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<document id="854A49FAB28848994FB581FFFC42118D" ID-DOI="10.5281/zenodo.5353844" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353844" ID-ZooBank="CA3621AC-1E54-464E-8096-5A17F605F67E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212686074" checkinUser="felipe" docAuthor="Riley, Edward G." docDate="2020" docId="03D80162FFDE7D63FF62EB8DFD61FD59" docLanguage="en" docName="0830.pdf" docOrigin="Insecta Mundi 2020 (830)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Colaspis suilla Fabricius 1801" docType="treatment" docVersion="4" lastPageNumber="7" masterDocId="FFE1791AFFD87D6BFFA2EA6EFF80FFC8" masterDocTitle="A review of the Colaspis suilla species group, with description of three new species from Florida (Coleoptera: Chrysomelidae: Eumolpinae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="5" updateTime="1699187982696" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="4C86ED34F4F3FF75E20BBC56E3F01FAF">A review of the Colaspis suilla species group, with description of three new species from Florida (Coleoptera: Chrysomelidae: Eumolpinae)</mods:title>
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<mods:namePart id="FABDFD2D4F024C046AFAE9C46A50B766">Riley, Edward G.</mods:namePart>
<mods:affiliation id="388CA9FF98D9A68B33AFB55FBA401150">Department of Entomology, Texas A &amp; M University, College Station, Texas 77843 - 2475 USA</mods:affiliation>
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<mods:title id="B1E3C562090C33DC50FA07ED5DDD3C93">Insecta Mundi</mods:title>
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<mods:date id="D21FDC8EF00F3A1F8238B35324C9ECEC">2020</mods:date>
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<treatment id="03D80162FFDE7D63FF62EB8DFD61FD59" ID-DOI="http://doi.org/10.5281/zenodo.5451797" ID-Zenodo-Dep="5451797" LSID="urn:lsid:plazi:treatment:03D80162FFDE7D63FF62EB8DFD61FD59" httpUri="http://treatment.plazi.org/id/03D80162FFDE7D63FF62EB8DFD61FD59" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5">
<subSubSection id="C36BE3FFFFDE7D6DFF62EB8DFE74FDC8" box="[192,500,483,512]" pageId="6" pageNumber="5" type="nomenclature">
<paragraph id="8BCEB074FFDE7D6DFF62EB8DFE74FDC8" blockId="6.[192,626,483,546]" box="[192,500,483,512]" pageId="6" pageNumber="5">
<heading id="D0860718FFDE7D6DFF62EB8DFE74FDC8" box="[192,500,483,512]" fontSize="12" level="2" pageId="6" pageNumber="5" reason="2">
<taxonomicName id="4C71CBF7FFDE7D6DFF62EB8DFE74FDC8" authority="Fabricius" authorityName="Fabricius" authorityYear="1801" box="[192,500,483,512]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFF62EB8DFE74FDC8" bold="true" box="[192,500,483,512]" pageId="6" pageNumber="5">
<emphasis id="B9056C66FFDE7D6DFF62EB8DFEF6FDC8" bold="true" box="[192,374,483,512]" italics="true" pageId="6" pageNumber="5">Colaspis suilla</emphasis>
Fabricius
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36BE3FFFFDE7D6DFF62E869FDF2FDE9" box="[192,626,519,546]" pageId="6" pageNumber="5" type="description">
<paragraph id="8BCEB074FFDE7D6DFF62E869FDF2FDE9" blockId="6.[192,626,483,546]" box="[192,626,519,546]" pageId="6" pageNumber="5">
(
<figureCitation id="134AACF1FFDE7D6DFF6AE869FEA5FDE9" box="[200,293,519,545]" captionStart="Figures 14" captionStartId="4.[192,279,1171,1197]" captionTargetBox="[233,1439,253,1139]" captionTargetId="figure-323@4.[192,1440,213,1140]" captionTargetPageId="4" captionText="Figures 14. Habitus of Colaspis species, dorsal and lateral. 1) Male, C. suilla F. (Lake Claiborne St. Pk., LA). 2) Female, C. suilla F. (Lake Claiborne St. Pk., LA). 3) Male, C. costipennis Crotch (Sand Hill St. Forest, SC). 4) Female, C. costipennis Crotch (Sand Hill St. Forest, SC). All figures to scale." figureDoi="http://doi.org/10.5281/zenodo.5353846" httpUri="https://zenodo.org/record/5353846/files/figure.png" pageId="6" pageNumber="5">Fig. 12</figureCitation>
,
<figureCitation id="134AACF1FFDE7D6DFE92E869FEF6FDE9" box="[304,374,519,545]" captionStart="Figures 1118" captionStartId="9.[192,279,1741,1767]" captionTargetBox="[192,1440,232,1710]" captionTargetId="figure-10@9.[192,1440,213,1710]" captionTargetPageId="9" captionText="Figures 1118. Forebodies of Colaspis suilla species group, anterodorsal and dorsolateral views. 11) Male C. ansa n. sp. (Port St. Joe, FL). 12) Male C. skelleyi n. sp. (Lake Marion Estates, FL). 13) Male C. thomasi n. sp. (Archbold Biol. Sta., FL). 14) Male C. suilla F. (Kisatchie Natl. Forest, LA). 15) Female C. suilla F. (Torreya St. Pk., FL). 16) Male C. suilla F. (Merritt Reservoir, NE). 17) Male C. costipennis Crotch (Gainesville, FL). 18) Male C. costipennis Crotch (Pensacola, FL). All figures to scale." figureDoi="http://doi.org/10.5281/zenodo.5353850" httpUri="https://zenodo.org/record/5353850/files/figure.png" pageId="6" pageNumber="5">1416</figureCitation>
,
