526 lines
67 KiB
XML
526 lines
67 KiB
XML
<document id="820B82E21DB9F1CBAECBC47F63D94876" ID-DOI="10.5281/zenodo.293962" ID-GBIF-Dataset="abcc806d-c833-4206-8841-cbeda487e635" ID-ISSN="1175-5326" ID-Zenodo-Dep="293962" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460227323868" checkinUser="plazi" docAuthor="Hill, Matthew A., Pfeiffer, John & Jacobus, Luke M." docDate="2010" docId="456D282AFFE2F463FF5EFB90FC5FCE4A" docLanguage="en" docName="zt02481p068.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Waynokiops dentatogriphus Hill, Pfeiffer & Jacobus, 2010, new species" docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="B9545052FFE3F465FFC9FFBFFFFFCC06" masterDocTitle="A new genus and new species of Baetidae (Ephemeroptera) from lakes and reservoirs in eastern North America" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698238654833" updateUser="plazi">
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<mods:title id="A9E8112E531A587DADD7B45369A5DC55">A new genus and new species of Baetidae (Ephemeroptera) from lakes and reservoirs in eastern North America</mods:title>
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</mods:titleInfo>
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<mods:name id="4AD5712EFE1A219F38955426ECFF6420" type="personal">
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<mods:namePart id="4655E10C769210CEF71A2DC96C6FC28C">Hill, Matthew A.</mods:namePart>
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<mods:namePart id="8F4A63040F802FF6AF71FF3A6F2662B1">Pfeiffer, John</mods:namePart>
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<mods:name id="F3D8117620B5FC43A4BAE624AB9E9489" type="personal">
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<mods:roleTerm id="5530F2A3A668F9599FC6D2BBB4E114B8">Author</mods:roleTerm>
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<mods:namePart id="26FAEC8F2A5BBB067FF79E09F629B572">Jacobus, Luke M.</mods:namePart>
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<mods:typeOfResource id="55529BEB6503016A45C12AC9182D238E">text</mods:typeOfResource>
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<mods:title id="D46E151CE101FB23238FCA15050CA050">Zootaxa</mods:title>
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<mods:part id="AF83E56200F98DF35839BE7850409F9C">
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<mods:date id="731E43D91536D5DC4750E6FB159A7529">2010</mods:date>
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<mods:detail id="FFA1300F4A48CEE8FAAFF288F3861FFF" type="volume">
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<mods:number id="552150A656E2976B781BFF6B73658BA6">2481</mods:number>
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<mods:classification id="039C3B44122C47D8B4B5A8FC6E9721A3">journal article</mods:classification>
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<mods:identifier id="A5F989CA32B6F8BC7D717455336C5BC8" type="DOI">10.5281/zenodo.293962</mods:identifier>
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<mods:identifier id="07603885B20C6594CAD1C239E453AB22" type="ISSN">1175-5326</mods:identifier>
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<treatment id="456D282AFFE2F463FF5EFB90FC5FCE4A" ID-DOI="http://doi.org/10.5281/zenodo.6203113" ID-GBIF-Taxon="119386718" ID-Zenodo-Dep="6203113" LSID="urn:lsid:plazi:treatment:456D282AFFE2F463FF5EFB90FC5FCE4A" httpUri="http://treatment.plazi.org/id/456D282AFFE2F463FF5EFB90FC5FCE4A" lastPageId="6" lastPageNumber="67" pageId="1" pageNumber="62">
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<subSubSection id="85DECAB7FFE2F464FF5EFB90FED9C86A" pageId="1" pageNumber="62" type="nomenclature">
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<paragraph id="CD7B993CFFE2F464FF5EFB90FD7BC84F" blockId="1.[151,644,1071,1132]" box="[151,644,1071,1097]" pageId="1" pageNumber="62">
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<heading id="96332E50FFE2F464FF5EFB90FD7BC84F" bold="true" box="[151,644,1071,1097]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
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<emphasis id="FFB0452EFFE2F464FF5EFB90FD7BC84F" bold="true" box="[151,644,1071,1097]" pageId="1" pageNumber="62">
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<taxonomicName id="0AC4E2BFFFE2F464FF5EFB90FE19C84F" ID-CoL="5BVH5" box="[151,486,1071,1097]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="dentatogriphus" status="sp. nov.">
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<emphasis id="FFB0452EFFE2F464FF5EFB90FE19C84F" bold="true" box="[151,486,1071,1097]" italics="true" pageId="1" pageNumber="62">Waynokiops dentatogriphus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="E483F855FFE2F464FE3CFB90FD7BC84F" box="[501,644,1071,1097]" pageId="1" pageNumber="62" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="CD7B993CFFE2F464FF5EFBEDFED9C86A" blockId="1.[151,644,1071,1132]" box="[151,294,1106,1132]" pageId="1" pageNumber="62">
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(
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||
<figureCitation id="55FF85B9FFE2F464FF56FBEDFEE3C86A" box="[159,284,1106,1132]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 9" captionStart-2="FIGURES 10 – 13" captionStartId-0="2.[151,275,1940,1964]" captionStartId-1="3.[151,270,1779,1803]" captionStartId-2="4.[151,271,1889,1913]" captionTargetBox-0="[209,1386,393,1898]" captionTargetBox-1="[292,1309,194,1754]" captionTargetBox-2="[193,1396,206,1857]" captionTargetId-0="figure@2.[199,1387,393,1905]" captionTargetId-1="figure@3.[281,1317,192,1757]" captionTargetId-2="figure@4.[190,1396,197,1865]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." captionText-1="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." captionText-2="FIGURES 10 – 13. Waynokiops dentatogriphus, new genus, new species. 10. Mesothoracic leg. 11. Tarsal claw; note double row of denticles restricted to proximal quarter of claw. 12. Gill VI. 13. Mesothoracic leg, patellotibial region." httpUri-0="https://zenodo.org/record/293963/files/figure.png" httpUri-1="https://zenodo.org/record/293964/files/figure.png" httpUri-2="https://zenodo.org/record/293965/files/figure.png" pageId="1" pageNumber="62">Figs. 1–13</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="85DECAB7FFE2F464FF5EFB22FDECC991" pageId="1" pageNumber="62" type="materials_examined">
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<paragraph id="CD7B993CFFE2F464FF5EFB22FB92C8DA" blockId="1.[151,1437,1181,2028]" pageId="1" pageNumber="62">
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<emphasis id="FFB0452EFFE2F464FF5EFB22FEB5C8B1" bold="true" box="[151,330,1181,1207]" pageId="1" pageNumber="62">
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<typeStatus id="127F279EFFE2F464FF5EFB22FF2DC8B1" box="[151,210,1181,1207]" pageId="1" pageNumber="62">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="127F279EFFE2F464FE9AFB22FE0DC8B1" box="[339,498,1181,1207]" pageId="1" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
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: nymph (preserved in ethanol), ARKANSAS, Johnson County, Horsehead Lake,
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<date id="B97ABFFCFFE2F464FF5EFB7DFECAC8DA" box="[151,309,1218,1244]" pageId="1" pageNumber="62" value="2007-08-26">26-VIII-2007</date>
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[
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<collectionCode id="ABD501F9FFE2F464FE8AFB7DFE5EC8DA" LSID="urn:lsid:biocol.org:col:34871" box="[323,417,1218,1244]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="62">USNM</collectionCode>
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].
