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<document id="FA0B517EE29360A3C573E2B526358A25" ID-DOI="10.1080/00222933.2019.1634772" ID-GBIF-Dataset="07592748-1046-4033-86f6-385de81795d7" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673463" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027693774" checkinUser="valdenar" docAuthor="St Laurent, Ryan A. &amp; Giusti, Alessandro" docDate="2019" docId="03F75269FFB9C84D4EAF11FCFF35DBFC" docLanguage="en" docName="JNATHIST.53.1920.12091246.pdf.imf" docOrigin="Journal of Natural History 53 (19)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Zaphanta fraterna Schausi 1912" docType="treatment" docVersion="8" lastPageNumber="1224" masterDocId="FFCE2A11FFB7C85C4F621005FFDEDB35" masterDocTitle="Revision of Zaphanta Dyarı 1910 (Lepidoptera: Mimallonidae: Zaphantinae) ı with descriptions of nine new species" masterLastPageNumber="1246" masterPageNumber="1209" pageNumber="1222" updateTime="1698718885971" updateUser="plazi">
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<mods:title id="88F4254ECE7474A729E69A937E1F8BE4">Revision of Zaphanta Dyarı 1910 (Lepidoptera: Mimallonidae: Zaphantinae) ı with descriptions of nine new species</mods:title>
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<mods:namePart id="BAC9CACE8816E1B4D6FEEA4BADA57A72">St Laurent, Ryan A.</mods:namePart>
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<mods:namePart id="DC37A72E5094EECEEC628D1D17383A63">Giusti, Alessandro</mods:namePart>
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<mods:title id="D4051229407D7780E014B864E14CDC64">Journal of Natural History</mods:title>
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<mods:date id="532142CC1BC1AB202970C9CEE16EBC4A">2019</mods:date>
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<treatment id="03F75269FFB9C84D4EAF11FCFF35DBFC" ID-DOI="http://doi.org/10.5281/zenodo.3679847" ID-GBIF-Taxon="162162017" ID-Zenodo-Dep="3679847" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC" httpUri="http://treatment.plazi.org/id/03F75269FFB9C84D4EAF11FCFF35DBFC" lastPageId="17" lastPageNumber="1224" pageId="14" pageNumber="1222">
<subSubSection id="C344B0F4FFB9C8524EAF11FCFC8BD926" box="[461,853,505,531]" pageId="14" pageNumber="1222" type="nomenclature">
<paragraph id="8BE1E37FFFB9C8524EAF11FCFC8BD926" blockId="14.[461,853,505,565]" box="[461,853,505,531]" pageId="14" pageNumber="1222">
<heading id="D0A95413FFB9C8524EAF11FCFC8BD926" box="[461,853,505,531]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="1222" reason="2">
<taxonomicName id="4C5E98FCFFB9C8524EAF11FCFC8BD926" ID-CoL="98PNC" authority="Schausi 1912" authorityName="Schausi" authorityYear="1912" box="[461,853,505,531]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524EAF11FCFD6CD927" bold="true" box="[461,690,505,530]" italics="true" pageId="14" pageNumber="1222">Zaphanta fraterna</emphasis>
Schaus, 1912
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C344B0F4FFB9C8524EBD1219FC9BD900" box="[479,837,540,565]" pageId="14" pageNumber="1222" type="description">
<paragraph id="8BE1E37FFFB9C8524EBD1219FC9BD900" blockId="14.[461,853,505,565]" box="[479,837,540,565]" pageId="14" pageNumber="1222">
<heading id="D0A95413FFB9C8524EBD1219FC9BD900" box="[479,837,540,565]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="1222" reason="5">
(
<figureCitation id="1365FFFAFFB9C8524E851219FD8DD900" box="[487,595,540,565]" captionStart="Figure 1" captionStartId="5.[160,225,1462,1485]" captionTargetBox="[164,1151,142,1445]" captionTargetId="figure@5.[163,1153,142,1445]" captionTargetPageId="5" captionText="Figure 1. Annotated Zaphanta fraterna male wing venation diagram. Adapted with permission from Forbes (1942, fig. 104). Note that the venation of the Zaphanta type species Z. infantilis is very similar, but Rs3 + Rs4 are shorter stalked." figureDoi="http://doi.org/10.5281/zenodo.3673465" httpUri="https://zenodo.org/record/3673465/files/figure.png" pageId="14" pageNumber="1222">Figures 1</figureCitation>
, 57, 10, 27, 40, 41)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C344B0F4FFB9C8524FC21254FE1FD9C1" pageId="14" pageNumber="1222" type="reference_group">
<paragraph id="8BE1E37FFFB9C8524FC21254FE1FD9C1" blockId="14.[160,1156,593,756]" pageId="14" pageNumber="1222">
<taxonomicName id="4C5E98FCFFB9C8524FC21254FDF5D95C" ID-CoL="98PNC" authority="Schausi 1912: 48" authorityName="Schausi" authorityPageNumber="48" authorityYear="1912" box="[160,555,593,617]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524FC21254FEB6D95C" box="[160,360,593,617]" italics="true" pageId="14" pageNumber="1222">Zaphanta fraterna</emphasis>
Schaus, 1912: 48
</taxonomicName>
. Original description;
<taxonomicName id="4C5E98FCFFB9C8524C7F1254FBA0D95C" ID-CoL="98PNC" authority=": Schaus 1928" authorityName=": Schaus" authorityYear="1928" box="[797,1150,593,617]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524C7F1254FC38D95C" box="[797,998,593,617]" italics="true" pageId="14" pageNumber="1222">Zaphanta fraterna</emphasis>
:
<bibRefCitation id="EFCF9E8EFFB9C8524C901254FBA0D95C" author="Schaus W" box="[1010,1150,593,617]" pageId="14" pageNumber="1222" pagination="635 - 672" refId="ref17433" refString="Schaus W. 1928. Familie Mimallonidae. In: Seitz Ai editor. Die Gross-Schmetterlinge der Erde 6 Die amerikanischen Spinn und Schwarmer. Stuttgart: A. Kernen; p. 635 - 672." type="book chapter" year="1928">Schaus 1928</bibRefCitation>
</taxonomicName>
, treated as a synonym of
<emphasis id="B92A3F6DFFB9C8524E881271FDB5D9B9" box="[490,619,628,652]" italics="true" pageId="14" pageNumber="1222">
<taxonomicName id="4C5E98FCFFB9C8524E881271FDBBD9B9" ID-CoL="5CWXQ" baseAuthorityName="Schaus" baseAuthorityYear="1928" box="[490,613,628,652]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="infantilis">Z. infantilis</taxonomicName>
;
</emphasis>
<taxonomicName id="4C5E98FCFFB9C8524D141271FC39D9B9" ID-CoL="5CWXQ" authority=": Forbes 1942" authorityName=": Forbes" authorityYear="1942" box="[630,999,628,652]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="infantilis">
<emphasis id="B92A3F6DFFB9C8524D141271FC97D9B9" box="[630,841,628,652]" italics="true" pageId="14" pageNumber="1222">Zaphanta infantilis</emphasis>
:
<bibRefCitation id="EFCF9E8EFFB9C8524C3B1271FC39D9B9" author="Forbes WTM" box="[857,999,628,652]" pageId="14" pageNumber="1222" pagination="265 - 406" refId="ref17149" refString="Forbes WTM. 1942. The Lepidoptera of Barro Colorado Islandi Panama (No. 2). Bull Museum Comp Zool. 90: 265 - 406." type="journal article" year="1942">Forbes 1942</bibRefCitation>
</taxonomicName>
, but
<collectingCountry id="F349A3EFFFB9C8524B4B1271FB5DD9B9" box="[1065,1155,628,652]" name="Panama" pageId="14" pageNumber="1222">Panama</collectingCountry>
records actually refer to
<emphasis id="B92A3F6DFFB9C8524E901292FDACD99A" box="[498,626,663,687]" italics="true" pageId="14" pageNumber="1222">
<taxonomicName id="4C5E98FCFFB9C8524E901292FDB5D99A" ID-CoL="98PNC" authorityName=": Schaus" authorityYear="1928" box="[498,619,663,687]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">Z. fraterna</taxonomicName>
;
</emphasis>
<taxonomicName id="4C5E98FCFFB9C8524DE31292FC29D99A" ID-CoL="98PNC" authority=": Becker 1996" authorityName="Becker" authorityYear="1996" box="[641,1015,663,687]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524DE31292FC8FD99A" box="[641,849,663,687]" italics="true" pageId="14" pageNumber="1222">Zaphanta fraterna</emphasis>
:
<bibRefCitation id="EFCF9E8EFFB9C8524C041292FC29D99A" author="Becker VO" box="[870,1015,663,687]" pageId="14" pageNumber="1222" pagination="17 - 19" refId="ref17001" refString="Becker VO. 1996. Mimallonidae. In: Heppner JBi editor. Atlas neotrop lepid checklist part 4 B Drepanoideai Bombycoideai Sphingoidea. Gainesville (Florida): Association for Tropical Lepidoptera &amp; Scientic Publishers; p. 17 - 19." type="book chapter" year="1996">Becker 1996</bibRefCitation>
</taxonomicName>
, treated as a synonym of
<emphasis id="B92A3F6DFFB9C8524E0612BCFE3DD9E4" box="[356,483,697,721]" italics="true" pageId="14" pageNumber="1222">
<taxonomicName id="4C5E98FCFFB9C8524E0612BCFE03D9E4" ID-CoL="5CWXQ" authorityName=": Forbes" authorityYear="1942" box="[356,477,697,721]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="infantilis">Z. infantilis</taxonomicName>
;
</emphasis>
<taxonomicName id="4C5E98FCFFB9C8524E8E12BCFBFDD9E4" ID-CoL="98PNC" authority=": St Laurent and Kawahara 2019" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[492,1059,697,721]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524E8E12BCFD68D9E4" box="[492,694,697,721]" italics="true" pageId="14" pageNumber="1222">Zaphanta fraterna</emphasis>
:
<bibRefCitation id="EFCF9E8EFFB9C8524DA712BCFBFDD9E4" author="St Laurent RAi Kawahara AY" box="[709,1059,697,721]" pageId="14" pageNumber="1222" pagination="1 - 114" refId="ref17584" refString="St Laurent RAi Kawahara AY. 2019. Reclassification of the sack-bearer moths (Lepidopterai Mimallonoideai Mimallonidae). Zookeys. 815: 1 - 114. doi: 10.3897 / zookeys. 815.27335." type="journal article" year="2019">St Laurent and Kawahara 2019</bibRefCitation>
</taxonomicName>
, revived status as valid species.
