8FFE6FF9AFD1C3132FCCB7578" box="[672,840,295,321]" pageId="2" pageNumber="393">A.A. Khaustov</collectorName>
coll., from forest soil-litter near the 
<collectorName id="4029D9A8FFE6FF9AFB613132FA1F7578" box="[1245,1436,295,321]" pageId="2" pageNumber="393">Thambadola Ella</collectorName>
waterfall
</materialsCitation>
. 
<materialsCitation id="5DB4B623FFE6FF9AFEB53159FA8E755F" ID-GBIF-Occurrence="3383508302" box="[265,1293,331,358]" collectingDate="2019-01-24" collectionCode="TUMZ" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="2" pageNumber="393" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sabaragamuwa" typeStatus="paratype">
<typeStatus id="326702DCFFE6FF9AFEB53159FEFA755F" box="[265,377,332,358]" pageId="2" pageNumber="393" type="paratype">Paratypes</typeStatus>
, 
<specimenCount id="FBDA77F7FFE6FF9AFE383159FD8C755C" box="[388,527,331,358]" pageId="2" pageNumber="393" type="female">two females</specimenCount>
, 
<specimenCount id="FBDA77F7FFE6FF9AFDA63159FDFC755C" box="[538,639,331,358]" pageId="2" pageNumber="393" type="male">one male</specimenCount>
and four protonymphs, same data as holotype (in 
<collectionCode id="8BCD24BBFFE6FF9AFB0C3159FA86755F" box="[1200,1285,332,358]" pageId="2" pageNumber="393">TUMZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="A5C6EFF5FFE6FF93FF7B317AFBF97594" lastPageId="11" lastPageNumber="402" pageId="2" pageNumber="393" type="description">
<paragraph id="ED63BC7EFFE6FF9AFF7B317AFCBC75B0" blockId="2.[151,1437,259,2014]" box="[199,831,367,393]" pageId="2" pageNumber="393">
<emphasis id="DFA8606CFFE6FF9AFF7B317AFE7A75B0" bold="true" box="[199,505,367,393]" pageId="2" pageNumber="393">Description. Adult female</emphasis>
(n=3) (
<figureCitation id="75E7A0FBFFE6FF9AFDF1317AFD6075B0" box="[589,739,367,393]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–11" captionStart-2="FIGURES 12–17" captionStartId-0="3.[151,264,1962,1986]" captionStartId-1="4.[151,264,1943,1967]" captionStartId-2="5.[151,264,1943,1967]" captionTargetBox-0="[211,1381,245,1927]" captionTargetBox-1="[179,1406,181,1920]" captionTargetBox-2="[184,1403,182,1918]" captionTargetId-0="figure-46@3.[192,1395,237,1939]" captionTargetId-1="figure-25@4.[179,1408,181,1920]" captionTargetId-2="figure-17@5.[184,1403,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." captionText-1="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." captionText-2="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5556471" figureDoi-1="http://doi.org/10.5281/zenodo.5556473" figureDoi-2="http://doi.org/10.5281/zenodo.5556475" httpUri-0="https://zenodo.org/record/5556471/files/figure.png" httpUri-1="https://zenodo.org/record/5556473/files/figure.png" httpUri-2="https://zenodo.org/record/5556475/files/figure.png" pageId="2" pageNumber="393">Figures 1–16</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFD4C317AFCB475B0" box="[752,823,367,393]" captionStart="FIGURES 18–21" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[174,1424,186,1915]" captionTargetId="figure-25@6.[151,1436,181,1919]" captionTargetPageId="6" captionText="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5556477" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="2" pageNumber="393">18–21</figureCitation>
)
</paragraph>
<paragraph id="ED63BC7EFFE6FF9AFF7B3186FDA57690" blockId="2.[151,1437,259,2014]" pageId="2" pageNumber="393">
<emphasis id="DFA8606CFFE6FF9AFF7B3186FE077594" box="[199,388,403,429]" italics="true" pageId="2" pageNumber="393">Dorsal idiosoma</emphasis>
(
<figureCitation id="75E7A0FBFFE6FF9AFE2F3186FE587594" box="[403,475,403,429]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 1</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFE553186FDA67594" box="[489,549,403,429]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">8, 10</figureCitation>
). Dorsal shield 683–721 long, 376–390 wide, covering entire dorsal idiosoma, oval-shaped, slightly extending onto ventrolateral aspect of idiosoma, faintly reticulated, with 38 pairs of setae, 21 pairs of podonotal setae, 
<emphasis id="DFA8606CFFE6FF9AFE0C31CEFE4B75CC" box="[432,456,475,501]" italics="true" pageId="2" pageNumber="393">z3</emphasis>
absent, 17 pairs of opisthonotal setae, including two pairs of 
<emphasis id="DFA8606CFFE6FF9AFBC431C9FB1775CC" box="[1144,1172,476,501]" italics="true" pageId="2" pageNumber="393">Zx</emphasis>
setae, a supernumerary seta 
<emphasis id="DFA8606CFFE6FF9AFF763215FF607620" box="[202,227,512,537]" italics="true" pageId="2" pageNumber="393">Jx</emphasis>
present between setae 
<emphasis id="DFA8606CFFE6FF9AFE5931EAFDE97620" box="[485,618,511,537]" italics="true" pageId="2" pageNumber="393">J2-J2-J3-J3</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFDC531EAFD127620" box="[633,657,511,537]" italics="true" pageId="2" pageNumber="393">r6</emphasis>
on lateral soft cuticle. All setae slender, smooth and pointed (22–27), much shorter than distance to following setal base, except 
<emphasis id="DFA8606CFFE6FF9AFC9E3236FCBB7604" box="[802,824,547,573]" italics="true" pageId="2" pageNumber="393">j1</emphasis>
(32–35) longer and thicker than other podonotal setae; 
<emphasis id="DFA8606CFFE6FF9AFF2B325DFF36765B" box="[151,181,584,610]" italics="true" pageId="2" pageNumber="393">Z5</emphasis>
(60–63) strongly thickened and elongated. Dorsal shield with about 23 pairs of discernible pore-like structures, including 16 poroids (
<emphasis id="DFA8606CFFE6FF9AFE24327EFE3F76BC" box="[408,444,619,645]" italics="true" pageId="2" pageNumber="393">id1</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFE71327EFE7376BC" box="[461,496,619,645]" italics="true" pageId="2" pageNumber="393">id2</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFDBD327EFD9476BC" box="[513,535,619,645]" italics="true" pageId="2" pageNumber="393">id</emphasis>
4– 
<emphasis id="DFA8606CFFE6FF9AFD8F327EFDD576BC" box="[563,598,619,645]" italics="true" pageId="2" pageNumber="393">id6</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFDD4327EFD1176BC" box="[616,658,619,645]" italics="true" pageId="2" pageNumber="393">idm</emphasis>
1– 
<emphasis id="DFA8606CFFE6FF9AFD12327EFD6676BC" box="[686,741,619,645]" italics="true" pageId="2" pageNumber="393">idm6</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFD4B327EFC9476BC" box="[759,791,619,645]" italics="true" pageId="2" pageNumber="393">is1</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFC94327EFCC976BC" box="[808,842,619,645]" italics="true" pageId="2" pageNumber="393">idx</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFCE7327EFC0476BC" box="[859,903,619,645]" italics="true" pageId="2" pageNumber="393">idl1</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFC24327EFC4076BC" box="[920,963,619,645]" italics="true" pageId="2" pageNumber="393">idl3</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFC68327EFC7C76BC" box="[980,1023,619,645]" italics="true" pageId="2" pageNumber="393">idl4</emphasis>
) and seven gland openings (
<emphasis id="DFA8606CFFE6FF9AFAF3327EFA1676BC" box="[1359,1429,619,646]" italics="true" pageId="2" pageNumber="393">gd1–2</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFF2B329AFF5E7690" box="[151,221,655,682]" italics="true" pageId="2" pageNumber="393">gd4–6</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFF57329AFECE7690" box="[235,333,655,681]" italics="true" pageId="2" pageNumber="393">gd8, gd9</emphasis>
) (
<figureCitation id="75E7A0FBFFE6FF9AFED9329AFE537690" box="[357,464,655,681]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figures 1</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFE62329AFD9A7690" box="[478,537,655,681]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">8, 10</figureCitation>
).