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<typeStatus id="127F279EFFE2F464FE7AFB7DFDA5C8DA" box="[435,602,1218,1244]" pageId="1" pageNumber="62" type="paratype">PARATYPES</typeStatus>
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: five nymphs, same data as
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<typeStatus id="127F279EFFE2F464FC69FB7DFBF9C8DA" box="[928,1030,1218,1244]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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[
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<collectionCode id="ABD501F9FFE2F464FBDDFB7DFB9EC8DA" LSID="urn:lsid:biocol.org:col:34797" box="[1044,1121,1218,1244]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="1" pageNumber="62">INHS</collectionCode>
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].
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</paragraph>
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<paragraph id="CD7B993CFFE2F464FF0DFB58FDECC991" blockId="1.[151,1437,1181,2028]" pageId="1" pageNumber="62">
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<emphasis id="FFB0452EFFE2F464FF0DFB58FDF3C907" bold="true" box="[196,524,1255,1281]" pageId="1" pageNumber="62">Other material examined.</emphasis>
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<collectingCountry id="B5D3D9ACFFE2F464FDDDFB57FD96C904" box="[532,617,1256,1282]" name="United States of America" pageId="1" pageNumber="62">U.S.A.</collectingCountry>
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: INDIANA, Bartholomew County: Tipton Lakes,
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<date id="B97ABFFCFFE2F464FB0BFB57FA9BC904" box="[1218,1380,1256,1282]" pageId="1" pageNumber="62" value="2007-08-13">13-VIII-2007</date>
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, one nymph [
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<collectionCode id="ABD501F9FFE2F464FF33FAB2FEB0C921" box="[250,335,1293,1319]" pageId="1" pageNumber="62">PERC</collectionCode>
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]; Steuben County: Lake James,
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<date id="B97ABFFCFFE2F464FD10FAB2FC89C921" box="[729,886,1293,1319]" pageId="1" pageNumber="62" value="2007-07-24">24-VII-2007</date>
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, one nymph [
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<collectionCode id="ABD501F9FFE2F464FBD2FAB2FB8FC921" box="[1051,1136,1293,1319]" pageId="1" pageNumber="62">PERC</collectionCode>
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||
].
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<collectionCode id="ABD501F9FFE2F464FB4DFAB2FB2CC921" box="[1156,1235,1293,1319]" pageId="1" pageNumber="62">OHIO</collectionCode>
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, Brown County: Lake Waynoka,
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<date id="B97ABFFCFFE2F464FE96FA8DFE07C94A" box="[351,504,1330,1356]" pageId="1" pageNumber="62" value="2007-08-02">2-VIII-2007</date>
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, three nymphs [
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<collectionCode id="ABD501F9FFE2F464FD72FA8DFCF2C94A" box="[699,781,1330,1356]" pageId="1" pageNumber="62">PERC</collectionCode>
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,
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<collectionCode id="ABD501F9FFE2F464FCD2FA8DFC96C94A" LSID="urn:lsid:biocol.org:col:34797" box="[795,873,1330,1356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="1" pageNumber="62">INHS</collectionCode>
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]. VIRGINIA, Suffolk/Isle of Wight Counties: Burnt Mills Lake,
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<date id="B97ABFFCFFE2F464FEBEFAE7FDF8C974" box="[375,519,1368,1394]" pageId="1" pageNumber="62" value="2007-09-19">19-IX-2007</date>
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, one nymph [
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<collectionCode id="ABD501F9FFE2F464FD62FAE7FCF4C974" LSID="urn:lsid:biocol.org:col:34871" box="[683,779,1368,1394]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="62">USNM</collectionCode>
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||
]. KENTUCKY, Christian County: Pennyrile Lake,
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<date id="B97ABFFCFFE2F464FABDFAE7FEEFC991" pageId="1" pageNumber="62" value="2007-08-17">17- VIII-2007</date>
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, two nymphs [
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<collectionCode id="ABD501F9FFE2F464FE72FAC2FDF7C991" LSID="urn:lsid:biocol.org:col:34797" box="[443,520,1405,1431]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="1" pageNumber="62">INHS</collectionCode>
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].
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="85DECAB7FFE2F460FF0FFA1DFE6FCDB1" lastPageId="5" lastPageNumber="66" pageId="1" pageNumber="62" type="description">
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<paragraph id="CD7B993CFFE2F464FF0FFA1DFB5DC9E4" blockId="1.[151,1437,1181,2028]" pageId="1" pageNumber="62">
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<emphasis id="FFB0452EFFE2F464FF0FFA1DFEA8C9BA" bold="true" box="[198,343,1442,1468]" pageId="1" pageNumber="62">Description</emphasis>
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.
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<emphasis id="FFB0452EFFE2F464FEA5FA1CFD21C9BA" box="[364,734,1443,1468]" italics="true" pageId="1" pageNumber="62">Nymph (preserved in ethanol)</emphasis>
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. Body length of final instar: 4.5 mm, excluding caudal filaments. Color: White with tiny, dark brown maculations scattered over dorsal surface.
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</paragraph>
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<paragraph id="CD7B993CFFE2F464FF0DFA52FB7DCB7A" blockId="1.[151,1437,1181,2028]" pageId="1" pageNumber="62">
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Head (
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<figureCitation id="55FF85B9FFE2F464FEDBFA52FEAACA01" box="[274,341,1517,1543]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
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) narrow, hypognathous, coronal suture acute. Eyes protuberant, protruding beyond outline of head in anterior view (
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<figureCitation id="55FF85B9FFE2F464FE63F9ADFE14CA2A" box="[426,491,1554,1580]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="1" pageNumber="62">Fig.1</figureCitation>
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||
). Antennae 2.5 times length of head. Pedicel and scape cylidrical, subequal in length. Labrum (
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<figureCitation id="55FF85B9FFE2F464FE93F987FE5FCA54" box="[346,416,1592,1618]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
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) with shallow median excavation on distal edge. Mandibles (
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||
<figureCitation id="55FF85B9FFE2F464FBA2F987FB26CA54" box="[1131,1241,1592,1618]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="1" pageNumber="62">Figs. 3, 4</figureCitation>
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) incisors cleft to base. Maxillae (
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||
<figureCitation id="55FF85B9FFE2F464FE9DF9E2FE64CA71" box="[340,411,1629,1655]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="1" pageNumber="62">Fig. 5</figureCitation>
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||
) with palp narrow, attenuate and two-segmented. Lacinia stout, pointed, with comb of thick setae on distal half of medial edge below apical tooth. Apical tooth of lacinia 0.4 times length of lacinia. Labium (
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||
<figureCitation id="55FF85B9FFE2F464FEC1F917FEABCAC4" box="[264,340,1704,1730]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="1" pageNumber="62">Fig. 6</figureCitation>
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||
) with paraglossae slightly longer than glossae. Glossae convex laterally, medial margin straight in proximal half, concave distally, apex bluntly pointed. Paraglossae round at apex. Labial palp (
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||
<figureCitation id="55FF85B9FFE2F464FAA5F972FF58CB0A" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="1" pageNumber="62">Fig. 7</figureCitation>
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||
) segment 1 rectangular, as long as segments 2 and 3 combined, and with peg-like setae on lateral surface. Segment two glabrous and 0.5 times length of segment 1. Segment 3 distally explanate and truncate; lateral and distal margins densely setose; subequal in length to segment 2, with distal width 1.5 times proximal width; apicolateral corner with blunt, triangular tooth; tooth length subequal to width.