</paragraph>
</subSubSection>
<subSubSection id="C344B0F4FFB9C8524FC21324FE44DFC9" pageId="14" pageNumber="1222" type="diagnosis">
<paragraph id="8BE1E37FFFB9C8524FC21324FECAD80F" blockId="14.[160,1156,801,1276]" box="[160,276,801,826]" pageId="14" pageNumber="1222">
<emphasis id="B92A3F6DFFB9C8524FC21324FECAD80F" bold="true" box="[160,276,801,826]" italics="true" pageId="14" pageNumber="1222">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE1E37FFFB9C8524FC21341FE44DFC9" blockId="14.[160,1156,801,1276]" pageId="14" pageNumber="1222">
As in the preceding species,
<taxonomicName id="4C5E98FCFFB9C8524D6E1341FD58D869" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[524,646,836,860]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524D6E1341FD58D869" box="[524,646,836,860]" italics="true" pageId="14" pageNumber="1222">Z. fraterna</emphasis>
</taxonomicName>
is not readily distinguishable from other
<taxonomicName id="4C5E98FCFFB9C8524FC21362FED5D84A" authorityName="Dyari" authorityYear="1910" box="[160,267,871,895]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="genus">
<emphasis id="B92A3F6DFFB9C8524FC21362FED5D84A" box="[160,267,871,895]" italics="true" pageId="14" pageNumber="1222">Zaphanta</emphasis>
</taxonomicName>
by external characters. This species is recognisable by the genitalia which in males display an elongated, narrow tegumen, truncated uncus, a well-sclerotised singular subuncus projection of variable width but which is always pointed mesally; a mostly smooth, simple phallus with the typical ventral distal sharpened extension which is more heavily sclerotised than the remainder of the phallus. The phallus lacks spines along its lateral side. These characters, combined with the absence of modified costal valvae apodemes, are unique to this species. The most similar species,
<taxonomicName id="4C5E98FCFFB9C8524B711432FB5DDF7A" authority="St Laurent &amp; Giusti, 2019" authorityName="St Laurent &amp; Giusti" authorityYear="2019" box="[1043,1155,1079,1103]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="rawlinsi" status="sp. nov.">
<emphasis id="B92A3F6DFFB9C8524B711432FB5DDF7A" box="[1043,1155,1079,1103]" italics="true" pageId="14" pageNumber="1222">Z. rawlinsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2198216FFB9C8524FC2145FFF21DF47" box="[160,255,1114,1138]" pageId="14" pageNumber="1222" rank="species">sp. nov.</taxonomicNameLabel>
, has a wider, blunter subuncus mesal projection, narrower valvae than any dissected
<taxonomicName id="4C5E98FCFFB9C8524E711479FE59DFA1" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[275,391,1148,1172]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524E711479FE59DFA1" box="[275,391,1148,1172]" italics="true" pageId="14" pageNumber="1222">Z. fraterna</emphasis>
</taxonomicName>
, and a shorter, broader phallus. As mentioned in the diagnosis of
<taxonomicName id="4C5E98FCFFB9C8524FC2149AFEC4DF82" authorityName=": Forbes" authorityYear="1942" box="[160,282,1183,1207]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="infantilis">
<emphasis id="B92A3F6DFFB9C8524FC2149AFEC4DF82" box="[160,282,1183,1207]" italics="true" pageId="14" pageNumber="1222">Z. infantilis</emphasis>
</taxonomicName>
, tergite VIII in the female genitalia is not deeply concave in
<taxonomicName id="4C5E98FCFFB9C8524CB4149AFB94DF82" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[982,1098,1183,1207]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB9C8524CB4149AFB94DF82" box="[982,1098,1183,1207]" italics="true" pageId="14" pageNumber="1222">Z. fraterna</emphasis>
</taxonomicName>
, and has a broad posteriorly directed lobe instead of a narrow posteriorly directed protrusion as seen in
<taxonomicName id="4C5E98FCFFB9C8524E7914E1FE4ADFC9" authorityName=": Forbes" authorityYear="1942" box="[283,404,1252,1276]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="1222" phylum="Arthropoda" rank="species" species="infantilis">
<emphasis id="B92A3F6DFFB9C8524E7914E1FE4ADFC9" box="[283,404,1252,1276]" italics="true" pageId="14" pageNumber="1222">Z. infantilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C344B0F4FFB9C8534FC2152CFD05D9E4" lastPageId="15" lastPageNumber="1223" pageId="14" pageNumber="1222" type="description">
<paragraph id="8BE1E37FFFB9C8524FC2152CFEF8DE77" blockId="14.[160,1156,1321,1727]" box="[160,294,1321,1346]" pageId="14" pageNumber="1222">
<emphasis id="B92A3F6DFFB9C8524FC2152CFEF8DE77" bold="true" box="[160,294,1321,1346]" italics="true" pageId="14" pageNumber="1222">Description</emphasis>
</paragraph>
<paragraph id="8BE1E37FFFB9C8534FC21549FC2EDBD9" blockId="14.[160,1156,1321,1727]" lastBlockId="15.[160,1155,142,236]" lastPageId="15" lastPageNumber="1223" pageId="14" pageNumber="1222">
<emphasis id="B92A3F6DFFB9C8524FC21549FF3FDE50" bold="true" box="[160,225,1356,1381]" italics="true" pageId="14" pageNumber="1222">Male.</emphasis>
Head: As for genus. Thorax: As for genus. Legs: As for genus. Fore wing dorsum: As for genus. Fore wing length: 7.511.0 mm, avg.: 9.6 mm, wingspan: 1922 mm, n = 7. Fore wing ventrum: As for genus. Hind wing dorsum: As for genus. Hind wing ventrum: As for genus. Abdomen: As for genus. Genitalia: (
<figureCitation id="1365FFFAFFB9C8524D8D15B0FC83DEF8" box="[751,861,1461,1485]" captionStart="Figures 2529" captionStartId="28.[160,235,879,902]" captionTargetBox="[174,1152,143,862]" captionTargetId="figure@28.[162,1153,142,862]" captionTargetPageId="28" captionText="Figures 2529. Zaphanta male genitalia; a = ventral, b = lateral, c = phallus, lateral aspect, d = phallus, ventral aspect. 25. Z. infantilis, French Guiana, St. Jean du Maroni, genitalia vial NHMUK 010402338 [note: phallus shown encased in its thick membrane; similar membranes are removed or were not present to the same degree in other preparations] (NHMUK). 26. Z. infantilis, Guyana, Potaro, genitalia vial NHMUK 010402340 [note: Figure 26(a) shows the uncus and subuncal sclerotisation to display the degree of variation in this character in Z. infantilis; Figure 26(c,d) shows the phallus with its membrane peeled back and also display a more apparent lateral spine cluster on the phallus in this particular individual] (NHMUK). 27. Z. fraterna, Costa Rica, Esperanza, genitalia vial NHMUK 010402344 (NHMUK). 28. Z. rawlinsi holotype, Ecuador, Sucumbios, Sacha Lodge, Laguna de Pilchicocha, 220 m, St Laurent diss.: 7-21-18:3 (CMNH). 29. Z. anas holotype, Brazil, Amazonas, Reserva Ducke, km 26, Manaus-Itacoatiara Highway, St Laurent diss.: 8-25-18:2 (CNC). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673477" httpUri="https://zenodo.org/record/3673477/files/figure.png" pageId="14" pageNumber="1222">Figure 27</figureCitation>
) n = 6. Vinculum ovoid, basally narrowed, squared. Tegumen elongate, narrower than vinculum. Uncus triangular and heavily sclerotised apically, apex truncated with squared tip. Subuncus projection forming a closed ring with lateral margins of tegumen, mesally more heavily sclerotised, forming triangular point; uncus-tegumen complex narrower relative to remainder of genitalia. Transtilla triangular, variable in width. Valvae variable in width but always narrowed distally and angled upward (when spread) with middle of saccular edge forming an elbow-like angle or curve; bases of valvae meet mesally, touching or nearly touching. Costal base of valvae (valva apodeme) not modified. Caecum of phallus shorter than half length of phallus, rounded. Phallus stout, cylindrical, ventrally extended as well-sclerotised, sharp point; phallus smooth or nearly smooth laterally.
</paragraph>
<paragraph id="8BE1E37FFFB8C8534FC2111DFD05D9E4" blockId="15.[160,1156,280,721]" pageId="15" pageNumber="1223">
<emphasis id="B92A3F6DFFB8C8534FC2111DFF23DA04" bold="true" box="[160,253,280,305]" italics="true" pageId="15" pageNumber="1223">Female.</emphasis>
Head: As for genus. Thorax: As for genus. Legs: As for genus. Fore wing dorsum: Fore wing length: 11.011.5 mm, avg.: 11.3 mm, wingspan: 21 mm, n = 2. As for male but wings slightly broader overall, marginal area in particular darker purple in two specimens examined than in most males. Fore wing ventrum: As for male, but antemedial shading and antemedial line less developed. Hind wing dorsum: As for male but with more darkly defined marginal area. Hind wing ventrum: As for genus. Abdomen: As for genus. Genitalia: (
<figureCitation id="1365FFFAFFB8C8534D2911ECFD65D934" box="[587,699,489,513]" captionStart="Figures 3840" captionStartId="32.[160,235,1449,1472]" captionTargetBox="[163,1153,142,1432]" captionTargetId="figure@32.[163,1153,142,1432]" captionTargetPageId="32" captionText="Figures 3840. Zaphanta female genitalia; a = ventral, b = dorsal, c = lateral. 38. Z. infantilis French Guiana, St. Jean du Maroni, genitalia vial NHMUK 010402342 [note: ductus and corpus bursae not shown in Figure 38b; Figure 38a reused with permission from St Laurent and Kawahara (2019), ZooKeys] (NHMUK). 39. Z. elephanta paratype, Brazil, Amazonas, Fonte Boa, genitalia vial NHMUK 010402348 [note: ductus corpus bursae not preserved] (NHMUK). 40. Z. fraterna Costa Rica, Alajuela, Potrero Argentina, 520 m, 12-SRNP-2303, St Laurent diss.: USNMENT 01373361 [note: Figure 40a not to scale] (USNM). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673485" httpUri="https://zenodo.org/record/3673485/files/figure.png" pageId="15" pageNumber="1223">Figure 40</figureCitation>
) n = 1. As for genus, but tergite VIII concave with posteriorly directed mesal lobe which is part of more heavily sclerotised mesal region of tergite VIII, region of VIII tergite proximal to base of papillae anales, covered in short setae; sternite VIII broadened and squared. Ostium bursae globular, bowl-like, more heavily sclerotised than surrounding VIII, width of sclerotised ostium region roughly half that of segment VIII. Ductus bursae and corpus bursae narrow, corpus bursae wider than ductus bursae, bag-like.