</paragraph>
<paragraph id="ED63BC7EFFE6FF9AFF7B32A6FCBD7268" blockId="2.[151,1437,259,2014]" pageId="2" pageNumber="393">
<emphasis id="DFA8606CFFE6FF9AFF7B32A6FE0476F4" box="[199,391,691,717]" italics="true" pageId="2" pageNumber="393">Ventral idiosoma</emphasis>
(
<figureCitation id="75E7A0FBFFE6FF9AFE2B32A6FE6376F4" box="[407,480,691,717]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 1</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFE5332A6FDAA76F4" box="[495,553,691,717]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">9, 11</figureCitation>
). Tritosternum with paired pilose laciniae (108–115), fused basally (4–6), columnar base 33–35 × 19–21 wide; pre-sternal platelets absent, apparently fused with sternal shield. Sternal shield length 180–189, maximum width 185–190, narrowest between coxae II (126–137), strongly desclerotised, represented by lateral and anterior margins, anterior margin undulating, median posterior margin absent, anterolateral edges fused to endopodal elements to form arms projecting between coxae I–II. Shield remnants capturing gland pores 
<emphasis id="DFA8606CFFE6FF9AFF603372FE8777B8" box="[220,260,871,897]" italics="true" pageId="2" pageNumber="393">gvb</emphasis>
; bearing three pairs of smooth setae (
<emphasis id="DFA8606CFFE6FF9AFD093372FD5577B8" box="[693,726,871,897]" italics="true" pageId="2" pageNumber="393">st1</emphasis>
58–60, 
<emphasis id="DFA8606CFFE6FF9AFC883372FCD777B8" box="[820,852,871,897]" italics="true" pageId="2" pageNumber="393">st2</emphasis>
52–54, 
<emphasis id="DFA8606CFFE6FF9AFC0F3372FC5777B8" box="[947,980,871,897]" italics="true" pageId="2" pageNumber="393">st3</emphasis>
47–49) on lateral remnants, setae short, not reaching base of next seta, one pair of reduced lyrifissures adjacent to setae 
<emphasis id="DFA8606CFFE6FF9AFB9D339EFBC1779C" box="[1057,1090,907,933]" italics="true" pageId="2" pageNumber="393">st1</emphasis>
, and a pair of reduced poroids between 
<emphasis id="DFA8606CFFE6FF9AFF4733BAFE9877F0" box="[251,283,943,969]" italics="true" pageId="2" pageNumber="393">st2</emphasis>
and 
<emphasis id="DFA8606CFFE6FF9AFEED33BAFEF177F0" box="[337,370,943,969]" italics="true" pageId="2" pageNumber="393">st3</emphasis>
; gland pores 
<emphasis id="DFA8606CFFE6FF9AFDB433BAFDB377F0" box="[520,560,943,969]" italics="true" pageId="2" pageNumber="393">gv1</emphasis>
apparently absent (
<figureCitation id="75E7A0FBFFE6FF9AFCB533BAFCCC77F0" box="[777,847,943,969]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 1</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFCE133BAFCEE77F0" box="[861,877,943,969]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">9</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFCC433BAFC1477F0" box="[888,919,943,969]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="2" pageNumber="393">12</figureCitation>
). Metasternal setae 
<emphasis id="DFA8606CFFE6FF9AFBC833BAFB1677F0" box="[1140,1173,943,969]" italics="true" pageId="2" pageNumber="393">st4</emphasis>
(40–42) and associated poroids located on soft integument. Endopodal plates II/III completely fused to sternal shield, endopodal plates III/IV elongate, narrow and curved. Genital shield tongue-shaped, slightly expanded posteriorly behind setae 
<emphasis id="DFA8606CFFE6FF9AFAC933EDFA16702B" box="[1397,1429,1016,1042]" italics="true" pageId="2" pageNumber="393">st5</emphasis>
, length 241–244, maximum width 100–107, anterior margin convex, abutting posterior area of sternal shield, posterior margin slightly tapered, surface reticulated with irregular longitudinal lines in anterior region, posteriorly with 6–8 irregular cells behind an inverted-V shaped ridge; bearing a pair of simple setae 
<emphasis id="DFA8606CFFE6FF9AFBF43471FBEA7047" box="[1096,1129,1124,1150]" italics="true" pageId="2" pageNumber="393">st5</emphasis>
(28–30) (
<figureCitation id="75E7A0FBFFE6FF9AFB6B3476FAA37044" box="[1239,1312,1123,1149]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 2</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFA973476FAE77044" box="[1323,1380,1123,1149]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">9, 11</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFACC3476FA0C7044" box="[1392,1423,1123,1149]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="2" pageNumber="393">13</figureCitation>
). Shield flanked by two pairs of minute, elongate paragenital platelets; paragenital poroids 
<emphasis id="DFA8606CFFE6FF9AFBC1349DFB1C709B" box="[1149,1183,1160,1186]" italics="true" pageId="2" pageNumber="393">iv5</emphasis>
located on soft cuticle lateral to shield near seta 
<emphasis id="DFA8606CFFE6FF9AFE7C34B9FE6270FF" box="[448,481,1196,1222]" italics="true" pageId="2" pageNumber="393">st5</emphasis>
. Anal shield elongate, somewhat truncate anteriorly, length 152–158, width 76–84, lineate-reticulate in anterior half, post-anal seta (47–51) longer and thicker than para-anal setae (27–31), cribrum consisting of a terminal tuft with 4–5 irregular rows of spicules and a pair of anterior arms extending slightly beyond post-anal seta; anal gland pores (
<emphasis id="DFA8606CFFE6FF9AFDB63502FDB17108" box="[522,562,1303,1329]" italics="true" pageId="2" pageNumber="393">gv3</emphasis>
) on anterolateral margin of anal shield (
<figureCitation id="75E7A0FBFFE6FF9AFC453502FBC17108" box="[1017,1090,1303,1329]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 2</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFBF33502FB0B7108" box="[1103,1160,1303,1329]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">9, 11</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFB293502FB377108" box="[1173,1204,1303,1329]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="2" pageNumber="393">14</figureCitation>
). Soft opisthogastric cuticle with one pair of small oval metapodal plates and 15 pairs of smooth and very thick setae (
<emphasis id="DFA8606CFFE6FF9AFB7D352EFA9D716F" box="[1217,1310,1339,1366]" italics="true" pageId="2" pageNumber="393">Jv1–Jv5</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFA96352EFAFD716C" box="[1322,1406,1339,1366]" italics="true" pageId="2" pageNumber="393">Zv1–Zv</emphasis>
5), 
<emphasis id="DFA8606CFFE6FF9AFF2B354AFF707140" box="[151,243,1375,1402]" italics="true" pageId="2" pageNumber="393">Jv1–Jv3</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFEBF354AFEE67140" box="[259,357,1375,1402]" italics="true" pageId="2" pageNumber="393">Zv1–Zv2</emphasis>
(20–30), 
<emphasis id="DFA8606CFFE6FF9AFE6B354AFD827140" box="[471,513,1375,1401]" italics="true" pageId="2" pageNumber="393">Zv3</emphasis>
(50–53), 
<emphasis id="DFA8606CFFE6FF9AFDCF354AFD4C7143" box="[627,719,1375,1402]" italics="true" pageId="2" pageNumber="393">Jv4–Jv5</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFD63354AFCC37143" box="[735,832,1375,1402]" italics="true" pageId="2" pageNumber="393">Zv4–Zv5</emphasis>
(60–70), including two and three pairs of 
<emphasis id="DFA8606CFFE6FF9AFA973575FABF7140" box="[1323,1340,1376,1401]" italics="true" pageId="2" pageNumber="393">R</emphasis>
and 
<emphasis id="DFA8606CFFE6FF9AFACB3575FA1F7140" box="[1399,1436,1376,1401]" italics="true" pageId="2" pageNumber="393">UR</emphasis>
setae (20–30), respectively (
<figureCitation id="75E7A0FBFFE6FF9AFE6A3596FDA371A4" box="[470,544,1411,1437]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 2</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFD913596FDE471A4" box="[557,615,1411,1437]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">9, 11</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFDCF3596FD1171A4" box="[627,658,1411,1437]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="2" pageNumber="393">14</figureCitation>
). Exopodal platelets between coxae II–III small, subtriangular; parapodal platelets well developed, curved, surrounding coxa IV, gland pores 
<emphasis id="DFA8606CFFE6FF9AFC7335B2FC7B71F8" box="[975,1016,1447,1473]" italics="true" pageId="2" pageNumber="393">gv2</emphasis>
on median apices of parapodal platelets. Peritreme narrow, maximum width 10–13 µm, extending anteriorly to mid-level of coxa I; peritrematal shield narrow, maximum width 15–17 µm, expanded anteriorly, fused to dorsal shield at level of 
<emphasis id="DFA8606CFFE6FF9AFB2735FAFB377230" box="[1179,1204,1519,1545]" italics="true" pageId="2" pageNumber="393">s1</emphasis>
, free from exopodal shields, bearing five pore-like structures, a lyrifissure 
<emphasis id="DFA8606CFFE6FF9AFD4B3601FC8E7214" box="[759,781,1556,1581]" italics="true" pageId="2" pageNumber="393">ip</emphasis>
and a gland pore 
<emphasis id="DFA8606CFFE6FF9AFC6B3601FC707214" box="[983,1011,1556,1581]" italics="true" pageId="2" pageNumber="393">gp</emphasis>
at level of coxa II, two lyrifissures 
<emphasis id="DFA8606CFFE6FF9AFA3A3601FA1F7214" box="[1414,1436,1556,1581]" italics="true" pageId="2" pageNumber="393">ip</emphasis>
and a gland pore 
<emphasis id="DFA8606CFFE6FF9AFEE4362DFEF77268" box="[344,372,1592,1617]" italics="true" pageId="2" pageNumber="393">gp</emphasis>
on post-stigmatic section (
<figureCitation id="75E7A0FBFFE6FF9AFD1E3622FD687268" box="[674,747,1591,1617]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 2</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFD4B3622FCB37268" box="[759,816,1591,1617]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[179,1406,181,1920]" captionTargetId="figure-25@4.[179,1408,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–11. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="2" pageNumber="393">9, 11</figureCitation>
).