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||
</paragraph>
|
||
<paragraph id="CD7B993CFFE2F467FF5EF837FEDDCD41" blockId="1.[151,1437,1181,2028]" lastBlockId="2.[151,1437,152,327]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
|
||
Pronotum (
|
||
<figureCitation id="55FF85B9FFE2F464FED4F837FE70CBA4" box="[285,399,1928,1954]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="1" pageNumber="62">Figs. 8, 9</figureCitation>
|
||
) width 3.5 times length, with two pairs of low mounds, one pair sublaterad of midline and one pair near lateral margins. Mesonotum (
|
||
<figureCitation id="55FF85B9FFE2F464FD08F812FCD0CBC1" box="[705,815,1965,1991]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="1" pageNumber="62">Figs. 8, 9</figureCitation>
|
||
) roughly sculptured, with median, longitudinal ridge slightly projected in lateral aspect.
|
||
<collectingCountry id="B5D3D9ACFFE2F464FDE7F86DFD96CBEA" box="[558,617,2002,2028]" name="India" pageId="1" pageNumber="62">Hind</collectingCountry>
|
||
wingpads absent. Legs (
|
||
<figureCitation id="55FF85B9FFE2F464FC41F86DFC22CBEA" box="[904,989,2002,2028]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,271,1889,1913]" captionTargetBox="[193,1396,206,1857]" captionTargetId="figure@4.[190,1396,197,1865]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Waynokiops dentatogriphus, new genus, new species. 10. Mesothoracic leg. 11. Tarsal claw; note double row of denticles restricted to proximal quarter of claw. 12. Gill VI. 13. Mesothoracic leg, patellotibial region." httpUri="https://zenodo.org/record/293965/files/figure.png" pageId="1" pageNumber="62">Fig. 10</figureCitation>
|
||
) with femur 5 times longer than wide; femur with ventral row of short, lanceolate setae and with sparse distribution of fine setae dorsally. Patellotibial region of mesothoracic leg as in
|
||
<figureCitation id="55FF85B9FFE1F467FD7CFF02FCD4CCD1" box="[693,811,189,215]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,271,1889,1913]" captionTargetBox="[193,1396,206,1857]" captionTargetId="figure@4.[190,1396,197,1865]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Waynokiops dentatogriphus, new genus, new species. 10. Mesothoracic leg. 11. Tarsal claw; note double row of denticles restricted to proximal quarter of claw. 12. Gill VI. 13. Mesothoracic leg, patellotibial region." httpUri="https://zenodo.org/record/293965/files/figure.png" pageId="2" pageNumber="63">Figure 13</figureCitation>
|
||
. Tibia and tarsus subequal in length and width, each bearing ventral row of pectinate, lanceolate setae, and with sparse distribution of dorsal fine setae. Tarsal claw slightly curved, subequal in length to tarsus, with two rows of tiny denticles (
|
||
<figureCitation id="55FF85B9FFE1F467FBFDFEB7FB75CD24" box="[1076,1162,264,290]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,271,1889,1913]" captionTargetBox="[193,1396,206,1857]" captionTargetId="figure@4.[190,1396,197,1865]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Waynokiops dentatogriphus, new genus, new species. 10. Mesothoracic leg. 11. Tarsal claw; note double row of denticles restricted to proximal quarter of claw. 12. Gill VI. 13. Mesothoracic leg, patellotibial region." httpUri="https://zenodo.org/record/293965/files/figure.png" pageId="2" pageNumber="63">Fig. 11</figureCitation>
|
||
) restricted to proximal 0.25 length.
|
||
</paragraph>
|
||
<caption id="99BBC9B4FFE1F467FF5EF82BFAAFCBCD" httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="2" pageNumber="63" targetBox="[209,1386,393,1898]" targetPageId="2">
|
||
<paragraph id="CD7B993CFFE1F467FF5EF82BFAAFCBCD" blockId="2.[151,1437,1940,1996]" pageId="2" pageNumber="63">
|
||
<emphasis id="FFB0452EFFE1F467FF5EF82BFEB1CBAA" bold="true" box="[151,334,1940,1964]" pageId="2" pageNumber="63">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="0AC4E2BFFFE1F467FE9CF82AFD6CCBAA" box="[341,659,1941,1964]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="dentatogriphus" status="gen. nov.">
|
||
<emphasis id="FFB0452EFFE1F467FE9CF82AFD6CCBAA" box="[341,659,1941,1964]" italics="true" pageId="2" pageNumber="63">Waynokiops dentatogriphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="FFB0452EFFE1F467FD6AF82BFC1CCBAA" bold="true" box="[675,995,1940,1964]" pageId="2" pageNumber="63">
|
||
<taxonomicNameLabel id="E483F855FFE1F467FD6AF82BFCDCCBAA" box="[675,803,1940,1964]" pageId="2" pageNumber="63" rank="genus">new genus</taxonomicNameLabel>
|
||
, new species. 1.
|
||
</emphasis>
|
||
Head, anterior view, cleared and with mouthparts removed.
|
||
<emphasis id="FFB0452EFFE1F467FE49F80BFE6BCBCA" bold="true" box="[384,404,1972,1996]" pageId="2" pageNumber="63">2.</emphasis>
|
||
Labrum.
|
||
<emphasis id="FFB0452EFFE1F467FE35F80BFDF0CBCA" bold="true" box="[508,527,1972,1996]" pageId="2" pageNumber="63">3.</emphasis>
|
||
Right (planate) mandible.
|
||
<emphasis id="FFB0452EFFE1F467FCE6F80BFCBDCBCA" bold="true" box="[815,834,1972,1996]" pageId="2" pageNumber="63">4.</emphasis>
|
||
Left (angulate) mandible.
|
||
<emphasis id="FFB0452EFFE1F467FBA9F80BFB8BCBCA" bold="true" box="[1120,1140,1972,1996]" pageId="2" pageNumber="63">5.</emphasis>
|
||
Maxilla.
|
||
<emphasis id="FFB0452EFFE1F467FB12F80BFB10CBCA" bold="true" box="[1243,1263,1972,1996]" pageId="2" pageNumber="63">6.</emphasis>
|
||
Labium.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="99BBC9B4FFE0F466FF5EF94CFEE0CB2D" httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="3" pageNumber="64" targetBox="[292,1309,194,1754]" targetPageId="3">
|
||
<paragraph id="CD7B993CFFE0F466FF5EF94CFEE0CB2D" blockId="3.[151,1435,1779,1835]" pageId="3" pageNumber="64">
|
||
<emphasis id="FFB0452EFFE0F466FF5EF94CFEBBCB0D" bold="true" box="[151,324,1779,1803]" pageId="3" pageNumber="64">FIGURES 7–9.</emphasis>
|
||
<taxonomicName id="0AC4E2BFFFE0F466FE82F94BFD8BCB0D" box="[331,628,1780,1803]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="dentatogriphus" status="gen. nov.">
|
||
<emphasis id="FFB0452EFFE0F466FE82F94BFD8BCB0D" box="[331,628,1780,1803]" italics="true" pageId="3" pageNumber="64">Waynokiops dentatogriphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="FFB0452EFFE0F466FD4BF94CFC53CB0D" bold="true" box="[642,940,1779,1803]" pageId="3" pageNumber="64">
|
||
<taxonomicNameLabel id="E483F855FFE0F466FD4BF94CFD05CB0D" box="[642,762,1779,1803]" pageId="3" pageNumber="64" rank="genus">new genus</taxonomicNameLabel>
|
||
, new species. 7.
|
||
</emphasis>
|
||
Labial palp.
|
||
<emphasis id="FFB0452EFFE0F466FBF5F94CFBAFCB0D" bold="true" box="[1084,1104,1779,1803]" pageId="3" pageNumber="64">8.</emphasis>
|
||
Dorsal view, whole.