</paragraph>
</subSubSection>
<subSubSection id="C344B0F4FFB8C84C4FC212FBFE1CD99A" lastPageId="16" lastPageNumber="1224" pageId="15" pageNumber="1223" type="materials_examined">
<paragraph id="8BE1E37FFFB8C8534FC212FBFE9BD822" blockId="15.[160,1155,766,895]" box="[160,325,766,791]" pageId="15" pageNumber="1223">
<emphasis id="B92A3F6DFFB8C8534FC212FBFE9BD822" bold="true" box="[160,325,766,791]" italics="true" pageId="15" pageNumber="1223">Type material</emphasis>
</paragraph>
<paragraph id="8BE1E37FFFB8C8534FC21325FBF1D84A" blockId="15.[160,1155,766,895]" pageId="15" pageNumber="1223">
<materialsCitation id="3B36E922FFB8C8534FC21325FC34D869" ID-GBIF-Occurrence="2565952733" collectorName="Rivi CR &amp; Type No." country="Costa Rica" location="Limon" pageId="15" pageNumber="1223" specimenCount="1" stateProvince="Limon" typeStatus="lectotype">
<emphasis id="B92A3F6DFFB8C8534FC21325FECCD80C" bold="true" box="[160,274,800,825]" italics="true" pageId="15" pageNumber="1223">
<typeStatus id="54E55DDDFFB8C8534FC21325FECCD80C" box="[160,274,800,825]" pageId="15" pageNumber="1223" type="lectotype">Lectotype</typeStatus>
</emphasis>
♂.
<collectingCountry id="F349A3EFFFB8C8534E2A1324FE1ED80C" box="[328,448,801,825]" name="Costa Rica" pageId="15" pageNumber="1223">Costa Rica</collectingCountry>
:
<collectingRegion id="499A2D9DFFB8C8534EAF1324FDC8D80C" box="[461,534,801,825]" country="Costa Rica" name="Limon" pageId="15" pageNumber="1223">Limón</collectingRegion>
: Sixola [Sixaola]
<collectorName id="26AB86A9FFB8C8534DB71324FCF7D80C" box="[725,809,801,825]" pageId="15" pageNumber="1223">Riv, CR</collectorName>
/March/
<taxonomicName id="4C5E98FCFFB8C8534C1E1324FB99D80C" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[892,1095,801,825]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="1223" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB8C8534C1E1324FB99D80C" box="[892,1095,801,825]" italics="true" pageId="15" pageNumber="1223">Zaphanta fraterna</emphasis>
</taxonomicName>
type Schaus/
<collectorName id="26AB86A9FFB8C8534F9A1341FEB6D869" box="[248,360,836,860]" pageId="15" pageNumber="1223">Type No.</collectorName>
17497/USNM-Mimal: 1104/St Laurent diss.: 5-16-18:6
</materialsCitation>
/
<materialsCitation id="3B36E922FFB8C8534C931341FBF1D84A" ID-GBIF-Occurrence="2565952773" pageId="15" pageNumber="1223" typeStatus="lectotype">
<typeStatus id="54E55DDDFFB8C8534C931341FB5DD869" box="[1009,1155,836,860]" pageId="15" pageNumber="1223" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C5E98FCFFB8C8534FC21362FEB4D84A" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[160,362,871,895]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="1223" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFB8C8534FC21362FEB4D84A" box="[160,362,871,895]" italics="true" pageId="15" pageNumber="1223">Zaphanta fraterna</emphasis>
</taxonomicName>
designated by St Laurent and Giusti 2019/(USNM, examined).
</materialsCitation>
</paragraph>
<paragraph id="8BE1E37FFFB8C8534FC213AEFDDBD8F1" blockId="15.[160,1155,939,1033]" box="[160,517,939,964]" pageId="15" pageNumber="1223">
<emphasis id="B92A3F6DFFB8C8534FC213AEFDDBD8F1" bold="true" box="[160,517,939,964]" italics="true" pageId="15" pageNumber="1223">Additional material examined</emphasis>
</paragraph>
<paragraph id="8BE1E37FFFB8C8534FC213CBFD1ADF3C" blockId="15.[160,1155,939,1033]" pageId="15" pageNumber="1223">
<materialsCitation id="3B36E922FFB8C8534FC213CBFD1ADF3C" ID-GBIF-Occurrence="2565952730" pageId="15" pageNumber="1223">
<emphasis id="B92A3F6DFFB8C8534FC213CBFF35D8D2" bold="true" box="[160,235,974,999]" italics="true" pageId="15" pageNumber="1223">
<collectingCountry id="F349A3EFFFB8C8534FC213CBFF39D8D2" box="[160,231,974,999]" name="Belize" pageId="15" pageNumber="1223">Belize</collectingCountry>
.
</emphasis>
<collectingRegion id="499A2D9DFFB8C8534F9913CAFE95D8D2" box="[251,331,975,999]" country="Belize" name="Toledo" pageId="15" pageNumber="1223">Toledo</collectingRegion>
: 1 ♂, no additional locality information [photo examined only] (
<collectionCode id="ED4F7BBAFFB8C8534B7513CAFBBBD8D2" box="[1047,1125,975,999]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">CMNH</collectionCode>
). 1 ♂, Punta Gorda [photo examined only] (
<collectionCode id="ED4F7BBAFFB8C8534D0E13F4FD64DF3C" box="[620,698,1009,1033]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">CMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE1E37FFFB8C8534FC21433FCE9DFA1" blockId="15.[160,1156,1078,1172]" pageId="15" pageNumber="1223">
<materialsCitation id="3B36E922FFB8C8534FC21433FDAEDF47" ID-GBIF-Occurrence="2565952729" collectionCode="USNM" collectorName="Di Quirigua &amp; Februaryi Schaus &amp; Barnes" country="Guatemala" location="Izabal" pageId="15" pageNumber="1223" specimenCount="1" stateProvince="Izabal">
<emphasis id="B92A3F6DFFB8C8534FC21433FEF4DF7A" bold="true" box="[160,298,1078,1103]" italics="true" pageId="15" pageNumber="1223">
<collectingCountry id="F349A3EFFFB8C8534FC21433FEFBDF7A" box="[160,293,1078,1103]" name="Guatemala" pageId="15" pageNumber="1223">Guatemala</collectingCountry>
.
</emphasis>
<collectingRegion id="499A2D9DFFB8C8534E581432FEA0DF7A" box="[314,382,1079,1103]" country="Guatemala" name="Izabal" pageId="15" pageNumber="1223">Izabal</collectingRegion>
: 1
<collectorName id="26AB86A9FFB8C8534EC81432FDEBDF7A" box="[426,565,1079,1103]" pageId="15" pageNumber="1223">♂, Quiriguá</collectorName>
:
<collectorName id="26AB86A9FFB8C8534D241432FCD5DF7A" box="[582,779,1079,1103]" pageId="15" pageNumber="1223">February, Schaus</collectorName>
and
<collectorName id="26AB86A9FFB8C8534C301432FC43DF7A" box="[850,925,1079,1103]" pageId="15" pageNumber="1223">Barnes</collectorName>
coll., USNM-Mimal:
<date id="FFE0C5BFFFB8C8534FC2145FFF00DF47" box="[160,222,1114,1138]" pageId="15" pageNumber="1223" value="2555-01">2555,</date>
St Laurent diss.: 5-16-18:5 (
<collectionCode id="ED4F7BBAFFB8C8534D78145FFDBBDF47" box="[538,613,1114,1138]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B36E922FFB8C8534D1A145FFCEDDFA1" ID-GBIF-Occurrence="2565952738" collectingDate="2553-01" collectionCode="USNM" collectorName="Di Quirigua &amp; Februaryi Schaus &amp; Barnes" country="Guatemala" location="Izabal" pageId="15" pageNumber="1223" specimenCount="1" stateProvince="Izabal">
2
<collectorName id="26AB86A9FFB8C8534DED145CFCD6DF47" box="[655,776,1113,1138]" pageId="15" pageNumber="1223">♂, Cayuga</collectorName>
:
<collectorName id="26AB86A9FFB8C8534C76145FFC25DF47" box="[788,1019,1114,1138]" pageId="15" pageNumber="1223">March, April, Schaus</collectorName>
and
<collectorName id="26AB86A9FFB8C8534B5A145FFB5DDF47" box="[1080,1155,1114,1138]" pageId="15" pageNumber="1223">Barnes</collectorName>
coll.,
<location id="8E81B5A4FFB8C8534FB91479FE73DFA1" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534FB91479FE73DFA1" box="[219,429,1148,1172]" country="Guatemala" name="Dognin Collectioni" pageId="15" pageNumber="1223">Dognin Collection,</location>
USNM-Mimal:
<date id="FFE0C5BFFFB8C8534D351479FD4BDFA1" box="[599,661,1148,1172]" pageId="15" pageNumber="1223" value="2553-01">
<collectingDate id="EFA43C57FFB8C8534D351479FD4BDFA1" box="[599,661,1148,1172]" pageId="15" pageNumber="1223" value="2553-01">2553,</collectingDate>
</date>
2556 (
<collectionCode id="ED4F7BBAFFB8C8534D831479FCF2DFA1" box="[737,812,1148,1172]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE1E37FFFB8C84C4FC214C4FC0BDBD9" blockId="15.[160,1157,1217,1727]" lastBlockId="16.[160,1155,142,236]" lastPageId="16" lastPageNumber="1224" pageId="15" pageNumber="1223">
<materialsCitation id="3B36E922FFB8C8534FC214C4FD1DDFEF" ID-GBIF-Occurrence="2565952743" box="[160,707,1217,1242]" collectorName="Di Bijaguai" country="Costa Rica" elevation="750" location="Alajuela" pageId="15" pageNumber="1223" specimenCount="1" stateProvince="Alajuela">
<emphasis id="B92A3F6DFFB8C8534FC214C4FEF8DFEF" bold="true" box="[160,294,1217,1242]" italics="true" pageId="15" pageNumber="1223">
<collectingCountry id="F349A3EFFFB8C8534FC214C4FEFCDFEF" box="[160,290,1217,1242]" name="Costa Rica" pageId="15" pageNumber="1223">Costa Rica</collectingCountry>
.