</paragraph>
<paragraph id="ED63BC7EFFE6FF9BFF7B364EFBB974EC" blockId="2.[151,1437,259,2014]" lastBlockId="3.[151,1437,151,213]" lastPageId="3" lastPageNumber="394" pageId="2" pageNumber="393">
<taxonomicName id="2ADCC7FDFFE6FF9AFF7B364EFED5724C" authorityName="Krefft" authorityYear="1871" box="[199,342,1627,1653]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="393" phylum="Nematoda" rank="genus">
<emphasis id="DFA8606CFFE6FF9AFF7B364EFED5724C" box="[199,342,1627,1653]" italics="true" pageId="2" pageNumber="393">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="75E7A0FBFFE6FF9AFED9364EFE49724C" box="[357,458,1627,1653]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 3–7</figureCitation>
, 
<figureCitation id="75E7A0FBFFE6FF9AFE6B364EFDA2724F" box="[471,545,1627,1654]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="2" pageNumber="393">15–16</figureCitation>
). Epistome subrectangular, anterior margin somewhat truncate, irregularly denticulate (
<figureCitation id="75E7A0FBFFE6FF9AFF47366AFEC272A3" box="[251,321,1663,1690]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Fig. 6</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 10–20 small denticles, with smooth anterior and posterior transverse lines, groove greatly wider anteriorly. Hypostome with four pairs of setae, rostral seta (
<emphasis id="DFA8606CFFE6FF9AFE3736D2FE2472D8" box="[395,423,1735,1761]" italics="true" pageId="2" pageNumber="393">h1</emphasis>
) longest, palpcoxal seta thickened, subulate, 
<emphasis id="DFA8606CFFE6FF9AFC1636D2FC4572D8" box="[938,966,1735,1761]" italics="true" pageId="2" pageNumber="393">h1</emphasis>
(39–41)&gt; 
<emphasis id="DFA8606CFFE6FF9AFBFB36D2FBE772D8" box="[1095,1124,1735,1761]" italics="true" pageId="2" pageNumber="393">h3</emphasis>
(28–30)&gt; 
<emphasis id="DFA8606CFFE6FF9AFB5936DDFB7C72D8" box="[1253,1279,1736,1761]" italics="true" pageId="2" pageNumber="393">pc</emphasis>
(19–21)&gt; 
<emphasis id="DFA8606CFFE6FF9AFA3C36D2FA1E72D8" box="[1408,1437,1735,1761]" italics="true" pageId="2" pageNumber="393">h2</emphasis>
(13–15) (
<figureCitation id="75E7A0FBFFE6FF9AFF4336FEFEC4733C" box="[255,327,1771,1797]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Figs 3</figureCitation>
&amp; 
<figureCitation id="75E7A0FBFFE6FF9AFED036FEFE08733F" box="[364,395,1771,1798]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="2" pageNumber="393">15</figureCitation>
). Corniculi robust, horn-like, extending slightly beyond mid-level of palpfemur. Supralabral process with a thick and undivided apical projection (
<figureCitation id="75E7A0FBFFE6FF9AFD48371AFCBA7313" box="[756,825,1807,1834]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Fig. 5</figureCitation>
). Internal malae shorter than corniculi, composed of a pair of smooth median projections, and lateral fimbriate elongate projections, labrum with pilose surface (
<figureCitation id="75E7A0FBFFE6FF9AFAF43726FA0D7374" box="[1352,1422,1843,1869]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Fig. 3</figureCitation>
). Chaetotaxy of palps (
<figureCitation id="75E7A0FBFFE6FF9AFE313742FE567348" box="[397,469,1879,1905]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Fig. 4</figureCitation>
): trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth and needle-like except setae 
<emphasis id="DFA8606CFFE6FF9AFE9B376EFEC773AC" box="[295,324,1915,1941]" italics="true" pageId="2" pageNumber="393">d1</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFEEE376EFEED73AC" box="[338,366,1915,1941]" italics="true" pageId="2" pageNumber="393">d2</emphasis>
, 
<emphasis id="DFA8606CFFE6FF9AFEC1376EFE1073AC" box="[381,403,1915,1941]" italics="true" pageId="2" pageNumber="393">al</emphasis>
on palpfemur and 
<emphasis id="DFA8606CFFE6FF9AFDD5376EFD0E73AC" box="[617,653,1915,1941]" italics="true" pageId="2" pageNumber="393">al1</emphasis>
on palpgenu thickened, 
<emphasis id="DFA8606CFFE6FF9AFC1D376EFC4673AC" box="[929,965,1915,1941]" italics="true" pageId="2" pageNumber="393">al2</emphasis>
on palpgenu paddle-like, palp tarsal claw two-tined (
<figureCitation id="75E7A0FBFFE6FF9AFEA8378AFED87380" box="[276,347,1951,1977]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="2" pageNumber="393">Fig. 4</figureCitation>
). Fixed and movable digits of chelicera bidentate, reduced dentition, fixed digit with short and relatively thick pilus dentilis, dorsal cheliceral seta conspicuous, prostrate, relatively thick, arthrodial membrane with a rounded flap and normal filaments; cheliceral lyrifissures indistinct (
<figureCitation id="75E7A0FBFFE7FF9BFC5F3082FBAA7488" box="[995,1065,151,177]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1962,1986]" captionTargetBox="[211,1381,245,1927]" captionTargetId="figure-46@3.[192,1395,237,1939]" captionTargetPageId="3" captionText="FIGURES 1–7. Hyposternus ceylonicus gen. nov, sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="3" pageNumber="394">Figs 7</figureCitation>
&amp; 
<figureCitation id="75E7A0FBFFE7FF9BFBF13082FBEF748B" box="[1101,1132,151,178]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="3" pageNumber="394">16</figureCitation>
); movable digit 46–49 µm, middle segment including fixed digit 134–138 µm, narrowing and tapered apically.
</paragraph>
<caption id="B9A3ECF6FFE7FF9BFF2B37BFFD3B73DE" ID-DOI="http://doi.org/10.5281/zenodo.5556471" ID-Zenodo-Dep="5556471" httpUri="https://zenodo.org/record/5556471/files/figure.png" pageId="3" pageNumber="394" startId="3.[151,264,1962,1986]" targetBox="[211,1381,245,1927]" targetPageId="3">
<paragraph id="ED63BC7EFFE7FF9BFF2B37BFFD3B73DE" blockId="3.[151,1436,1962,2023]" pageId="3" pageNumber="394">
<emphasis id="DFA8606CFFE7FF9BFF2B37BFFEB973FA" bold="true" box="[151,314,1962,1987]" pageId="3" pageNumber="394">FIGURES 1–7.</emphasis>
<taxonomicName id="2ADCC7FDFFE7FF9BFE8337BEFDB373FA" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[319,560,1963,1987]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="ceylonicus" status="gen. nov, sp. nov.">
<emphasis id="DFA8606CFFE7FF9BFE8337BEFDB373FA" box="[319,560,1963,1987]" italics="true" pageId="3" pageNumber="394">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49BDD17FFE7FF9BFD8A37BEFD6873FA" box="[566,747,1963,1987]" pageId="3" pageNumber="394" rank="species">
<emphasis id="DFA8606CFFE7FF9BFD8A37BEFD0D73FA" bold="true" box="[566,654,1963,1987]" pageId="3" pageNumber="394">gen. nov</emphasis>
, 
<emphasis id="DFA8606CFFE7FF9BFD2737BEFD6873FA" bold="true" box="[667,747,1963,1987]" pageId="3" pageNumber="394">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, palp; 5, supralabral process; 6, epistome; 7, chelicera.
</paragraph>
</caption>
<caption id="B9A3ECF6FFE0FF9CFF2B3782FC0173ED" ID-DOI="http://doi.org/10.5281/zenodo.5556473" ID-Zenodo-Dep="5556473" httpUri="https://zenodo.org/record/5556473/files/figure.png" pageId="4" pageNumber="395" startId="4.[151,264,1943,1967]" targetBox="[179,1406,181,1920]" targetPageId="4">
<paragraph id="ED63BC7EFFE0FF9CFF2B3782FC0173ED" blockId="4.[151,1437,1943,2004]" pageId="4" pageNumber="395">
<emphasis id="DFA8606CFFE0FF9CFF2B3782FEC47396" bold="true" box="[151,327,1943,1967]" pageId="4" pageNumber="395">FIGURES 8–11.</emphasis>
DIC micrographs of 
<taxonomicName id="2ADCC7FDFFE0FF9CFD9E378DFC977389" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[546,788,1944,1968]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="ceylonicus" status="gen. nov, sp. nov.">
<emphasis id="DFA8606CFFE0FF9CFD9E378DFC977389" box="[546,788,1944,1968]" italics="true" pageId="4" pageNumber="395">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49BDD17FFE0FF9CFCA7378DFC577389" box="[795,980,1944,1968]" pageId="4" pageNumber="395" rank="species">
<emphasis id="DFA8606CFFE0FF9CFCA7378DFCF67389" bold="true" box="[795,885,1944,1968]" pageId="4" pageNumber="395">gen. nov</emphasis>
, 
<emphasis id="DFA8606CFFE0FF9CFC3F378DFC577389" bold="true" box="[899,980,1944,1968]" pageId="4" pageNumber="395">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 8, general view dorsally; 9, general view ventrally; 10, idiosoma in dorsal view; 11, idiosoma in ventral view.
