treatments-xml/data/03/EE/FD/03EEFD2AFFA1FF85FEB31988D6973B19.xml
2024-06-21 12:22:17 +02:00

343 lines
35 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://doi.org/10.5281/zenodo.3795980" ID-GBIF-Dataset="b8147d5e-955a-4e04-8862-0a30cbfc084b" ID-GBIF-Taxon="163877337" ID-ISSN="1734-1833" ID-Zenodo-Dep="3795980" approvalRequired="21" approvalRequired_for_textStreams="21" checkinTime="1588190682237" checkinUser="felipe" docAuthor="Pang, Hong, Tang, Xue-Fei, Booth, Roger G., Vandenberg, Natalia, Forrester, Juanita, Mchugh, Joseph &amp; Ślipiński, Adam" docDate="2020" docId="03EEFD2AFFA1FF85FEB31988D6973B19" docLanguage="en" docName="AnnalesZoologici.70.1.1-24.pdf.imf" docOrigin="Annales Zoologici 70 (1)" docStyle="DocumentStyle{}" docStyleName="annaleszoologici.2020.journal_article" docTitle="Novius Mulsant 1846" docType="treatment" docVersion="6" lastPageId="4" lastPageNumber="5" masterDocId="FFD78552FFA2FF81FFFF1B16D44F3960" masterDocTitle="Revision Of The Australian Coccinellidae (Coleoptera). Genus Novius Mulsant Of Tribe Noviini" masterLastPageNumber="24" masterPageNumber="1" pageId="3" pageNumber="4" updateTime="1643445081607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision Of The Australian Coccinellidae (Coleoptera). Genus Novius Mulsant Of Tribe Noviini</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pang, Hong</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocontrol, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, China; e-mails: Lsshpang @ mail. sysu. edu. cn, Tangxf 5 @ mail 2. sysu. edu. cn</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tang, Xue-Fei</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocontrol, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, China; e-mails: Lsshpang @ mail. sysu. edu. cn, Tangxf 5 @ mail 2. sysu. edu. cn</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Booth, Roger G.</mods:namePart>
<mods:affiliation>The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK; e-mail: r. booth @ nhm. ac. uk</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vandenberg, Natalia</mods:namePart>
<mods:affiliation>Systematic Entomology Lab, Agricultural Research Service, USDA c / o National Museum of Natural History, Smithsonian Institution P. O. Box 37012, MRC- 168, Washington, DC 20013 - 7012, U. S. A; e-mails: VandenbergN @ si. edu or Natalia. Vandenberg @ usda. gov</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Forrester, Juanita</mods:namePart>
<mods:affiliation>Division of Science, Chattahoochee Technical College, 1645 Bluffs Parkway, Canton, Georgia, 30114, U. S. A; e-mail: juanita. forrester @ gmail. com</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mchugh, Joseph</mods:namePart>
<mods:affiliation>Department of Entomology, University of Georgia, 413 Biological Sciences Building, Athens, Georgia, 30602 - 2603, U. S. A; e-mail: mchugh. jv @ gmail. com</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ślipiński, Adam</mods:namePart>
<mods:affiliation>Australian National Insect Collection, CSIRO, GPO Box 1700, Canberra ACT 2601, Australia; e-mail: Adam. Slipinski @ csiro. au</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Annales Zoologici</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-03-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>70</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>24</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3161/00034541ANZ2020.70.1.001</mods:identifier>
<mods:identifier type="GBIF-Dataset">b8147d5e-955a-4e04-8862-0a30cbfc084b</mods:identifier>
<mods:identifier type="ISSN">1734-1833</mods:identifier>
<mods:identifier type="Zenodo-Dep">3776582</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3795980" ID-GBIF-Taxon="163877337" ID-Zenodo-Dep="3795980" LSID="urn:lsid:plazi:treatment:03EEFD2AFFA1FF85FEB31988D6973B19" httpUri="http://treatment.plazi.org/id/03EEFD2AFFA1FF85FEB31988D6973B19" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection box="[332,583,670,692]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph blockId="3.[332,583,670,692]" box="[332,583,670,692]" pageId="3" pageNumber="4">