</emphasis>
<collectingRegion id="499A2D9DFFB8C8534E5414C7FE4CDFEF" box="[310,402,1218,1242]" country="Costa Rica" name="Alajuela" pageId="15" pageNumber="1223">Alajuela</collectingRegion>
: 1
<collectorName id="26AB86A9FFB8C8534ED514C7FDEDDFEF" box="[439,563,1218,1242]" pageId="15" pageNumber="1223">♂, Bijagua,</collectorName>
<quantity id="4CA64E9AFFB8C8534D5C14C7FD57DFEF" box="[574,649,1218,1242]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="1223" unit="m" value="750.0">
<elevation id="0073044CFFB8C8534D5C14C7FD57DFEF" box="[574,649,1218,1242]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="1223" unit="m" value="750.0">750 m</elevation>
</quantity>
: 34
</materialsCitation>
.
<materialsCitation id="3B36E922FFB8C8534DB214C7FE49DFC8" ID-GBIF-Occurrence="2565952756" collectingDate="2000-01" collectionCode="VOB" country="Costa Rica" isTailEnumeration="true" location="November" pageId="15" pageNumber="1223" specimenCount="1">
<location id="8E81B5A4FFB8C8534DB214C7FC98DFEF" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534DB214C7FC98DFEF" box="[720,838,1218,1242]" country="Costa Rica" name="November" pageId="15" pageNumber="1223">November</location>
<date id="FFE0C5BFFFB8C8534C3314C7FC51DFEF" box="[849,911,1218,1242]" pageId="15" pageNumber="1223" value="2000-01">
<collectingDate id="EFA43C57FFB8C8534C3314C7FC51DFEF" box="[849,911,1218,1242]" pageId="15" pageNumber="1223" value="2000-01">2000,</collectingDate>
</date>
V.O. Becker col., Col.
<location id="8E81B5A4FFB8C8534FC214E0FF34DFC8" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534FC214E0FF34DFC8" box="[160,234,1253,1277]" country="Costa Rica" name="Becker" pageId="15" pageNumber="1223">Becker</location>
129398 (
<collectionCode id="ED4F7BBAFFB8C8534E3914E0FE4FDFC8" box="[347,401,1253,1277]" pageId="15" pageNumber="1223">VOB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B36E922FFB8C8534EC414E0FD71DEB2" ID-GBIF-Occurrence="2565952728" collectingDate="2009-01" collectorName="Di Vochysia &amp; de Conservacion &amp; March" country="Costa Rica" county="Di Vochysia" elevation="320" location="Wi" municipality="Ni" pageId="15" pageNumber="1223" specimenCount="1" stateProvince="Guanacaste">
1
<collectorName id="26AB86A9FFB8C8534EA314E1FD97DFC8" box="[449,585,1252,1277]" pageId="15" pageNumber="1223">
<collectingCounty id="62809BF3FFB8C8534EA314E1FD97DFC8" box="[449,585,1252,1277]" pageId="15" pageNumber="1223">♂, Vochysia</collectingCounty>
</collectorName>
[note: this is the name of the site in Área
<collectorName id="26AB86A9FFB8C8534B0514E0FEE9DE2A" pageId="15" pageNumber="1223">de Conservación</collectorName>
<collectingRegion id="499A2D9DFFB8C8534E5D1502FE1FDE2A" box="[319,449,1287,1311]" country="Costa Rica" name="Guanacaste" pageId="15" pageNumber="1223">Guanacaste</collectingRegion>
where the specimen was collected,
<emphasis id="B92A3F6DFFB8C8534C3C1502FC5DDE2A" box="[862,899,1287,1311]" italics="true" pageId="15" pageNumber="1223">not</emphasis>
the host plant genus], 10.86666°
<collectingMunicipality id="6B857905FFB8C8534E72152FFEFBDE77" box="[272,293,1322,1346]" pageId="15" pageNumber="1223">N,</collectingMunicipality>
85.24528°
<location id="8E81B5A4FFB8C8534EC4152FFE1FDE77" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534EC4152FFE1FDE77" box="[422,449,1322,1346]" country="Costa Rica" county="Di Vochysia" municipality="Ni" name="Wi" pageId="15" pageNumber="1223" stateProvince="Guanacaste">W,</location>
<quantity id="4CA64E9AFFB8C8534EB6152FFDF9DE77" box="[468,551,1322,1346]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="1223" unit="m" value="320.0">
<elevation id="0073044CFFB8C8534EB6152FFDF9DE77" box="[468,551,1322,1346]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="1223" unit="m" value="320.0">320 m</elevation>
</quantity>
: larva collected 13
<collectorName id="26AB86A9FFB8C8534C49152FFCAFDE77" box="[811,881,1322,1346]" pageId="15" pageNumber="1223">March</collectorName>
<date id="FFE0C5BFFFB8C8534CE1152FFC1FDE77" box="[899,961,1322,1346]" pageId="15" pageNumber="1223" value="2009-01">
<collectingDate id="EFA43C57FFB8C8534CE1152FFC1FDE77" box="[899,961,1322,1346]" pageId="15" pageNumber="1223" value="2009-01">2009,</collectingDate>
</date>
adult emerged 26 April
<date id="FFE0C5BFFFB8C8534E661548FE9CDE50" box="[260,322,1357,1381]" pageId="15" pageNumber="1223" value="2009-01">
<collectingDate id="EFA43C57FFB8C8534E661548FE9CDE50" box="[260,322,1357,1381]" pageId="15" pageNumber="1223" value="2009-01">2009,</collectingDate>
</date>
food plant:
<taxonomicName id="4C5E98FCFFB8C8534EB01548FD72DE50" authorityName="A. Chev" authorityYear="1909" box="[466,684,1357,1381]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="1223" phylum="Tracheophyta" rank="species" species="ivorensis">
<emphasis id="B92A3F6DFFB8C8534EB01548FD72DE50" box="[466,684,1357,1381]" italics="true" pageId="15" pageNumber="1223">Terminalia ivorensis</emphasis>
</taxonomicName>
(introduced plant), 09-SRNP-40335, DNA and genital prep. USNMENT 01373360 (
<collectionCode id="ED4F7BBAFFB8C8534D3C156AFD77DEB2" box="[606,681,1391,1415]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B36E922FFB8C8534DD8156AFCD7DEF8" ID-GBIF-Occurrence="2565952775" collectingDate="2012-01" collectingDateMax="2303-01" collectingDateMin="2012-01" collectionCode="USNM" collectorName="Di &amp; June" country="Costa Rica" county="Di" elevation="520" location="Ni" municipality="Potrero Argentinai" pageId="15" pageNumber="1223" specimenCount="4" specimenCount-female="4">
1
<collectingCounty id="62809BF3FFB8C8534DAD156AFD37DEB2" box="[719,745,1391,1415]" pageId="15" pageNumber="1223">♂,</collectingCounty>
<specimenCount id="9D5828F6FFB8C8534D92156AFCC4DEB2" box="[752,794,1391,1415]" pageId="15" pageNumber="1223" type="female">2 ♀</specimenCount>
<collectingMunicipality id="6B857905FFB8C8534C40156AFC2FDEB2" box="[802,1009,1391,1415]" pageId="15" pageNumber="1223">Potrero Argentina,</collectingMunicipality>
10.89,021°
<location id="8E81B5A4FFB8C8534B0F156AFB5ADEB2" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534B0F156AFB5ADEB2" box="[1133,1156,1391,1415]" country="Costa Rica" county="Di" municipality="Potrero Argentinai" name="Ni" pageId="15" pageNumber="1223">N,</location>
85.38,803°
<location id="8E81B5A4FFB8C8534E411597FEE1DE9F" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534E411597FEE1DE9F" box="[291,319,1426,1450]" country="Costa Rica" county="Di" municipality="Potrero Argentinai" name="Wi" pageId="15" pageNumber="1223">W,</location>
<quantity id="4CA64E9AFFB8C8534E281597FE4BDE9F" box="[330,405,1426,1450]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="15" pageNumber="1223" unit="m" value="520.0">
<elevation id="0073044CFFB8C8534E281597FE4BDE9F" box="[330,405,1426,1450]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="15" pageNumber="1223" unit="m" value="520.0">520 m</elevation>
</quantity>
: 1
<collectorName id="26AB86A9FFB8C8534EDE1597FE08DE9F" box="[444,470,1426,1450]" pageId="15" pageNumber="1223">♂,</collectorName>
larva collected 5
<collectorName id="26AB86A9FFB8C8534DCA1597FD03DE9F" box="[680,733,1426,1450]" pageId="15" pageNumber="1223">June</collectorName>
<date id="FFE0C5BFFFB8C8534D851597FCFBDE9F" box="[743,805,1426,1450]" pageId="15" pageNumber="1223" value="2012-01">
<collectingDate id="EFA43C57FFB8C8534D851597FCFBDE9F" box="[743,805,1426,1450]" pageId="15" pageNumber="1223" value="2012-01">2012,</collectingDate>
</date>
adult emerged 25
<collectorName id="26AB86A9FFB8C8534B6A1597FBE3DE9F" box="[1032,1085,1426,1450]" pageId="15" pageNumber="1223">June</collectorName>
<date id="FFE0C5BFFFB8C8534B251597FB5BDE9F" box="[1095,1157,1426,1450]" pageId="15" pageNumber="1223" value="2012-01">
<collectingDate id="EFA43C57FFB8C8534B251597FB5BDE9F" box="[1095,1157,1426,1450]" pageId="15" pageNumber="1223" value="2012-01">2012,</collectingDate>
</date>
food plant:
<taxonomicName id="4C5E98FCFFB8C8534E4615B0FE24DEF8" box="[292,506,1461,1485]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="1223" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="B92A3F6DFFB8C8534E4615B0FE24DEF8" box="[292,506,1461,1485]" italics="true" pageId="15" pageNumber="1223">Terminalia oblonga</emphasis>
</taxonomicName>
, 12-SRNP-2306 (
<collectionCode id="ED4F7BBAFFB8C8534DD115B0FD20DEF8" box="[691,766,1461,1485]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">USNM</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B36E922FFB8C8534C7215B0FDB9DD27" ID-GBIF-Occurrence="2565952754" collectingDate="2012-01" collectingDateMax="2303-01" collectingDateMin="2012-01" collectionCode="USNM" collectorName="Di &amp; June" country="Costa Rica" county="Di" elevation="520" location="Ni" municipality="Potrero Argentinai" pageId="15" pageNumber="1223" specimenCount="4" specimenCount-female="4">
<specimenCount id="9D5828F6FFB8C8534C7215B0FCE4DEF9" box="[784,826,1460,1485]" pageId="15" pageNumber="1223" type="female">1 ♀</specimenCount>
, larva collected 5 June
<date id="FFE0C5BFFFB8C8534B2515B0FB5BDEF8" box="[1095,1157,1461,1485]" pageId="15" pageNumber="1223" value="2012-01">
<collectingDate id="EFA43C57FFB8C8534B2515B0FB5BDEF8" box="[1095,1157,1461,1485]" pageId="15" pageNumber="1223" value="2012-01">2012,</collectingDate>
</date>
adult emerged 22 June
<date id="FFE0C5BFFFB8C8534ED715D2FE2DDEDA" box="[437,499,1495,1519]" pageId="15" pageNumber="1223" value="2012-01">
<collectingDate id="EFA43C57FFB8C8534ED715D2FE2DDEDA" box="[437,499,1495,1519]" pageId="15" pageNumber="1223" value="2012-01">2012,</collectingDate>
</date>
food plant:
<taxonomicName id="4C5E98FCFFB8C8534DE015D2FC87DEDA" box="[642,857,1495,1519]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="1223" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="B92A3F6DFFB8C8534DE015D2FC87DEDA" box="[642,857,1495,1519]" italics="true" pageId="15" pageNumber="1223">Terminalia oblonga</emphasis>
</taxonomicName>
, 12-SRNP-2304 [abdomen missing, no genital preparation] (
<collectionCode id="ED4F7BBAFFB8C8534D7015FFFD83DD27" box="[530,605,1530,1554]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">USNM</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B36E922FFB8C8534D0F15FFFDDEDD62" ID-GBIF-Occurrence="2565952781" collectingDate="2012-01" collectingDateMax="2303-01" collectingDateMin="2012-01" collectionCode="USNM" collectorName="Di &amp; June" country="Costa Rica" county="Di" elevation="520" location="Ni" municipality="Potrero Argentinai" pageId="15" pageNumber="1223" specimenCount="4" specimenCount-female="4">
<specimenCount id="9D5828F6FFB8C8534D0F15FFFD4BDD27" box="[621,661,1530,1554]" pageId="15" pageNumber="1223" type="female">1 ♀</specimenCount>
, larva collected 5 June
<date id="FFE0C5BFFFB8C8534CF915FFFC07DD27" box="[923,985,1530,1554]" pageId="15" pageNumber="1223" value="2012-01">
<collectingDate id="EFA43C57FFB8C8534CF915FFFC07DD27" box="[923,985,1530,1554]" pageId="15" pageNumber="1223" value="2012-01">2012,</collectingDate>
</date>
adult emerged 1 July
<date id="FFE0C5BFFFB8C8534F941618FEEADD00" box="[246,308,1565,1589]" pageId="15" pageNumber="1223" value="2012-01">
<collectingDate id="EFA43C57FFB8C8534F941618FEEADD00" box="[246,308,1565,1589]" pageId="15" pageNumber="1223" value="2012-01">2012,</collectingDate>
</date>
food plant:
<taxonomicName id="4C5E98FCFFB8C8534EB31618FD73DD00" box="[465,685,1565,1589]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="1223" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="B92A3F6DFFB8C8534EB31618FD73DD00" box="[465,685,1565,1589]" italics="true" pageId="15" pageNumber="1223">Terminalia oblonga</emphasis>
</taxonomicName>
, 12-SRNP-
<date id="FFE0C5BFFFB8C8534C491618FCB8DD00" box="[811,870,1565,1589]" pageId="15" pageNumber="1223" value="2303-01">
<collectingDate id="EFA43C57FFB8C8534C491618FCB8DD00" box="[811,870,1565,1589]" pageId="15" pageNumber="1223" value="2303-01">2303,</collectingDate>
</date>
DNA and genital prep. USNMENT 01373361 (
<collectionCode id="ED4F7BBAFFB8C8534EC9163AFE28DD62" box="[427,502,1599,1623]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B36E922FFB8C8534D70163AFB5EDD62" ID-GBIF-Occurrence="2565952783" box="[530,1152,1599,1623]" collectionCode="USNM" country="Costa Rica" location="Di No" pageId="15" pageNumber="1223" specimenCount="1">
1
<location id="8E81B5A4FFB8C8534D51163AFDA1DD62" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534D51163AFDA1DD62" box="[563,639,1599,1623]" country="Costa Rica" name="Di No" pageId="15" pageNumber="1223">♂, No</location>
collecting data: 06-SRNP-108831 (
<collectionCode id="ED4F7BBAFFB8C8534B4D163AFBA4DD62" box="[1071,1146,1599,1623]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="1223" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B36E922FFB8C8534FC21667FE59DDA8" ID-GBIF-Occurrence="2565952707" collectingDate="1972-01" collectingDateMax="1972-01-25" collectingDateMin="1972-01" collectionCode="VOB" collectorName="Di Turrialbai &amp; May &amp; June" country="Costa Rica" elevation="600" isTailEnumeration="true" location="Cartago" pageId="15" pageNumber="1223" specimenCount="1" stateProvince="Cartago">
<collectingRegion id="499A2D9DFFB8C8534FC21667FF25DD4F" box="[160,251,1634,1658]" country="Costa Rica" name="Cartago" pageId="15" pageNumber="1223">Cartago</collectingRegion>
: 2
<collectorName id="26AB86A9FFB8C8534E451667FE69DD4F" box="[295,439,1634,1658]" pageId="15" pageNumber="1223">♂, Turrialba,</collectorName>
<quantity id="4CA64E9AFFB8C8534EA71667FDCADD4F" box="[453,532,1634,1658]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="1223" unit="m" value="600.0">
<elevation id="0073044CFFB8C8534EA71667FDCADD4F" box="[453,532,1634,1658]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="1223" unit="m" value="600.0">600 m</elevation>
</quantity>
: 20
<collectorName id="26AB86A9FFB8C8534D2D1667FDA0DD4F" box="[591,638,1634,1658]" pageId="15" pageNumber="1223">May</collectorName>
<date id="FFE0C5BFFFB8C8534DEE1667FD2FDD4F" box="[652,753,1634,1658]" pageId="15" pageNumber="1223" value="1972-01-25">
<collectingDate id="EFA43C57FFB8C8534DEE1667FD2FDD4F" box="[652,753,1634,1658]" pageId="15" pageNumber="1223" value="1972-01-25">1972, 25</collectingDate>
</date>
<collectorName id="26AB86A9FFB8C8534C621667FCEBDD4F" box="[768,821,1634,1658]" pageId="15" pageNumber="1223">June</collectorName>
<date id="FFE0C5BFFFB8C8534C211667FC5EDD4F" box="[835,896,1634,1658]" pageId="15" pageNumber="1223" value="1972-01">
<collectingDate id="EFA43C57FFB8C8534C211667FC5EDD4F" box="[835,896,1634,1658]" pageId="15" pageNumber="1223" value="1972-01">1972,</collectingDate>
</date>
V.O. Becker col., Col.
<location id="8E81B5A4FFB8C8534FC21680FF34DDA8" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534FC21680FF34DDA8" box="[160,234,1669,1693]" country="Costa Rica" name="Becker" pageId="15" pageNumber="1223">Becker</location>
55965 (
<collectionCode id="ED4F7BBAFFB8C8534E291680FE5FDDA8" box="[331,385,1669,1693]" pageId="15" pageNumber="1223">VOB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B36E922FFB8C8534EF71680FC6FDDA8" ID-GBIF-Occurrence="2565952784" box="[405,945,1668,1693]" collectorName="Di Sixola &amp; Sixaola &amp; River &amp; Marchi W" country="Costa Rica" location="Limon" pageId="15" pageNumber="1223" specimenCount="1" stateProvince="Limon">
<collectingRegion id="499A2D9DFFB8C8534EF71680FE00DDA8" box="[405,478,1669,1693]" country="Costa Rica" name="Limon" pageId="15" pageNumber="1223">Limón</collectingRegion>
: 1
<collectorName id="26AB86A9FFB8C8534D671681FDB2DDA8" box="[517,620,1668,1693]" pageId="15" pageNumber="1223">♂, Sixola</collectorName>
[
<collectorName id="26AB86A9FFB8C8534D1E1680FD0EDDA8" box="[636,720,1669,1693]" pageId="15" pageNumber="1223">Sixaola</collectorName>
]
<collectorName id="26AB86A9FFB8C8534D821680FCC4DDA8" box="[736,794,1669,1693]" pageId="15" pageNumber="1223">River</collectorName>
: 3
<collectorName id="26AB86A9FFB8C8534C231680FC6FDDA8" box="[833,945,1669,1693]" pageId="15" pageNumber="1223">March, W</collectorName>
</materialsCitation>
.
<materialsCitation id="3B36E922FFB8C8534CDD1680FE2EDD8A" ID-GBIF-Occurrence="2565952758" collectionCode="NHMUK" country="Costa Rica" location="Schausi" pageId="15" pageNumber="1223" specimenCode="NHMUK 010890497" specimenCount="1">
<location id="8E81B5A4FFB8C8534CDD1680FBCDDDA8" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534CDD1680FBCDDDA8" box="[959,1043,1669,1693]" country="Costa Rica" name="Schausi" pageId="15" pageNumber="1223">Schaus,</location>
191132,
<specimenCode id="DBF84B04FFB8C8534FC216A2FEA2DD8A" box="[160,380,1703,1727]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="1223" type="Museum">NHMUK 010890497</specimenCode>
(
<collectionCode id="ED4F7BBAFFB8C8534EE916A2FE34DD8A" box="[395,490,1703,1727]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="1223" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B36E922FFB8C84C4E9916A2FE02DB93" ID-GBIF-Occurrence="2565952769" collectionCode="USNM" collectorName="Di Sixola &amp; River &amp; Marchi Collection Wm Schausi" country="Costa Rica" lastPageId="16" lastPageNumber="1224" location="Sixaola" pageId="15" pageNumber="1223" specimenCount="1">
1
<collectorName id="26AB86A9FFB8C8534D7016A2FDA9DD8A" box="[530,631,1703,1727]" pageId="15" pageNumber="1223">♂, Sixola</collectorName>
[
<location id="8E81B5A4FFB8C8534DE616A2FD06DD8A" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFB8C8534DE616A2FD06DD8A" box="[644,728,1703,1727]" country="Costa Rica" name="Sixaola" pageId="15" pageNumber="1223">Sixaola</location>
]
<collectorName id="26AB86A9FFB8C8534D8416A2FCFEDD8A" box="[742,800,1703,1727]" pageId="15" pageNumber="1223">River</collectorName>
:
<collectorName id="26AB86A9FFB8C8534C4E16A2FB5ADD8A" box="[812,1156,1703,1727]" pageId="15" pageNumber="1223">March, Collection Wm Schaus,</collectorName>
USNM-Mimal: 2552 (
<collectionCode id="ED4F7BBAFFA7C84C4EE8108BFE0BDB93" box="[394,469,142,166]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="1224" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B36E922FFA7C84C4E88108BFEA1DBFC" ID-GBIF-Occurrence="2565952731" collectionCode="CMNH" collectorName="River" country="Costa Rica" location="Sixola" pageId="16" pageNumber="1224" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D5828F6FFA7C84C4E88108BFDC8DB93" box="[490,534,142,166]" pageId="16" pageNumber="1224" type="female">1 ♀</specimenCount>
,
<location id="8E81B5A4FFA7C84C4D44108BFDB6DB93" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFA7C84C4D44108BFDB6DB93" box="[550,616,142,166]" country="Costa Rica" name="Sixola" pageId="16" pageNumber="1224">Sixola</location>
[Sixaola]
<collectorName id="26AB86A9FFA7C84C4DB9108BFCCCDB93" box="[731,786,142,166]" pageId="16" pageNumber="1224">River</collectorName>
[photo examined only, sex not confirmed] (
<collectionCode id="ED4F7BBAFFA7C84C4E4E10B4FEA4DBFC" box="[300,378,177,201]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="16" pageNumber="1224" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B36E922FFA7C84C4EEC10B4FC0CDBD9" ID-GBIF-Occurrence="2565952735" collectingDate="1939-01-01" collectionCode="NHMUK" collectorName="Di Esperanza &amp; Mayi Rothschild Bequest BM" country="Costa Rica" location="Unknown province" pageId="16" pageNumber="1224" specimenCode="NHMUK 010890496, NHMUK 010402344" specimenCount="1">
<location id="8E81B5A4FFA7C84C4EEC10B4FDB4DBFC" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFA7C84C4EEC10B4FDB4DBFC" box="[398,618,177,201]" country="Costa Rica" name="Unknown province" pageId="16" pageNumber="1224">Unknown province</location>
: 1
<collectorName id="26AB86A9FFA7C84C4DF010B4FCF5DBFC" box="[658,811,177,201]" pageId="16" pageNumber="1224">♂, Esperanza</collectorName>
:
<collectorName id="26AB86A9FFA7C84C4C5B10B4FB5DDBFC" box="[825,1155,177,201]" pageId="16" pageNumber="1224">May, Rothschild Bequest BM</collectorName>
<date id="FFE0C5BFFFA7C84C4FC210D1FF29DBD9" box="[160,247,211,236]" pageId="16" pageNumber="1224" value="1939-01-01">
<collectingDate id="EFA43C57FFA7C84C4FC210D1FF29DBD9" box="[160,247,211,236]" pageId="16" pageNumber="1224" value="1939-01-01">19391,</collectingDate>
</date>
<specimenCode id="DBF84B04FFA7C84C4E6210D1FE3EDBD9" box="[256,480,212,236]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="1224" type="Museum">NHMUK 010890496</specimenCode>
, genitalia vial
<specimenCode id="DBF84B04FFA7C84C4DE010D1FC80DBD9" box="[642,862,212,236]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="1224" type="Museum">NHMUK 010402344</specimenCode>
(
<collectionCode id="ED4F7BBAFFA7C84C4C0F10D1FC12DBD9" box="[877,972,212,236]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="1224" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE1E37FFFA7C84C4FC2111DFECADA89" blockId="16.[160,1156,280,444]" pageId="16" pageNumber="1224">
<materialsCitation id="3B36E922FFA7C84C4FC2111DFD24DA61" ID-GBIF-Occurrence="2565952776" collectingDate="1935-12" collectingDateMax="1936-01" collectingDateMin="1935-12" collectionCode="CUIC" collectorName="Di No" country="Panama" location="Panama" pageId="16" pageNumber="1224" specimenCount="1" stateProvince="Panama">
<emphasis id="B92A3F6DFFA7C84C4FC2111DFED4DA04" bold="true" box="[160,266,280,305]" italics="true" pageId="16" pageNumber="1224">
<collectingCountry id="F349A3EFFFA7C84C4FC2111DFEDBDA04" box="[160,261,280,305]" name="Panama" pageId="16" pageNumber="1224">Panama</collectingCountry>
.
</emphasis>
2
<collectorName id="26AB86A9FFA7C84C4E5E111CFE54DA04" box="[316,394,281,305]" pageId="16" pageNumber="1224">♂, No</collectorName>
specific locality,
<date id="FFE0C5BFFFA7C84C4D13111CFCEFDA04" box="[625,817,281,305]" pageId="16" pageNumber="1224" value="1935-12">
<collectingDate id="EFA43C57FFA7C84C4D13111CFCEFDA04" box="[625,817,281,305]" pageId="16" pageNumber="1224" value="1935-12">December 1935</collectingDate>
</date>
January
<date id="FFE0C5BFFFA7C84C4CCB111CFC39DA04" box="[937,999,281,305]" pageId="16" pageNumber="1224" value="1936-01">
<collectingDate id="EFA43C57FFA7C84C4CCB111CFC39DA04" box="[937,999,281,305]" pageId="16" pageNumber="1224" value="1936-01">1936,</collectingDate>
</date>
L.M.
<collectorName id="26AB86A9FFA7C84C4B5E111CFEF3DA61" pageId="16" pageNumber="1224">Smith, Franclemont</collectorName>
dissection 1771 (
<collectionCode id="ED4F7BBAFFA7C84C4E9F1139FDE4DA61" box="[509,570,316,340]" country="USA" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="16" pageNumber="1224" type="University or college">CUIC</collectionCode>
).
<collectingRegion id="499A2D9DFFA7C84C4D2C1139FD76DA61" box="[590,680,316,340]" country="Panama" name="Panama" pageId="16" pageNumber="1224">Panamá</collectingRegion>
<collectorName id="26AB86A9FFA7C84C4DD11139FD28DA61" box="[691,758,316,340]" pageId="16" pageNumber="1224">Oeste</collectorName>
:
</materialsCitation>
<materialsCitation id="3B36E922FFA7C84C4C661139FF03DA43" ID-GBIF-Occurrence="2565952734" collectionCode="CUIC" collectorName="Smithi Franclemont" country="Panama" location="Di Barro Colorado Island" pageId="16" pageNumber="1224" specimenCount="1" stateProvince="Panama">
6
<location id="8E81B5A4FFA7C84C4C7C113EFBE3DA61" LSID="urn:lsid:plazi:treatment:03F75269FFB9C84D4EAF11FCFF35DBFC:8E81B5A4FFA7C84C4C7C113EFBE3DA61" box="[798,1085,315,340]" country="Panama" name="Di Barro Colorado Island" pageId="16" pageNumber="1224" stateProvince="Panama">♂, Barro Colorado Island</location>
(4
<collectorName id="26AB86A9FFA7C84C4B08113EFB5ADA61" box="[1130,1156,315,340]" pageId="16" pageNumber="1224">♂,</collectorName>
<collectionCode id="ED4F7BBAFFA7C84C4FC2115BFF07DA43" box="[160,217,350,374]" country="USA" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="16" pageNumber="1224" type="University or college">CUIC</collectionCode>
;
</materialsCitation>
<materialsCitation id="3B36E922FFA7C84C4F8A115BFECFDA89" ID-GBIF-Occurrence="2565952726" collectionCode="CUIC" collectorName="Smithi Franclemont" country="Panama" location="Di Barro Colorado Island" pageId="16" pageNumber="1224" specimenCount="1" stateProvince="Panama">
2 ♂
<collectionCode id="ED4F7BBAFFA7C84C4E42115BFE8DDA43" box="[288,339,350,374]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="1224">CNC</collectionCode>
) [an additional 20+ specimens from Barro Colorado Island are in the Museum of Comparative Zoology, Harvard University, USA, although we did not examine them]
</materialsCitation>
.
</paragraph>
<paragraph id="8BE1E37FFFA7C84C4FC211EDFD03D911" blockId="16.[160,1155,488,548]" pageId="16" pageNumber="1224">
<emphasis id="B92A3F6DFFA7C84C4FC211EDFEC7D934" bold="true" box="[160,281,488,513]" italics="true" pageId="16" pageNumber="1224">
<collectingCountry id="F349A3EFFFA7C84C4FC211EDFECAD934" box="[160,276,488,513]" name="Colombia" pageId="16" pageNumber="1224">Colombia</collectingCountry>
.
</emphasis>
<collectingRegion id="499A2D9DFFA7C84C4E4A11ECFE79D934" box="[296,423,489,513]" country="Colombia" name="Magdalena" pageId="16" pageNumber="1224">Magdalena</collectingRegion>
: 1 ♂, Don Amo,
<quantity id="4CA64E9AFFA7C84C4D1911ECFD0ED934" box="[635,720,489,513]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="16" pageNumber="1224" unit="ft" value="2000.0">2000 ft</quantity>
: July, H.H. Smith [leg.], genitalia vial
<collectionCode id="ED4F7BBAFFA7C84C4FC21209FF26D911" box="[160,248,524,548]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="1224" type="Museum">NHMUK</collectionCode>
010402343,
<collectionCode id="ED4F7BBAFFA7C84C4EE81209FE3CD911" box="[394,482,524,548]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="1224" type="Museum">NHMUK</collectionCode>
010890495 (
<collectionCode id="ED4F7BBAFFA7C84C4D171209FD0AD911" box="[629,724,524,548]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="1224" type="Museum">NHMUK</collectionCode>
).
</paragraph>
<paragraph id="8BE1E37FFFA7C84C4FC21254FD99D95F" blockId="16.[160,1155,593,687]" box="[160,583,593,618]" pageId="16" pageNumber="1224">
<emphasis id="B92A3F6DFFA7C84C4FC21254FD99D95F" bold="true" box="[160,583,593,618]" italics="true" pageId="16" pageNumber="1224">Photo of living specimen examined</emphasis>
</paragraph>
<paragraph id="8BE1E37FFFA7C84C4FC21271FE1CD99A" blockId="16.[160,1155,593,687]" pageId="16" pageNumber="1224">
1 ♂,
<collectingCountry id="F349A3EFFFA7C84C4FB91271FE8AD9B9" box="[219,340,628,652]" name="Costa Rica" pageId="16" pageNumber="1224">Costa Rica</collectingCountry>
:
<collectingRegion id="499A2D9DFFA7C84C4E031271FE65D9B9" box="[353,443,628,652]" country="Costa Rica" name="Heredia" pageId="16" pageNumber="1224">Heredia</collectingRegion>
: La Selva Biological Station:
<date id="FFE0C5BFFFA7C84C4C671271FC6CD9B9" box="[773,946,628,652]" pageId="16" pageNumber="1224" value="2015-03-16">16 March 2015</date>
(
<figureCitation id="1365FFFAFFA7C84C4CA11271FBF1D9B9" box="[963,1071,628,652]" captionStart="Figure 10" captionStartId="10.[160,225,753,776]" captionTargetBox="[163,1153,142,736]" captionTargetId="figure@10.[163,1153,142,736]" captionTargetPageId="10" captionText="Figure 10. Zaphanta fraterna in nature; Costa Rica, Heredia, La Selva Biological Station (Photo courtesy of Lena Struwe, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.3673469" httpUri="https://zenodo.org/record/3673469/files/figure.png" pageId="16" pageNumber="1224">Figure 10</figureCitation>
, photo courtesy of Lena Struwe).
</paragraph>
</subSubSection>
<subSubSection id="C344B0F4FFA7C84C4FC212DEFC76D869" pageId="16" pageNumber="1224" type="distribution">
<paragraph id="8BE1E37FFFA7C84C4FC212DEFEF2D9C1" blockId="16.[160,1156,731,860]" box="[160,300,731,756]" pageId="16" pageNumber="1224">
<emphasis id="B92A3F6DFFA7C84C4FC212DEFEF2D9C1" bold="true" box="[160,300,731,756]" italics="true" pageId="16" pageNumber="1224">Distribution</emphasis>
</paragraph>
<paragraph id="8BE1E37FFFA7C84C4FC212FAFC76D869" blockId="16.[160,1156,731,860]" pageId="16" pageNumber="1224">
This species is distributed from
<collectingCountry id="F349A3EFFFA7C84C4D6A12FAFD5FD822" box="[520,641,767,791]" name="Guatemala" pageId="16" pageNumber="1224">Guatemala</collectingCountry>
and Belize, to the south apparently throughout Central America, although records are lacking between
<collectingCountry id="F349A3EFFFA7C84C4C241327FC1ED80F" box="[838,960,802,826]" name="Guatemala" pageId="16" pageNumber="1224">Guatemala</collectingCountry>
/
<collectingCountry id="F349A3EFFFA7C84C4CAA1327FBD4D80F" box="[968,1034,802,826]" name="Belize" pageId="16" pageNumber="1224">Belize</collectingCountry>
and
<collectingCountry id="F349A3EFFFA7C84C4B271327FF0ED869" name="Costa Rica" pageId="16" pageNumber="1224">Costa Rica</collectingCountry>
/
<collectingCountry id="F349A3EFFFA7C84C4FBA1341FEEDD869" box="[216,307,836,860]" name="Panama" pageId="16" pageNumber="1224">Panama</collectingCountry>
.
<taxonomicName id="4C5E98FCFFA7C84C4E5D1341FDD7D869" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[319,521,836,860]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFA7C84C4E5D1341FDD7D869" box="[319,521,836,860]" italics="true" pageId="16" pageNumber="1224">Zaphanta fraterna</emphasis>
</taxonomicName>
is also found in northern
<collectingCountry id="F349A3EFFFA7C84C4C541341FC7AD869" box="[822,932,836,860]" name="Colombia" pageId="16" pageNumber="1224">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C344B0F4FFA7C84D4FC2138CFF35DBFC" lastPageId="17" lastPageNumber="1225" pageId="16" pageNumber="1224" type="discussion">
<paragraph id="8BE1E37FFFA7C84C4FC2138CFED8D897" blockId="16.[160,1156,905,1727]" box="[160,262,905,930]" pageId="16" pageNumber="1224">
<emphasis id="B92A3F6DFFA7C84C4FC2138CFED8D897" bold="true" box="[160,262,905,930]" italics="true" pageId="16" pageNumber="1224">Remarks</emphasis>
</paragraph>
<paragraph id="8BE1E37FFFA7C84C4FC213A8FDD9DF82" blockId="16.[160,1156,905,1727]" pageId="16" pageNumber="1224">
Since
<bibRefCitation id="EFCF9E8EFFA7C84C4F8113A8FEA0D8F0" author="Schaus W" box="[227,382,941,965]" pageId="16" pageNumber="1224" pagination="635 - 672" refId="ref17433" refString="Schaus W. 1928. Familie Mimallonidae. In: Seitz Ai editor. Die Gross-Schmetterlinge der Erde 6 Die amerikanischen Spinn und Schwarmer. Stuttgart: A. Kernen; p. 635 - 672." type="book chapter" year="1928">Schaus (1928)</bibRefCitation>
and until
<bibRefCitation id="EFCF9E8EFFA7C84C4E9313A8FC86D8F0" author="St Laurent RAi Kawahara AY" box="[497,856,941,965]" pageId="16" pageNumber="1224" pagination="1 - 114" refId="ref17584" refString="St Laurent RAi Kawahara AY. 2019. Reclassification of the sack-bearer moths (Lepidopterai Mimallonoideai Mimallonidae). Zookeys. 815: 1 - 114. doi: 10.3897 / zookeys. 815.27335." type="journal article" year="2019">St Laurent and Kawahara (2019)</bibRefCitation>
, this species was regarded as a synonym of
<taxonomicName id="4C5E98FCFFA7C84C4E0113CAFE02D8D2" authorityName=": Forbes" authorityYear="1942" box="[355,476,975,999]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="infantilis">
<emphasis id="B92A3F6DFFA7C84C4E0113CAFE02D8D2" box="[355,476,975,999]" italics="true" pageId="16" pageNumber="1224">Z. infantilis</emphasis>
</taxonomicName>
. However, these two species occupy different regions, with
<taxonomicName id="4C5E98FCFFA7C84C4FC213F7FECADF3F" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[160,276,1010,1034]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFA7C84C4FC213F7FECADF3F" box="[160,276,1010,1034]" italics="true" pageId="16" pageNumber="1224">Z. fraterna</emphasis>
</taxonomicName>
in north-western South America and Central America and
<taxonomicName id="4C5E98FCFFA7C84C4CDE13F7FBE8DF3F" authorityName=": Forbes" authorityYear="1942" box="[956,1078,1010,1034]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="infantilis">
<emphasis id="B92A3F6DFFA7C84C4CDE13F7FBE8DF3F" box="[956,1078,1010,1034]" italics="true" pageId="16" pageNumber="1224">Z. infantilis</emphasis>
</taxonomicName>
in the Guianas and
<collectingCountry id="F349A3EFFFA7C84C4E561410FE4DDF18" box="[308,403,1045,1069]" name="Trinidad and Tobago" pageId="16" pageNumber="1224">Trinidad</collectingCountry>
. The genitalia differ markedly between these two species, particularly by the presence of spined valvae apodemes in
<taxonomicName id="4C5E98FCFFA7C84C4D981432FCAADF7A" authorityName=": Forbes" authorityYear="1942" box="[762,884,1079,1103]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="infantilis">
<emphasis id="B92A3F6DFFA7C84C4D981432FCAADF7A" box="[762,884,1079,1103]" italics="true" pageId="16" pageNumber="1224">Z. infantilis</emphasis>
</taxonomicName>
and unmodified valvae apodemes in
<taxonomicName id="4C5E98FCFFA7C84C4E5F145FFE6FDF47" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[317,433,1114,1138]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFA7C84C4E5F145FFE6FDF47" box="[317,433,1114,1138]" italics="true" pageId="16" pageNumber="1224">Z. fraterna</emphasis>
</taxonomicName>
. The Peruvian and Ecuadorian
<taxonomicName id="4C5E98FCFFA7C84C4C7A145FFC56DF47" authority="St Laurent &amp; Giusti, 2019" authorityName="St Laurent &amp; Giusti" authorityYear="2019" box="[792,904,1114,1138]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="rawlinsi" status="sp. nov.">
<emphasis id="B92A3F6DFFA7C84C4C7A145FFC56DF47" box="[792,904,1114,1138]" italics="true" pageId="16" pageNumber="1224">Z. rawlinsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2198216FFA7C84C4CF0145FFC35DF47" box="[914,1003,1114,1138]" pageId="16" pageNumber="1224" rank="species">sp. nov.</taxonomicNameLabel>
has genitalia very similar to
<taxonomicName id="4C5E98FCFFA7C84C4E331478FE19DFA0" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[337,455,1149,1173]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFA7C84C4E331478FE19DFA0" box="[337,455,1149,1173]" italics="true" pageId="16" pageNumber="1224">Z. fraterna</emphasis>
</taxonomicName>
but differs in subuncus and phallus structure and occupies a distant biogeographic region.
</paragraph>
<paragraph id="8BE1E37FFFA7C84C4FA214C7FC2EDE2A" blockId="16.[160,1156,905,1727]" pageId="16" pageNumber="1224">
<taxonomicName id="4C5E98FCFFA7C84C4FA214C7FE54DFEF" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[192,394,1218,1242]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFA7C84C4FA214C7FE54DFEF" box="[192,394,1218,1242]" italics="true" pageId="16" pageNumber="1224">Zaphanta fraterna</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFCF9E8EFFA7C84C4D3D14C7FD25DFEF" author="Schaus W" box="[607,763,1218,1242]" pageId="16" pageNumber="1224" pagination="34 - 57" refId="ref17393" refString="Schaus W. 1912. II. - New species of Heterocera from Costa Rica. - XII. Ann Mag Nat Hist Incl Zool Bot Geol. 9: 34 - 57. doi: 10.1080 / 00222931208693102." type="journal article" year="1912">Schaus (1912)</bibRefCitation>
from an indeterminate number of specimens, although we presume the specimen labelled as the type in the USNM is in fact the sole type. Regardless, we designate a
<typeStatus id="54E55DDDFFA7C84C4DD01502FCC5DE2A" box="[690,795,1287,1311]" pageId="16" pageNumber="1224" type="lectotype">lectotype</typeStatus>
for this specimen.
</paragraph>
<paragraph id="8BE1E37FFFA7C84D4FA2152FFF35DBFC" blockId="16.[160,1156,905,1727]" lastBlockId="17.[160,1155,142,201]" lastPageId="17" lastPageNumber="1225" pageId="16" pageNumber="1224">
<bibRefCitation id="EFCF9E8EFFA7C84C4FA2152FFDCBDE77" author="Janzen DHi Hallwachs W" box="[192,533,1322,1346]" pageId="16" pageNumber="1224" refId="ref17209" refString="Janzen DHi Hallwachs W 2017. Dynamic database for an inventory of the macrocaterpillar faunai and its food plants and parasitoidsi of Area de Conservacion Guanacaste (ACG) i northwestern Costa Rica. [accessed 2019 Jun 15] http: // janzen. sas. upenn. edu." type="journal volume" year="2017">Janzen and Hallwachs (2017)</bibRefCitation>
reared
<taxonomicName id="4C5E98FCFFA7C84C4D15152FFD30DE77" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[631,750,1322,1346]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFA7C84C4D15152FFD30DE77" box="[631,750,1322,1346]" italics="true" pageId="16" pageNumber="1224">Z. fraterna</emphasis>
</taxonomicName>
from
<taxonomicName id="4C5E98FCFFA7C84C4C5C152FFE8CDE50" authority="A. Chev" authorityName="A. Chev" authorityYear="1909" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="1224" phylum="Tracheophyta" rank="species" species="ivorensis">
<emphasis id="B92A3F6DFFA7C84C4C5C152FFBC5DE77" box="[830,1051,1322,1346]" italics="true" pageId="16" pageNumber="1224">Terminalia ivorensis</emphasis>
A. Chev (Combretaceae)
</taxonomicName>
(voucher 09-SRNP-40335) and
<taxonomicName id="4C5E98FCFFA7C84C4DCC1548FBC5DE50" authority="(Ruiz and Pav.) Steud." authorityName="Steud." baseAuthorityName="Ruiz and Pav." box="[686,1051,1357,1381]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="1224" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="B92A3F6DFFA7C84C4DCC1548FCFFDE50" box="[686,801,1357,1381]" italics="true" pageId="16" pageNumber="1224">T. oblonga</emphasis>
(Ruiz and Pav.) Steud.
</taxonomicName>
(voucher numbers 12-SRNP-
<date id="FFE0C5BFFFA7C84C4EE3156AFE3BDEB2" box="[385,485,1391,1415]" pageId="16" pageNumber="1224" value="2303-01-12">2303, 12</date>
-SRNP-
<date id="FFE0C5BFFFA7C84C4D55156AFD45DEB2" box="[567,667,1391,1415]" pageId="16" pageNumber="1224" value="2304-01-12">2304, 12</date>
-SRNP-2306) in
<collectingCountry id="F349A3EFFFA7C84C4C06156AFC3BDEB2" box="[868,997,1391,1415]" name="Costa Rica" pageId="16" pageNumber="1224">Costa Rica</collectingCountry>
. The former
<taxonomicName id="4C5E98FCFFA7C84C4FC21597FECDDE9F" box="[160,275,1426,1450]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="1224" phylum="Tracheophyta" rank="genus">
<emphasis id="B92A3F6DFFA7C84C4FC21597FECDDE9F" box="[160,275,1426,1450]" italics="true" pageId="16" pageNumber="1224">Terminalia</emphasis>
</taxonomicName>
species is introduced in
<collectingCountry id="F349A3EFFFA7C84C4D591597FD6FDE9F" box="[571,689,1426,1450]" name="Costa Rica" pageId="16" pageNumber="1224">Costa Rica</collectingCountry>
as per
<bibRefCitation id="EFCF9E8EFFA7C84C4C6C1597FB80DE9F" author="Janzen DHi Hallwachs W" box="[782,1118,1426,1450]" pageId="16" pageNumber="1224" refId="ref17209" refString="Janzen DHi Hallwachs W 2017. Dynamic database for an inventory of the macrocaterpillar faunai and its food plants and parasitoidsi of Area de Conservacion Guanacaste (ACG) i northwestern Costa Rica. [accessed 2019 Jun 15] http: // janzen. sas. upenn. edu." type="journal volume" year="2017">Janzen and Hallwachs (2017)</bibRefCitation>
.
<bibRefCitation id="EFCF9E8EFFA7C84C4B0F1597FE54DEF8" author="St Laurent RAi Hamilton CAi Kawahara AY" pageId="16" pageNumber="1224" pagination="729 - 761" refId="ref17537" refString="St Laurent RAi Hamilton CAi Kawahara AY. 2018. Museum specimens provide phylogenomic data to resolve relationships of sack-bearer moths (Lepidopterai Mimallonoideai Mimallonidae). Syst Entomol. 43: 729 - 761. doi: 10.1111 / syen. 2018.43. issue- 4." type="journal article" year="2018">St Laurent et al. (2018)</bibRefCitation>
mentioned that another Costa Rican specimen of
<taxonomicName id="4C5E98FCFFA7C84C4CBB15B0FB90DEF8" authorityName=": St Laurent and Kawahara" authorityYear="2019" box="[985,1102,1461,1485]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B92A3F6DFFA7C84C4CBB15B0FB90DEF8" box="[985,1102,1461,1485]" italics="true" pageId="16" pageNumber="1224">Z. fraterna</emphasis>
</taxonomicName>
was reared from
<taxonomicName id="4C5E98FCFFA7C84C4E4E15D2FDFADEDA" authority="(Sw.)" baseAuthorityName="Sw." box="[300,548,1495,1519]" class="Magnoliopsida" family="Myrtaceae" genus="Myrcia" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="1224" phylum="Tracheophyta" rank="species" species="splendens">
<emphasis id="B92A3F6DFFA7C84C4E4E15D2FE39DEDA" box="[300,487,1495,1519]" italics="true" pageId="16" pageNumber="1224">Myrcia splendens</emphasis>
(Sw.)
</taxonomicName>
DC. (
<taxonomicName id="4C5E98FCFFA7C84C4D3D15D2FD08DEDA" box="[607,726,1495,1519]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="1224" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) (11-SRNP-32214), but we believe this identification was in error and therefore only know
<taxonomicName id="4C5E98FCFFA7C84C4C6815FFFCABDD27" authorityName="Dyari" authorityYear="1910" box="[778,885,1530,1554]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="genus">
<emphasis id="B92A3F6DFFA7C84C4C6815FFFCABDD27" box="[778,885,1530,1554]" italics="true" pageId="16" pageNumber="1224">Zaphanta</emphasis>
</taxonomicName>
to feed on
<taxonomicName id="4C5E98FCFFA7C84C4B6815FFFBA3DD27" box="[1034,1149,1530,1554]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="1224" phylum="Tracheophyta" rank="genus">
<emphasis id="B92A3F6DFFA7C84C4B6815FFFBA3DD27" box="[1034,1149,1530,1554]" italics="true" pageId="16" pageNumber="1224">Terminalia</emphasis>
</taxonomicName>
. Unfortunately, images of the larvae are not available of any of these specimens, although the following description was offered for 09-SRNP-40335: crema con puntitos cafes which translates to cream with brown dots. It is not known whether
<taxonomicName id="4C5E98FCFFA7C84C4B7A1667FB5DDD4F" authorityName="Dyari" authorityYear="1910" box="[1048,1155,1634,1658]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="genus">
<emphasis id="B92A3F6DFFA7C84C4B7A1667FB5DDD4F" box="[1048,1155,1634,1658]" italics="true" pageId="16" pageNumber="1224">Zaphanta</emphasis>
</taxonomicName>
larvae are case builders as is typical for
<taxonomicName id="4C5E98FCFFA7C84C4D0F1680FCD6DDA8" authorityName="Burmeisteri" authorityYear="1878" box="[621,776,1669,1693]" class="Insecta" family="Mimallonidae" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="1224" phylum="Arthropoda" rank="family">Mimallonidae</taxonomicName>
. Such life history traits would be extremely useful in understanding the evolutionary history and ancestral natural history of Mimallonidae, considering the sister relationship of
<taxonomicName id="4C5E98FCFFA6C84D4C65108BFCACDB93" authorityName="Dyari" authorityYear="1910" box="[775,882,142,166]" class="Insecta" family="Mimallonidae" genus="Zaphanta" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="1225" phylum="Arthropoda" rank="genus">
<emphasis id="B92A3F6DFFA6C84D4C65108BFCACDB93" box="[775,882,142,166]" italics="true" pageId="17" pageNumber="1225">Zaphanta</emphasis>
</taxonomicName>
to the remainder of the family.
</paragraph>
</subSubSection>
</treatment>
</document>