</paragraph>
</caption>
<caption id="B9A3ECF6FFE1FF9DFF2B3782FC0973CE" ID-DOI="http://doi.org/10.5281/zenodo.5556475" ID-Zenodo-Dep="5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="5" pageNumber="396" startId="5.[151,264,1943,1967]" targetBox="[184,1403,182,1918]" targetPageId="5">
<paragraph id="ED63BC7EFFE1FF9DFF2B3782FC0973CE" blockId="5.[151,1436,1943,2039]" pageId="5" pageNumber="396">
<emphasis id="DFA8606CFFE1FF9DFF2B3782FED07396" bold="true" box="[151,339,1943,1967]" pageId="5" pageNumber="396">FIGURES 12–17.</emphasis>
DIC micrographs of 
<taxonomicName id="2ADCC7FDFFE1FF9DFD95378DFC9A7396" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[553,793,1943,1967]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="396" phylum="Arthropoda" rank="species" species="ceylonicus" status="gen. nov, sp. nov.">
<emphasis id="DFA8606CFFE1FF9DFD95378DFC9A7396" box="[553,793,1943,1967]" italics="true" pageId="5" pageNumber="396">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49BDD17FFE1FF9DFCA33782FC577396" box="[799,980,1943,1967]" pageId="5" pageNumber="396" rank="species">
<emphasis id="DFA8606CFFE1FF9DFCA33782FCFB7396" bold="true" box="[799,888,1943,1967]" pageId="5" pageNumber="396">gen. nov</emphasis>
, 
<emphasis id="DFA8606CFFE1FF9DFC383782FC577396" bold="true" box="[900,980,1943,1967]" pageId="5" pageNumber="396">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view.
</paragraph>
</caption>
<caption id="B9A3ECF6FFE2FF9EFF2B3782FA1E7396" ID-DOI="http://doi.org/10.5281/zenodo.5556477" ID-Zenodo-Dep="5556477" box="[151,1437,1943,1967]" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="6" pageNumber="397" startId="6.[151,264,1943,1967]" targetBox="[174,1424,186,1915]" targetPageId="6">
<paragraph id="ED63BC7EFFE2FF9EFF2B3782FA1E7396" blockId="6.[151,1437,1943,1967]" box="[151,1437,1943,1967]" pageId="6" pageNumber="397">
<emphasis id="DFA8606CFFE2FF9EFF2B3782FED17396" bold="true" box="[151,338,1943,1967]" pageId="6" pageNumber="397">FIGURES 18–21.</emphasis>
<taxonomicName id="2ADCC7FDFFE2FF9EFEEB378DFDC57396" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[343,582,1943,1967]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="397" phylum="Arthropoda" rank="species" species="ceylonicus" status="gen. nov, sp. nov.">
<emphasis id="DFA8606CFFE2FF9EFEEB378DFDC57396" box="[343,582,1943,1967]" italics="true" pageId="6" pageNumber="397">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49BDD17FFE2FF9EFDF73782FD7E7396" box="[587,765,1943,1967]" pageId="6" pageNumber="397" rank="species">
<emphasis id="DFA8606CFFE2FF9EFDF73782FD207396" bold="true" box="[587,675,1943,1967]" pageId="6" pageNumber="397">gen. nov</emphasis>
, 
<emphasis id="DFA8606CFFE2FF9EFD123782FD7E7396" bold="true" box="[686,765,1943,1967]" pageId="6" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV.
</paragraph>
</caption>
<caption id="B9A3ECF6FFE3FF9FFF2B3692FB397299" ID-DOI="http://doi.org/10.5281/zenodo.5556479" ID-Zenodo-Dep="5556479" box="[151,1210,1671,1696]" httpUri="https://zenodo.org/record/5556479/files/figure.png" pageId="7" pageNumber="398" startId="7.[151,264,1671,1695]" targetBox="[169,1420,199,1632]" targetPageId="7">
<paragraph id="ED63BC7EFFE3FF9FFF2B3692FB397299" blockId="7.[151,1210,1671,1696]" box="[151,1210,1671,1696]" pageId="7" pageNumber="398">
<emphasis id="DFA8606CFFE3FF9FFF2B3692FED772A6" bold="true" box="[151,340,1671,1695]" pageId="7" pageNumber="398">FIGURES 22–23.</emphasis>
<taxonomicName id="2ADCC7FDFFE3FF9FFEE6369DFDCF7299" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[346,588,1672,1696]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="398" phylum="Arthropoda" rank="species" species="ceylonicus" status="gen. nov, sp. nov.">
<emphasis id="DFA8606CFFE3FF9FFEE6369DFDCF7299" box="[346,588,1672,1696]" italics="true" pageId="7" pageNumber="398">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49BDD17FFE3FF9FFDEE369DFC8A7299" box="[594,777,1672,1696]" pageId="7" pageNumber="398" rank="species">
<emphasis id="DFA8606CFFE3FF9FFDEE369DFD287299" bold="true" box="[594,683,1672,1696]" pageId="7" pageNumber="398">gen. nov</emphasis>
, 
<emphasis id="DFA8606CFFE3FF9FFD05369DFC8A7299" bold="true" box="[697,777,1672,1696]" pageId="7" pageNumber="398">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. 22, ventral idiosoma; 23, chelicera.
</paragraph>
</caption>
<paragraph id="ED63BC7EFFE3FF90FF7B36DAFC6575B0" blockId="7.[151,1436,1742,2021]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="399" pageId="7" pageNumber="398">
<emphasis id="DFA8606CFFE3FF9FFF7B36DAFF7F72D1" box="[199,252,1743,1768]" italics="true" pageId="7" pageNumber="398">Legs</emphasis>
(
<figureCitation id="75E7A0FBFFE3FF9FFEB036DBFE1372D1" box="[268,400,1742,1768]" captionStart="FIGURES 18–21" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[174,1424,186,1915]" captionTargetId="figure-25@6.[151,1436,181,1919]" captionTargetPageId="6" captionText="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5556477" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="7" pageNumber="398">Figs 18–21</figureCitation>
). Legs II (500–521) and III (515–547) short, I (710–730) and IV (713–744) longer. Length of segments: coxa I 80–84, trochanter I 49–53, femur I 135–139, genu I 116–119, tibia I 130–134, tarsus I 200– 205, coxa II 48–51, trochanter II 49–53, femur II 100–103, genu I 75–78, tibia 76–79, tarsus I 142–147, coxa III 48–51, trochanter III 112–115, genu III 71–73, tibia III 70–73, tarsus III 155–161, coxa IV 53–57, trochanter IV 94–97, femur 153–157, genu IV 101–104, tibia IV 113–117, tarsus IV 200–206. Chaetotaxy normal for free-living 
<taxonomicName id="2ADCC7FDFFE3FF9FFF2B3797FE9573A5" authorityName="A.Berlese" authorityYear="1892" box="[151,278,1922,1948]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="398" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="75E7A0FBFFE3FF9FFED13797FE4273A5" box="[365,449,1922,1948]" captionStart="FIGURES 18–21" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[174,1424,186,1915]" captionTargetId="figure-25@6.[151,1436,181,1919]" captionTargetPageId="6" captionText="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5556477" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="7" pageNumber="398">Fig. 18</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 1/1, 0/2 1 (
<emphasis id="DFA8606CFFE3FF9FFC123797FC4973A5" box="[942,970,1922,1948]" italics="true" pageId="7" pageNumber="398">ad</emphasis>
thickened), femur 2 3/1, 2/3 2 (
<emphasis id="DFA8606CFFE3FF9FFA8F3797FADE73A5" box="[1331,1373,1922,1948]" italics="true" pageId="7" pageNumber="398">ad1</emphasis>
, 
<emphasis id="DFA8606CFFE3FF9FFAD73797FA1673A5" box="[1387,1429,1922,1948]" italics="true" pageId="7" pageNumber="398">pd2</emphasis>
, 
<emphasis id="DFA8606CFFE3FF9FFF2B37B3FF3873F9" box="[151,187,1958,1984]" italics="true" pageId="7" pageNumber="398">al1</emphasis>
slightly thickened, 
<emphasis id="DFA8606CFFE3FF9FFE2537B3FE4073F9" box="[409,451,1958,1984]" italics="true" pageId="7" pageNumber="398">ad1</emphasis>
longest, 51–54), genu 2 3/2, 3/1 2 (almost all setae elongated, 50–62), tibia 2 3/2, 3/1 2 (almost all setae elongated, 70–87). Leg II (
<figureCitation id="75E7A0FBFFE3FF9FFD3C37DFFD5073DD" box="[640,723,1994,2020]" captionStart="FIGURES 18–21" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[174,1424,186,1915]" captionTargetId="figure-25@6.[151,1436,181,1919]" captionTargetPageId="6" captionText="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5556477" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="7" pageNumber="398">Fig. 19</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 0/1, 0/2 1 (
<emphasis id="DFA8606CFFE3FF9FFB0537DFFB4C73DD" box="[1209,1231,1994,2020]" italics="true" pageId="7" pageNumber="398">al</emphasis>
thickened), femur 2 3/1, 2/2 1 (
<emphasis id="DFA8606CFFECFF90FE963082FED77488" box="[298,340,151,177]" italics="true" pageId="8" pageNumber="399">ad1</emphasis>
long, 66–70), genu 2 3/1, 2/1 2, tibia 2 2/1, 2/1 2. Leg III (
<figureCitation id="75E7A0FBFFECFF90FC4F3082FBCA7488" box="[1011,1097,151,177]" captionStart="FIGURES 18–21" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[174,1424,186,1915]" captionTargetId="figure-25@6.[151,1436,181,1919]" captionTargetPageId="6" captionText="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5556477" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="8" pageNumber="399">Fig. 20</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 1/1, 0/2 0, femur 1 2/1, 1/0 1 (
<emphasis id="DFA8606CFFECFF90FDBF30AEFDAE74EC" box="[515,557,187,213]" italics="true" pageId="8" pageNumber="399">ad1</emphasis>
long, 66–71), genu 2 2/1, 2/1 1 (
<emphasis id="DFA8606CFFECFF90FC1E30AEFC4F74EC" box="[930,972,187,213]" italics="true" pageId="8" pageNumber="399">pd2</emphasis>
thickened), tibia: 2 1/1, 2/1 1 (all ventral setae, 
<emphasis id="DFA8606CFFECFF90FF6130CAFE8274C0" box="[221,257,223,249]" italics="true" pageId="8" pageNumber="399">al1</emphasis>
&amp; 
<emphasis id="DFA8606CFFECFF90FE9830CAFEB974C0" box="[292,314,223,249]" italics="true" pageId="8" pageNumber="399">pl</emphasis>
thickened). Leg IV (