<heading box="[332,583,670,692]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName authority="Mulsant, 1846" authorityName="Mulsant" authorityYear="1846" box="[332,583,670,692]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[332,416,670,691]" italics="true" pageId="3" pageNumber="4">Novius</emphasis>
<bibRefCitation author="Mulsant, E." box="[423,583,670,692]" pageId="3" pageNumber="4" refId="ref15936" refString="Mulsant, E. 1846. Histoire Naturelle des Coleopteres de France. Sulcicolles-Securipalpes. Paris: Maison xxiv + 26 pp. + 280 pp. + 1 pl." type="book" year="1846">Mulsant, 1846</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="reference_group">
<paragraph blockId="3.[144,772,726,1183]" pageId="3" pageNumber="4">
<treatmentCitationGroup pageId="3" pageNumber="4">
<treatmentCitation author="Mulsant, E." box="[144,397,726,743]" page="213" pageId="3" pageNumber="4" year="1846">
<taxonomicName authority="Mulsant, 1846: 213" authorityName="Mulsant. Treated" authorityPageNumber="213" authorityYear="1846" box="[144,397,726,743]" class="Insecta" family="Carabidae" genus="Nomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[144,219,726,742]" italics="true" pageId="3" pageNumber="4">Nomius</emphasis>
<bibRefCitation author="Mulsant, E." box="[226,397,726,743]" pageId="3" pageNumber="4" refId="ref15936" refString="Mulsant, E. 1846. Histoire Naturelle des Coleopteres de France. Sulcicolles-Securipalpes. Paris: Maison xxiv + 26 pp. + 280 pp. + 1 pl." type="book" year="1846">Mulsant, 1846: 213</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[407,451,726,742]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName authority="Mulsant, 1846" authorityName="Mulsant" authorityYear="1846" class="Insecta" family="Carabidae" genus="Nomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cruentatus">
<emphasis box="[536,724,726,742]" italics="true" pageId="3" pageNumber="4">Nomius cruentatus</emphasis>
<bibRefCitation author="Mulsant, E." pageId="3" pageNumber="4" refId="ref15936" refString="Mulsant, E. 1846. Histoire Naturelle des Coleopteres de France. Sulcicolles-Securipalpes. Paris: Maison xxiv + 26 pp. + 280 pp. + 1 pl." type="book" year="1846">Mulsant, 1846</bibRefCitation>
</taxonomicName>
, by monotypy. Junior homonym of
<taxonomicName authority="Laporte, 1835" authorityName="Laporte" authorityYear="1835" class="Insecta" family="Carabidae" genus="Nomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[611,686,752,768]" italics="true" pageId="3" pageNumber="4">Nomius</emphasis>
<bibRefCitation author="Laporte, F. L. de" pageId="3" pageNumber="4" pagination="95 - 159" refId="ref15419" refString="Laporte, F. L. de 1835. Etudes entomologiques ou description d'Insectes nouveaux et observations sur leur synonymie. Premiere partie. Paris: Mequignon-Marvis Pere et Fils, pp. 95 - 159." type="book chapter" year="1835">Laporte, 1835</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="3.[144,772,726,1183]" pageId="3" pageNumber="4">
<taxonomicName authority="Mulsant, 1846" authorityName="Mulsant" authorityYear="1846" box="[144,343,803,820]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[144,212,804,820]" italics="true" pageId="3" pageNumber="4">Novius</emphasis>
<bibRefCitation author="Mulsant, E." box="[217,343,803,820]" pageId="3" pageNumber="4" refId="ref15936" refString="Mulsant, E. 1846. Histoire Naturelle des Coleopteres de France. Sulcicolles-Securipalpes. Paris: Maison xxiv + 26 pp. + 280 pp. + 1 pl." type="book" year="1846">Mulsant, 1846</bibRefCitation>
</taxonomicName>
: addenda et errata [p. 4]. Replacement name for