|
||
<emphasis id="FFB0452EFFE0F466FAFCF94CFAB6CB0D" bold="true" box="[1333,1353,1779,1803]" pageId="3" pageNumber="64">9.</emphasis>
|
||
Lateral view, whole.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CD7B993CFFE0F460FF0DF8EBFA7CCD94" blockId="3.[151,1436,1876,2014]" lastBlockId="5.[151,1437,152,1036]" lastPageId="5" lastPageNumber="66" pageId="3" pageNumber="64">
|
||
Abdomen dorsoventrally compressed. Segments 1–9 each with dark, transverse band on lateral margins (most apparent on later instars). Triangular, acute, posteriorly-directed lateral expansions present on segments 1–7 and parallel-sided lateral flanges present on segments 8–9 (
|
||
<figureCitation id="55FF85B9FFE0F466FC45F821FC2BCBBE" box="[908,980,1950,1976]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="3" pageNumber="64">Fig. 8</figureCitation>
|
||
). Lateral processes on segment 7 with stout spinules on posterior margins. Lateral expansions of segments 8 and 9 with stout spinules on posterior half of lateral margins and on free posterior margins. Such stout spinules sometimes present on more anterior segments. Single, median, erect, posteriorly hooked spines present on terga 1–6, decreasing in relative development posteriorly (
|
||
<figureCitation id="55FF85B9FFE6F460FE1CFF5DFDDECCFA" box="[469,545,226,252]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="5" pageNumber="66">Fig. 9</figureCitation>
|
||
). Broad, bilamellate gills (
|
||
<figureCitation id="55FF85B9FFE6F460FCA7FF5DFC37CCFA" box="[878,968,226,252]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,271,1889,1913]" captionTargetBox="[193,1396,206,1857]" captionTargetId="figure@4.[190,1396,197,1865]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Waynokiops dentatogriphus, new genus, new species. 10. Mesothoracic leg. 11. Tarsal claw; note double row of denticles restricted to proximal quarter of claw. 12. Gill VI. 13. Mesothoracic leg, patellotibial region." httpUri="https://zenodo.org/record/293965/files/figure.png" pageId="5" pageNumber="66">Fig. 12</figureCitation>
|
||
) present on abdominal segments 1–7. Dorsal lamella of gills round, approximately half size of lower lamella. Lower lamella triangular, with broken, branched venation. Cerci and median filament (
|
||
<figureCitation id="55FF85B9FFE6F460FD25FE92FCC8CD41" box="[748,823,301,327]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="5" pageNumber="66">Fig. 8</figureCitation>
|
||
) pale with numerous, narrow faint darker bands. Median filament with lateral setal fringes continuous throughout length. Cerci with medial setal fringe continuous throughout length. Caudal filaments subequal in length to each other, and to length of abdomen.
|
||
</paragraph>
|
||
<caption id="99BBC9B4FFE7F461FF5EF8DEFA8DCB9E" httpUri="https://zenodo.org/record/293965/files/figure.png" pageId="4" pageNumber="65" targetBox="[193,1396,206,1857]" targetPageId="4">
|
||
<paragraph id="CD7B993CFFE7F461FF5EF8DEFA8DCB9E" blockId="4.[151,1436,1889,1945]" pageId="4" pageNumber="65">
|
||
<emphasis id="FFB0452EFFE7F461FF5EF8DEFE9DCB7F" bold="true" box="[151,354,1889,1913]" pageId="4" pageNumber="65">FIGURES 10–13.</emphasis>
|
||
<taxonomicName id="0AC4E2BFFFE7F461FEA0F8DDFD67CB7F" box="[361,664,1890,1913]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="dentatogriphus" status="gen. nov.">
|
||
<emphasis id="FFB0452EFFE7F461FEA0F8DDFD67CB7F" box="[361,664,1890,1913]" italics="true" pageId="4" pageNumber="65">Waynokiops dentatogriphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="FFB0452EFFE7F461FD6EF8DEFC1ACB7F" bold="true" box="[679,997,1889,1913]" pageId="4" pageNumber="65">
|
||
<taxonomicNameLabel id="E483F855FFE7F461FD6EF8DEFCDECB7F" box="[679,801,1889,1913]" pageId="4" pageNumber="65" rank="genus">new genus</taxonomicNameLabel>
|
||
, new species. 10.
|
||
</emphasis>
|
||
Mesothoracic leg.
|
||
<emphasis id="FFB0452EFFE7F461FB71F8DEFB26CB7F" bold="true" box="[1208,1241,1889,1913]" pageId="4" pageNumber="65">11.</emphasis>
|
||
Tarsal claw; note double row of denticles restricted to proximal quarter of claw.
|
||
<emphasis id="FFB0452EFFE7F461FCF8F83EFCADCB9F" bold="true" box="[817,850,1921,1945]" pageId="4" pageNumber="65">12.</emphasis>
|
||
Gill VI.
|
||
<emphasis id="FFB0452EFFE7F461FC7AF83EFC2BCB9F" bold="true" box="[947,980,1921,1945]" pageId="4" pageNumber="65">13.</emphasis>
|
||
Mesothoracic leg, patellotibial region.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CD7B993CFFE6F460FF0FFE22FE6FCDB1" blockId="5.[151,1437,152,1036]" box="[198,400,413,439]" pageId="5" pageNumber="66">
|
||
<emphasis id="FFB0452EFFE6F460FF0FFE22FEF1CDB0" box="[198,270,413,438]" italics="true" pageId="5" pageNumber="66">Adult.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="85DECAB7FFE6F460FF0DFE7DFE69CF2A" pageId="5" pageNumber="66" type="etymology">
|
||
<paragraph id="CD7B993CFFE6F460FF0DFE7DFA98CE21" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">
|
||
<emphasis id="FFB0452EFFE6F460FF0DFE7DFEA8CDDA" bold="true" box="[196,343,450,476]" pageId="5" pageNumber="66">Etymology.</emphasis>
|
||
The specific epithet is a combination of the Latin word
|
||
<emphasis id="FFB0452EFFE6F460FBDEFE7DFB7FCDDD" box="[1047,1152,450,475]" italics="true" pageId="5" pageNumber="66">dentatus</emphasis>
|
||
, meaning toothed, and
|
||
<emphasis id="FFB0452EFFE6F460FF5EFE57FF0DCE07" box="[151,242,488,513]" italics="true" pageId="5" pageNumber="66">griphus</emphasis>
|
||
, meaning riddle. This name is a reference to the prominent median protuberances on the abdominal terga, and to the puzzling morphology and phylogenetic position of the species. The gender is masculine.