<figureCitation id="75E7A0FBFFECFF90FD9E30CAFDF574C0" box="[546,630,223,249]" captionStart="FIGURES 18–21" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[174,1424,186,1915]" captionTargetId="figure-25@6.[151,1436,181,1919]" captionTargetPageId="6" captionText="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5556477" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="8" pageNumber="399">Fig. 21</figureCitation>
): coxa 0 0/1, 0/0 0, trochanter 1 1/1, 0/1 1 (
<emphasis id="DFA8606CFFECFF90FBE430CAFBF774C0" box="[1112,1140,223,249]" italics="true" pageId="8" pageNumber="399">ad</emphasis>
slightly thickened), femur 1 2/1, 1/0 1 (
<emphasis id="DFA8606CFFECFF90FE9A3116FED27524" box="[294,337,259,285]" italics="true" pageId="8" pageNumber="399">ad2</emphasis>
thickened, 
<emphasis id="DFA8606CFFECFF90FE6E3116FE7F7524" box="[466,508,259,285]" italics="true" pageId="8" pageNumber="399">ad1</emphasis>
macroseta, 97–101), genu 2 2/1, 3/0 1 (
<emphasis id="DFA8606CFFECFF90FC063116FC577525" box="[954,980,259,284]" italics="true" pageId="8" pageNumber="399">av</emphasis>
slightly thickened), tibia 2 1/1, 3/1 2 (
<emphasis id="DFA8606CFFECFF90FA3E3116FA1F7525" box="[1410,1436,259,284]" italics="true" pageId="8" pageNumber="399">av</emphasis>
thickened). Tarsi II–IV with 18 setae (3 3/2, 3/2 3 + 
<emphasis id="DFA8606CFFECFF90FD503132FC8E7579" box="[748,781,295,320]" italics="true" pageId="8" pageNumber="399">mv</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FCA03132FCBD7578" box="[796,830,295,321]" italics="true" pageId="8" pageNumber="399">md</emphasis>
); most setae on ventral and lateral faces thickened or spine-like (
<figureCitation id="75E7A0FBFFECFF90FEAF315EFE3A755C" box="[275,441,331,357]" captionStart="FIGURES 18–21" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[174,1424,186,1915]" captionTargetId="figure-25@6.[151,1436,181,1919]" captionTargetPageId="6" captionText="FIGURES 18–21. Hyposternus ceylonicus gen. nov, sp. nov., female.18, leg I (trochanter-tibia); 19, leg II; 20, leg III; 21, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5556477" httpUri="https://zenodo.org/record/5556477/files/figure.png" pageId="8" pageNumber="399">Figures 19–21</figureCitation>
). Setae 
<emphasis id="DFA8606CFFECFF90FDB1315EFDD0755C" box="[525,595,331,357]" italics="true" pageId="8" pageNumber="399">ad2–3</emphasis>
and 
<emphasis id="DFA8606CFFECFF90FD3B315EFD4E755C" box="[647,717,331,357]" italics="true" pageId="8" pageNumber="399">pd2–3</emphasis>
(75–80) slightly longer than others on tarsus IV.All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B3186FCF07594" blockId="8.[151,1437,151,2013]" box="[199,883,403,429]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B3186FE587594" bold="true" box="[199,475,403,429]" italics="true" pageId="8" pageNumber="399">Insemination structures</emphasis>
. Not seen, apparently unsclerotised.
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B31A3FD4C75E8" blockId="8.[151,1437,151,2013]" box="[199,719,438,465]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B31A3FED275E9" bold="true" box="[199,337,438,464]" pageId="8" pageNumber="399">Description</emphasis>
. 
<emphasis id="DFA8606CFFECFF90FEDC31A2FE6875E9" bold="true" box="[352,491,438,465]" pageId="8" pageNumber="399">Male (n= 1)</emphasis>
(
<figureCitation id="75E7A0FBFFECFF90FE4531A2FDF775E8" box="[505,628,439,465]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="8" pageNumber="399">Figures 17</figureCitation>
, 
<figureCitation id="75E7A0FBFFECFF90FD3C31A2FD4475E8" box="[640,711,439,465]" captionStart="FIGURES 22–23" captionStartId="7.[151,264,1671,1695]" captionTargetBox="[169,1420,199,1632]" captionTargetId="figure-25@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURES 22–23. Hyposternus ceylonicus gen. nov, sp. nov., male. 22, ventral idiosoma; 23, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556479" httpUri="https://zenodo.org/record/5556479/files/figure.png" pageId="8" pageNumber="399">22–23</figureCitation>
)
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B31CEFF687620" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B31CEFE0775CC" box="[199,388,475,501]" italics="true" pageId="8" pageNumber="399">Dorsal idiosoma</emphasis>
. Dorsal shield 391 long, 205 wide; ornamentation, chaetotaxy, length and form of setae as in female.
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B3236FD247764" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B3236FE097604" box="[199,394,547,573]" italics="true" pageId="8" pageNumber="399">Ventral idiosoma</emphasis>
(
<figureCitation id="75E7A0FBFFECFF90FE203236FE757604" box="[412,502,547,573]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="8" pageNumber="399">Figs 17</figureCitation>
, 
<figureCitation id="75E7A0FBFFECFF90FDBA3236FDA57604" box="[518,550,547,573]" captionStart="FIGURES 22–23" captionStartId="7.[151,264,1671,1695]" captionTargetBox="[169,1420,199,1632]" captionTargetId="figure-25@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURES 22–23. Hyposternus ceylonicus gen. nov, sp. nov., male. 22, ventral idiosoma; 23, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556479" httpUri="https://zenodo.org/record/5556479/files/figure.png" pageId="8" pageNumber="399">22</figureCitation>
). Sternal, genital, endopodal, ventral and anal shields fused into holoventral shield, 301 long from anterior to posterior margins of shield, 88 wide at level of 
<emphasis id="DFA8606CFFECFF90FBAA3252FBB47658" box="[1046,1079,583,609]" italics="true" pageId="8" pageNumber="399">st2</emphasis>
, 94 at 
<emphasis id="DFA8606CFFECFF90FB3E3252FB207658" box="[1154,1187,583,609]" italics="true" pageId="8" pageNumber="399">st3</emphasis>
level and 94 at broadest point (behind parapodals), anterolateral edges of shield fused to endopodals between coxae I-II; shield reticulate throughout, except smooth median area between setae 
<emphasis id="DFA8606CFFECFF90FC87329AFCCD7691" box="[827,846,655,680]" italics="true" pageId="8" pageNumber="399">st</emphasis>
2– 
<emphasis id="DFA8606CFFECFF90FCD5329AFCF47690" box="[873,887,655,681]" italics="true" pageId="8" pageNumber="399">4</emphasis>
; with 10 pairs of smooth setae (
<emphasis id="DFA8606CFFECFF90FB51329AFA837691" box="[1261,1280,655,680]" italics="true" pageId="8" pageNumber="399">st</emphasis>
1– 
<emphasis id="DFA8606CFFECFF90FAA0329AFAA97690" box="[1308,1322,655,681]" italics="true" pageId="8" pageNumber="399">5</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FA86329AFAD07691" box="[1338,1363,655,680]" italics="true" pageId="8" pageNumber="399">Jv</emphasis>
1– 
<emphasis id="DFA8606CFFECFF90FAD2329AFA167690" box="[1390,1429,655,681]" italics="true" pageId="8" pageNumber="399">Jv3</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FF2B32A6FF3076F5" box="[151,179,691,716]" italics="true" pageId="8" pageNumber="399">Zv</emphasis>
1– 
<emphasis id="DFA8606CFFECFF90FF7332A6FF7976F4" box="[207,250,691,717]" italics="true" pageId="8" pageNumber="399">Zv2</emphasis>
) excluding circumanals, post-anal seta (28) longer and thicker than para-anals (21), bearing four pairs of poroids and three pairs of gland pores (
<emphasis id="DFA8606CFFECFF90FDE132C2FD0676C8" box="[605,645,727,753]" italics="true" pageId="8" pageNumber="399">gvb</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FD2A32C2FD7576C8" box="[662,758,727,753]" italics="true" pageId="8" pageNumber="399">gv2, gv3</emphasis>
), 
<emphasis id="DFA8606CFFECFF90FCAC32C2FCB176C8" box="[784,818,727,753]" italics="true" pageId="8" pageNumber="399">iv1</emphasis>
slit-like; cribrum with 3–4 irregular rows of spicules, restricted to region posterior to post-anal seta; metapodal platelets narrow and fused to shield. Soft opisthogastric cuticle with five pairs of short and thick setae, shorter and finer than those of female. Peritremes and peritrematal shields similar to those in female (
<figureCitation id="75E7A0FBFFECFF90FDAB3356FDED7764" box="[535,622,835,861]" captionStart="FIGURES 12–17" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[184,1403,182,1918]" captionTargetId="figure-17@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 12–17. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., female. 12, sternal shield; 13, genital shield; 14, anal shield and portion of opisthogaster; 15, subcapitulum; 16, chelicera (note that the movable digit of left chelicera has been moved from its natural position). Male: 17, idiosoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5556475" httpUri="https://zenodo.org/record/5556475/files/figure.png" pageId="8" pageNumber="399">Figs 17</figureCitation>
, 
<figureCitation id="75E7A0FBFFECFF90FDC63356FD197764" box="[634,666,835,861]" captionStart="FIGURES 22–23" captionStartId="7.[151,264,1671,1695]" captionTargetBox="[169,1420,199,1632]" captionTargetId="figure-25@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURES 22–23. Hyposternus ceylonicus gen. nov, sp. nov., male. 22, ventral idiosoma; 23, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556479" httpUri="https://zenodo.org/record/5556479/files/figure.png" pageId="8" pageNumber="399">22</figureCitation>
).