<figureCitation id="134AACF1FFDE7D6DFE23E869FE1EFDE9" box="[385,414,519,545]" captionStart="Figures 3340" captionStartId="13.[191,278,1454,1480]" captionTargetBox="[192,1440,213,1423]" captionTargetId="figure-94@13.[192,1440,213,1423]" captionTargetPageId="13" captionText="Figures 3340. Colaspis suilla species group, male median lobe en-face view and male genitalia lateral view. 33) C. ansa n. sp., a. Port St. Joe, FL; b. Eastpoint, FL; c. Port St. Joe, FL. 34) C. ansa n. sp. (Eastpoint, FL). 35) C. skelleyi n. sp., a. Tiger Creek Preserve, FL; b. Lake Marion Estates, FL; c. Hopkins Prairie, FL. 36) C. skelleyi n. sp. (Lake Marion Estates, FL). 37) C. thomasi n. sp., a. Archbold Biol. Sta., FL; b. Lake Jackson, FL. 38) C. thomasi n. sp. (Archbold Biol. Sta., FL). 39) C. suilla F. a. Kirby St. Forest, TX; b Merritt Reservoir, NE; c. Hope Mills, NC; d. Schenectady, NY. 40) C. costipennis Crotch, a. Sand Hill St. Forest, SC; b. Gainesville (airport), FL; c. Gadsden County, FL; d. Gainesville, FL. En-face views (dry preparations) to scale, lateral views (wet preparations) to scale. AS = apical sclerite." figureDoi="http://doi.org/10.5281/zenodo.5353856" httpUri="https://zenodo.org/record/5353856/files/figure.png" pageId="6" pageNumber="5">39</figureCitation>
, 42, 4748, Map 1)
</paragraph>
</subSubSection>
<subSubSection id="C36BE3FFFFDE7D6DFF42E841FBFFFD4C" pageId="6" pageNumber="5" type="reference_group">
<paragraph id="8BCEB074FFDE7D6DFF42E841FD9CFDAE" blockId="6.[224,1440,559,645]" pageId="6" pageNumber="5">
<treatmentCitationGroup id="AB61975AFFDE7D6DFF42E841FD9CFDAE" pageId="6" pageNumber="5">
<treatmentCitation id="0AD09665FFDE7D6DFF42E841FDB6FD80" author="Fabricius JC" box="[224,566,559,584]" page="417" pageId="6" pageNumber="5" year="1801">
<taxonomicName id="4C71CBF7FFDE7D6DFF42E841FDB6FD80" authority="Fabricius 1801: 417" authorityName="Fabricius" authorityPageNumber="417" authorityYear="1801" box="[224,566,559,584]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFF42E841FEEDFD8F" box="[224,365,559,583]" italics="true" pageId="6" pageNumber="5">Colaspis suilla</emphasis>
<bibRefCitation id="EFE0CD85FFDE7D6DFED0E841FDB6FD80" author="Fabricius JC" box="[370,566,559,584]" pageId="6" pageNumber="5" refId="ref13371" refString="Fabricius JC. 1801. Systema Eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Tom. I. Bibliopolii Academici Novi; Kiliae. xxiv + 506 p." type="book" year="1801">Fabricius 1801: 417</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; type locality: Carolina.
<treatmentCitation id="0AD09665FFDE7D6DFC8EE841FC50FD8F" author="Lefevre E." box="[812,976,559,583]" page="52" pageId="6" pageNumber="5" year="1885">
<bibRefCitation id="EFE0CD85FFDE7D6DFC8EE841FC50FD8F" author="Lefevre E." box="[812,976,559,583]" pageId="6" pageNumber="5" pagination="1 - 172" refId="ref13623" refString="Lefevre E. 1885. Eumolpidarum hucusque cognitarum catalogus, sectionum conspectu systematico, generum sicut et specierum nonnullarum novarum descriptionibus adjunctis. Memoires de la Societe Royale des Sciences de Liege (ser. 2) 11: 1 - 172." type="journal article" year="1885">Lefèvre 1885: 52</bibRefCitation>
</treatmentCitation>
[catalogue].
<treatmentCitation id="0AD09665FFDE7D6DFBF0E841FB57FD80" author="Blake DH" box="[1106,1239,559,584]" page="8" pageId="6" pageNumber="5" year="1974">
<bibRefCitation id="EFE0CD85FFDE7D6DFBF0E841FB57FD80" author="Blake DH" box="[1106,1239,559,584]" pageId="6" pageNumber="5" pagination="1 - 24" refId="ref12952" refString="Blake DH. 1974. The costate species of Colaspis in the United States (Coleoptera: Chrysomelidae). Smithsonian Contributions to Zoology 181: i - iii + 1 - 24." type="journal article" year="1974">Blake 1974: 8</bibRefCitation>
</treatmentCitation>
, fig. 79 [redescription,
<typeStatus id="54CA0ED6FFDE7D6DFE97E820FE11FDAE" box="[309,401,590,614]" pageId="6" pageNumber="5" type="lectotype">lectotype</typeStatus>
designation].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCEB074FFDE7D6DFF42E802FBFFFD4C" blockId="6.[224,1440,559,645]" box="[224,1151,620,645]" pageId="6" pageNumber="5">
<treatmentCitationGroup id="AB61975AFFDE7D6DFF42E802FBFFFD4C" box="[224,1151,620,645]" pageId="6" pageNumber="5">
<treatmentCitation id="0AD09665FFDE7D6DFF42E802FDCCFD4D" author="Blake DH" box="[224,588,620,645]" page="9" pageId="6" pageNumber="5" year="1974">
<taxonomicName id="4C71CBF7FFDE7D6DFF42E802FDCCFD4D" authority="Blake 1974: 9" authorityName="Blake" authorityPageNumber="9" authorityYear="1974" box="[224,588,620,645]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="suilla" subSpecies="borealis">
<emphasis id="B9056C66FFDE7D6DFF42E802FE3FFD4C" box="[224,447,620,644]" italics="true" pageId="6" pageNumber="5">Colaspis suilla borealis</emphasis>
<bibRefCitation id="EFE0CD85FFDE7D6DFE67E802FDCCFD4D" author="Blake DH" box="[453,588,620,645]" pageId="6" pageNumber="5" pagination="1 - 24" refId="ref12952" refString="Blake DH. 1974. The costate species of Colaspis in the United States (Coleoptera: Chrysomelidae). Smithsonian Contributions to Zoology 181: i - iii + 1 - 24." type="journal article" year="1974">Blake 1974: 9</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 11;
<materialsCitation id="3B19BA29FFDE7D6DFD3DE803FBFFFD4C" box="[671,1151,620,645]" country="Canada" location="Toronto" municipality="New Synonymy" pageId="6" pageNumber="5" specimenCount="1">
<typeStatus id="54CA0ED6FFDE7D6DFD3DE803FD4BFD4D" box="[671,715,621,645]" pageId="6" pageNumber="5">type</typeStatus>
locality:
<location id="8EAEE6AFFFDE7D6DFC85E803FCFAFD4C" LSID="urn:lsid:plazi:treatment:03D80162FFDE7D63FF62EB8DFD61FD59:8EAEE6AFFFDE7D6DFC85E803FCFAFD4C" box="[807,890,621,644]" country="Canada" municipality="New Synonymy" name="Toronto" pageId="6" pageNumber="5">Toronto</location>
,
<collectingCountry id="F366F0E4FFDE7D6DFC21E802FC53FD4C" box="[899,979,620,644]" name="Canada" pageId="6" pageNumber="5">Canada</collectingCountry>
.