|
||
</paragraph>
|
||
<paragraph id="CD7B993CFFE6F460FF0DFD8DFE69CF2A" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">
|
||
<emphasis id="FFB0452EFFE6F460FF0DFD8DFDD8CE4A" bold="true" box="[196,551,562,588]" pageId="5" pageNumber="66">Morphological variability.</emphasis>
|
||
The stout spinules of the lateral and sometimes posterior margins of abdominal terga vary in the size and number per segment. The spinules are smaller and fewer in number on more anterior segments. One specimen (
|
||
<collectingCountry id="B5D3D9ACFFE6F460FDBFFDC2FD36CE91" box="[630,713,637,663]" name="United States of America" pageId="5" pageNumber="66">U.S.A.</collectingCountry>
|
||
, Indiana, Bartholomew County) has such spinules present as far anteriorly as segment two; however, most specimens do not have them appear this far anteriorly. The spinules usually begin to appear around segment 6. Later instars have larger and more numerous spinules. Likewise, the single, median, erect, posteriorly hooked spines present on the abdominal terga vary in their relative development.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="85DECAB7FFE6F460FF0DFC88FA63C8F2" pageId="5" pageNumber="66" type="diagnosis">
|
||
<paragraph id="CD7B993CFFE6F460FF0DFC88FC9ECFC4" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">
|
||
<emphasis id="FFB0452EFFE6F460FF0DFC88FEBBCF57" bold="true" box="[196,324,823,849]" pageId="5" pageNumber="66">Diagnosis.</emphasis>
|
||
The monotypic new genus is distinguishable from all other
|
||
<taxonomicName id="0AC4E2BFFFE6F460FBD9FC87FB88CF54" box="[1040,1143,824,850]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
by having the following combination of characters: somewhat truncate and expanded labial palp segment 3 (
|
||
<figureCitation id="55FF85B9FFE6F460FB09FCE2FAF1CF71" box="[1216,1294,861,887]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="5" pageNumber="66">Fig. 7</figureCitation>
|
||
), variously developed lateral spinules on the abdomen (
|
||
<bibRefCitation id="A955E4CDFFE6F460FD55FC3DFC4BCF9A" author="Gillies" box="[668,948,898,924]" pageId="5" pageNumber="67" refString="Gillies, M. T. (1990) A revision of the African species of Centroptilum Eaton (Baetidae, Ephemeroptera). Aquatic Insects, 12, 97 - 128." type="journal article" year="1990">
|
||
Gillies, 1990:
|
||
<figureCitation id="55FF85B9FFE6F460FC8CFC3DFC4BCF9A" box="[837,948,898,924]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="5" pageNumber="66">Figs. 2, 3</figureCitation>
|
||
</bibRefCitation>
|
||
), laterally expanded abdominal segments (
|
||
<figureCitation id="55FF85B9FFE6F460FF56FC17FF1ACFC4" box="[159,229,936,962]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="5" pageNumber="66">Fig. 8</figureCitation>
|
||
) and medial armature on the abdominal terga (
|
||
<figureCitation id="55FF85B9FFE6F460FCC2FC17FCACCFC4" box="[779,851,936,962]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="5" pageNumber="66">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CD7B993CFFE6F460FF0DFC72FA9FC80A" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">
|
||
We modify a recent and widely-used key to genera of North American mayfly nymphs (
|
||
<bibRefCitation id="A955E4CDFFE6F460FB11FC72FF2CC80A" author="Waltz" pageId="5" pageNumber="68" refString="Waltz, R. D. & Burian, S. K. (2008) Chapter 11, Ephemeroptera. In: Merritt, R. W., Cummins, K. W. & Berg, M. B. (Eds.). An Introduction to the Aquatic Insects of North America, Fouth Edition. Kendall / Hunt, Dubuque, Iowa, pp. 181 - 236." type="book chapter" year="2008">Waltz & Burian, 2008</bibRefCitation>
|
||
) to accommodate
|
||
<taxonomicName id="0AC4E2BFFFE6F460FE62FC4DFDC8C80D" box="[427,567,1010,1035]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FFB0452EFFE6F460FE62FC4DFDC8C80D" box="[427,567,1010,1035]" italics="true" pageId="5" pageNumber="66">Waynokiops</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E483F855FFE6F460FD8FFC4DFD3AC80A" box="[582,709,1010,1036]" pageId="5" pageNumber="66" rank="genus">new genus</taxonomicNameLabel>
|
||
, as follows, because an impasse is reached at couplet 24.
|
||
</paragraph>
|
||
<paragraph id="CD7B993CFFE6F460FF5EFB82FA63C892" blockId="5.[151,1436,1085,1268]" pageId="5" pageNumber="66">24 (23’). Median tubercles present on anterior abdominal terga..................................................................................... 24a 24’. No median tubercles on abdominal terga ............................................................................................................ 25 24a. Abdominal gills (or gill sockets) present on segments 1–5 only; gills extend ventrally from the pleura; South-</paragraph>
|
||
<paragraph id="CD7B993CFFE6F460FF3CFB22FA67C8D2" blockId="5.[151,1436,1085,1268]" pageId="5" pageNumber="66">
|
||
west, Texas, Utah ...................................................................................................
|
||
<taxonomicName id="0AC4E2BFFFE6F460FB9CFB22FA63C8B2" authority="Needham & Murphy" authorityName="Needham & Murphy" box="[1109,1436,1181,1204]" class="Insecta" family="Baetidae" genus="Baetodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FB9CFB22FB48C8B2" box="[1109,1207,1181,1204]" italics="true" pageId="5" pageNumber="66">Baetodes</emphasis>
|
||
Needham & Murphy
|
||
</taxonomicName>
|
||
24a’. Abdominal gills (or gill sockets) present on segments 1–7; gills held dorsally; Central and East .........................
|
||
</paragraph>
|
||
<paragraph id="CD7B993CFFE6F460FF34FB62FA63C8F2" blockId="5.[151,1436,1085,1268]" box="[253,1436,1245,1268]" pageId="5" pageNumber="66">
|
||
<emphasis id="FFB0452EFFE6F460FF34FB62FAE2C8F2" box="[253,1309,1245,1268]" italics="true" pageId="5" pageNumber="66">
|
||
..........................................................................................................................................
|
||
<taxonomicName id="0AC4E2BFFFE6F460FB54FB62FAE2C8F2" box="[1181,1309,1245,1268]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">Waynokiops</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicNameLabel id="E483F855FFE6F460FAE3FB62FA63C8F2" box="[1322,1436,1245,1268]" pageId="5" pageNumber="66" rank="genus">new genus</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="85DECAB7FFE6F463FF0FFA9DFC5FCE4A" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="discussion">
|
||
<paragraph id="CD7B993CFFE6F460FF0FFA9DFE2DCB41" blockId="5.[151,1437,1314,2012]" pageId="5" pageNumber="66">
|
||
<emphasis id="FFB0452EFFE6F460FF0FFA9DFEADC93A" bold="true" box="[198,338,1314,1340]" pageId="5" pageNumber="66">Discussion.</emphasis>
|
||
The new genus is part of the
|
||
<taxonomicName id="0AC4E2BFFFE6F460FD71FA9CFCA2C93A" authority="Leach" authorityName="Leach" box="[696,861,1314,1340]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FD71FA9CFCF3C93A" box="[696,780,1315,1340]" italics="true" pageId="5" pageNumber="66">Cloeon</emphasis>
|
||
Leach
|
||
</taxonomicName>
|
||
complex of genera. This possibly monophyletic group (reviewed in parts by
|
||