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B3372FD2877D4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="399">
<taxonomicName id="2ADCC7FDFFECFF90FF7B3372FED577B8" authorityName="Krefft" authorityYear="1871" box="[199,342,871,897]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="8" pageNumber="399" phylum="Nematoda" rank="genus">
<emphasis id="DFA8606CFFECFF90FF7B3372FED577B8" box="[199,342,871,897]" italics="true" pageId="8" pageNumber="399">Gnathosoma</emphasis>
</taxonomicName>
. Epistome and subcapitulum similar to female. Fixed digit of chelicera with one tooth, a thorn-like pilus dentilis, dorsal seta conspicuous, relatively thick. Movable digit of chelicera with one blunt tooth, spermatodactyl stout and slightly curved dorsally, with truncate tip, free portion of spermatodactyl about as long as movable digit (
<figureCitation id="75E7A0FBFFECFF90FF6633C6FEAD77D4" box="[218,302,979,1005]" captionStart="FIGURES 22–23" captionStartId="7.[151,264,1671,1695]" captionTargetBox="[169,1420,199,1632]" captionTargetId="figure-25@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURES 22–23. Hyposternus ceylonicus gen. nov, sp. nov., male. 22, ventral idiosoma; 23, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556479" httpUri="https://zenodo.org/record/5556479/files/figure.png" pageId="8" pageNumber="399">Fig. 23</figureCitation>
). Palps similar to those of female.
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B33E2FDA37028" blockId="8.[151,1437,151,2013]" box="[199,544,1015,1041]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B33E2FF7F7029" box="[199,252,1015,1040]" italics="true" pageId="8" pageNumber="399">Legs</emphasis>
. Chaetotaxy as in female.
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B340FFCB4700C" blockId="8.[151,1437,151,2013]" box="[199,823,1050,1077]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B340FFED2700D" bold="true" box="[199,337,1050,1076]" pageId="8" pageNumber="399">Description</emphasis>
. 
<emphasis id="DFA8606CFFECFF90FEDC340EFDC7700D" bold="true" box="[352,580,1050,1077]" pageId="8" pageNumber="399">Protonymph (n= 4)</emphasis>
(
<figureCitation id="75E7A0FBFFECFF90FDEF340EFD57700C" box="[595,724,1051,1077]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="8" pageNumber="399">Figs 24–29</figureCitation>
, 
<figureCitation id="75E7A0FBFFECFF90FD5C340EFCA9700C" box="[736,810,1051,1077]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">30–34</figureCitation>
).
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B342AFC637140" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B342AFE067060" box="[199,389,1087,1113]" italics="true" pageId="8" pageNumber="399">Dorsal idiosoma</emphasis>
(
<figureCitation id="75E7A0FBFFECFF90FE29342AFE6F7060" box="[405,492,1087,1113]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="8" pageNumber="399">Figs 24</figureCitation>
&amp; 
<figureCitation id="75E7A0FBFFECFF90FDAF342AFDB07060" box="[531,563,1087,1113]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">30</figureCitation>
). Whitish, oval, with podonotal (252–269 long 214–226 wide) and pygidial (118–130 long 169–175 wide) shields, three pairs of mesonotal scutellae, the largest 33–37 long 9–13 wide, and seven pairs of setae located between podonotal and pygidial shields (
<emphasis id="DFA8606CFFECFF90FC283493FC2E7099" box="[916,941,1158,1184]" italics="true" pageId="8" pageNumber="399">s6</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FC063492FC577098" box="[954,980,1159,1185]" italics="true" pageId="8" pageNumber="399">J1</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FC5D3492FC787098" box="[993,1019,1159,1185]" italics="true" pageId="8" pageNumber="399">J2</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FBB53492FBA57098" box="[1033,1062,1159,1185]" italics="true" pageId="8" pageNumber="399">Z1</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FB883492FBD27098" box="[1076,1105,1159,1185]" italics="true" pageId="8" pageNumber="399">Z2</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FBE23492FBF97098" box="[1118,1146,1159,1185]" italics="true" pageId="8" pageNumber="399">S2</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FB343492FB207098" box="[1160,1187,1159,1185]" italics="true" pageId="8" pageNumber="399">S3</emphasis>
) (
<figureCitation id="75E7A0FBFFECFF90FB073492FAB87098" box="[1211,1339,1158,1185]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="8" pageNumber="399">Figs 24–25</figureCitation>
, 
<figureCitation id="75E7A0FBFFECFF90FAFA3492FA127098" box="[1350,1425,1159,1185]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">30–31</figureCitation>
). Podonotal shield weakly reticulated, only distinct in lateral regions, with 11 pairs of simple setae (
<emphasis id="DFA8606CFFECFF90FB6034BEFB7170FC" box="[1244,1266,1195,1221]" italics="true" pageId="8" pageNumber="399">j1</emphasis>
–6, 
<emphasis id="DFA8606CFFECFF90FAA034BEFAD370FC" box="[1308,1360,1195,1221]" italics="true" pageId="8" pageNumber="399">z2–5</emphasis>
, s 
<emphasis id="DFA8606CFFECFF90FAD534BEFA1070FC" box="[1385,1427,1195,1221]" italics="true" pageId="8" pageNumber="399">4–5</emphasis>
) (
<emphasis id="DFA8606CFFECFF90FF1D34DAFF5670D0" box="[161,213,1231,1257]" italics="true" pageId="8" pageNumber="399">r2–3</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FF5834DAFF7E70D0" box="[228,253,1231,1257]" italics="true" pageId="8" pageNumber="399">r5</emphasis>
on lateral soft cuticle) and six pairs of pore-like structures, including three pairs of poroids (
<emphasis id="DFA8606CFFECFF90FAA034DAFAD870D1" box="[1308,1371,1230,1257]" italics="true" pageId="8" pageNumber="399">id4–6</emphasis>
), and three pairs of gland pores (
<emphasis id="DFA8606CFFECFF90FE7D34E6FD847134" box="[449,519,1267,1293]" italics="true" pageId="8" pageNumber="399">gd1–2</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FDA834E6FDBC7134" box="[532,575,1267,1293]" italics="true" pageId="8" pageNumber="399">gd4</emphasis>
). All setae similar in length (23–30) except 
<emphasis id="DFA8606CFFECFF90FB9B34E6FBBE7134" box="[1063,1085,1267,1293]" italics="true" pageId="8" pageNumber="399">j1</emphasis>
slightly longer (30–33). Pygidial shield faintly reticulated throughout, with eight pairs of simple setae (
<emphasis id="DFA8606CFFECFF90FC013502FC5B7108" box="[957,984,1303,1329]" italics="true" pageId="8" pageNumber="399">J3</emphasis>
–5, 
<emphasis id="DFA8606CFFECFF90FBBD3502FBB97108" box="[1025,1082,1303,1329]" italics="true" pageId="8" pageNumber="399">Z3–5</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FBFB3502FBFC7108" box="[1095,1151,1303,1329]" italics="true" pageId="8" pageNumber="399">S4–5</emphasis>
) and pore-like structures, including seven pairs of poroids (
<emphasis id="DFA8606CFFECFF90FDB1352EFDE2716D" box="[525,609,1338,1365]" italics="true" pageId="8" pageNumber="399">idm3–6</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FDD3352EFD12716C" box="[623,657,1339,1365]" italics="true" pageId="8" pageNumber="399">idx</emphasis>
, 
<emphasis id="DFA8606CFFECFF90FD23352EFD65716C" box="[671,742,1339,1365]" italics="true" pageId="8" pageNumber="399">idl3–4</emphasis>
), and one pair of gland pores (
<emphasis id="DFA8606CFFECFF90FB81352EFBE4716C" box="[1085,1127,1339,1365]" italics="true" pageId="8" pageNumber="399">gd8</emphasis>
). All setae similar in length (23–30) except 
<emphasis id="DFA8606CFFECFF90FEFB354AFEE67140" box="[327,357,1375,1401]" italics="true" pageId="8" pageNumber="399">Z5</emphasis>
about twice as long (58–61) as other setae (
<figureCitation id="75E7A0FBFFECFF90FCED354AFC247140" box="[849,935,1375,1401]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="8" pageNumber="399">Figs 24</figureCitation>
, 
<figureCitation id="75E7A0FBFFECFF90FC0F354AFC507140" box="[947,979,1375,1401]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">30</figureCitation>
).