<emphasis id="B9056C66FFDE7D6DFC7EE803FBFFFD4C" bold="true" box="[988,1151,620,645]" pageId="6" pageNumber="5">
<collectingMunicipality id="6BAA2A0EFFDE7D6DFC7EE803FBFFFD4C" box="[988,1151,620,645]" pageId="6" pageNumber="5">New Synonymy</collectingMunicipality>
</emphasis>
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36BE3FFFFDE7D6DFF62E8FBFAB0FC75" pageId="6" pageNumber="5" type="materials_examined">
<paragraph id="8BCEB074FFDE7D6DFF62E8FBFE37FD18" blockId="6.[191,1443,660,957]" pageId="6" pageNumber="5">
<emphasis id="B9056C66FFDE7D6DFF62E8FBFEE9FD66" bold="true" box="[192,361,660,687]" pageId="6" pageNumber="5">Type material.</emphasis>
Of
<taxonomicName id="4C71CBF7FFDE7D6DFE36E8FBFE6EFD66" authorityName="Fabricius" authorityYear="1801" box="[404,494,660,687]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFE36E8FBFE25FD67" box="[404,421,661,687]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B9056C66FFDE7D6DFE11E8FAFE6EFD66" box="[435,494,660,686]" italics="true" pageId="6" pageNumber="5">suilla</emphasis>
</taxonomicName>
, male
<typeStatus id="54CA0ED6FFDE7D6DFD9AE8FAFD1EFD66" box="[568,670,660,686]" pageId="6" pageNumber="5" type="lectotype">lectotype</typeStatus>
[Fabricius collection, Copenhagen], designated and figured by
<bibRefCitation id="EFE0CD85FFDE7D6DFAC1E8FAFE8BFD18" author="Blake DH" pageId="6" pageNumber="5" pagination="1 - 24" refId="ref12952" refString="Blake DH. 1974. The costate species of Colaspis in the United States (Coleoptera: Chrysomelidae). Smithsonian Contributions to Zoology 181: i - iii + 1 - 24." type="journal article" year="1974">Blake (1974)</bibRefCitation>
, not examined.
</paragraph>
<paragraph id="8BCEB074FFDE7D6DFEA2E8B6FAB0FC75" blockId="6.[191,1443,660,957]" pageId="6" pageNumber="5">
Of
<taxonomicName id="4C71CBF7FFDE7D6DFE81E8B6FE54FD3A" authorityName="Blake" authorityYear="1974" box="[291,468,728,755]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="suilla" subSpecies="borealis">
<emphasis id="B9056C66FFDE7D6DFE81E8B6FEB5FD3A" box="[291,309,728,754]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B9056C66FFDE7D6DFEE2E8B6FE54FD3A" box="[320,468,728,754]" italics="true" pageId="6" pageNumber="5">suilla borealis</emphasis>
</taxonomicName>
,
<typeStatus id="54CA0ED6FFDE7D6DFE43E8B6FDC3FD3A" box="[481,579,728,754]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
[
<collectionCode id="ED6028B1FFDE7D6DFDEDE8B9FD27FD3A" box="[591,679,727,754]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="5" type="Museum">USNM</collectionCode>
], examined, a male labeled “Toronto
<collectingCountry id="F366F0E4FFDE7D6DFB9CE8B6FB14FD3A" box="[1086,1172,728,754]" name="Canada" pageId="6" pageNumber="5">Canada</collectingCountry>
[hand-written, black on white] || Type no. 26913
<collectingCountry id="F366F0E4FFDE7D6DFE6DE894FD81FCDC" box="[463,513,762,788]" name="United States of America" pageId="6" pageNumber="5">U. S.</collectingCountry>
N. M. [printed and hand-written, black on red] ||
<typeStatus id="54CA0ED6FFDE7D6DFB88E894FB12FCDC" box="[1066,1170,762,788]" pageId="6" pageNumber="5" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C71CBF7FFDE7D6DFB3AE894FEA9FCFD" authority="Blake" authorityName="Blake" authorityYear="1974" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="suilla" subSpecies="borealis">
<emphasis id="B9056C66FFDE7D6DFB3AE894FAB4FCDC" box="[1176,1332,762,788]" italics="true" pageId="6" pageNumber="5">Colaspis suilla</emphasis>
ssp.
<emphasis id="B9056C66FFDE7D6DFAC5E894FF65FCFD" italics="true" pageId="6" pageNumber="5">borealis</emphasis>
Blake
</taxonomicName>
[printed and hand-written, red and black on white with red border]. The specimen is point-mounted and was dissected through the abdominal tergites. It is in poor condition, missing various legs and tarsi, with ventral areas mostly obscured by glue. One hind tibia and one antenna are viewable. The shaft of the median lobe is glued to the base of the point with distal portion viewable. The length of the
<typeStatus id="54CA0ED6FFDE7D6DFB82E9EFFB02FC53" box="[1056,1154,897,923]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
was measured at
<quantity id="4C891D91FFDE7D6DFAE6E9EFFA1BFC53" box="[1348,1435,897,923]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="5" unit="mm" value="3.9">3.9 mm</quantity>
, a detail Blake omitted from her original description. Two
<typeStatus id="54CA0ED6FFDE7D6DFCE3E9CDFC2DFC75" box="[833,941,931,957]" pageId="6" pageNumber="5" type="paratype">paratypes</typeStatus>
at the
<collectionCode id="ED6028B1FFDE7D6DFC5AE9CDFBC7FC75" box="[1016,1095,931,957]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="5" type="Museum">USNM</collectionCode>
were also examined.