<bibRefCitation id="A955E4CDFFE6F460FE29FAF7FCF3C964" author="Kluge" box="[480,780,1352,1378]" pageId="5" pageNumber="67" refString="Kluge, N. & Novikova, E. A. (1992) Revision of Palearctic genera and subgenera of mayflies in the subfamily Cloeoninae (Ephemeroptera, Baetidae) with descriptions of new species from the USSR. Entomologicheskoye Obozreniye, 71, 60 - 83." type="journal article" year="1992">Kluge & Novikova, 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE6F460FCDEFAF7FBDFC964" author="Monaghan" box="[791,1056,1352,1378]" pageId="5" pageNumber="67" refString="Monaghan, M. T., Gattolliat, J. - L., Sartori, M., Elouard, J. - M., James, H., Derleth, P., Glaizot, O., de Moor, P. F., & Vogler, A. P. (2005) Trans-oceanic and endemic origins of the small minnow mayflies (Ephemeroptera, Baetidae) of Madagascar. Proceedings of the Royal Society, B, 272, 1829 - 1836." type="journal article" year="2005">Monaghan et al., 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE6F460FBE5FAF7FAD9C964" author="Gattolliat" box="[1068,1318,1352,1378]" pageId="5" pageNumber="67" refString="Gattolliat, J. - L., Monaghan, M. T., Sartori, M., Elouard, J. - M., Barber-James, H., Derleth, P., Glaizot, O., de Moor, F. & Vogler, A. P. (2008) A Molecular analysis of the Afrotropical Baetidae. In: Hauer, F. R., Stanford, J. A., & Newell, R. L. (Eds.), International Advances in the Ecology, Zoogeography and Systematics of Mayflies and Stoneflies. University of California Publications in Entomology, vol. 128, pp. 219 - 232." type="journal article" year="2008">Gattolliat et al., 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE6F460FAFBFAF7FEF1C981" author="Ogden" lastPageId="5" lastPageNumber="68" pageId="5" pageNumber="67" refString="Ogden, T. H., Gattolliat, J. L., Sartori, M., Staniczek, A. H., Soldan, T. & Whiting, M. F. (2009) Towards a new paradigm in mayfly phylogeny (Ephemeroptera): Combined analysis of morphological and molecular data. Systematic Entomology, 34, 616 - 634." type="journal article" year="2009">Ogden et al., 2009</bibRefCitation>
|
||
:
|
||
<figureCitation id="55FF85B9FFE6F460FED3FAD2FE69C981" box="[282,406,1389,1415]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="5" pageNumber="66">Figs. 3, 6</figureCitation>
|
||
) is defined here to include those
|
||
<taxonomicName id="0AC4E2BFFFE6F460FC80FAD2FC46C981" box="[841,953,1389,1415]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
that have nymphs with the following synapomorphies (
|
||
<bibRefCitation id="A955E4CDFFE6F460FEBDFA2DFD2FC9AA" author="Kluge" box="[372,720,1426,1452]" pageId="5" pageNumber="67" refString="Kluge, N. & Novikova, E. A. (1992) Revision of Palearctic genera and subgenera of mayflies in the subfamily Cloeoninae (Ephemeroptera, Baetidae) with descriptions of new species from the USSR. Entomologicheskoye Obozreniye, 71, 60 - 83." type="journal article" year="1992">Kluge and Novikova, 1992</bibRefCitation>
|
||
): an at least somewhat truncate and expanded labial palp segment 3 (similar to
|
||
<figureCitation id="55FF85B9FFE6F460FE51FA07FE03C9D4" box="[408,508,1464,1490]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,270,1779,1803]" captionTargetBox="[292,1309,194,1754]" captionTargetId="figure@3.[281,1317,192,1757]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." httpUri="https://zenodo.org/record/293964/files/figure.png" pageId="5" pageNumber="66">Figure 7</figureCitation>
|
||
) and variously developed lateral spinules on the abdomen (
|
||
<bibRefCitation id="A955E4CDFFE6F460FB70FA07FF3BC9F1" author="Gillies" pageId="5" pageNumber="67" refString="Gillies, M. T. (1990) A revision of the African species of Centroptilum Eaton (Baetidae, Ephemeroptera). Aquatic Insects, 12, 97 - 128." type="journal article" year="1990">
|
||
Gillies, 1990:
|
||
<figureCitation id="55FF85B9FFE6F460FAABFA07FF3BC9F1" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="5" pageNumber="66">Figs. 2, 3</figureCitation>
|
||
</bibRefCitation>
|
||
). These spinules sometimes are minute and restricted to only one or two segments. Caudal filaments with dark rings on about every fourth segment may be another synapomorphy of the
|
||
<taxonomicName id="0AC4E2BFFFE6F460FB49F9BCFB25CA1A" box="[1152,1242,1539,1564]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FB49F9BCFB25CA1A" box="[1152,1242,1539,1564]" italics="true" pageId="5" pageNumber="66">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
complex genus groups, but our few nymphs with intact caudal filaments were faded, and we could not evaluate the character with confidence. Worldwide, the
|
||
<taxonomicName id="0AC4E2BFFFE6F460FD80F9F2FD5ACA60" box="[585,677,1613,1638]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FD80F9F2FD5ACA60" box="[585,677,1613,1638]" italics="true" pageId="5" pageNumber="66">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
complex includes
|
||
<taxonomicName id="0AC4E2BFFFE6F460FC52F9F2FBCDCA60" box="[923,1074,1613,1638]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FFB0452EFFE6F460FC52F9F2FBCDCA60" box="[923,1074,1613,1638]" italics="true" pageId="5" pageNumber="66">Waynokiops</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E483F855FFE6F460FB8FF9F2FB2DCA61" box="[1094,1234,1613,1639]" pageId="5" pageNumber="66" rank="genus">new genus</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FB28F9F2FA67CA61" authority="Leach" authorityName="Leach" box="[1249,1432,1613,1639]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FB28F9F2FAC2CA60" box="[1249,1341,1613,1638]" italics="true" pageId="5" pageNumber="66">Cloeon</emphasis>
|
||
Leach
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FF5EF9CCFE7ECA8A" authority="Eaton" authorityName="Eaton" box="[151,385,1650,1676]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FF5EF9CCFECBCA8A" box="[151,308,1651,1676]" italics="true" pageId="5" pageNumber="66">Centroptilum</emphasis>
|
||
Eaton
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="0AC4E2BFFFE6F460FE6AF9CCFD81CA8A" authority="Traver" authorityName="Traver" box="[419,638,1650,1676]" class="Insecta" family="Baetidae" genus="Neocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FE6AF9CCFDDDCA8A" box="[419,546,1651,1676]" italics="true" pageId="5" pageNumber="66">Neocloeon</emphasis>
|
||
Traver
|
||
</taxonomicName>
|
||
],
|
||
<taxonomicName id="0AC4E2BFFFE6F460FD58F9CCFBEACA8A" authority="Kluge & Novikova" authorityName="Kluge & Novikova" box="[657,1045,1650,1676]" class="Insecta" family="Baetidae" genus="Similicloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FD58F9CCFCDACA8A" box="[657,805,1651,1676]" italics="true" pageId="5" pageNumber="66">Similicloeon</emphasis>
|
||
Kluge & Novikova
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FBE8F9CCFA6BCA8A" authority="Kluge & Novikova" authorityName="Kluge & Novikova" box="[1057,1428,1650,1676]" class="Insecta" family="Baetidae" genus="Intercloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FBE8F9CCFB57CA8A" box="[1057,1192,1651,1676]" italics="true" pageId="5" pageNumber="66">Intercloeon</emphasis>
|
||
Kluge & Novikova
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FF5EF927FDDBCAB4" authority="Bogoescu" authorityName="Bogoescu" box="[151,548,1688,1714]" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FF5EF927FE66CAB7" box="[151,409,1688,1713]" italics="true" pageId="5" pageNumber="66">Pseudocentroptilum</emphasis>
|
||
Bogoescu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FDFBF927FCB7CAB4" authority="Bengtsson" authorityName="Bengtsson" box="[562,840,1688,1714]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FDFBF927FD49CAB7" box="[562,694,1688,1713]" italics="true" pageId="5" pageNumber="66">Procloeon</emphasis>
|
||