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B3596FB6E72D8" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="399">
<emphasis id="DFA8606CFFECFF90FF7B3596FE0571A4" box="[199,390,1411,1437]" italics="true" pageId="8" pageNumber="399">Ventral idiosoma</emphasis>
(
<figureCitation id="75E7A0FBFFECFF90FE283596FE6B71A4" box="[404,488,1410,1437]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="8" pageNumber="399">Figs 25</figureCitation>
&amp; 
<figureCitation id="75E7A0FBFFECFF90FDB03596FDA871A4" box="[524,555,1411,1437]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">31</figureCitation>
). Tritosternum with paired pilose laciniae (89–95), fused basally (4–6), columnar base 23–26 16–19 wide. Sternal shield indistinct, sternal area with three pairs of smooth setae (
<emphasis id="DFA8606CFFECFF90FB4335B2FA9C71F8" box="[1279,1311,1447,1473]" italics="true" pageId="8" pageNumber="399">st1</emphasis>
31–33, 
<emphasis id="DFA8606CFFECFF90FAC035B2FA1F71F8" box="[1404,1436,1447,1473]" italics="true" pageId="8" pageNumber="399">st2</emphasis>
26–28, 
<emphasis id="DFA8606CFFECFF90FF5035DEFE8E71DC" box="[236,269,1483,1509]" italics="true" pageId="8" pageNumber="399">st3</emphasis>
23–25) and sternal poroids located on soft integument, metasternal setae 
<emphasis id="DFA8606CFFECFF90FBF135DEFBED71DC" box="[1101,1134,1483,1509]" italics="true" pageId="8" pageNumber="399">st4</emphasis>
absent. Anal shield subtriangular, elongate, length 96–99, width 58–61, post-anal seta (36–39) much longer and thicker than para-anal setae (23–26), cribrum consisting of irregular rows of spicules extending slightly beyond post-anal seta; anal gland pores (
<emphasis id="DFA8606CFFECFF90FF1D3622FF4A7268" box="[161,201,1591,1617]" italics="true" pageId="8" pageNumber="399">gv3</emphasis>
) on anterolateral margin of anal shield (
<figureCitation id="75E7A0FBFFECFF90FD2D3622FD647268" box="[657,743,1590,1617]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="8" pageNumber="399">Figs 25</figureCitation>
&amp; 
<figureCitation id="75E7A0FBFFECFF90FCB13622FCAF7268" box="[781,812,1591,1617]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">31</figureCitation>
). Opisthogastric cuticle with five pairs of thorn-shaped setae (
<emphasis id="DFA8606CFFECFF90FF5D364EFE82724C" box="[225,257,1627,1653]" italics="true" pageId="8" pageNumber="399">st5</emphasis>
11–13, 
<emphasis id="DFA8606CFFECFF90FEE2364EFE22724C" box="[350,417,1627,1653]" italics="true" pageId="8" pageNumber="399">Jv1–2</emphasis>
16–19, 
<emphasis id="DFA8606CFFECFF90FE43364EFDA5724C" box="[511,550,1627,1653]" italics="true" pageId="8" pageNumber="399">Jv5</emphasis>
51–53, 
<emphasis id="DFA8606CFFECFF90FD38364EFD2D724C" box="[644,686,1627,1653]" italics="true" pageId="8" pageNumber="399">Zv2</emphasis>
16–19), five pairs of 
<emphasis id="DFA8606CFFECFF90FC1B364EFC4A724D" box="[935,969,1627,1652]" italics="true" pageId="8" pageNumber="399">ivo</emphasis>
, a pair of 
<emphasis id="DFA8606CFFECFF90FBFE364EFBE7724D" box="[1090,1124,1627,1652]" italics="true" pageId="8" pageNumber="399">ivp</emphasis>
and 
<emphasis id="DFA8606CFFECFF90FB21364EFB46724C" box="[1181,1221,1627,1653]" italics="true" pageId="8" pageNumber="399">gv2</emphasis>
, and three pairs of small platelets. Peritreme short, located in the region of coxae III/IV (
<figureCitation id="75E7A0FBFFECFF90FC28366AFC6572A0" box="[916,998,1662,1689]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="8" pageNumber="399">Fig. 25</figureCitation>
), peritrematal shield lightly sclerotised, reduced to a small fragment between coxae II-III, bearing a pore (
<emphasis id="DFA8606CFFECFF90FC2E36B6FC2D7285" box="[914,942,1699,1724]" italics="true" pageId="8" pageNumber="399">gp</emphasis>
), and a distinguishable platelet bearing the peritreme. Post-stigmatic section not developed, a lyrifissure 
<emphasis id="DFA8606CFFECFF90FCFA36D2FCDF72D9" box="[838,860,1735,1760]" italics="true" pageId="8" pageNumber="399">ip</emphasis>
and a gland pore 
<emphasis id="DFA8606CFFECFF90FB9F36D2FBBC72D9" box="[1059,1087,1735,1760]" italics="true" pageId="8" pageNumber="399">gp</emphasis>
on soft cuticle.
</paragraph>
<paragraph id="ED63BC7EFFECFF90FF7B36FEFA9473E4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="399">
<taxonomicName id="2ADCC7FDFFECFF90FF7B36FEFED5733C" authorityName="Krefft" authorityYear="1871" box="[199,342,1771,1797]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="8" pageNumber="399" phylum="Nematoda" rank="genus">
<emphasis id="DFA8606CFFECFF90FF7B36FEFED5733C" box="[199,342,1771,1797]" italics="true" pageId="8" pageNumber="399">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="75E7A0FBFFECFF90FED936FEFE6B733C" box="[357,488,1771,1797]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">Figs 32–34</figureCitation>
). Epistome subrectangular, anterior margin straight, irregularly denticulate (
<figureCitation id="75E7A0FBFFECFF90FA8736FEFA0C733C" box="[1339,1423,1771,1797]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">Fig. 33</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 13–20 tiny denticles (
<figureCitation id="75E7A0FBFFECFF90FB10371AFB7C7310" box="[1196,1279,1807,1833]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">Fig. 32</figureCitation>
), with smooth anterior and posterior transverse lines, groove greatly wider anteriorly (
<figureCitation id="75E7A0FBFFECFF90FC103726FC7C7374" box="[940,1023,1843,1869]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">Fig. 32</figureCitation>
). Hypostome with four pairs of setae, 
<emphasis id="DFA8606CFFECFF90FF2B3742FF307348" box="[151,179,1879,1905]" italics="true" pageId="8" pageNumber="399">h3</emphasis>
(23–25), 
<emphasis id="DFA8606CFFECFF90FEA33742FEB87348" box="[287,315,1879,1905]" italics="true" pageId="8" pageNumber="399">h1</emphasis>
(25–29), 
<emphasis id="DFA8606CFFECFF90FE1B3742FE407348" box="[423,451,1879,1905]" italics="true" pageId="8" pageNumber="399">h2</emphasis>
(12–14), palpcoxal 
<emphasis id="DFA8606CFFECFF90FD1D3742FD387349" box="[673,699,1879,1904]" italics="true" pageId="8" pageNumber="399">pc</emphasis>
(14–16) thickened, subulate (
<figureCitation id="75E7A0FBFFECFF90FBBA3742FBDA7348" box="[1030,1113,1879,1905]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">Fig. 32</figureCitation>
). Corniculi robust, horn-like, extending slightly beyond mid-level palpfemur. Supralabral process with a thick and undivided apical projection (
<figureCitation id="75E7A0FBFFECFF90FF1C378AFF717380" box="[160,242,1951,1977]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">Fig. 33</figureCitation>
). Internal malae shorter than corniculi, composed of a pair of smooth median projections, and lateral fimbriate elongated projections, labrum with pilose surface. Palps and chelicera similar to the adult (
<figureCitation id="75E7A0FBFFECFF90FB0A37D6FA8873E4" box="[1206,1291,1987,2013]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1908,1932]" captionTargetBox="[164,1426,182,1885]" captionTargetId="figure-25@10.[161,1426,181,1885]" captionTargetPageId="10" captionText="FIGURES 30–34. DIC micrographs of Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="8" pageNumber="399">Fig. 34</figureCitation>
).
</paragraph>
<caption id="B9A3ECF6FFEDFF91FF2B3782FB3173ED" ID-DOI="http://doi.org/10.5281/zenodo.5556481" ID-Zenodo-Dep="5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="9" pageNumber="400" startId="9.[151,264,1943,1967]" targetBox="[192,1407,185,1914]" targetPageId="9">
<paragraph id="ED63BC7EFFEDFF91FF2B3782FB3173ED" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="400">
<emphasis id="DFA8606CFFEDFF91FF2B3782FED77396" bold="true" box="[151,340,1943,1967]" pageId="9" pageNumber="400">FIGURES 24–29.</emphasis>
<taxonomicName id="2ADCC7FDFFEDFF91FEE5378DFDC97389" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[345,586,1944,1968]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="400" phylum="Arthropoda" rank="species" species="ceylonicus" status="gen. nov, sp. nov.">
<emphasis id="DFA8606CFFEDFF91FEE5378DFDC97389" box="[345,586,1944,1968]" italics="true" pageId="9" pageNumber="400">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49BDD17FFEDFF91FDEC378DFC857389" box="[592,774,1944,1968]" pageId="9" pageNumber="400" rank="species">
<emphasis id="DFA8606CFFEDFF91FDEC378DFD2A7389" bold="true" box="[592,681,1944,1968]" pageId="9" pageNumber="400">gen. nov</emphasis>
, 
<emphasis id="DFA8606CFFEDFF91FD0A378DFC857389" bold="true" box="[694,774,1944,1968]" pageId="9" pageNumber="400">sp. nov.</emphasis>
</taxonomicNameLabel>
, protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia).
</paragraph>
</caption>
<caption id="B9A3ECF6FFEEFF92FF2B3761FBFE7388" ID-DOI="http://doi.org/10.5281/zenodo.5556483" ID-Zenodo-Dep="5556483" httpUri="https://zenodo.org/record/5556483/files/figure.png" pageId="10" pageNumber="401" startId="10.[151,264,1908,1932]" targetBox="[164,1426,182,1885]" targetPageId="10">
<paragraph id="ED63BC7EFFEEFF92FF2B3761FBFE7388" blockId="10.[151,1436,1908,1969]" pageId="10" pageNumber="401">
<emphasis id="DFA8606CFFEEFF92FF2B3761FED673B4" bold="true" box="[151,341,1908,1933]" pageId="10" pageNumber="401">FIGURES 30–34.</emphasis>
DIC micrographs of 
<taxonomicName id="2ADCC7FDFFEEFF92FD8C3760FCA173B4" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[560,802,1909,1933]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="401" phylum="Arthropoda" rank="species" species="ceylonicus" status="gen. nov, sp. nov.">
<emphasis id="DFA8606CFFEEFF92FD8C3760FCA173B4" box="[560,802,1909,1933]" italics="true" pageId="10" pageNumber="401">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49BDD17FFEEFF92FC953760FC6073B4" box="[809,995,1909,1933]" pageId="10" pageNumber="401" rank="species">
<emphasis id="DFA8606CFFEEFF92FC953760FC0073B4" bold="true" box="[809,899,1909,1933]" pageId="10" pageNumber="401">gen. nov</emphasis>
, 
<emphasis id="DFA8606CFFEEFF92FC2D3760FC6073B4" bold="true" box="[913,995,1909,1933]" pageId="10" pageNumber="401">sp. nov.</emphasis>
</taxonomicNameLabel>
, protonymph. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, subcapitulum; 33, supralabral process &amp; epistome; 34, chelicera.
</paragraph>
</caption>
<paragraph id="ED63BC7EFFEFFF93FF7B3082FBF97594" blockId="11.[151,1437,150,681]" pageId="11" pageNumber="402">
<emphasis id="DFA8606CFFEFFF93FF7B3082FF7F7489" box="[199,252,151,176]" italics="true" pageId="11" pageNumber="402">Legs</emphasis>
(
<figureCitation id="75E7A0FBFFEFFF93FEB73082FE047488" box="[267,391,151,177]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="11" pageNumber="402">Figs 26 29</figureCitation>
). Legs II (372 382) and III (347 360) short, I (525 535) and IV (460 489) longer. Chaetotaxy: Leg I (
<figureCitation id="75E7A0FBFFEFFF93FF5E30AEFEB574EF" box="[226,310,187,214]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="11" pageNumber="402">Fig. 26</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 0/1, 0/1 1, femur 2 2/1, 2/1 2 (
<emphasis id="DFA8606CFFEFFF93FC4E30AEFB9F74EC" box="[1010,1052,187,213]" italics="true" pageId="11" pageNumber="402">ad1</emphasis>
longest), genu 1 2/1, 2/1 1, tibia 1 2/1, 2/1 1. Leg II (
<figureCitation id="75E7A0FBFFEFFF93FED930CAFE3A74C0" box="[357,441,223,249]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="11" pageNumber="402">Fig. 27</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 0/0, 0/2 1 (
<emphasis id="DFA8606CFFEFFF93FC1C30CAFC3574C0" box="[928,950,223,249]" italics="true" pageId="11" pageNumber="402">al</emphasis>
thickened), femur 1 2/1, 2/1 1 (
<emphasis id="DFA8606CFFEFFF93FAA030CAFAC574C0" box="[1308,1350,223,249]" italics="true" pageId="11" pageNumber="402">ad1</emphasis>
macroseta), genu 1 2/0, 2/0 1, tibia 1 1/1, 2/1 2. Leg III (
<figureCitation id="75E7A0FBFFEFFF93FD6E3116FCA47524" box="[722,807,259,285]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="11" pageNumber="402">Fig. 28</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 1/1, 0/1 0, femur 1 2/1, 1/0 0 (
<emphasis id="DFA8606CFFEFFF93FF5F3132FE8E7578" box="[227,269,295,321]" italics="true" pageId="11" pageNumber="402">ad1</emphasis>
macroseta), genu 1 2/0, 2/0 1, tibia: 1 1/1, 2/1 1. Leg IV (
<figureCitation id="75E7A0FBFFEFFF93FC1B3132FC7F7578" box="[935,1020,295,321]" captionStart="FIGURES 24–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[192,1407,185,1914]" captionTargetId="figure-17@9.[164,1424,181,1919]" captionTargetPageId="9" captionText="FIGURES 24–29. Hyposternus ceylonicus gen. nov, sp. nov., protonymph. 24, dorsal idiosoma; 25, ventral idiosoma; 26, leg I (trochanter-tibia); 27, leg II (trochanter-tibia); 28, leg III (trochanter-tibia); 29, leg IV (trochanter-tibia)." figureDoi="http://doi.org/10.5281/zenodo.5556481" httpUri="https://zenodo.org/record/5556481/files/figure.png" pageId="11" pageNumber="402">Fig. 29</figureCitation>
): coxa 0 0/1, 0/0 0, trochanter 1 1/1, 0/1 0 (
<emphasis id="DFA8606CFFEFFF93FF5C315EFF7F755C" box="[224,252,331,357]" italics="true" pageId="11" pageNumber="402">ad</emphasis>
thickened), femur 1 2/0, 1/0 0 (
<emphasis id="DFA8606CFFEFFF93FDDC315EFD09755C" box="[608,650,331,357]" italics="true" pageId="11" pageNumber="402">ad1</emphasis>
, 
<emphasis id="DFA8606CFFEFFF93FD24315EFD41755C" box="[664,706,331,357]" italics="true" pageId="11" pageNumber="402">ad2</emphasis>
thickened &amp; 
<emphasis id="DFA8606CFFEFFF93FCE5315EFC00755C" box="[857,899,331,357]" italics="true" pageId="11" pageNumber="402">ad1</emphasis>
macroseta), genu 1 2/0, 2/0 0, tibia 1 1/1, 2/1 1. Tarsi II-IV with 17 setae (3 3/2, 3/2 3 + 
<emphasis id="DFA8606CFFEFFF93FDEB317AFDFA75B0" box="[599,633,367,393]" italics="true" pageId="11" pageNumber="402">md</emphasis>
); most setae on ventral and lateral aspects thickened (or spine-like). All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
</subSubSection>
<subSubSection id="A5C6EFF5FFEFFF93FF7B31A2FE0475CC" pageId="11" pageNumber="402" type="etymology">
<paragraph id="ED63BC7EFFEFFF93FF7B31A2FE0475CC" blockId="11.[151,1437,150,681]" pageId="11" pageNumber="402">
<emphasis id="DFA8606CFFEFFF93FF7B31A2FECD75E8" bold="true" box="[199,334,439,465]" pageId="11" pageNumber="402">Etymology.</emphasis>
The species name refers to the name of 
<collectingCountry id="95CBFCEEFFEFFF93FCA331A2FC0D75E8" box="[799,910,439,465]" name="Sri Lanka" pageId="11" pageNumber="402">Sri Lanka</collectingCountry>
(previously known as 
<collectingCountry id="95CBFCEEFFEFFF93FB2F31A2FB6B75EB" box="[1171,1256,439,466]" name="Sri Lanka" pageId="11" pageNumber="402">Ceylon</collectingCountry>
), the country of origin of this species.
</paragraph>
</subSubSection>
<subSubSection id="A5C6EFF5FFEFFF93FF7B31EAFD097690" pageId="11" pageNumber="402" type="discussion">
<paragraph id="ED63BC7EFFEFFF93FF7B31EAFD097690" blockId="11.[151,1437,150,681]" pageId="11" pageNumber="402">
<emphasis id="DFA8606CFFEFFF93FF7B31EAFEB77620" bold="true" box="[199,308,511,537]" pageId="11" pageNumber="402">Remarks</emphasis>
. 
<taxonomicName id="2ADCC7FDFFEFFF93FEF831EAFDD37620" authority="Joharchi &amp; Halliday, 2021" authorityName="Joharchi &amp; Halliday" authorityYear="2021" box="[324,592,511,537]" class="Arachnida" family="Laelapidae" genus="Hyposternus" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="402" phylum="Arthropoda" rank="species" species="ceylonicus" status="sp. nov.">
<emphasis id="DFA8606CFFEFFF93FEF831EAFDD37620" box="[324,592,511,537]" italics="true" pageId="11" pageNumber="402">Hyposternus ceylonicus</emphasis>
</taxonomicName>
<emphasis id="DFA8606CFFEFFF93FDE531EAFD377620" bold="true" box="[601,692,511,537]" pageId="11" pageNumber="402">
<taxonomicNameLabel id="C49BDD17FFEFFF93FDE531EAFD377620" box="[601,692,511,537]" pageId="11" pageNumber="402" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may be distinguished from other laelapids by the diagnostic features of the genus mentioned above and further emphasized in the Discussion below. In addition, the short setae of the dorsal shield (especially in contrast to elongate Z5), including a single Jx seta and two pairs of Zx setae, and a faintly ornamented dorsal shield may aid in identifying the new species, including from any undescribed related species that may be discovered in the future.
</paragraph>
</subSubSection>
</treatment>
</document>