</paragraph>
</subSubSection>
<subSubSection id="C36BE3FFFFDE7D63FF62E9A3FD61FD59" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5" type="discussion">
<paragraph id="8BCEB074FFDE7D6DFF62E9A3FCB3FB7A" blockId="6.[191,1441,972,1607]" pageId="6" pageNumber="5">
<emphasis id="B9056C66FFDE7D6DFF62E9A3FEAFFC2F" bold="true" box="[192,303,973,999]" pageId="6" pageNumber="5">Remarks.</emphasis>
The uniformly developed and narrow primary elytral costae, dark pronotum, flat surface of male ventrite V, shape of the male genitalia, and shape of the spermatheca will distinguish this species. The median lobe in en-face view is unlike other members of the group, having the tip tapered with a relatively long post-orifical length (
<figureCitation id="134AACF1FFDE7D6DFEB5EE5CFEE8FB84" box="[279,360,1074,1100]" captionStart="Figures 3340" captionStartId="13.[191,278,1454,1480]" captionTargetBox="[192,1440,213,1423]" captionTargetId="figure-94@13.[192,1440,213,1423]" captionTargetPageId="13" captionText="Figures 3340. Colaspis suilla species group, male median lobe en-face view and male genitalia lateral view. 33) C. ansa n. sp., a. Port St. Joe, FL; b. Eastpoint, FL; c. Port St. Joe, FL. 34) C. ansa n. sp. (Eastpoint, FL). 35) C. skelleyi n. sp., a. Tiger Creek Preserve, FL; b. Lake Marion Estates, FL; c. Hopkins Prairie, FL. 36) C. skelleyi n. sp. (Lake Marion Estates, FL). 37) C. thomasi n. sp., a. Archbold Biol. Sta., FL; b. Lake Jackson, FL. 38) C. thomasi n. sp. (Archbold Biol. Sta., FL). 39) C. suilla F. a. Kirby St. Forest, TX; b Merritt Reservoir, NE; c. Hope Mills, NC; d. Schenectady, NY. 40) C. costipennis Crotch, a. Sand Hill St. Forest, SC; b. Gainesville (airport), FL; c. Gadsden County, FL; d. Gainesville, FL. En-face views (dry preparations) to scale, lateral views (wet preparations) to scale. AS = apical sclerite." figureDoi="http://doi.org/10.5281/zenodo.5353856" httpUri="https://zenodo.org/record/5353856/files/figure.png" pageId="6" pageNumber="5">Fig. 39</figureCitation>
). The spermatheca is also unique within the group, and appears to be a stable, reliable diagnostic character (Fig. 4748). The unique spermatheca shape will not only distinguish females of
<taxonomicName id="4C71CBF7FFDE7D6DFB02EE3AFB7AFBA5" authorityName="Fabricius" authorityYear="1801" box="[1184,1274,1107,1134]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFB02EE3AFB32FBA6" box="[1184,1202,1108,1134]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B9056C66FFDE7D6DFB1DEE3DFB7AFBA5" box="[1215,1274,1107,1133]" italics="true" pageId="6" pageNumber="5">suilla</emphasis>
</taxonomicName>
from the other members of the
<taxonomicName id="4C71CBF7FFDE7D6DFEDAEE18FE52FB47" authorityName="Fabricius" authorityYear="1801" box="[376,466,1141,1168]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFEDAEE18FE09FB58" box="[376,393,1142,1168]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B9056C66FFDE7D6DFE35EE1BFE52FB47" box="[407,466,1141,1167]" italics="true" pageId="6" pageNumber="5">suilla</emphasis>
</taxonomicName>
species group, but also from those females of similar-appearing, co-occurring
<taxonomicName id="4C71CBF7FFDE7D6DFAE4EE1BFE8FFB79" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9056C66FFDE7D6DFAE4EE1BFA21FB47" box="[1350,1441,1141,1167]" italics="true" pageId="6" pageNumber="5">Colaspis</emphasis>
species
</taxonomicName>
where the males possess a tubercle on ventrite V.
</paragraph>
<paragraph id="8BCEB074FFDE7D6DFEA2EED7FBE3F98E" blockId="6.[191,1441,972,1607]" pageId="6" pageNumber="5">
Most specimens examined from the southeastern states of
<collectingRegion id="49B57E96FFDE7D6DFC23EED7FC5BFB1B" box="[897,987,1209,1235]" country="United States of America" name="Georgia" pageId="6" pageNumber="5">Georgia</collectingRegion>
and
<collectingRegion id="49B57E96FFDE7D6DFBADEED7FBE0FB1B" box="[1039,1120,1209,1235]" country="United States of America" name="Florida" pageId="6" pageNumber="5">Florida</collectingRegion>
are darker, with the punctate strial areas darkened (as in
<figureCitation id="134AACF1FFDE7D6DFE4DEEB4FDBDFB3D" box="[495,573,1242,1269]" captionStart="Figures 1118" captionStartId="9.[192,279,1741,1767]" captionTargetBox="[192,1440,232,1710]" captionTargetId="figure-10@9.[192,1440,213,1710]" captionTargetPageId="9" captionText="Figures 1118. Forebodies of Colaspis suilla species group, anterodorsal and dorsolateral views. 11) Male C. ansa n. sp. (Port St. Joe, FL). 12) Male C. skelleyi n. sp. (Lake Marion Estates, FL). 13) Male C. thomasi n. sp. (Archbold Biol. Sta., FL). 14) Male C. suilla F. (Kisatchie Natl. Forest, LA). 15) Female C. suilla F. (Torreya St. Pk., FL). 16) Male C. suilla F. (Merritt Reservoir, NE). 17) Male C. costipennis Crotch (Gainesville, FL). 18) Male C. costipennis Crotch (Pensacola, FL). All figures to scale." figureDoi="http://doi.org/10.5281/zenodo.5353850" httpUri="https://zenodo.org/record/5353850/files/figure.png" pageId="6" pageNumber="5">Fig. 15</figureCitation>
). In specimens from other regions, these intercostal areas are rarely darkened, except usually immediately posterior to the scutellum.
<taxonomicName id="4C71CBF7FFDE7D6DFCB8EE92FC37FADE" authorityName="Fabricius" authorityYear="1801" box="[794,951,1276,1302]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFCB8EE92FC37FADE" box="[794,951,1276,1302]" italics="true" pageId="6" pageNumber="5">Colaspis suilla</emphasis>
</taxonomicName>
exhibits considerable variation in size, more so than other
<taxonomicName id="4C71CBF7FFDE7D6DFEFDEF70FD8FFAF0" box="[351,527,1310,1336]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9056C66FFDE7D6DFEFDEF70FE3AFAF0" box="[351,442,1310,1336]" italics="true" pageId="6" pageNumber="5">Colaspis</emphasis>
species
</taxonomicName>
occurring in the
<collectingCountry id="F366F0E4FFDE7D6DFD77EF70FC8DFAF0" box="[725,781,1310,1336]" name="United States of America" pageId="6" pageNumber="5">USA</collectingCountry>
. Blake noted that her specimens from
<collectingRegion id="49B57E96FFDE7D6DFB1DEF70FAA3FAF0" box="[1215,1315,1310,1336]" country="United States of America" name="Alabama" pageId="6" pageNumber="5">Alabama</collectingRegion>
and
<collectingRegion id="49B57E96FFDE7D6DFAFEEF70FEA2FA92" country="United States of America" name="North Carolina" pageId="6" pageNumber="5">North Carolina</collectingRegion>
(both examined) were small, and this is true for some other specimens examined in the present study, with males as small as
<quantity id="4C891D91FFDE7D6DFE1BEF0CFD91FAB4" box="[441,529,1378,1404]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="5" unit="mm" value="3.7">3.7 mm</quantity>
. Blake reported the length of the Fabrician
<typeStatus id="54CA0ED6FFDE7D6DFC4CEF0FFBD4FAB3" box="[1006,1108,1377,1403]" pageId="6" pageNumber="5" type="lectotype">lectotype</typeStatus>
at
<quantity id="4C891D91FFDE7D6DFBD6EF0FFB4BFAB4" box="[1140,1227,1377,1404]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="5" unit="mm" value="3.5">3.5 mm</quantity>
, the smallest specimen she examined. Series of specimens from
<collectingRegion id="49B57E96FFDE7D6DFD65EFEDFCB0FA55" box="[711,816,1411,1437]" country="United States of America" name="Nebraska" pageId="6" pageNumber="5">Nebraska</collectingRegion>
and
<collectingRegion id="49B57E96FFDE7D6DFCC8EFEDFB86FA55" box="[874,1030,1411,1437]" country="United States of America" name="South Dakota" pageId="6" pageNumber="5">South Dakota</collectingRegion>
show considerable variation in size, with some females up to
<quantity id="4C891D91FFDE7D6DFE77EFCBFDAAFA08" box="[469,554,1445,1472]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="6" pageNumber="5" unit="mm" value="5.7">5.7 mm</quantity>
and some males as small as
<quantity id="4C891D91FFDE7D6DFCC0EFCBFC3AFA08" box="[866,954,1445,1472]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="5" unit="mm" value="3.7">3.7 mm</quantity>
. A male from Schenectady,
<collectingRegion id="49B57E96FFDE7D6DFB4EEFC8FADCFA77" box="[1260,1372,1445,1472]" country="United States of America" name="New York" pageId="6" pageNumber="5">New York</collectingRegion>
, at
<quantity id="4C891D91FFDE7D6DFA22EFCBFF70FA29" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="5" unit="mm" value="3.7">3.7 mm</quantity>
, is also among the smallest specimens observed in this study. The size is more stable in series from
<collectingRegion id="49B57E96FFDE7D6DFA8EEFA9FA1BFA29" box="[1324,1435,1479,1505]" country="United States of America" name="Louisiana" pageId="6" pageNumber="5">Louisiana</collectingRegion>
,
<collectingRegion id="49B57E96FFDE7D6DFF1DEF87FEA5F9CB" box="[191,293,1513,1539]" country="United States of America" name="Missouri" pageId="6" pageNumber="5">Missouri</collectingRegion>
, and
<collectingRegion id="49B57E96FFDE7D6DFEC2EF87FE21F9CB" box="[352,417,1513,1539]" country="United States of America" name="Texas" pageId="6" pageNumber="5">Texas</collectingRegion>
. Blake named
<taxonomicName id="4C71CBF7FFDE7D6DFDE6EF87FD78F9CB" authorityName="Blake" authorityYear="1974" box="[580,760,1513,1539]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="suilla" subSpecies="borealis">
<emphasis id="B9056C66FFDE7D6DFDE6EF87FDD5F9CB" box="[580,597,1513,1539]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B9056C66FFDE7D6DFDC1EF87FD78F9CB" box="[611,760,1513,1539]" italics="true" pageId="6" pageNumber="5">suilla borealis</emphasis>
</taxonomicName>
based on specimens from Toronto,
<collectingCountry id="F366F0E4FFDE7D6DFB2BEF87FB61F9CB" box="[1161,1249,1513,1539]" name="Canada" pageId="6" pageNumber="5">Canada</collectingCountry>
, stating that they were somewhat larger and more rotund than those from the Southeast. Given the amount of observed variation in body size, there seems little justification for maintaining a subspecies distinction.
</paragraph>
<paragraph id="8BCEB074FFDE7D6DFF62EC38FE03F8F2" blockId="6.[191,1441,1622,1850]" pageId="6" pageNumber="5">
<emphasis id="B9056C66FFDE7D6DFF62EC38FE91F9B8" bold="true" box="[192,273,1622,1648]" pageId="6" pageNumber="5">Range.</emphasis>
Map 1. This is the only species of the
<taxonomicName id="4C71CBF7FFDE7D6DFD0DEC38FC88F9B8" authorityName="Fabricius" authorityYear="1801" box="[687,776,1622,1648]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFD0DEC38FD41F9B8" box="[687,705,1622,1648]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B9056C66FFDE7D6DFD6FEC38FC88F9B8" box="[717,776,1622,1648]" italics="true" pageId="6" pageNumber="5">suilla</emphasis>
</taxonomicName>
species group whose range extends broadly inland from the Atlantic coastal plain. Its range extends as far north as
<collectingRegion id="49B57E96FFDE7D6DFC8EEC16FC06F95A" box="[812,902,1656,1682]" country="Canada" name="Ontario" pageId="6" pageNumber="5">Ontario</collectingRegion>
,
<collectingRegion id="49B57E96FFDE7D6DFC30EC16FC7EF95A" box="[914,1022,1656,1682]" country="Canada" name="Manitoba" pageId="6" pageNumber="5">Manitoba</collectingRegion>
and Maine, and as far west as North Dakota and Texas.
<bibRefCitation id="EFE0CD85FFDE7D6DFE3FECF7FDACF97C" author="Blake DH" box="[413,556,1689,1716]" pageId="6" pageNumber="5" pagination="1 - 24" refId="ref12952" refString="Blake DH. 1974. The costate species of Colaspis in the United States (Coleoptera: Chrysomelidae). Smithsonian Contributions to Zoology 181: i - iii + 1 - 24." type="journal article" year="1974">Blake (1974)</bibRefCitation>
reported very little locality data for this species, specifically mentioning only Alabama, North Carolina and
<collectingRegion id="49B57E96FFDE7D6DFDBCECD5FDF9F91D" box="[542,633,1723,1749]" country="Canada" name="Ontario" pageId="6" pageNumber="5">Ontario</collectingRegion>
. I have seen specimens from
<collectingCountry id="F366F0E4FFDE7D6DFC6CECD5FBA7F91D" box="[974,1063,1723,1749]" name="Canada" pageId="6" pageNumber="5">Canada</collectingCountry>
: MB, ON; and the
<collectingCountry id="F366F0E4FFDE7D6DFAA1ECD5FA1DF91D" box="[1283,1437,1723,1749]" name="United States of America" pageId="6" pageNumber="5">United States</collectingCountry>
: AL, FL, GA, IA, IN, IL, LA, MA, MD, ME, MI, MN, MO, MS, ND, NE, NH, NJ, NY, SD, TX, WI.
<bibRefCitation id="EFE0CD85FFDE7D6DFAA2ECB3FA20F93F" author="Ciegler JC" box="[1280,1440,1757,1783]" pageId="6" pageNumber="5" refId="ref13189" refString="Ciegler JC. 2007. Leaf and seed beetles of South Carolina (Coleoptera: Chrysomelidae and Orsodacnidae). Biota of South Carolina. Vol. 5. Clemson University; Clemson, SC. 246 p." type="book" year="2007">Ciegler (2007)</bibRefCitation>
reported
<taxonomicName id="4C71CBF7FFDE7D6DFE85EC91FE00F8D1" authorityName="Fabricius" authorityYear="1801" box="[295,384,1791,1817]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="suilla">
<emphasis id="B9056C66FFDE7D6DFE85EC91FEB9F8D1" box="[295,313,1791,1817]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B9056C66FFDE7D6DFEE7EC91FE00F8D1" box="[325,384,1791,1817]" italics="true" pageId="6" pageNumber="5">suilla</emphasis>
</taxonomicName>
from
<collectingRegion id="49B57E96FFDE7D6DFE66EC91FDEFF8D1" box="[452,623,1791,1817]" country="United States of America" name="South Carolina" pageId="6" pageNumber="5">South Carolina</collectingRegion>
,
<bibRefCitation id="EFE0CD85FFDE7D6DFDDBEC91FC87F8D1" author="Clark SM" box="[633,775,1791,1817]" pageId="6" pageNumber="5" pagination="1 - 93" refId="ref13228" refString="Clark SM. 2000. An annotated list of the leaf beetles of West Virginia (Coleoptera: Orsodacnidae, Megalopodidae, Chrysomelidae exclusive of Bruchinae). Occasional Publications of the West Virginia Department of Agriculture no. 1: 1 - 93." type="journal article" year="2000">Clark (2000)</bibRefCitation>
reported it from
<collectingRegion id="49B57E96FFDE7D6DFC65EC91FBA5F8D1" box="[967,1061,1791,1817]" country="United States of America" name="Virginia" pageId="6" pageNumber="5">Virginia</collectingRegion>
, and
<bibRefCitation id="EFE0CD85FFDE7D6DFBC2EC91FABAF8D1" author="Barney RJ &amp; Clark SM &amp; Riley EG" box="[1120,1338,1791,1817]" pageId="6" pageNumber="5" pagination="3 - 18" refId="ref12860" refString="Barney RJ, Clark SM, Riley EG. 2010. Annotated list of leaf beetles (Coleoptera: Chrysomelidae) of Kentucky: Subfamily Eumolpinae. Journal of the Kentucky Academy of Science 71 (1 - 2): 3 - 18." type="journal article" year="2010">Barney et al. (2010)</bibRefCitation>
reported it from
<collectingRegion id="49B57E96FFDE7D6DFEB6ED4EFEFFF8F2" box="[276,383,1824,1850]" country="United States of America" name="Kentucky" pageId="6" pageNumber="5">Kentucky</collectingRegion>
.
</paragraph>
<paragraph id="8BCEB074FFDE7D63FF62ED24FD72FE74" blockId="6.[192,1440,1866,1927]" lastBlockId="8.[191,1442,214,616]" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5">
<emphasis id="B9056C66FFDE7D6DFF62ED24FE05F8AC" bold="true" box="[192,389,1866,1892]" pageId="6" pageNumber="5">Biological notes.</emphasis>
The species appears to be uncommonly encountered throughout most of its range, and this seems to be especially true for the southeastern states. In most collections, it is represented by few specimens or singletons. However, it can be locally abundant at times and appears most abundant in the northwestern portion of its range. Habitats with sandy soils or deposits seem to be preferred. I have collected it in fair numbers at locations with sandy soils in
<collectingRegion id="49B57E96FFD07D63FD86EB74FC85FEFC" box="[548,773,282,308]" isEnumeration="true" pageId="8" pageNumber="7">Louisiana, Missouri</collectingRegion>
, and
<collectingRegion id="49B57E96FFD07D63FCE0EB74FC03FEFC" box="[834,899,282,308]" country="United States of America" name="Texas" pageId="8" pageNumber="7">Texas</collectingRegion>
. The
<collectingRegion id="49B57E96FFD07D63FC1DEB74FB23FEFC" box="[959,1187,282,308]" isEnumeration="true" pageId="8" pageNumber="7">Louisiana and Texas</collectingRegion>
localities are long-leaf pine communities;
<materialsCitation id="3B19BA29FFD07D63FE35EB52FD6DFE74" collectorName="Crowley's Ridge &amp; It &amp; Some" county="Stoddard County" location="Stoddard County" pageId="8" pageNumber="7" specimenCount="1" stateProvince="Missouri">
the southeastern
<collectingRegion id="49B57E96FFD07D63FDF7EB52FD39FE9E" box="[597,697,316,342]" country="United States of America" name="Missouri" pageId="8" pageNumber="7">Missouri</collectingRegion>
locality (
<collectingCounty id="62AFC8F8FFD07D63FCBCEB52FC62FE9E" box="[798,994,316,342]" pageId="8" pageNumber="7">Stoddard County</collectingCounty>
) is at
<collectorName id="2684D5A2FFD07D63FB83EB52FB56FE9E" box="[1057,1238,316,342]" pageId="8" pageNumber="7">Crowleys Ridge</collectorName>
, an area with sand deposits.
<collectorName id="2684D5A2FFD07D63FE89EB31FEBCFEB1" box="[299,316,351,377]" pageId="8" pageNumber="7">It</collectorName>
is frequently encountered and sometimes abundant in the sandhill areas of western
<collectingRegion id="49B57E96FFD07D63FAA6EB30FAEDFEB0" box="[1284,1389,350,376]" country="United States of America" name="Nebraska" pageId="8" pageNumber="7">Nebraska</collectingRegion>
and adjacent
<collectingRegion id="49B57E96FFD07D63FE81EBEEFE3FFE52" box="[291,447,384,410]" country="United States of America" name="South Dakota" pageId="8" pageNumber="7">South Dakota</collectingRegion>
.
<collectorName id="2684D5A2FFD07D63FE6AEBEEFD86FE52" box="[456,518,384,410]" pageId="8" pageNumber="7">Some</collectorName>
specimens from
<collectingRegion id="49B57E96FFD07D63FD60EBEEFCB8FE52" box="[706,824,384,410]" country="United States of America" name="Wisconsin" pageId="8" pageNumber="7">Wisconsin</collectingRegion>
are labeled “Sandy Urban RR Pr.” [sandy urban railroad prairie], “sand prairie”, and “… sandy oak barrens”
</materialsCitation>
.
</paragraph>
<caption id="DF0EE0FCFFDF7D6CFF62EC17FAD5F91F" ID-DOI="http://doi.org/10.5281/zenodo.5353848" ID-Zenodo-Dep="5353848" httpUri="https://zenodo.org/record/5353848/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,279,1657,1683]" targetBox="[192,1431,246,1614]" targetPageId="7">
<paragraph id="8BCEB074FFDF7D6CFF62EC17FAD5F91F" blockId="7.[192,1440,1657,1752]" pageId="7" pageNumber="6">
<emphasis id="B9056C66FFDF7D6CFF62EC17FEDDF95C" bold="true" box="[192,349,1657,1684]" pageId="7" pageNumber="6">Figures 510.</emphasis>
Habitus of
<taxonomicName id="4C71CBF7FFDF7D6CFE40EC14FD15F95C" box="[482,661,1658,1684]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9056C66FFDF7D6CFE40EC14FDBDF95C" box="[482,573,1658,1684]" italics="true" pageId="7" pageNumber="6">Colaspis</emphasis>
species
</taxonomicName>
, dorsal and lateral.
<emphasis id="B9056C66FFDF7D6CFCD0EC17FC0AF95B" bold="true" box="[882,906,1657,1683]" pageId="7" pageNumber="6">5)</emphasis>
Male holotype
<taxonomicName id="4C71CBF7FFDF7D6CFB98EC14FB0CF95C" authority="Riley, 2020" authorityName="Riley" authorityYear="2020" box="[1082,1164,1658,1684]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ansa" status="sp. nov.">
<emphasis id="B9056C66FFDF7D6CFB98EC14FBCBF95C" box="[1082,1099,1658,1684]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B9056C66FFDF7D6CFBFBEC15FB0CF95C" box="[1113,1164,1659,1684]" italics="true" pageId="7" pageNumber="6">ansa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236D11DFFDF7D6CFB31EC14FB4FF95C" box="[1171,1231,1658,1684]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B9056C66FFDF7D6CFB75EC17FB6FF95B" bold="true" box="[1239,1263,1657,1683]" pageId="7" pageNumber="6">6)</emphasis>
Female
<taxonomicName id="4C71CBF7FFDF7D6CFAEFEC14FA20F95C" authority="Riley, 2020" authorityName="Riley" authorityYear="2020" box="[1357,1440,1658,1684]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ansa" status="sp. nov.">
<emphasis id="B9056C66FFDF7D6CFAEFEC14FADFF95C" box="[1357,1375,1658,1684]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B9056C66FFDF7D6CFACFEC15FA20F95C" box="[1389,1440,1659,1684]" italics="true" pageId="7" pageNumber="6">ansa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236D11DFFDF7D6CFF62ECF2FF7BF97E" box="[192,251,1692,1718]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(Eastpoint, FL).
<emphasis id="B9056C66FFDF7D6CFE15ECF5FE4FF97D" bold="true" box="[439,463,1691,1717]" pageId="7" pageNumber="6">7)</emphasis>
Male holotype
<taxonomicName id="4C71CBF7FFDF7D6CFDDEECF2FD69F97D" authority="Riley, 2020" authorityName="Riley" authorityYear="2020" box="[636,745,1691,1718]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="skelleyi" status="sp. nov.">
<emphasis id="B9056C66FFDF7D6CFDDEECF2FD0EF97E" box="[636,654,1692,1718]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B9056C66FFDF7D6CFD39ECF5FD69F97D" box="[667,745,1691,1717]" italics="true" pageId="7" pageNumber="6">skelleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236D11DFFDF7D6CFD4DECF2FCAAF97E" box="[751,810,1692,1718]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B9056C66FFDF7D6CFC92ECF5FCC8F97D" bold="true" box="[816,840,1691,1717]" pageId="7" pageNumber="6">8)</emphasis>
Female
<taxonomicName id="4C71CBF7FFDF7D6CFC04ECF2FB92F97D" authority="Riley, 2020" authorityName="Riley" authorityYear="2020" box="[934,1042,1691,1718]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="skelleyi" status="sp. nov.">
<emphasis id="B9056C66FFDF7D6CFC04ECF2FC38F97E" box="[934,952,1692,1718]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B9056C66FFDF7D6CFC66ECF5FB92F97D" box="[964,1042,1691,1717]" italics="true" pageId="7" pageNumber="6">skelleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236D11DFFDF7D6CFBBBECF2FBD4F97E" box="[1049,1108,1692,1718]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(Lake Marion Estates, FL).
<emphasis id="B9056C66FFDF7D6CFA2AECF5FA20F97D" bold="true" box="[1416,1440,1691,1717]" pageId="7" pageNumber="6">9)</emphasis>
Male holotype
<taxonomicName id="4C71CBF7FFDF7D6CFEC5ECD3FE5EF91F" authority="Riley, 2020" authorityName="Riley" authorityYear="2020" box="[359,478,1725,1752]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="B9056C66FFDF7D6CFEC5ECD3FEF9F91F" box="[359,377,1725,1751]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B9056C66FFDF7D6CFE24ECD3FE5EF91F" box="[390,478,1725,1751]" italics="true" pageId="7" pageNumber="6">thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236D11DFFDF7D6CFE46ECD0FD9EF910" box="[484,542,1726,1752]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B9056C66FFDF7D6CFD86ECD3FDCAF91F" bold="true" box="[548,586,1725,1751]" pageId="7" pageNumber="6">10)</emphasis>
Female
<taxonomicName id="4C71CBF7FFDF7D6CFD05ECD3FC9EF91F" authority="Riley, 2020" authorityName="Riley" authorityYear="2020" box="[679,798,1725,1752]" class="Insecta" family="Chrysomelidae" genus="Colaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="B9056C66FFDF7D6CFD05ECD3FD39F91F" box="[679,697,1725,1751]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B9056C66FFDF7D6CFD64ECD3FC9EF91F" box="[710,798,1725,1751]" italics="true" pageId="7" pageNumber="6">thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236D11DFFDF7D6CFC86ECD0FCDFF910" box="[804,863,1726,1752]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(Archbold Biol. Sta., FL). All figures to scale.
</paragraph>
</caption>
<paragraph id="8BCEB074FFD07D63FEA2EBAAFBFFFDA0" blockId="8.[191,1442,214,616]" pageId="8" pageNumber="7">
Plant data from specimen labels indicate multiple associations with legumes (
<taxonomicName id="4C71CBF7FFD07D63FBF9EBAAFB47FE17" box="[1115,1223,452,479]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
):
<taxonomicName id="4C71CBF7FFD07D63FB7BEBAAFAE7FE17" box="[1241,1383,452,479]" class="Magnoliopsida" family="Fabaceae" genus="Tephrosia" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9056C66FFD07D63FB7BEBAAFAC3FE16" box="[1241,1347,452,478]" italics="true" pageId="8" pageNumber="7">Tephrosia</emphasis>
sp.
</taxonomicName>
(
<specimenCount id="9D777BFDFFD07D63FAD7EBABFEADFDC8" pageId="8" pageNumber="7" type="generic">one specimen</specimenCount>
,
<collectingRegion id="49B57E96FFD07D63FE9AEB88FE29FDC9" box="[312,425,486,513]" country="United States of America" name="Louisiana" pageId="8" pageNumber="7">Louisiana</collectingRegion>
), soybean (
<specimenCount id="9D777BFDFFD07D63FD8AEB89FD47FDC8" box="[552,711,486,513]" pageId="8" pageNumber="7" type="generic">one specimen</specimenCount>
,
<collectingRegion id="49B57E96FFD07D63FD70EB88FCCCFDC9" box="[722,844,486,513]" country="United States of America" name="Wisconsin" pageId="8" pageNumber="7">Wisconsin</collectingRegion>
), flower of
<taxonomicName id="4C71CBF7FFD07D63FC6EEB88FA20FDC9" authority="(Michx.) Greene" authorityName="Greene" baseAuthorityName="Michx." box="[972,1440,486,513]" class="Magnoliopsida" family="Fabaceae" genus="Chamaecrista" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="species" species="fasciculata">
<emphasis id="B9056C66FFD07D63FC6EEB88FB5EFDC8" box="[972,1246,486,512]" italics="true" pageId="8" pageNumber="7">Chamaecrista fasciculata</emphasis>
(Michx.) Greene
</taxonomicName>
(
<specimenCount id="9D777BFDFFD07D63FF6BE867FEF3FDEB" box="[201,371,521,547]" pageId="8" pageNumber="7" type="generic">two specimens</specimenCount>
,
<collectingRegion id="49B57E96FFD07D63FEDDE867FE54FDEC" box="[383,468,521,548]" country="United States of America" name="Florida" pageId="8" pageNumber="7">Florida</collectingRegion>
),
<taxonomicName id="4C71CBF7FFD07D63FE4BE867FD13FDEB" box="[489,659,521,547]" class="Magnoliopsida" family="Fabaceae" genus="Lespedeza" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="species" species="hirta">
<emphasis id="B9056C66FFD07D63FE4BE867FD13FDEB" box="[489,659,521,547]" italics="true" pageId="8" pageNumber="7">Lespedeza hirta</emphasis>
</taxonomicName>
(L.) Hornem. (
<specimenCount id="9D777BFDFFD07D63FCE3E867FC61FDEB" box="[833,993,521,547]" pageId="8" pageNumber="7" type="generic">one specimen</specimenCount>
,
<collectingRegion id="49B57E96FFD07D63FC4EE867FBDEFDEC" box="[1004,1118,521,548]" country="United States of America" name="Maryland" pageId="8" pageNumber="7">Maryland</collectingRegion>
), and “at night on
<taxonomicName id="4C71CBF7FFD07D63FA91E867FF60FD8D" class="Magnoliopsida" family="Fabaceae" genus="Lespedeza" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9056C66FFD07D63FA91E867FA21FDEB" box="[1331,1441,521,547]" italics="true" pageId="8" pageNumber="7">Lespedeza</emphasis>
sp.
</taxonomicName>
, either
<taxonomicName id="4C71CBF7FFD07D63FE93E845FE03FD8D" box="[305,387,555,581]" class="Magnoliopsida" family="Fabaceae" genus="Lespedeza" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="species" species="hirta">
<emphasis id="B9056C66FFD07D63FE93E845FEC0FD8C" box="[305,320,555,580]" italics="true" pageId="8" pageNumber="7">L</emphasis>
.
<emphasis id="B9056C66FFD07D63FEEFE845FE03FD8D" box="[333,387,555,581]" italics="true" pageId="8" pageNumber="7">hirta</emphasis>
</taxonomicName>
(L.) Hornem. or
<taxonomicName id="4C71CBF7FFD07D63FDE2E845FD65FD8D" authority="Nutt." authorityName="Nutt." box="[576,741,555,581]" class="Magnoliopsida" family="Fabaceae" genus="Lespedeza" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="species" species="stuevei">
<emphasis id="B9056C66FFD07D63FDE2E845FDCFFD8C" box="[576,591,555,580]" italics="true" pageId="8" pageNumber="7">L</emphasis>
.
<emphasis id="B9056C66FFD07D63FDFEE845FD27FD8C" box="[604,679,555,580]" italics="true" pageId="8" pageNumber="7">stuevei</emphasis>
Nutt.
</taxonomicName>
” (
<specimenCount id="9D777BFDFFD07D63FD59E845FC29FD8D" box="[763,937,555,581]" pageId="8" pageNumber="7" type="generic">four specimens</specimenCount>
,
<collectingRegion id="49B57E96FFD07D63FC11E845FBE3FD8E" box="[947,1123,555,582]" country="United States of America" name="North Carolina" pageId="8" pageNumber="7">North Carolina</collectingRegion>
). Label data also cite traps in oak savanna (multiple collections,
<collectingRegion id="49B57E96FFD07D63FDE3E823FD39FDA0" box="[577,697,589,616]" country="United States of America" name="Wisconsin" pageId="8" pageNumber="7">Wisconsin</collectingRegion>
) and from oak (
<specimenCount id="9D777BFDFFD07D63FCCEE823FB8BFDAF" box="[876,1035,589,615]" pageId="8" pageNumber="7" type="generic">one specimen</specimenCount>
,
<collectingRegion id="49B57E96FFD07D63FBB7E823FBF4FDA0" box="[1045,1140,589,616]" country="United States of America" name="Georgia" pageId="8" pageNumber="7">Georgia</collectingRegion>
).
</paragraph>
<paragraph id="8BCEB074FFD07D63FF62E819FD61FD59" blockId="8.[192,737,631,658]" box="[192,737,631,658]" pageId="8" pageNumber="7">
<emphasis id="B9056C66FFD07D63FF62E819FE3CFD59" bold="true" box="[192,444,631,657]" pageId="8" pageNumber="7">Specimens examined.</emphasis>
517 total, see Appendix 1.
</paragraph>
</subSubSection>
</treatment>
</document>