<taxonomicName authority="Mulsant. Treated" authorityName="Mulsant. Treated" authorityYear="1846" box="[181,416,829,846]" class="Insecta" family="Carabidae" genus="Nomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[181,256,830,846]" italics="true" pageId="3" pageNumber="4">Nomius</emphasis>
Mulsant. Treated
</taxonomicName>
as synonym of
<taxonomicName authorityName="Mulsant" authorityYear="1850" box="[556,633,829,846]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[556,633,829,846]" italics="true" pageId="3" pageNumber="4">Rodolia</emphasis>
</taxonomicName>
by
<bibRefCitation author="Iqbal, Z. &amp; M. F. Nasir &amp; I. Bodlah &amp; R. Qureshi" pageId="3" pageNumber="4" pagination="1103 - 1111" refId="ref14927" refString="Iqbal, Z., M. F. Nasir, I. Bodlah and R. Qureshi. 2018. A Contribution to the Genus Rodolia Mulsant, 1850 (Coleoptera: Coccinellidae) from Pothwar Plateau of Pakistan. The Journal of Animal &amp; Plant Sciences, 28 (4): 1103 - 1111." type="journal article" year="2018">
Iqbal
<emphasis box="[717,761,829,846]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
., 2018:1104
</bibRefCitation>
.
</paragraph>
<paragraph blockId="3.[144,772,726,1183]" pageId="3" pageNumber="4">
<treatmentCitationGroup pageId="3" pageNumber="4">
<treatmentCitation author="Mulsant, E." box="[144,403,881,898]" page="902" pageId="3" pageNumber="4" year="1850">
<taxonomicName authority="Mulsant, 1850: 902" authorityName="Mulsant" authorityPageNumber="902" authorityYear="1850" box="[144,403,881,898]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[144,221,881,898]" italics="true" pageId="3" pageNumber="4">Rodolia</emphasis>
<bibRefCitation author="Mulsant, E." box="[230,403,881,898]" pageId="3" pageNumber="4" pagination="1 - 1104" refId="ref15967" refString="Mulsant, E. 1850. Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2: xv + 1 - 1104 pp." type="book chapter" year="1850">Mulsant, 1850: 902</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[414,458,881,897]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis box="[546,723,881,898]" italics="true" pageId="3" pageNumber="4">Rodolia ruficollis</emphasis>
<bibRefCitation author="Mulsant, E." pageId="3" pageNumber="4" pagination="1 - 1104" refId="ref15967" refString="Mulsant, E. 1850. Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2: xv + 1 - 1104 pp." type="book chapter" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<treatmentCitation author="Crotch, G. R." box="[561,719,907,924]" page="280" pageId="3" pageNumber="4" year="1874">
<bibRefCitation author="Crotch, G. R." box="[561,719,907,924]" pageId="3" pageNumber="4" refId="ref14207" refString="Crotch, G. R. 1874. A Revision of the Coleopterous Family Coccinellidae. London: E. W. Janson, xv + 311 pp." type="book" year="1874">Crotch, 1874: 280</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation author="Korschefsky, R." page="98" pageId="3" pageNumber="4" year="1931">
<bibRefCitation author="Korschefsky, R." pageId="3" pageNumber="4" pagination="1 - 224" refId="ref15197" refString="Korschefsky, R. 1931. Pars 118, Coccinellidae I. In: Coleoptorum Catalogus, vol. 16: 1 - 224." type="journal article" year="1931">Korschefsky, 1931: 98</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Sasaji, H." box="[362,519,933,950]" page="233" pageId="3" pageNumber="4" year="1971">
<bibRefCitation author="Sasaji, H." box="[362,519,933,950]" pageId="3" pageNumber="4" refId="ref16461" refString="Sasaji, H. 1971. Coccinellidae (Insecta: Coleoptera). Fauna Japonica Academic Press of Japan, Keigaku Publishing, Tokyo, ix + 344 pp." type="book" year="1971">Sasaji, 1971: 233</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Gordon, R. D." box="[531,688,933,950]" page="25" pageId="3" pageNumber="4" year="1972">
<bibRefCitation author="Gordon, R. D." box="[531,688,933,950]" pageId="3" pageNumber="4" pagination="23 - 31" refId="ref14645" refString="Gordon, R. D. 1972. The tribe Noviini in the New World (Coleoptera: Coccinellidae). Journal of the Washington Academy of Sciences, 62 (1): 23 - 31." type="journal article" year="1972">Gordon, 1972: 25</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Gordon, R. D." page="665" pageId="3" pageNumber="4" year="1985">
<bibRefCitation author="Gordon, R. D." pageId="3" pageNumber="4" pagination="654 - 678" refId="ref14683" refString="Gordon, R. D. 1985. The Coccinellidae (Coleoptera) of America north of Mexico. Journal of the New York Entomological Society, 93: 654 - 678." type="journal article" year="1985">Gordon, 1985: 665</bibRefCitation>
</treatmentCitation>
; Gordon, 1990: 291;
<bibRefCitation author="Chazeau" box="[450,582,959,976]" pageId="3" pageNumber="4" pagination="151 - 159" refId="ref14048" refString="Chazeau J. 1991. Coleoptera Coccinellidae Noviini de Nouvelle-Caledonie. Genre Rodolia Mulsant. Memoires du Museum National d'Histoire Naturelle, Serie A Zoologie, 149: 151 - 159." type="journal article" year="1991">Chazeau, 1991</bibRefCitation>
;
<treatmentCitation author="Slipinski, S. A." box="[592,768,957,976]" page="141" pageId="3" pageNumber="4" year="2007">
<bibRefCitation author="Slipinski, S. A." box="[592,768,957,976]" pageId="3" pageNumber="4" refId="ref16790" refString="Slipinski, S. A. 2007. Australian Ladybird Beetles (Coleoptera: Coccinellidae): Their Biology and Classification. ABRS, Canberra. 286 pp." type="book" year="2007">Ślipiński, 2007: 141</bibRefCitation>
</treatmentCitation>
. Synonymized by
<treatmentCitation author="Iqbal, Z. &amp; M. F. Nasir &amp; I. Bodlah &amp; R. Qureshi" box="[333,538,984,1002]" page="1104" pageId="3" pageNumber="4" year="2018">
<bibRefCitation author="Iqbal, Z. &amp; M. F. Nasir &amp; I. Bodlah &amp; R. Qureshi" box="[333,538,984,1002]" pageId="3" pageNumber="4" pagination="1103 - 1111" refId="ref14927" refString="Iqbal, Z., M. F. Nasir, I. Bodlah and R. Qureshi. 2018. A Contribution to the Genus Rodolia Mulsant, 1850 (Coleoptera: Coccinellidae) from Pothwar Plateau of Pakistan. The Journal of Animal &amp; Plant Sciences, 28 (4): 1103 - 1111." type="journal article" year="2018">
Iqbal
<emphasis box="[383,427,984,1001]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
., 2018: 1104
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="3.[144,772,726,1183]" pageId="3" pageNumber="4">
<treatmentCitationGroup pageId="3" pageNumber="4">
<treatmentCitation author="Weise, J." box="[144,441,1010,1027]" page="63" pageId="3" pageNumber="4" year="1885">
<taxonomicName authority="Weise, 1885: 63" authorityName="Weise" authorityPageNumber="63" authorityYear="1885" box="[144,441,1010,1027]" class="Insecta" family="Coccinellidae" genus="Macronovius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[144,273,1011,1027]" italics="true" pageId="3" pageNumber="4">Macronovius</emphasis>
<bibRefCitation author="Weise, J." box="[286,441,1010,1027]" pageId="3" pageNumber="4" refId="ref16856" refString="Weise, J. 1885. Bestimmungs-Tabellen der Eeuropaischen Coleopteren. II. Heft. Coccinellidae. II. Auflage mit Beruck- sichtigung der Arten aus dem Nordlichen Asien. Modling, 83 pp." type="book" year="1885">Weise, 1885: 63</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[457,502,1011,1027]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName authority="Motschulsky, 1866" authorityName="Motschulsky" authorityYear="1866" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis box="[600,771,1010,1027]" italics="true" pageId="3" pageNumber="4">Novius limbatus</emphasis>
Motschulsky, 1866
</taxonomicName>
(subgenus of
<taxonomicName authorityName="Mulsant" authorityYear="1850" box="[474,551,1036,1053]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[474,551,1036,1053]" italics="true" pageId="3" pageNumber="4">Rodolia</emphasis>
</taxonomicName>
).
<treatmentCitation author="Weise, J." box="[571,720,1036,1053]" page="149" pageId="3" pageNumber="4" year="1895">
<bibRefCitation author="Weise, J." box="[571,720,1036,1053]" pageId="3" pageNumber="4" pagination="147 - 151" refId="ref16916" refString="Weise, J. 1895. Uber die mit Novius Mulsant. Annales de la Societe Entomologique de Belgique, 39: 147 - 151." type="journal article" year="1895">Weise, 1895: 149</bibRefCitation>
</treatmentCitation>
(synonym of
<taxonomicName authorityName="Mulsant" authorityYear="1850" box="[258,335,1062,1079]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[258,335,1062,1079]" italics="true" pageId="3" pageNumber="4">Rodolia</emphasis>
</taxonomicName>
).
<treatmentCitation author="Sicard, A." box="[354,497,1062,1079]" page="68" pageId="3" pageNumber="4" year="1907">
<bibRefCitation author="Sicard, A." box="[354,497,1062,1079]" pageId="3" pageNumber="4" pagination="132 - 136" refId="ref16554" refString="Sicard, A. 1907. Notes sur quelques Coccinellides Palearctiques (Col.). Bulletin de la Societe entomologique de France, 1907: 132 - 136." type="journal article" year="1907">Sicard, 1907: 68</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="3.[144,772,726,1183]" pageId="3" pageNumber="4">
<treatmentCitationGroup pageId="3" pageNumber="4">
<treatmentCitation author="Weise, J." box="[144,421,1088,1105]" page="149" pageId="3" pageNumber="4" year="1895">
<taxonomicName authority="Weise, 1895: 149" authorityName="Severini Weise" authorityPageNumber="149" authorityYear="1895" box="[144,421,1088,1105]" class="Insecta" family="Coccinellidae" genus="Eurodolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[144,244,1088,1105]" italics="true" pageId="3" pageNumber="4">Eurodolia</emphasis>
<bibRefCitation author="Weise, J." box="[257,421,1088,1105]" pageId="3" pageNumber="4" pagination="147 - 151" refId="ref16916" refString="Weise, J. 1895. Uber die mit Novius Mulsant. Annales de la Societe Entomologique de Belgique, 39: 147 - 151." type="journal article" year="1895">Weise, 1895: 149</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[436,481,1088,1104]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName authority="Weise, 1895" authorityName="Weise" authorityYear="1895" class="Insecta" family="Coccinellidae" genus="Eurodolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="severini">
<emphasis box="[577,771,1088,1105]" italics="true" pageId="3" pageNumber="4">Eurodolia severini</emphasis>
<bibRefCitation author="Weise, J." box="[181,289,1114,1131]" pageId="3" pageNumber="4" pagination="147 - 151" refId="ref16916" refString="Weise, J. 1895. Uber die mit Novius Mulsant. Annales de la Societe Entomologique de Belgique, 39: 147 - 151." type="journal article" year="1895">Weise, 1895</bibRefCitation>
</taxonomicName>
, by monotypy. Synonymized by
<treatmentCitation author="Slipinski, S. A." box="[574,747,1112,1131]" page="141" pageId="3" pageNumber="4" year="2007">
<bibRefCitation author="Slipinski, S. A." box="[574,747,1112,1131]" pageId="3" pageNumber="4" refId="ref16790" refString="Slipinski, S. A. 2007. Australian Ladybird Beetles (Coleoptera: Coccinellidae): Their Biology and Classification. ABRS, Canberra. 286 pp." type="book" year="2007">Ślipiński, 2007: 141</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="3.[144,772,726,1183]" pageId="3" pageNumber="4">
<treatmentCitationGroup pageId="3" pageNumber="4">
<treatmentCitation author="Casey, T. L." box="[144,375,1140,1157]" page="408" pageId="3" pageNumber="4" year="1908">
<taxonomicName authority="Casey, 1908: 408" authorityName="Casey" authorityPageNumber="408" authorityYear="1908" box="[144,375,1140,1157]" class="Insecta" family="Coccinellidae" genus="Anovia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[144,214,1140,1156]" italics="true" pageId="3" pageNumber="4">Anovia</emphasis>
<bibRefCitation author="Casey, T. L." box="[222,375,1140,1157]" pageId="3" pageNumber="4" pagination="393 - 421" refId="ref13786" refString="Casey, T. L. 1908. Notes on the Coccinellidae. Canadian Entomologist, 40: 393 - 421." type="journal article" year="1908">Casey, 1908: 408</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[385,429,1140,1156]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName authority="Wickham, 1905" authorityName="Wickham" authorityYear="1905" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="virginalis">
<emphasis box="[515,712,1140,1157]" italics="true" pageId="3" pageNumber="4">Scymnus virginalis</emphasis>
<bibRefCitation author="Wickham, H. F." pageId="3" pageNumber="4" pagination="165 - 171" refId="ref17218" refString="Wickham, H. F. 1905. New species of Coleoptera from the western United States II. Canadian Entomologist, 37: 165 - 171." type="journal article" year="1905">Wickham, 1905</bibRefCitation>
</taxonomicName>
, by monotypy. Synonymized by
<treatmentCitation author="Iqbal, Z. &amp; M. F. Nasir &amp; I. Bodlah &amp; R. Qureshi" box="[559,764,1165,1183]" page="1104" pageId="3" pageNumber="4" year="2018">
<bibRefCitation author="Iqbal, Z. &amp; M. F. Nasir &amp; I. Bodlah &amp; R. Qureshi" box="[559,764,1165,1183]" pageId="3" pageNumber="4" pagination="1103 - 1111" refId="ref14927" refString="Iqbal, Z., M. F. Nasir, I. Bodlah and R. Qureshi. 2018. A Contribution to the Genus Rodolia Mulsant, 1850 (Coleoptera: Coccinellidae) from Pothwar Plateau of Pakistan. The Journal of Animal &amp; Plant Sciences, 28 (4): 1103 - 1111." type="journal article" year="2018">
Iqbal
<emphasis box="[609,653,1165,1182]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
., 2018: 1104
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="description">
<paragraph blockId="3.[144,772,1224,1901]" lastBlockId="3.[816,1444,180,1901]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[181,332,1224,1245]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[181,325,1224,1245]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
.
</emphasis>
Length 2.55.0 mm. Body oval or elongate-oval, winged; dorsum densely hairy without distinct rows of punctures. Elytra red, yellowish, brown or black, often with colour pattern. Head broad with eyes large, finely facetted, not emarginate, densely setose, broadly separated on vertex. Antenna with 8 antennomeres, much shorter than head capsule, with scape enlarged; antennal club with 3 antennomeres, asymmetrical, with divisions often indistinct. Mandible bifid apically; molar part with basal tooth; prostheca distinct. Terminal maxillary palpomere large, strongly securiform; labial palp with 2 palpomeres. Pronotum with anterior angles not thickened, blunt, indistinct; lateral edge smooth; submarginal carina absent; hind border rarely with marginal bead. Prosternal process about 0.30.6 times coxal diameter, surface without carinae but densely setose. Prosternum in front of coxae strongly reduced; anterior margin continuing as a straight or arcuate line. Procoxal cavity distinctly transverse. Meso- metaventral process narrow, narrower than coxal diameter, junction forming a straight line, with suture visible. Metaventral postcoxal lines separated at middle, complete and recurved, very close to coxal cavities. Tibiae without apical spurs. Protibia flattened and angulate externally; protarsus trimerous; tarsal claws in female bifid, tarsal claws in male appendiculate on mid and hind legs.
<emphasis bold="true" box="[1172,1287,303,324]" pageId="3" pageNumber="4">Abdomen</emphasis>
with six ventrites in both sexes. Abdominal postcoxal lines separate medially, recurved, complete, posteriorly distinctly separated from hind margin of ventrite; ventrite
<quantity box="[1401,1444,396,418]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="4" unit="in" value="5.0">5 in</quantity>
male emarginate medially.
<emphasis bold="true" box="[1150,1367,427,449]" pageId="3" pageNumber="4">Male Terminalia:</emphasis>
parameres and phallobase symmetrical; penis guide symmetrical; parameres articulated with phallobase. Penis slender or stout; apex abbreviated or drawn out into elongate flagellum; basal capsule with more or less distinct inner and outer arms forming T-shaped junction at base of penis, except inner arm vestigial or absent in Australian species. Apodeme of male sternum IX absent.
<emphasis bold="true" box="[912,1158,674,696]" pageId="3" pageNumber="4">Female Terminalia:</emphasis>
Coxites distinctly elongate, triangular; styli absent; sperm duct simple, flexible, relatively uniform in diameter; infundibulum absent. Spermathecal capsule (
<figureCitation box="[1186,1342,767,789]" captionStart="Figures 137142" captionStartId="17.[144,212,826,843]" captionTargetBox="[151,765,182,797]" captionTargetId="figure@17.[145,775,177,807]" captionTargetPageId="17" captionText="Figures 137142. Novius species, spermatheca: (137) N. pumilus (Weise); (138) N. koebelei; (139) N. breviusculus (Weise); (140142) N. cardinalis." figureDoi="http://doi.org/10.5281/zenodo.3776606" httpUri="https://zenodo.org/record/3776606/files/figure.png" pageId="3" pageNumber="4">Figs 137142</figureCitation>
) usually with well developed cornu, sessile ramus, small tapered or indistinct nodulus; general form of capsule variable, strongly inflated, reniform to oval in many species, or slender, vermiform, with distinct bend near middle, forming hairpin loop; some species with capsule of intermediate form, slender basally with distal part bulbous to strongly inflated. Spermathecal accessory gland present, but often lost in dissections.
</paragraph>
</subSubSection>
<subSubSection box="[854,1347,1045,1067]" pageId="3" pageNumber="4" type="distribution">
<paragraph blockId="3.[816,1444,180,1901]" box="[854,1347,1045,1067]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[854,1011,1045,1067]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[854,1003,1045,1067]" italics="true" pageId="3" pageNumber="4">Distribution</emphasis>
.
</emphasis>
Same as for the tribe, above.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="discussion">
<paragraph blockId="3.[816,1444,180,1901]" lastBlockId="4.[144,771,180,633]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis box="[854,972,1076,1098]" italics="true" pageId="3" pageNumber="4">
<emphasis bold="true" box="[854,965,1076,1098]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
.
</emphasis>
Noviine lady beetles can be found on every major land mass except
<collectingCountry box="[1199,1323,1107,1129]" name="Antarctica" pageId="3" pageNumber="4">Antarctica</collectingCountry>
, yet their morphologies are surprisingly consistent throughout this range and defy our ability to subdivide the tribe into multiple genera based on external characters alone. Possibly this is due to the group having optimized their morphologies early within their evolutionary history (Zhang
<emphasis box="[1040,1109,1292,1314]" italics="true" pageId="3" pageNumber="4">et. al.</emphasis>
unpublished), presumably in response to their specializing on a particular group of prey (scale insects in the family
<taxonomicName box="[1215,1378,1354,1376]" class="Insecta" family="Margarodidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Margarodidae</taxonomicName>
) that occur on woody plants. This specialization may further correlate with a predictable set of biotic and abiotic factors which continued to exert selection pressure towards certain established morphologies. We might speculate, for example, that the compact and densely hairy forms of these noviine beetles, along with the use of carminic acid as a chemical repellant (
<bibRefCitation author="Forrester, J. A." pageId="3" pageNumber="4" refId="ref14265" refString="Forrester, J. A. 2008. Sacred systematics: the Noviini of the world (Coleoptera: Coccinellidae). A Dissertation Submitted to the Graduate Faculty of the University of Georgia in Partial Fulfillment of the Requirements for the Degree." type="book" year="2008">Forrester 2008</bibRefCitation>
), arose as a defense against ants that are drawn to the copious secretions of sugary honeydew produced by their sap-feeding scale prey. Most lady beetle tribes contain a greater array of habitat and trophic specializations than do noviines and exhibit correspondingly diverse beetle morphologies, which facilitates their classification into multiple genera. On the otherhand, internal characters within
<taxonomicName authorityName="Mulsant" authorityYear="1846" box="[1122,1205,1817,1839]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Noviini">Noviini</taxonomicName>
, such as the configuration of the male and female genitalia (see generic description, above), show greater diversity than the external characters and suggest that multiple lineages may exist within our broadly defined
<taxonomicName authorityName="Mulsant" authorityYear="1846" box="[580,668,211,232]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[580,668,211,232]" italics="true" pageId="4" pageNumber="5">Novius</emphasis>
</taxonomicName>
. A much more in depth study of molecular variation and internal anatomy is needed to provide a meaningful reconstruction of the phylogeny of this group.
</paragraph>
<paragraph blockId="4.[144,771,180,633]" pageId="4" pageNumber="5">
<bibRefCitation author="Forrester, J. A." box="[181,383,334,356]" pageId="4" pageNumber="5" refId="ref14265" refString="Forrester, J. A. 2008. Sacred systematics: the Noviini of the world (Coleoptera: Coccinellidae). A Dissertation Submitted to the Graduate Faculty of the University of Georgia in Partial Fulfillment of the Requirements for the Degree." type="book" year="2008">Forrester (2008)</bibRefCitation>
had suggested conserving the name
<taxonomicName authorityName="Mulsant" authorityYear="1850" box="[215,314,365,386]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[215,314,365,386]" italics="true" pageId="4" pageNumber="5">Rodolia</emphasis>
</taxonomicName>
because of more than 100 years of use in many papers dealing with the biological control of scales. However, since the senior synonym
<taxonomicName authorityName="Mulsant" authorityYear="1846" box="[636,724,427,448]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[636,724,427,448]" italics="true" pageId="4" pageNumber="5">Novius</emphasis>
</taxonomicName>
has been commonly used in European literature, its suppression under article 23.9.3 requires a ruling by the International Commission on Zoological Nomenclature. Therefore, given the uncertain outcome of such an application, and after some discussion, we have jointly decided to use the senior synonym,
<taxonomicName authorityName="Mulsant" authorityYear="1846" box="[634,722,612,633]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[634,722,612,633]" italics="true" pageId="4" pageNumber="5">Novius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>