Bengtsson
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FC9EF927FB36CAB4" authority="Jacob" authorityName="Jacob" box="[855,1225,1688,1714]" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FC9EF927FB88CAB7" box="[855,1143,1688,1713]" italics="true" pageId="5" pageNumber="66">Pseudocentroptiloides</emphasis>
|
||
Jacob
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="0AC4E2BFFFE6F460FB39F927FEDBCAD1" authority="Glazaczow" authorityName="Glazaczow" class="Insecta" family="Baetidae" genus="Psammonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FB39F927FA62CAB7" box="[1264,1437,1688,1713]" italics="true" pageId="5" pageNumber="66">Psammonella</emphasis>
|
||
Glazaczow
|
||
</taxonomicName>
|
||
] and
|
||
<taxonomicName id="0AC4E2BFFFE6F460FEA0F902FC45CAD1" authority="Jacobus, McCafferty & Gattolliat. In" authorityName="Jacobus, McCafferty & Gattolliat. In" box="[361,954,1725,1751]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FEA0F902FE1BCAD0" box="[361,484,1725,1750]" italics="true" pageId="5" pageNumber="66">Securiops</emphasis>
|
||
Jacobus, McCafferty & Gattolliat. In
|
||
</taxonomicName>
|
||
North
|
||
<collectingCountry id="B5D3D9ACFFE6F460FBDAF902FB7ECAD1" box="[1043,1153,1725,1751]" name="United States of America" pageId="5" pageNumber="66">America</collectingCountry>
|
||
, species of the
|
||
<taxonomicName id="0AC4E2BFFFE6F460FA8AF902FA63CAD0" box="[1347,1436,1725,1750]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FA8AF902FA63CAD0" box="[1347,1436,1725,1750]" italics="true" pageId="5" pageNumber="66">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
complex are classified into the genera
|
||
<taxonomicName id="0AC4E2BFFFE6F460FDB9F95CFD38CAFA" box="[624,711,1763,1788]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FDB9F95CFD38CAFA" box="[624,711,1763,1788]" italics="true" pageId="5" pageNumber="66">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FD11F95CFC86CAFA" box="[728,889,1763,1788]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FD11F95CFC86CAFA" box="[728,889,1763,1788]" italics="true" pageId="5" pageNumber="66">Centroptilum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FC40F95CFBF9CAFA" box="[905,1030,1763,1788]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FC40F95CFBF9CAFA" box="[905,1030,1763,1788]" italics="true" pageId="5" pageNumber="66">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FBDEF95CFAD9CAFA" box="[1047,1318,1763,1788]" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FBDEF95CFAD9CAFA" box="[1047,1318,1763,1788]" italics="true" pageId="5" pageNumber="66">Pseudocentroptiloides</emphasis>
|
||
</taxonomicName>
|
||
and now,
|
||
<taxonomicName id="0AC4E2BFFFE6F460FF5EF8B7FECCCB27" box="[151,307,1800,1825]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FFB0452EFFE6F460FF5EF8B7FECCCB27" box="[151,307,1800,1825]" italics="true" pageId="5" pageNumber="66">Waynokiops</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E483F855FFE6F460FE8FF8B7FE29CB24" box="[326,470,1800,1826]" pageId="5" pageNumber="66" rank="genus">new genus</taxonomicNameLabel>
|
||
. Globally, the rank and application of these names varies among current
|
||
<taxonomicName id="0AC4E2BFFFE6F460FF5EF892FEB5CB41" box="[151,330,1837,1863]" class="Insecta" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="order">Ephemeroptera</taxonomicName>
|
||
specialists.
|
||
</paragraph>
|
||
<paragraph id="CD7B993CFFE6F463FF0FF8EDFF1ECCFA" blockId="5.[151,1437,1314,2012]" lastBlockId="6.[151,1437,152,588]" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
|
||
Our new genus has characteristics that
|
||
<bibRefCitation id="A955E4CDFFE6F460FD50F8EDFC2BCB6A" author="Kluge" box="[665,980,1874,1900]" pageId="5" pageNumber="67" refString="Kluge, N. & Novikova, E. A. (1992) Revision of Palearctic genera and subgenera of mayflies in the subfamily Cloeoninae (Ephemeroptera, Baetidae) with descriptions of new species from the USSR. Entomologicheskoye Obozreniye, 71, 60 - 83." type="journal article" year="1992">Kluge & Novikova (1992)</bibRefCitation>
|
||
considered separately apomorphic for the genus groups
|
||
<taxonomicName id="0AC4E2BFFFE6F460FEAFF8C7FE44CB97" box="[358,443,1912,1937]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FEAFF8C7FE44CB97" box="[358,443,1912,1937]" italics="true" pageId="5" pageNumber="66">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0AC4E2BFFFE6F460FE3CF8C7FD6FCB97" box="[501,656,1912,1937]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE6F460FE3CF8C7FD6FCB97" box="[501,656,1912,1937]" italics="true" pageId="5" pageNumber="66">Centroptilum</emphasis>
|
||
</taxonomicName>
|
||
, namely the expanded gills, mandibles with incisors cleft to bases and shapes of labial palps. Thus, we advocate critical evaluation of these characters with regards to their use and interpretation in phylogenetic inference. We also call attention to the apical projection on labial palp segment 3 and its similarity to a potentially a homologous structure in
|
||
<taxonomicName id="0AC4E2BFFFE5F463FC04FF27FBBFCCB7" box="[973,1088,152,177]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FC04FF27FBBFCCB7" box="[973,1088,152,177]" italics="true" pageId="6" pageNumber="67">Securiops</emphasis>
|
||
</taxonomicName>
|
||
(Jacobus et al., 2006:
|
||
<figureCitation id="55FF85B9FFE5F463FA8CFF27FA68CCB4" box="[1349,1431,152,178]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,275,1940,1964]" captionTargetBox="[209,1386,393,1898]" captionTargetId="figure@2.[199,1387,393,1905]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Waynokiops dentatogriphus, new genus, new species. 1. Head, anterior view, cleared and with mouthparts removed. 2. Labrum. 3. Right (planate) mandible. 4. Left (angulate) mandible. 5. Maxilla. 6. Labium." httpUri="https://zenodo.org/record/293963/files/figure.png" pageId="6" pageNumber="67">Figs. 6</figureCitation>
|
||
, 14). Notably,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FEF3FF02FE51CCD0" box="[314,430,189,214]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FEF3FF02FE51CCD0" box="[314,430,189,214]" italics="true" pageId="6" pageNumber="67">Securiops</emphasis>
|
||
</taxonomicName>
|
||
also has mandibles with deeply cleft mandibular incisors (Jacobus et al., 2006:
|
||
<figureCitation id="55FF85B9FFE5F463FAABFF02FF2BCCFA" captionStart-0="FIGURES 7 – 9" captionStart-1="FIGURES 10 – 13" captionStartId-0="3.[151,270,1779,1803]" captionStartId-1="4.[151,271,1889,1913]" captionTargetBox-0="[292,1309,194,1754]" captionTargetBox-1="[193,1396,206,1857]" captionTargetId-0="figure@3.[281,1317,192,1757]" captionTargetId-1="figure@4.[190,1396,197,1865]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 7 – 9. Waynokiops dentatogriphus, new genus, new species. 7. Labial palp. 8. Dorsal view, whole. 9. Lateral view, whole." captionText-1="FIGURES 10 – 13. Waynokiops dentatogriphus, new genus, new species. 10. Mesothoracic leg. 11. Tarsal claw; note double row of denticles restricted to proximal quarter of claw. 12. Gill VI. 13. Mesothoracic leg, patellotibial region." httpUri-0="https://zenodo.org/record/293964/files/figure.png" httpUri-1="https://zenodo.org/record/293965/files/figure.png" pageId="6" pageNumber="67">Figs. 8–11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CD7B993CFFE5F463FF0DFEB7FC5FCE4A" blockId="6.[151,1437,152,588]" pageId="6" pageNumber="67">
|
||
Our new genus demonstrates autapomorphies, including the expanded abdominal segments 1–7 and the dorsal abdominal armature. Dorsal abdominal armature on a baetid nymph from lentic habitats is of particular interest, because thusfar, such armature has been found only within genera whose nymphs are associated with lotic habitats. These genera include
|
||
<taxonomicName id="0AC4E2BFFFE5F463FD86FEC7FCA9CD94" authority="Bengtsson" authorityName="Bengtsson" box="[591,854,376,402]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FD86FEC7FD33CD97" box="[591,716,376,401]" italics="true" pageId="6" pageNumber="67">Acentrella</emphasis>
|
||
Bengtsson
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FCAAFEC7FB11CD94" authority="Waltz & McCafferty" authorityName="Waltz & McCafferty" box="[867,1262,376,402]" class="Insecta" family="Baetidae" genus="Acanthiops" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FCAAFEC7FC14CD97" box="[867,1003,376,401]" italics="true" pageId="6" pageNumber="67">Acanthiops</emphasis>
|
||
Waltz & McCafferty
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FB35FEC7FEEECDB1" authority="Demoulin" authorityName="Demoulin" class="Insecta" family="Baetidae" genus="Afrobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FB35FEC7FA63CD97" box="[1276,1436,376,401]" italics="true" pageId="6" pageNumber="67">Afrobaetodes</emphasis>
|
||
Demoulin
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FED4FE22FE2BCDB1" authority="Ueno" authorityName="Ueno" box="[285,468,413,439]" class="Insecta" family="Baetidae" genus="Baetiella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FED4FE22FE76CDB0" box="[285,393,413,438]" italics="true" pageId="6" pageNumber="67">Baetiella</emphasis>
|
||
Ueno
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FE28FE22FCAFCDB1" authority="Needham & Murphy" authorityName="Needham & Murphy" box="[481,848,413,439]" class="Insecta" family="Baetidae" genus="Baetodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FE28FE22FDB1CDB0" box="[481,590,413,438]" italics="true" pageId="6" pageNumber="67">Baetodes</emphasis>
|
||
Needham & Murphy
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FC95FE22FB24CDB1" authority="Wuillot & Gillies" authorityName="Wuillot & Gillies" box="[860,1243,413,439]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FC95FE22FC00CDB0" box="[860,1023,413,438]" italics="true" pageId="6" pageNumber="67">Cheleocloeon</emphasis>
|
||
Wuillot & Gillies
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FB21FE22FE98CDDA" authority="Wuillot & Gillies" authorityName="Wuillot & Gillies" class="Insecta" family="Baetidae" genus="Dicentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FB21FE22FA63CDB0" box="[1256,1436,413,438]" italics="true" pageId="6" pageNumber="67">Dicentroptilum</emphasis>
|
||
Wuillot & Gillies
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FEBDFE7DFDB6CDDA" authority="Mol" authorityName="Mol" box="[372,585,450,476]" class="Insecta" family="Baetidae" genus="Echinobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FEBDFE7DFDF1CDDD" box="[372,526,450,475]" italics="true" pageId="6" pageNumber="67">Echinobaetis</emphasis>
|
||
Mol
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FD9FFE7DFCF1CDDA" authority="Thomas" authorityName="Thomas" box="[598,782,450,476]" class="Insecta" family="Baetidae" genus="Gratia" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FD9FFE7DFD5BCDDD" box="[598,676,450,475]" italics="true" pageId="6" pageNumber="67">Gratia</emphasis>
|
||
Thomas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FCD2FE7DFB97CDDA" authority="Muller-Liebenau" authorityName="Muller-Liebenau" box="[795,1128,450,476]" class="Insecta" family="Baetidae" genus="Jubabaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FCD2FE7DFC65CDDD" box="[795,922,450,475]" italics="true" pageId="6" pageNumber="67">Jubabaetis</emphasis>
|
||
Muller-Liebenau
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FBB1FE7DFEBFCE04" authority="Lugo-Ortiz & McCafferty" authorityName="Lugo-Ortiz & McCafferty" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FBB1FE7DFAEFCDDD" box="[1144,1296,450,475]" italics="true" pageId="6" pageNumber="67">Papuanatula</emphasis>
|
||
Lugo-Ortiz & McCafferty
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AC4E2BFFFE5F463FE85FE57FCD5CE04" authority="McCafferty & Barber-James" authorityName="McCafferty & Barber-James" box="[332,810,488,514]" class="Insecta" family="Baetidae" genus="Tanzaniops" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FE85FE57FE2ECE07" box="[332,465,488,513]" italics="true" pageId="6" pageNumber="67">Tanzaniops</emphasis>
|
||
McCafferty & Barber-James
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0AC4E2BFFFE5F463FCACFE57FADBCE04" authority="Elouard and Hideux" authorityName="Elouard and Hideux" box="[869,1316,488,514]" class="Insecta" family="Baetidae" genus="Thraulobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FFB0452EFFE5F463FCACFE57FBD2CE07" box="[869,1069,488,513]" italics="true" pageId="6" pageNumber="67">Thraulobaetodes</emphasis>
|
||
Elouard and Hideux
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="A955E4CDFFE5F463FAFDFE57FF26CE21" author="Thomas" pageId="6" pageNumber="68" refString="Thomas, A. (1992) Gratia sororculaenadinae n. gen., n. sp., Ephemeroptere nouveau de Thailande (Ephemeroptera, Baetidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 128, 47 - 51." type="journal article" year="1992">Thomas, 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE5F463FF2EFDB2FDC4CE21" author="Waltz" box="[231,571,525,551]" pageId="6" pageNumber="68" refString="Waltz, R. D. & McCafferty. W. P. (1987) Systematics of Pseudocloeon, Acentrella, Baetiella, and Liebebiella, new genus (Ephemeroptera: Baetidae). Journal of the New York Entomological Society, 95, 553 - 568." type="journal article" year="1987">Waltz & McCafferty, 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE5F463FD80FDB2FC8FCE21" author="Wuillot" box="[585,880,525,551]" pageId="6" pageNumber="68" refString="Wuillot, J. & Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26, 213 - 217." type="journal article" year="1993">Wuillot & Gllies, 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE5F463FCB7FDB2FB58CE21" author="Boonsoong" box="[894,1191,525,551]" pageId="6" pageNumber="67" refString="Boonsoong, B., Thomas, A. & Sangpradub, N. (2002) Gratia narumonae n. sp., a new mayfly from Thailand (Ephemeroptera, Baetidae). Ephemera, 4, 1 - 9." type="journal article" year="2002">Boonsoong et al., 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE5F463FB7CFDB2FA68CE21" author="McCafferty" box="[1205,1431,525,551]" pageId="6" pageNumber="67" refString="McCafferty, W. P. (2003) Gose's African Ephemeroptera (Baetidae, Heptageniidae). Entomological News, 113, 294 - 302." type="journal article" year="2003">McCafferty, 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A955E4CDFFE5F463FF5EFD8DFDCECE4A" author="McCafferty" box="[151,561,562,588]" pageId="6" pageNumber="67" refString="McCafferty, W. P., & Barber-James, H. M. (2005) Tanzaniops, replacement name for certain African Baetidae (Ephmeroptera). Proceedings of the Entomological Society of Washington, 107, 238." type="journal article" year="2005">McCafferty & Barber-James, 2005</bibRefCitation>
|
||
; Jacobus & McCafferty, 2006).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |