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<document id="6FD6D4AF5D1AE3227E8AC3814B55C1C6" ID-CLB-Dataset="4945" ID-DOI="10.5852/ejt.2021.762.1457" ID-GBIF-Dataset="f37caa88-763d-4dbb-8ed3-5d5daccd6104" ID-ISSN="2118-9773" ID-Zenodo-Dep="5211725" ID-ZooBank="253B791F-78FF-42AC-8047-E92449BA4E47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629206175651" checkinUser="felipe" docAuthor="Rosenmejer, Trine, Enghoff, Henrik, Moritz, Leif &amp; Wesener, Thomas" docDate="2021" docId="03D487ADFFBBFFC995D0FE94A7A3FE0B" docLanguage="en" docName="ejt-762_108-132.pdf" docOrigin="European Journal of Taxonomy 762 (1)" docSource="http://zoobank.org/253b791f-78ff-42ac-8047-e92449ba4e47" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zephronia viridisoma Rosenmejer &amp; Enghoff &amp; Moritz &amp; Wesener 2021, sp. nov." docType="treatment" docUuid="272E06C8-2DE5-4722-A7D5-D85002D68424" docUuidSource="ZooBank" docVersion="8" lastPageNumber="127" masterDocId="FFEDFFD5FFB6FFDA9408FF9CA333FFC5" masterDocTitle="Integrative description of new giant pill-millipedes from southern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="132" masterPageNumber="108" pageNumber="121" updateTime="1699086988907" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B918D8A37DE92C4061F38BEBB83A6282">Integrative description of new giant pill-millipedes from southern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="1026C81906D0DB68DFB66BF4F44F3B6A">Rosenmejer, Trine</mods:namePart>
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<mods:affiliation id="ACFEBE29696DD7E6E493A14FE4C271D4">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen OE, Denmark.</mods:affiliation>
<mods:nameIdentifier id="95503F7D32242FD6AFC0397F86F23A51" type="email">trinehansen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="60AF45BD39BC8B4612E25A3CBA929A8A">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="B10E2DB7D0C0F8325234BF87B46DCDFC">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen OE, Denmark.</mods:affiliation>
<mods:nameIdentifier id="7A9121803F8D535729F703E18BA7D21A" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="217F9301D340704A90BACA0CB4204B02">Moritz, Leif</mods:namePart>
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<mods:affiliation id="899DFDCB5EC83290A0C4596018C35978">Zoological Research Museum Alexander Koenig, Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; Institute of Evolutionary Biology and Ecology, University of Bonn, An der Immenburg 1, 53121 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="07F1D771196C74E39653D1154621D2AA" type="email">moritz.leif@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0346DD6D2217CDA85992178041D7AB43">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="A5EE208988E049228C190649B8A171A6">Zoological Research Museum Alexander Koenig, Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<treatment id="03D487ADFFBBFFC995D0FE94A7A3FE0B" ID-DOI="http://doi.org/10.5281/zenodo.5213033" ID-GBIF-Taxon="184570867" ID-Zenodo-Dep="5213033" LSID="urn:lsid:zoobank.org:act:272E06C8-2DE5-4722-A7D5-D85002D68424" httpUri="http://treatment.plazi.org/id/03D487ADFFBBFFC995D0FE94A7A3FE0B" lastPageId="19" lastPageNumber="127" pageId="13" pageNumber="121">
<subSubSection id="C3676530FFBBFFD795D0FE94A76FFEE6" box="[472,1116,264,291]" pageId="13" pageNumber="121" type="nomenclature">
<paragraph id="8BC236BBFFBBFFD795D0FE94A76FFEE6" blockId="13.[371,1217,264,361]" box="[472,1116,264,291]" pageId="13" pageNumber="121">
<heading id="D08A81D7FFBBFFD795D0FE94A76FFEE6" box="[472,1116,264,291]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="121" reason="2">
<taxonomicName id="4C7D4D38FFBBFFD795D0FE94A0C9FEE6" ID-CoL="9BDFN" authority="Rosenmejer &amp; Wesener" authorityName="Rosenmejer &amp; Enghoff &amp; Moritz &amp; Wesener" authorityYear="2021" box="[472,1018,264,291]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="121" phylum="Arthropoda" rank="species" species="viridisoma" status="sp. nov.">
<emphasis id="B909EAA9FFBBFFD795D0FE94A1EEFEE6" bold="true" box="[472,733,264,291]" italics="true" pageId="13" pageNumber="121">Zephronia viridisoma</emphasis>
Rosenmejer &amp; Wesener
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<taxonomicNameLabel id="A23A57D2FFBBFFD79009FE95A76FFEE6" box="[1025,1116,265,291]" pageId="13" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="8BC236BBFFBBFFD796B5FED3A045FEAC" blockId="13.[371,1217,264,361]" box="[701,886,335,361]" pageId="13" pageNumber="121">
<figureCitation id="13462A3EFFBBFFD796B5FED3A02FFEAC" box="[701,796,335,361]" captionStart="Fig" captionStartId="7.[189,232,1882,1908]" captionTargetBox="[202,1381,333,1766]" captionTargetId="figure-16@7.[189,1398,321,1784]" captionTargetPageId="7" captionText="Fig. 2. Map of Thailand, collection localities of Sphaerobelum aesculus Rosenmejer &amp; Wesener sp. nov. and Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., and habitus photographs: S. aesculus Rosenmejer &amp; Wesener sp. nov., ♀ (NHMD 621694); Z. viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="13" pageNumber="121">Figs 2B</figureCitation>
,
<figureCitation id="13462A3EFFBBFFD79720FED3A07AFEAC" box="[808,841,335,361]" captionStart="Fig" captionStartId="12.[189,232,1916,1942]" captionTargetBox="[194,1394,1255,1873]" captionTargetId="figure-181@12.[189,1398,1247,1878]" captionTargetPageId="12" captionText="Fig. 6. Endoterga of midbody tergites, scanning electron micrographs. A. Sphaerobelum aesculus Rosenmejer &amp; Wesener sp. nov., holotype (NHMD 621693). B. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695). Abbreviations: cp = cuticular patterns; iA = inner area." figureDoi="http://doi.org/10.5281/zenodo.5211743" httpUri="https://zenodo.org/record/5211743/files/figure.png" pageId="13" pageNumber="121">6B</figureCitation>
–
<figureCitation id="13462A3EFFBBFFD79752FED3A045FEAC" box="[858,886,335,361]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="13" pageNumber="121">12</figureCitation>
</paragraph>
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<subSubSection id="C3676530FFBBFFD794B5FE0CA2BFFD36" pageId="13" pageNumber="121" type="diagnosis">
<paragraph id="8BC236BBFFBBFFD794B5FE0CA205FE6F" blockId="13.[189,310,400,426]" box="[189,310,400,426]" pageId="13" pageNumber="121">
<heading id="D08A81D7FFBBFFD794B5FE0CA205FE6F" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="13" pageNumber="121" reason="3">
<emphasis id="B909EAA9FFBBFFD794B5FE0CA205FE6F" bold="true" box="[189,310,400,426]" pageId="13" pageNumber="121">Diagnosis</emphasis>
</heading>
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<paragraph id="8BC236BBFFBBFFD794B5FE23A2BFFD36" blockId="13.[187,1399,447,755]" pageId="13" pageNumber="121">
Posterior telopod typical for the genus, not differing from those of other
<taxonomicName id="4C7D4D38FFBBFFD79019FE23A7DDFE1C" box="[1041,1262,447,473]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="121" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B909EAA9FFBBFFD79019FE23A7B9FE1C" box="[1041,1162,447,473]" italics="true" pageId="13" pageNumber="121">Zephronia</emphasis>
species.
</taxonomicName>
Small (
<quantity id="4C859B5EFFBBFFD79141FE23A220FE38" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="13" pageNumber="121" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">25– 28 mm</quantity>
long) green species (
<figureCitation id="13462A3EFFBBFFD79612FE7EA149FE38" box="[538,634,482,509]" captionStart="Fig" captionStartId="7.[189,232,1882,1908]" captionTargetBox="[202,1381,333,1766]" captionTargetId="figure-16@7.[189,1398,321,1784]" captionTargetPageId="7" captionText="Fig. 2. Map of Thailand, collection localities of Sphaerobelum aesculus Rosenmejer &amp; Wesener sp. nov. and Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., and habitus photographs: S. aesculus Rosenmejer &amp; Wesener sp. nov., ♀ (NHMD 621694); Z. viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="13" pageNumber="121">Fig. 2B</figureCitation>
), surface appearing glabrous, dull, with a single medium sized locking carina at the anal shield and a strongly projecting pleurite 1 (
<figureCitation id="13462A3EFFBBFFD797D1FD9AA704FDE5" box="[985,1079,518,544]" captionStart="Fig" captionStartId="16.[189,232,1852,1878]" captionTargetBox="[200,1389,266,1810]" captionTargetId="figure-12@16.[197,1390,265,1813]" captionTargetPageId="16" captionText="Fig. 9. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), legs and pleurite (laterotergite), scanning electron micrographs. A. First pleurite. B. Left midbody leg, posterior view. C. Leg, femur, detail of sclerotized teeth. D. Surface of midbody tergite. Abbreviations: Cx = coxa; Fe = femur; Led = ledge; Po = postfemur; Pr = protuberance; Pre = prefemur; Sp = small triangular spines; Ta = tarsus; Tib = tibia." figureDoi="http://doi.org/10.5281/zenodo.5211751" httpUri="https://zenodo.org/record/5211751/files/figure.png" pageId="13" pageNumber="121">Fig. 9A</figureCitation>
). One of the few
<taxonomicName id="4C7D4D38FFBBFFD790F4FD9AA221FD86" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="121" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B909EAA9FFBBFFD790F4FD9AA646FDE5" box="[1276,1397,518,544]" italics="true" pageId="13" pageNumber="121">Zephronia</emphasis>
species
</taxonomicName>
with just a single apical spine on the legs (
<figureCitation id="13462A3EFFBBFFD79729FDB5A04CFD86" box="[801,895,553,579]" captionStart="Fig" captionStartId="16.[189,232,1852,1878]" captionTargetBox="[200,1389,266,1810]" captionTargetId="figure-12@16.[197,1390,265,1813]" captionTargetPageId="16" captionText="Fig. 9. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), legs and pleurite (laterotergite), scanning electron micrographs. A. First pleurite. B. Left midbody leg, posterior view. C. Leg, femur, detail of sclerotized teeth. D. Surface of midbody tergite. Abbreviations: Cx = coxa; Fe = femur; Led = ledge; Po = postfemur; Pr = protuberance; Pre = prefemur; Sp = small triangular spines; Ta = tarsus; Tib = tibia." figureDoi="http://doi.org/10.5281/zenodo.5211751" httpUri="https://zenodo.org/record/5211751/files/figure.png" pageId="13" pageNumber="121">Fig. 9B</figureCitation>
), differing in this character from all other described Thai
<taxonomicName id="4C7D4D38FFBBFFD7957FFDD0A163FDA3" box="[375,592,588,614]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="121" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B909EAA9FFBBFFD7957FFDD0A2C3FDA3" box="[375,496,588,614]" italics="true" pageId="13" pageNumber="121">Zephronia</emphasis>
species
</taxonomicName>
which have 2–5 apical tarsal spines. Male antennomere 6 swollen (
<figureCitation id="13462A3EFFBBFFD794CEFDF3A213FD4F" box="[198,288,623,650]" captionStart="Fig" captionStartId="14.[189,232,1803,1829]" captionTargetBox="[198,1391,322,1712]" captionTargetId="figure-11@14.[189,1398,310,1722]" captionTargetPageId="14" captionText="Fig. 7. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), head, scanning elelctron micrographs. A. Lateral view. B. Right eye. C. Organ of Tömösváry. D. Left antenna, lateral view. Abbreviations: Ant = antenna; IL = incisura lateralis; O = ommatidia; TO = organ of Tömösváry." figureDoi="http://doi.org/10.5281/zenodo.5211745" httpUri="https://zenodo.org/record/5211745/files/figure.png" pageId="13" pageNumber="121">Fig. 7A</figureCitation>
) but not axe-shaped, with &lt;50 apical cones. Endotergum with three dense rows of long marginal setae (
<figureCitation id="13462A3EFFBBFFD7950EFD0EA251FD68" box="[262,354,658,685]" captionStart="Fig" captionStartId="12.[189,232,1916,1942]" captionTargetBox="[194,1394,1255,1873]" captionTargetId="figure-181@12.[189,1398,1247,1878]" captionTargetPageId="12" captionText="Fig. 6. Endoterga of midbody tergites, scanning electron micrographs. A. Sphaerobelum aesculus Rosenmejer &amp; Wesener sp. nov., holotype (NHMD 621693). B. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695). Abbreviations: cp = cuticular patterns; iA = inner area." figureDoi="http://doi.org/10.5281/zenodo.5211743" httpUri="https://zenodo.org/record/5211743/files/figure.png" pageId="13" pageNumber="121">Fig. 6B</figureCitation>
). Palpi of gnathochilarium with sensory cones arranged in clusters (
<figureCitation id="13462A3EFFBBFFD79076FD0EA7CCFD68" box="[1150,1279,658,685]" captionStart="Fig" captionStartId="15.[189,232,1671,1697]" captionTargetBox="[189,1397,302,1580]" captionTargetId="figure-15@15.[189,1398,302,1581]" captionTargetPageId="15" captionText="Fig. 8. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), mouthparts, scanning electron micrographs. A. Epipharynx, ventral view.B. Gnathochilarium, underside. C. Gnathochilarium frontal view at lateral palps and central pads. D. Detail of sensory cones at lateral palps. E. Right mandible, dorsal view. F. Right mandible, mesal view. Abbreviations: 3iT = 3-combed inner tooth; Co = condylus; cP = central pads; Endo = endochilarium; eT = external tooth; iA = inner area; LL = lamellae linguales; LP = lateral palpi; Me = mentum; Mp = molar plate; pL = pectinate lamellae; St = stipes; T = tooth of epipharynx." figureDoi="http://doi.org/10.5281/zenodo.5211749" httpUri="https://zenodo.org/record/5211749/files/figure.png" pageId="13" pageNumber="121">Fig. 8C–D</figureCitation>
). Anterior telopod podomere 3 with an elevated process at posterior side carrying sclerotized teeth. Podomere 4 short and narrow.
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFBBFFD794B5FC86A7C8FCA6" pageId="13" pageNumber="121" type="etymology">
<paragraph id="8BC236BBFFBBFFD794B5FC86A2AEFCF1" blockId="13.[189,413,794,820]" box="[189,413,794,820]" pageId="13" pageNumber="121">
<heading id="D08A81D7FFBBFFD794B5FC86A2AEFCF1" bold="true" box="[189,413,794,820]" fontSize="11" level="3" pageId="13" pageNumber="121" reason="3">
<emphasis id="B909EAA9FFBBFFD794B5FC86A2AEFCF1" bold="true" box="[189,413,794,820]" pageId="13" pageNumber="121">Derivatio nominis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC236BBFFBBFFD794B5FCD5A7C8FCA6" blockId="13.[189,1275,841,867]" box="[189,1275,841,867]" pageId="13" pageNumber="121">Named after the overall green colour of living individuals of the species, noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="C3676530FFBBFFD794B5FC16A2A8FB2D" pageId="13" pageNumber="121" type="materials_examined">
<paragraph id="8BC236BBFFBBFFD794B5FC16A299FC61" blockId="13.[189,426,906,932]" box="[189,426,906,932]" pageId="13" pageNumber="121">
<heading id="D08A81D7FFBBFFD794B5FC16A299FC61" bold="true" box="[189,426,906,932]" fontSize="11" level="3" pageId="13" pageNumber="121" reason="3">
<emphasis id="B909EAA9FFBBFFD794B5FC16A299FC61" bold="true" box="[189,426,906,932]" pageId="13" pageNumber="121">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BC236BBFFBBFFD794E2FC25A146FBF8" blockId="13.[189,1399,953,1085]" pageId="13" pageNumber="121">
<materialsCitation id="3B153CE6FFBBFFD794E2FC25A146FBF8" ID-GBIF-Occurrence="3335033301" collectedFrom="dense jungle on limestone" collectingDate="2007-08-25" collectionCode="NHMD" collectorName="Chulalongkorn University expedition of millipede workshop" country="Thailand" latitude="9.051667" location="Khao Lark Waterfall" longLatPrecision="21" longitude="99.79" municipality="Sichon District" pageId="13" pageNumber="121" specimenCode="NHMD 621695" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Si Thammarat Province" typeStatus="holotype">
<emphasis id="B909EAA9FFBBFFD794E2FC25A26FFC16" bold="true" box="[234,348,953,979]" pageId="13" pageNumber="121">
<typeStatus id="54C68819FFBBFFD794E2FC25A26FFC16" box="[234,348,953,979]" pageId="13" pageNumber="121" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F36A762BFFBBFFD794B5FC40A266FC33" box="[189,341,988,1014]" name="Thailand" pageId="13" pageNumber="121">THAILAND</collectingCountry>
•
<specimenCount id="9D7BFD32FFBBFFD7957AFC41A2B8FC32" box="[370,395,989,1015]" pageId="13" pageNumber="121" type="male">♂</specimenCount>
;
<collectingRegion id="49B9F859FFBBFFD79592FC40A02EFC33" box="[410,797,988,1014]" country="Thailand" name="Nakhon Si Thammarat" pageId="13" pageNumber="121">Nakhon Si Thammarat Province</collectingRegion>
,
<collectingMunicipality id="6BA6ACC1FFBBFFD79724FC40A0D1FC33" box="[812,994,988,1014]" pageId="13" pageNumber="121">Sichon District</collectingMunicipality>
,
<location id="8EA26060FFBBFFD797F8FC40A7DAFC33" LSID="urn:lsid:plazi:treatment:03D487ADFFBBFFC995D0FE94A7A3FE0B:8EA26060FFBBFFD797F8FC40A7DAFC33" box="[1008,1257,988,1014]" country="Thailand" latitude="9.051667" longLatPrecision="21" longitude="99.79" municipality="Sichon District" name="Khao Lark Waterfall" pageId="13" pageNumber="121" stateProvince="Nakhon Si Thammarat Province">Khao Lark Waterfall</location>
;
<geoCoordinate id="EE49507CFFBBFFD790FFFC46A641FC32" box="[1271,1394,986,1015]" degrees="9" direction="north" minutes="03" orientation="latitude" pageId="13" pageNumber="121" precision="15" seconds="6" value="9.051667">9°03′6″ N</geoCoordinate>
,
<geoCoordinate id="EE49507CFFBBFFD794B5FC62A261FBDF" box="[189,338,1022,1050]" degrees="99" direction="east" minutes="47" orientation="longitude" pageId="13" pageNumber="121" precision="15" seconds="24" value="99.79">99°47′24″ E</geoCoordinate>
;
<date id="FFC3107BFFBBFFD79569FC63A13AFBDC" box="[353,521,1023,1050]" pageId="13" pageNumber="121" value="2007-08-25">
<collectingDate id="EF87E993FFBBFFD79569FC63A13AFBDC" box="[353,521,1023,1050]" pageId="13" pageNumber="121" value="2007-08-25">25 Aug. 2007</collectingDate>
</date>
;
<collectorName id="2688536DFFBBFFD79610FC63A7DEFBDC" box="[536,1261,1023,1049]" pageId="13" pageNumber="121">Chulalongkorn University expedition of millipede workshop</collectorName>
leg.;
<collectedFrom id="38617546FFBBFFD7913CFC63A29BFBF9" pageId="13" pageNumber="121">dense jungle on limestone</collectedFrom>
;
<specimenCode id="DBDB9EC0FFBBFFD795BCFBBFA142FBF8" box="[436,625,1058,1085]" collectionCode="NHMD" pageId="13" pageNumber="121">NHMD 621695</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC236BBFFBBFFD794E2FBF8A2A8FB2D" blockId="13.[189,1399,1124,1256]" pageId="13" pageNumber="121">
<materialsCitation id="3B153CE6FFBBFFD794E2FBF8A743FB64" ID-GBIF-Occurrence="3335033305" collectedFrom="dense jungle on limestone" collectingDate="2007-08-25" collectionCode="NHMD" collectorName="Chulalongkorn University expedition of millipede workshop" country="Thailand" latitude="9.051667" location="THAILAND" longLatPrecision="21" longitude="99.79" municipality="Sichon District" pageId="13" pageNumber="121" specimenCode="NHMD 621696" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Nakhon Si Thammarat" typeStatus="paratype">
<emphasis id="B909EAA9FFBBFFD794E2FBF8A254FBBB" bold="true" box="[234,359,1124,1150]" pageId="13" pageNumber="121">
<typeStatus id="54C68819FFBBFFD794E2FBF8A254FBBB" box="[234,359,1124,1150]" pageId="13" pageNumber="121" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F36A762BFFBBFFD794B5FB1BA266FB64" box="[189,341,1159,1185]" name="Thailand" pageId="13" pageNumber="121">THAILAND</collectingCountry>
•
<specimenCount id="9D7BFD32FFBBFFD79563FB1BA29DFB67" box="[363,430,1159,1186]" pageId="13" pageNumber="121" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D7BFD32FFBBFFD795B2FB1BA2CFFB67" box="[442,508,1159,1186]" pageId="13" pageNumber="121" type="female">4 ♀♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBDB9EC0FFBBFFD797B0FB14A743FB64" box="[952,1136,1159,1186]" collectionCode="NHMD" pageId="13" pageNumber="121">NHMD 621696</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B153CE6FFBBFFD7908FFB1BA1A3FB01" ID-GBIF-Occurrence="3335033303" collectedFrom="dense jungle on limestone" collectingDate="2007-08-25" collectionCode="ZFMK" collectorName="Chulalongkorn University expedition of millipede workshop" country="Thailand" latitude="9.051667" location="Khao Lark Waterfall" longLatPrecision="21" longitude="99.79" municipality="Sichon District" pageId="13" pageNumber="121" specimenCode="MYR8786" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Si Thammarat" typeStatus="paratype">
<specimenCount id="9D7BFD32FFBBFFD7908FFB1BA786FB67" box="[1159,1205,1159,1186]" pageId="13" pageNumber="121" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED6CAE7EFFBBFFD795B1FB37A139FB00" box="[441,522,1195,1221]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="13" pageNumber="121" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBDB9EC0FFBBFFD7961BFB36A1A3FB01" box="[531,656,1194,1220]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="13" pageNumber="121" type="Museum">MYR8786</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B153CE6FFBBFFD796A3FB36A2A4FB2D" ID-GBIF-Occurrence="3335033304" collectedFrom="dense jungle on limestone" collectingDate="2007-08-25" collectionCode="ZFMK" collectorName="Chulalongkorn University expedition of millipede workshop" country="Thailand" latitude="9.051667" location="Khao Lark Waterfall" longLatPrecision="21" longitude="99.79" municipality="Sichon District" pageId="13" pageNumber="121" specimenCode="MYR8787" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Si Thammarat" typeStatus="paratype">
<specimenCount id="9D7BFD32FFBBFFD796A3FB36A1E9FB00" box="[683,730,1194,1221]" pageId="13" pageNumber="121" type="female">1 ♀</specimenCount>
; same collection data as for holotype; CT scan voucher;
<collectionCode id="ED6CAE7EFFBBFFD794B5FB52A23DFB2D" box="[189,270,1230,1256]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="13" pageNumber="121" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBDB9EC0FFBBFFD7951EFB52A2A4FB2D" box="[278,407,1230,1256]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="13" pageNumber="121" type="Museum">MYR8787</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3676530FFBBFFC994B5FA93A7A3FE0B" lastPageId="19" lastPageNumber="127" pageId="13" pageNumber="121" type="description">
<paragraph id="8BC236BBFFBBFFD794B5FA93A10DFAEF" blockId="13.[189,574,1295,1322]" box="[189,574,1295,1322]" pageId="13" pageNumber="121">
<emphasis id="B909EAA9FFBBFFD794B5FA93A27DFAEC" bold="true" box="[189,334,1295,1321]" pageId="13" pageNumber="121">Description</emphasis>
(based on
<typeStatus id="54C68819FFBBFFD795C6FA93A104FAEF" box="[462,567,1295,1322]" pageId="13" pageNumber="121" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BC236BBFFBBFFD794B5FAA2A07BFABE" blockId="13.[189,1398,1341,1403]" pageId="13" pageNumber="121">
SIZE. Length
<quantity id="4C859B5EFFBBFFD79556FAA1A2FEFA9D" box="[350,461,1341,1368]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="13" pageNumber="121" unit="mm" value="25.6">25.6 mm</quantity>
. Width of thoracic shield
<quantity id="4C859B5EFFBBFFD79703FAA2A049FA9D" box="[779,890,1342,1368]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="13" pageNumber="121" unit="mm" value="12.1">12.1 mm</quantity>
, of widest segment (8)
<quantity id="4C859B5EFFBBFFD79094FAA2A7C6FA9D" box="[1180,1269,1342,1368]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="121" unit="mm" value="13.0">13 mm</quantity>
. Height of thoracic shield
<quantity id="4C859B5EFFBBFFD79578FAFDA2FEFABE" box="[368,461,1377,1403]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="13" pageNumber="121" unit="mm" value="7.1">7.1 mm</quantity>
, of highest segment (8)
<quantity id="4C859B5EFFBBFFD796EFFAFDA077FABE" box="[743,836,1377,1403]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="13" pageNumber="121" unit="mm" value="7.7">7.7 mm</quantity>
.
</paragraph>
<paragraph id="8BC236BBFFBBFFD794B5FA3EA21CF9E3" blockId="13.[188,1399,1442,1574]" pageId="13" pageNumber="121">
COLOUR. Head medium brown, faded from green. Antennae medium brown. Legs medium brown, tarsal claws apically dark brown. Tergites light brown, faded green medially, with darker brown posterior margin. Paratergite tips medium brown with darker edges. Anal shield dark green, edges faded to brown (
<figureCitation id="13462A3EFFBBFFD794CDF990A212F9E3" box="[197,289,1548,1574]" captionStart="Fig" captionStartId="7.[189,232,1882,1908]" captionTargetBox="[202,1381,333,1766]" captionTargetId="figure-16@7.[189,1398,321,1784]" captionTargetPageId="7" captionText="Fig. 2. Map of Thailand, collection localities of Sphaerobelum aesculus Rosenmejer &amp; Wesener sp. nov. and Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., and habitus photographs: S. aesculus Rosenmejer &amp; Wesener sp. nov., ♀ (NHMD 621694); Z. viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="13" pageNumber="121">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8BC236BBFFBBFFD794B5F9D2A7F9F94F" blockId="13.[189,1399,1613,1675]" pageId="13" pageNumber="121">
HEAD (
<figureCitation id="13462A3EFFBBFFD79518F9D1A2A1F9A2" box="[272,402,1613,1639]" captionStart="Fig" captionStartId="14.[189,232,1803,1829]" captionTargetBox="[198,1391,322,1712]" captionTargetId="figure-11@14.[189,1398,310,1722]" captionTargetPageId="14" captionText="Fig. 7. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), head, scanning elelctron micrographs. A. Lateral view. B. Right eye. C. Organ of Tömösváry. D. Left antenna, lateral view. Abbreviations: Ant = antenna; IL = incisura lateralis; O = ommatidia; TO = organ of Tömösváry." figureDoi="http://doi.org/10.5281/zenodo.5211745" httpUri="https://zenodo.org/record/5211745/files/figure.png" pageId="13" pageNumber="121">Fig. 7A–C</figureCitation>
). Number of ommatidia 50 (
<figureCitation id="13462A3EFFBBFFD796EDF9D1A070F9AD" box="[741,835,1613,1640]" captionStart="Fig" captionStartId="14.[189,232,1803,1829]" captionTargetBox="[198,1391,322,1712]" captionTargetId="figure-11@14.[189,1398,310,1722]" captionTargetPageId="14" captionText="Fig. 7. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), head, scanning elelctron micrographs. A. Lateral view. B. Right eye. C. Organ of Tömösváry. D. Left antenna, lateral view. Abbreviations: Ant = antenna; IL = incisura lateralis; O = ommatidia; TO = organ of Tömösváry." figureDoi="http://doi.org/10.5281/zenodo.5211745" httpUri="https://zenodo.org/record/5211745/files/figure.png" pageId="13" pageNumber="121">Fig. 7B</figureCitation>
). Organ of Tömösváry placed midway between ocelli and antennal groove (
<figureCitation id="13462A3EFFBBFFD7960DF9ECA151F94E" box="[517,610,1648,1675]" captionStart="Fig" captionStartId="14.[189,232,1803,1829]" captionTargetBox="[198,1391,322,1712]" captionTargetId="figure-11@14.[189,1398,310,1722]" captionTargetPageId="14" captionText="Fig. 7. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), head, scanning elelctron micrographs. A. Lateral view. B. Right eye. C. Organ of Tömösváry. D. Left antenna, lateral view. Abbreviations: Ant = antenna; IL = incisura lateralis; O = ommatidia; TO = organ of Tömösváry." figureDoi="http://doi.org/10.5281/zenodo.5211745" httpUri="https://zenodo.org/record/5211745/files/figure.png" pageId="13" pageNumber="121">Fig. 7A</figureCitation>
), with its typical coral-like inner structure (
<figureCitation id="13462A3EFFBBFFD79057F9ECA78FF94F" box="[1119,1212,1648,1674]" captionStart="Fig" captionStartId="14.[189,232,1803,1829]" captionTargetBox="[198,1391,322,1712]" captionTargetId="figure-11@14.[189,1398,310,1722]" captionTargetPageId="14" captionText="Fig. 7. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), head, scanning elelctron micrographs. A. Lateral view. B. Right eye. C. Organ of Tömösváry. D. Left antenna, lateral view. Abbreviations: Ant = antenna; IL = incisura lateralis; O = ommatidia; TO = organ of Tömösváry." figureDoi="http://doi.org/10.5281/zenodo.5211745" httpUri="https://zenodo.org/record/5211745/files/figure.png" pageId="13" pageNumber="121">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="8BC236BBFFBBFFD794B5F92EA094F92A" blockId="13.[185,1399,1714,1776]" pageId="13" pageNumber="121">
ANTENNAE (
<figureCitation id="13462A3EFFBBFFD7954AF92EA28FF909" box="[322,444,1714,1740]" captionStart="Fig" captionStartId="14.[189,232,1803,1829]" captionTargetBox="[198,1391,322,1712]" captionTargetId="figure-11@14.[189,1398,310,1722]" captionTargetPageId="14" captionText="Fig. 7. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), head, scanning elelctron micrographs. A. Lateral view. B. Right eye. C. Organ of Tömösváry. D. Left antenna, lateral view. Abbreviations: Ant = antenna; IL = incisura lateralis; O = ommatidia; TO = organ of Tömösváry." figureDoi="http://doi.org/10.5281/zenodo.5211745" httpUri="https://zenodo.org/record/5211745/files/figure.png" pageId="13" pageNumber="121">Fig.7A,D</figureCitation>
). Reaching leg pair3,6visible antennomeres.Antennomere lengths:6&gt;1&gt;2=3&gt;4=5. Antennomere 6 apically swollen, number of apical cones 32/44.
</paragraph>
<paragraph id="8BC236BBFFBBFFD794B5F88BA098F8F5" blockId="13.[189,939,1814,1841]" box="[189,939,1814,1841]" pageId="13" pageNumber="121">
EPIPHARYNX (
<figureCitation id="13462A3EFFBBFFD7955EF88AA287F8F4" box="[342,436,1814,1841]" captionStart="Fig" captionStartId="15.[189,232,1671,1697]" captionTargetBox="[189,1397,302,1580]" captionTargetId="figure-15@15.[189,1398,302,1581]" captionTargetPageId="15" captionText="Fig. 8. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), mouthparts, scanning electron micrographs. A. Epipharynx, ventral view.B. Gnathochilarium, underside. C. Gnathochilarium frontal view at lateral palps and central pads. D. Detail of sensory cones at lateral palps. E. Right mandible, dorsal view. F. Right mandible, mesal view. Abbreviations: 3iT = 3-combed inner tooth; Co = condylus; cP = central pads; Endo = endochilarium; eT = external tooth; iA = inner area; LL = lamellae linguales; LP = lateral palpi; Me = mentum; Mp = molar plate; pL = pectinate lamellae; St = stipes; T = tooth of epipharynx." figureDoi="http://doi.org/10.5281/zenodo.5211749" httpUri="https://zenodo.org/record/5211749/files/figure.png" pageId="13" pageNumber="121">Fig. 8A</figureCitation>
). With an extraordinarily large inner tooth.
</paragraph>
<paragraph id="8BC236BBFFBBFFD794B5F8C4A10BF850" blockId="13.[189,1398,1880,1942]" pageId="13" pageNumber="121">
MANDIBLE (
<figureCitation id="13462A3EFFBBFFD7954CF8C4A28CF8B7" box="[324,447,1880,1906]" captionStart="Fig" captionStartId="15.[189,232,1671,1697]" captionTargetBox="[189,1397,302,1580]" captionTargetId="figure-15@15.[189,1398,302,1581]" captionTargetPageId="15" captionText="Fig. 8. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), mouthparts, scanning electron micrographs. A. Epipharynx, ventral view.B. Gnathochilarium, underside. C. Gnathochilarium frontal view at lateral palps and central pads. D. Detail of sensory cones at lateral palps. E. Right mandible, dorsal view. F. Right mandible, mesal view. Abbreviations: 3iT = 3-combed inner tooth; Co = condylus; cP = central pads; Endo = endochilarium; eT = external tooth; iA = inner area; LL = lamellae linguales; LP = lateral palpi; Me = mentum; Mp = molar plate; pL = pectinate lamellae; St = stipes; T = tooth of epipharynx." figureDoi="http://doi.org/10.5281/zenodo.5211749" httpUri="https://zenodo.org/record/5211749/files/figure.png" pageId="13" pageNumber="121">Fig. 8E–F</figureCitation>
). Inner tooth 3-cusped. Pectinate lamellae with 6 or 7 rows of teeth. Condylus at anterior margin with two ridges.
</paragraph>
<caption id="DF026633FFB8FFD494B5F897A27AF84A" ID-DOI="http://doi.org/10.5281/zenodo.5211745" ID-Zenodo-Dep="5211745" httpUri="https://zenodo.org/record/5211745/files/figure.png" pageId="14" pageNumber="122" startId="14.[189,232,1803,1829]" targetBox="[198,1391,322,1712]" targetPageId="14">
<paragraph id="8BC236BBFFB8FFD494B5F897A27AF84A" blockId="14.[188,1398,1803,1935]" pageId="14" pageNumber="122">
<emphasis id="B909EAA9FFB8FFD494B5F897A23EF8E0" bold="true" box="[189,269,1803,1829]" pageId="14" pageNumber="122">Fig. 7.</emphasis>
<taxonomicName id="4C7D4D38FFB8FFD4951DF897A006F8E3" authority="Rosenmejer &amp; Wesener" authorityName="Rosenmejer &amp; Enghoff &amp; Moritz &amp; Wesener" authorityYear="2021" box="[277,821,1803,1830]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="122" phylum="Arthropoda" rank="species" species="viridisoma" status="sp. nov.">
<emphasis id="B909EAA9FFB8FFD4951DF897A127F8E0" box="[277,532,1803,1829]" italics="true" pageId="14" pageNumber="122">Zephronia viridisoma</emphasis>
Rosenmejer &amp; Wesener
</taxonomicName>
<taxonomicNameLabel id="A23A57D2FFB8FFD49736F890A0AFF8E3" box="[830,924,1804,1830]" pageId="14" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (ZFMK MYR8786), head, scanning elelctron micrographs.
<emphasis id="B909EAA9FFB8FFD49630F8B2A17EF88D" bold="true" box="[568,589,1838,1864]" pageId="14" pageNumber="122">A</emphasis>
. Lateral view.
<emphasis id="B909EAA9FFB8FFD496F0F8B3A03FF88C" bold="true" box="[760,780,1839,1865]" pageId="14" pageNumber="122">B</emphasis>
. Right eye.
<emphasis id="B909EAA9FFB8FFD4979FF8B2A09FF88D" bold="true" box="[919,940,1838,1864]" pageId="14" pageNumber="122">C</emphasis>
. Organ of Tömösváry.
<emphasis id="B909EAA9FFB8FFD490B1F8B3A7FDF88C" bold="true" box="[1209,1230,1839,1865]" pageId="14" pageNumber="122">D</emphasis>
. Left antenna, lateral view. Abbreviations: Ant = antenna; IL = incisura lateralis; O = ommatidia; TO = organ of Tömösváry.
</paragraph>
</caption>
<caption id="DF026633FFB9FFD594B5F91BA1EAF8B1" ID-DOI="http://doi.org/10.5281/zenodo.5211749" ID-Zenodo-Dep="5211749" httpUri="https://zenodo.org/record/5211749/files/figure.png" pageId="15" pageNumber="123" startId="15.[189,232,1671,1697]" targetBox="[189,1397,302,1580]" targetPageId="15">
<paragraph id="8BC236BBFFB9FFD594B5F91BA1EAF8B1" blockId="15.[186,1400,1671,1909]" pageId="15" pageNumber="123">
<emphasis id="B909EAA9FFB9FFD594B5F91BA221F964" bold="true" box="[189,274,1671,1697]" pageId="15" pageNumber="123">Fig. 8.</emphasis>
<taxonomicName id="4C7D4D38FFB9FFD59529F91BA06BF967" authority="Rosenmejer &amp; Wesener" authorityName="Rosenmejer &amp; Enghoff &amp; Moritz &amp; Wesener" authorityYear="2021" box="[289,856,1671,1698]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="123" phylum="Arthropoda" rank="species" species="viridisoma" status="sp. nov.">
<emphasis id="B909EAA9FFB9FFD59529F91BA115F964" box="[289,550,1671,1697]" italics="true" pageId="15" pageNumber="123">Zephronia viridisoma</emphasis>
Rosenmejer &amp; Wesener
</taxonomicName>
<taxonomicNameLabel id="A23A57D2FFB9FFD5976FF914A0FFF967" box="[871,972,1672,1698]" pageId="15" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (ZFMK MYR8786), mouthparts, scanning electron micrographs.
<emphasis id="B909EAA9FFB9FFD596B4F936A1E2F901" bold="true" box="[700,721,1706,1732]" pageId="15" pageNumber="123">A</emphasis>
. Epipharynx, ventral view.
<emphasis id="B909EAA9FFB9FFD59000F936A72FF901" bold="true" box="[1032,1052,1706,1732]" pageId="15" pageNumber="123">B</emphasis>
. Gnathochilarium, underside.
<emphasis id="B909EAA9FFB9FFD594B2F951A3FCF922" bold="true" box="[186,207,1741,1767]" pageId="15" pageNumber="123">C</emphasis>
. Gnathochilarium frontal view at lateral palps and central pads.
<emphasis id="B909EAA9FFB9FFD597C6F951A0D1F922" bold="true" box="[974,994,1741,1767]" pageId="15" pageNumber="123">D</emphasis>
. Detail of sensory cones at lateral palps.
<emphasis id="B909EAA9FFB9FFD59502F96DA22DF8CE" bold="true" box="[266,286,1777,1803]" pageId="15" pageNumber="123">E</emphasis>
. Right mandible, dorsal view.
<emphasis id="B909EAA9FFB9FFD59682F96DA1A8F8CE" bold="true" box="[650,667,1777,1803]" pageId="15" pageNumber="123">F</emphasis>
. Right mandible, mesal view. Abbreviations: 3iT = 3-combed inner tooth; Co = condylus; cP = central pads; Endo = endochilarium; eT = external tooth; iA = inner area; LL = lamellae linguales; LP = lateral palpi; Me = mentum; Mp = molar plate; pL = pectinate lamellae; St = stipes; T = tooth of epipharynx.
</paragraph>
</caption>
<caption id="DF026633FFA6FFCA94B5F8A0A115F826" ID-DOI="http://doi.org/10.5281/zenodo.5211751" ID-Zenodo-Dep="5211751" httpUri="https://zenodo.org/record/5211751/files/figure.png" pageId="16" pageNumber="124" startId="16.[189,232,1852,1878]" targetBox="[200,1389,266,1810]" targetPageId="16">
<paragraph id="8BC236BBFFA6FFCA94B5F8A0A115F826" blockId="16.[188,1399,1852,2020]" pageId="16" pageNumber="124">
<emphasis id="B909EAA9FFA6FFCA94B5F8A0A239F893" bold="true" box="[189,266,1852,1878]" pageId="16" pageNumber="124">Fig. 9.</emphasis>
<taxonomicName id="4C7D4D38FFA6FFCA9518F8A0A015F892" authority="Rosenmejer &amp; Wesener" authorityName="Rosenmejer &amp; Enghoff &amp; Moritz &amp; Wesener" authorityYear="2021" box="[272,806,1852,1879]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="124" phylum="Arthropoda" rank="species" species="viridisoma" status="sp. nov.">
<emphasis id="B909EAA9FFA6FFCA9518F8A0A13FF893" box="[272,524,1852,1878]" italics="true" pageId="16" pageNumber="124">Zephronia viridisoma</emphasis>
Rosenmejer &amp; Wesener
</taxonomicName>
<taxonomicNameLabel id="A23A57D2FFA6FFCA9722F8A1A0BBF892" box="[810,904,1853,1879]" pageId="16" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (ZFMK MYR8786), legs and pleurite (laterotergite), scanning electron micrographs.
<emphasis id="B909EAA9FFA6FFCA974EF8C3A068F8BC" bold="true" box="[838,859,1887,1913]" pageId="16" pageNumber="124">A</emphasis>
. First pleurite.
<emphasis id="B909EAA9FFA6FFCA9019F8FCA717F8BF" bold="true" box="[1041,1060,1888,1914]" pageId="16" pageNumber="124">B</emphasis>
. Left midbody leg, posterior view.
<emphasis id="B909EAA9FFA6FFCA950EF81EA228F859" bold="true" box="[262,283,1922,1948]" pageId="16" pageNumber="124">C</emphasis>
. Leg, femur, detail of sclerotized teeth.
<emphasis id="B909EAA9FFA6FFCA970DF81FA029F858" bold="true" box="[773,794,1923,1949]" pageId="16" pageNumber="124">D</emphasis>
. Surface of midbody tergite. Abbreviations: Cx = coxa; Fe = femur; Led = ledge; Po = postfemur; Pr = protuberance; Pre = prefemur; Sp = small triangular spines; Ta = tarsus; Tib = tibia.
</paragraph>
</caption>
<paragraph id="8BC236BBFFA7FFCB94B5FE95A2A7FEAC" blockId="17.[188,1399,265,361]" pageId="17" pageNumber="125">
GNATHOCHILARIUM (
<figureCitation id="13462A3EFFA7FFCB9597FE95A12CFEE6" box="[415,543,265,291]" captionStart="Fig" captionStartId="15.[189,232,1671,1697]" captionTargetBox="[189,1397,302,1580]" captionTargetId="figure-15@15.[189,1398,302,1581]" captionTargetPageId="15" captionText="Fig. 8. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), mouthparts, scanning electron micrographs. A. Epipharynx, ventral view.B. Gnathochilarium, underside. C. Gnathochilarium frontal view at lateral palps and central pads. D. Detail of sensory cones at lateral palps. E. Right mandible, dorsal view. F. Right mandible, mesal view. Abbreviations: 3iT = 3-combed inner tooth; Co = condylus; cP = central pads; Endo = endochilarium; eT = external tooth; iA = inner area; LL = lamellae linguales; LP = lateral palpi; Me = mentum; Mp = molar plate; pL = pectinate lamellae; St = stipes; T = tooth of epipharynx." figureDoi="http://doi.org/10.5281/zenodo.5211749" httpUri="https://zenodo.org/record/5211749/files/figure.png" pageId="17" pageNumber="125">Fig. 8B–D</figureCitation>
). Lamellae linguales with numerous long setae, medially glabrous. Stipes and mentum with numerous long setae in a regular patter. Sensory cones on gnathochilarium palps in clusters (
<figureCitation id="13462A3EFFA7FFCB9520FED3A2B4FEAC" box="[296,391,335,361]" captionStart="Fig" captionStartId="15.[189,232,1671,1697]" captionTargetBox="[189,1397,302,1580]" captionTargetId="figure-15@15.[189,1398,302,1581]" captionTargetPageId="15" captionText="Fig. 8. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), mouthparts, scanning electron micrographs. A. Epipharynx, ventral view.B. Gnathochilarium, underside. C. Gnathochilarium frontal view at lateral palps and central pads. D. Detail of sensory cones at lateral palps. E. Right mandible, dorsal view. F. Right mandible, mesal view. Abbreviations: 3iT = 3-combed inner tooth; Co = condylus; cP = central pads; Endo = endochilarium; eT = external tooth; iA = inner area; LL = lamellae linguales; LP = lateral palpi; Me = mentum; Mp = molar plate; pL = pectinate lamellae; St = stipes; T = tooth of epipharynx." figureDoi="http://doi.org/10.5281/zenodo.5211749" httpUri="https://zenodo.org/record/5211749/files/figure.png" pageId="17" pageNumber="125">Fig. 8D</figureCitation>
).
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FE0DA0AEFE6E" blockId="17.[189,925,401,427]" box="[189,925,401,427]" pageId="17" pageNumber="125">COLLUM. With short setation, evenly distributed across surface.</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FE4FA211FDD5" blockId="17.[189,1398,466,528]" pageId="17" pageNumber="125">THORACIC SHIELD. Thoracic shield grooves wide and deep, with 9 sclerotized ledges along inner ridge of grooves.</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FDABA654FDB1" blockId="17.[189,1399,567,629]" pageId="17" pageNumber="125">
TERGITES (
<figureCitation id="13462A3EFFA7FFCB9531FDABA2A9FD94" box="[313,410,567,593]" captionStart="Fig" captionStartId="16.[189,232,1852,1878]" captionTargetBox="[200,1389,266,1810]" captionTargetId="figure-12@16.[197,1390,265,1813]" captionTargetPageId="16" captionText="Fig. 9. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), legs and pleurite (laterotergite), scanning electron micrographs. A. First pleurite. B. Left midbody leg, posterior view. C. Leg, femur, detail of sclerotized teeth. D. Surface of midbody tergite. Abbreviations: Cx = coxa; Fe = femur; Led = ledge; Po = postfemur; Pr = protuberance; Pre = prefemur; Sp = small triangular spines; Ta = tarsus; Tib = tibia." figureDoi="http://doi.org/10.5281/zenodo.5211751" httpUri="https://zenodo.org/record/5211751/files/figure.png" pageId="17" pageNumber="125">Fig. 9D</figureCitation>
). Tergites glabrous, with dull orange skin like surface. At high magnification tiny setae and knobs become visible. Paratergite tips on posterior half projecting backwards (
<figureCitation id="13462A3EFFA7FFCB90C2FDC6A668FDB1" box="[1226,1371,602,628]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="17" pageNumber="125">Fig. 12A–B</figureCitation>
).
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FD00A25CFCE5" blockId="17.[189,1399,667,800]" pageId="17" pageNumber="125">
ENDOTERGUM (
<figureCitation id="13462A3EFFA7FFCB9562FD07A2F8FD73" box="[362,459,667,694]" captionStart="Fig" captionStartId="12.[189,232,1916,1942]" captionTargetBox="[194,1394,1255,1873]" captionTargetId="figure-181@12.[189,1398,1247,1878]" captionTargetPageId="12" captionText="Fig. 6. Endoterga of midbody tergites, scanning electron micrographs. A. Sphaerobelum aesculus Rosenmejer &amp; Wesener sp. nov., holotype (NHMD 621693). B. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695). Abbreviations: cp = cuticular patterns; iA = inner area." figureDoi="http://doi.org/10.5281/zenodo.5211743" httpUri="https://zenodo.org/record/5211743/files/figure.png" pageId="17" pageNumber="125">Fig. 6B</figureCitation>
). With a regular flat margin. Outer zone with three dense rows of irregular marginal setae, some of them extending beyond posterior margin. A single row of rounded cuticular impressions present next to marginal ridge. Intersegmental membrane smooth, without cones and with very few setae.
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FCDBA78BFC41" blockId="17.[188,1398,838,900]" pageId="17" pageNumber="125">
STIGMATIC PLATE (
<figureCitation id="13462A3EFFA7FFCB958EFCDAA2C1FCA4" box="[390,498,838,865]" captionStart="Fig" captionStartId="18.[189,232,1818,1844]" captionTargetBox="[270,1319,280,1770]" captionTargetId="figure-11@18.[253,1335,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695), drawings. A. Coxa of first left leg with stigmatic plate, posterior view. B. Coxa of second left leg with gonopore, posterior view. C. Left anterior telopod, lateral view on podomeres 2–4. D. Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Right posterior telopod, posterior view. G. Right posteiror telopod, anterior view. Abbreviations: cr-t = crenulated teeth; Cx = coxa; Go = gonopore; St = stigmatic plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211753" httpUri="https://zenodo.org/record/5211753/files/figure.png" pageId="17" pageNumber="125">Fig. 10A</figureCitation>
). First plate with a rounded sub-triangular apex. More posterior plates similar to those of other representatives of the family, half-covered by the pleurites (
<figureCitation id="13462A3EFFA7FFCB9037FCF6A798FC41" box="[1087,1195,874,900]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="17" pageNumber="125">Fig. 12C</figureCitation>
).
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FC30A61DFC2D" blockId="17.[189,1398,939,1001]" pageId="17" pageNumber="125">
PLEURITES (
<figureCitation id="13462A3EFFA7FFCB9549FC37A293FC00" box="[321,416,939,965]" captionStart="Fig" captionStartId="16.[189,232,1852,1878]" captionTargetBox="[200,1389,266,1810]" captionTargetId="figure-12@16.[197,1390,265,1813]" captionTargetPageId="16" captionText="Fig. 9. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), legs and pleurite (laterotergite), scanning electron micrographs. A. First pleurite. B. Left midbody leg, posterior view. C. Leg, femur, detail of sclerotized teeth. D. Surface of midbody tergite. Abbreviations: Cx = coxa; Fe = femur; Led = ledge; Po = postfemur; Pr = protuberance; Pre = prefemur; Sp = small triangular spines; Ta = tarsus; Tib = tibia." figureDoi="http://doi.org/10.5281/zenodo.5211751" httpUri="https://zenodo.org/record/5211751/files/figure.png" pageId="17" pageNumber="125">Figs 9A</figureCitation>
,
<figureCitation id="13462A3EFFA7FFCB95A7FC37A2D1FC03" box="[431,482,939,966]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="17" pageNumber="125">12D</figureCitation>
). Pleurite 1 projecting strongly, with sharp apex. Pleurite 2 projecting slightly less than 1 and with more rounded apex (
<figureCitation id="13462A3EFFA7FFCB96A8FC52A03DFC2D" box="[672,782,974,1000]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="17" pageNumber="125">Fig. 12D</figureCitation>
). Projection absent from pleurite 3 (
<figureCitation id="13462A3EFFA7FFCB90BCFC52A611FC2D" box="[1204,1314,974,1000]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="17" pageNumber="125">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FB8CA0C0FB56" blockId="17.[188,1398,1040,1172]" pageId="17" pageNumber="125">
LEGS. Ventral spines on leg 1 1/1, on leg 2 3/2, on leg 3 4/4. Apical spine on leg 3 absent. A single apical spine and 5 or 6 ventral spines on midbody legs (
<figureCitation id="13462A3EFFA7FFCB96F9FBAFA07FFB88" box="[753,844,1075,1101]" captionStart="Fig" captionStartId="16.[189,232,1852,1878]" captionTargetBox="[200,1389,266,1810]" captionTargetId="figure-12@16.[197,1390,265,1813]" captionTargetPageId="16" captionText="Fig. 9. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), legs and pleurite (laterotergite), scanning electron micrographs. A. First pleurite. B. Left midbody leg, posterior view. C. Leg, femur, detail of sclerotized teeth. D. Surface of midbody tergite. Abbreviations: Cx = coxa; Fe = femur; Led = ledge; Po = postfemur; Pr = protuberance; Pre = prefemur; Sp = small triangular spines; Ta = tarsus; Tib = tibia." figureDoi="http://doi.org/10.5281/zenodo.5211751" httpUri="https://zenodo.org/record/5211751/files/figure.png" pageId="17" pageNumber="125">Fig. 9B</figureCitation>
). Femur regularly shaped, 2.2 times longer than wide, sclerotized ledge of medium length, inner margin apically with 5 or 6 small rounded triangular spines on ventral side (
<figureCitation id="13462A3EFFA7FFCB95C4FBE5A17FFB51" box="[460,588,1145,1172]" captionStart="Fig" captionStartId="16.[189,232,1852,1878]" captionTargetBox="[200,1389,266,1810]" captionTargetId="figure-12@16.[197,1390,265,1813]" captionTargetPageId="16" captionText="Fig. 9. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♂ (ZFMK MYR8786), legs and pleurite (laterotergite), scanning electron micrographs. A. First pleurite. B. Left midbody leg, posterior view. C. Leg, femur, detail of sclerotized teeth. D. Surface of midbody tergite. Abbreviations: Cx = coxa; Fe = femur; Led = ledge; Po = postfemur; Pr = protuberance; Pre = prefemur; Sp = small triangular spines; Ta = tarsus; Tib = tibia." figureDoi="http://doi.org/10.5281/zenodo.5211751" httpUri="https://zenodo.org/record/5211751/files/figure.png" pageId="17" pageNumber="125">Fig. 9B–C</figureCitation>
). Tarsus 4.2 times longer than wide.
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FB27A090FB3D" blockId="17.[189,1398,1211,1273]" pageId="17" pageNumber="125">
ANAL SHIELD. Well-rounded and glabrous. Ventral side with dark coloured. A medium-sized locking carina placed slightly closer to pleurite than margin (
<figureCitation id="13462A3EFFA7FFCB9720FB42A0A5FB3D" box="[808,918,1246,1272]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="17" pageNumber="125">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FABCA250FA98" blockId="17.[189,1398,1311,1373]" pageId="17" pageNumber="125">
MALE GONOPORE (
<figureCitation id="13462A3EFFA7FFCB9581FA83A2C2FAFF" box="[393,497,1311,1338]" captionStart="Fig" captionStartId="18.[189,232,1818,1844]" captionTargetBox="[270,1319,280,1770]" captionTargetId="figure-11@18.[253,1335,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695), drawings. A. Coxa of first left leg with stigmatic plate, posterior view. B. Coxa of second left leg with gonopore, posterior view. C. Left anterior telopod, lateral view on podomeres 2–4. D. Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Right posterior telopod, posterior view. G. Right posteiror telopod, anterior view. Abbreviations: cr-t = crenulated teeth; Cx = coxa; Go = gonopore; St = stigmatic plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211753" httpUri="https://zenodo.org/record/5211753/files/figure.png" pageId="17" pageNumber="125">Fig. 10B</figureCitation>
). Inconspicuous, consisting of large membranous opening located directly at mesal margin.
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5FA18A027F98B" blockId="17.[188,1399,1412,1615]" pageId="17" pageNumber="125">
ANTERIOR TELOPODS (
<figureCitation id="13462A3EFFA7FFCB95A7FA18A108FA5B" box="[431,571,1412,1438]" captionStart="Fig" captionStartId="18.[189,232,1818,1844]" captionTargetBox="[270,1319,280,1770]" captionTargetId="figure-11@18.[253,1335,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695), drawings. A. Coxa of first left leg with stigmatic plate, posterior view. B. Coxa of second left leg with gonopore, posterior view. C. Left anterior telopod, lateral view on podomeres 2–4. D. Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Right posterior telopod, posterior view. G. Right posteiror telopod, anterior view. Abbreviations: cr-t = crenulated teeth; Cx = coxa; Go = gonopore; St = stigmatic plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211753" httpUri="https://zenodo.org/record/5211753/files/figure.png" pageId="17" pageNumber="125">Fig. 10C–E</figureCitation>
). With long setae medially on the first 2 podomeres, covering posterior part on podomere 2. Podomere 1 rectangular, as long as wide. Podomere
<quantity id="4C859B5EFFA7FFCB902AFA3BA763FA04" box="[1058,1104,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="17" pageNumber="125" unit="in" value="2.0">2 in</quantity>
anterior view as wide as but slightly narrower than podomere 1. Immovable finger barely visible in anterior view (
<figureCitation id="13462A3EFFA7FFCB90F2FA56A65BFA20" box="[1274,1384,1482,1509]" captionStart="Fig" captionStartId="18.[189,232,1818,1844]" captionTargetBox="[270,1319,280,1770]" captionTargetId="figure-11@18.[253,1335,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695), drawings. A. Coxa of first left leg with stigmatic plate, posterior view. B. Coxa of second left leg with gonopore, posterior view. C. Left anterior telopod, lateral view on podomeres 2–4. D. Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Right posterior telopod, posterior view. G. Right posteiror telopod, anterior view. Abbreviations: cr-t = crenulated teeth; Cx = coxa; Go = gonopore; St = stigmatic plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211753" httpUri="https://zenodo.org/record/5211753/files/figure.png" pageId="17" pageNumber="125">Fig. 10D</figureCitation>
), short, not protruding up to podomere 4 (
<figureCitation id="13462A3EFFA7FFCB969CFA72A033F9CD" box="[660,768,1518,1544]" captionStart="Fig" captionStartId="18.[189,232,1818,1844]" captionTargetBox="[270,1319,280,1770]" captionTargetId="figure-11@18.[253,1335,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695), drawings. A. Coxa of first left leg with stigmatic plate, posterior view. B. Coxa of second left leg with gonopore, posterior view. C. Left anterior telopod, lateral view on podomeres 2–4. D. Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Right posterior telopod, posterior view. G. Right posteiror telopod, anterior view. Abbreviations: cr-t = crenulated teeth; Cx = coxa; Go = gonopore; St = stigmatic plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211753" httpUri="https://zenodo.org/record/5211753/files/figure.png" pageId="17" pageNumber="125">Fig. 10C</figureCitation>
). Podomere 3 long and wide, with rounded projection on posterior side, bearing 3 crenulated teeth. Podomere 4 very narrow and short, with 3 large spines near apex, and a smaller spine near edge to podomere 3.
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5F9EAA7D6F8D9" blockId="17.[189,1400,1653,1821]" pageId="17" pageNumber="125">
POSTERIOR TELOPODS (
<figureCitation id="13462A3EFFA7FFCB95B6F9E9A17CF955" box="[446,591,1653,1680]" captionStart="Fig" captionStartId="18.[189,232,1818,1844]" captionTargetBox="[270,1319,280,1770]" captionTargetId="figure-11@18.[253,1335,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., holotype, ♂ (NHMD 621695), drawings. A. Coxa of first left leg with stigmatic plate, posterior view. B. Coxa of second left leg with gonopore, posterior view. C. Left anterior telopod, lateral view on podomeres 2–4. D. Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Right posterior telopod, posterior view. G. Right posteiror telopod, anterior view. Abbreviations: cr-t = crenulated teeth; Cx = coxa; Go = gonopore; St = stigmatic plate. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211753" httpUri="https://zenodo.org/record/5211753/files/figure.png" pageId="17" pageNumber="125">Fig. 10F–G</figureCitation>
). Podomere 2 setose, immovable finger straight, apically tapering, with rows of small circular sclerotized spots. Membranous lobe between podomere 3 and 4 bearing 2 elongated processes, fused at base. Podomere 3 with 8 or 9 small crenulated teeth on posterior side. Membranous lobe on podomere 3 with single spine. Podomere 4 slightly curved towards immovable finger, membranous lobe with 2 spines. Setation on podomere 2 primarily on anterior side.
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94E2F8DFA15EF89B" blockId="17.[189,1400,1859,1957]" box="[234,621,1859,1886]" pageId="17" pageNumber="125">
<emphasis id="B909EAA9FFA7FFCB94E2F8DFA15EF89B" bold="true" box="[234,621,1859,1886]" pageId="17" pageNumber="125">
Description of female
<typeStatus id="54C68819FFA7FFCB95F5F8D8A15EF89B" box="[509,621,1860,1886]" pageId="17" pageNumber="125" type="paratype">paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC236BBFFA7FFCB94B5F8FBA64BF861" blockId="17.[189,1400,1859,1957]" pageId="17" pageNumber="125">
Length
<quantity id="4C859B5EFFA7FFCB9511F8FBA2B6F847" box="[281,389,1895,1922]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="17" pageNumber="125" unit="mm" value="25.6">25.6 mm</quantity>
. Width of thoracic shield
<quantity id="4C859B5EFFA7FFCB96B0F8FBA016F847" box="[696,805,1895,1922]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="125" unit="mm" value="12.3">12.3 mm</quantity>
, of widest tergite (8)
<quantity id="4C859B5EFFA7FFCB902DF8FBA7A1F847" box="[1061,1170,1895,1922]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="17" pageNumber="125" unit="mm" value="13.7">13.7 mm</quantity>
. Height of thoracic shield:
<quantity id="4C859B5EFFA7FFCB951FF816A244F860" box="[279,375,1930,1957]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="17" pageNumber="125" unit="mm" value="8.4">8.4 mm</quantity>
, of highest tergite (8)
<quantity id="4C859B5EFFA7FFCB9680F816A1DBF860" box="[648,744,1930,1957]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="17" pageNumber="125" unit="mm" value="9.3">9.3 mm</quantity>
. Antennae reaching leg pair 3. Apical cones left/right:
</paragraph>
<caption id="DF026633FFA4FFC894B5F886A023F820" ID-DOI="http://doi.org/10.5281/zenodo.5211753" ID-Zenodo-Dep="5211753" httpUri="https://zenodo.org/record/5211753/files/figure.png" pageId="18" pageNumber="126" startId="18.[189,232,1818,1844]" targetBox="[270,1319,280,1770]" targetPageId="18">
<paragraph id="8BC236BBFFA4FFC894B5F886A023F820" blockId="18.[189,1399,1818,2021]" pageId="18" pageNumber="126">
<emphasis id="B909EAA9FFA4FFC894B5F886A210F8F1" bold="true" box="[189,291,1818,1844]" pageId="18" pageNumber="126">Fig. 10.</emphasis>
<taxonomicName id="4C7D4D38FFA4FFC8953AF886A05DF8F0" authority="Rosenmejer &amp; Wesener" authorityName="Rosenmejer &amp; Enghoff &amp; Moritz &amp; Wesener" authorityYear="2021" box="[306,878,1818,1845]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="126" phylum="Arthropoda" rank="species" species="viridisoma" status="sp. nov.">
<emphasis id="B909EAA9FFA4FFC8953AF886A10BF8F1" box="[306,568,1818,1844]" italics="true" pageId="18" pageNumber="126">Zephronia viridisoma</emphasis>
Rosenmejer &amp; Wesener
</taxonomicName>
<taxonomicNameLabel id="A23A57D2FFA4FFC89776F887A0D0F8F0" box="[894,995,1819,1845]" pageId="18" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621695), drawings.
<emphasis id="B909EAA9FFA4FFC89530F8A1A27EF892" bold="true" box="[312,333,1853,1879]" pageId="18" pageNumber="126">A</emphasis>
. Coxa of first left leg with stigmatic plate, posterior view.
<emphasis id="B909EAA9FFA4FFC8900CF8A2A72BF89D" bold="true" box="[1028,1048,1854,1880]" pageId="18" pageNumber="126">B</emphasis>
. Coxa of second left leg with gonopore, posterior view.
<emphasis id="B909EAA9FFA4FFC895F2F8FCA13CF8BF" bold="true" box="[506,527,1888,1914]" pageId="18" pageNumber="126">C</emphasis>
. Left anterior telopod, lateral view on podomeres 2–4.
<emphasis id="B909EAA9FFA4FFC890B0F8FDA7FEF8BE" bold="true" box="[1208,1229,1889,1915]" pageId="18" pageNumber="126">D</emphasis>
. Left anterior telopod, anterior view.
<emphasis id="B909EAA9FFA4FFC895DEF818A2D9F85B" bold="true" box="[470,490,1924,1950]" pageId="18" pageNumber="126">E</emphasis>
. Left anterior telopod, posterior view.
<emphasis id="B909EAA9FFA4FFC897CAF818A0E7F85B" bold="true" box="[962,980,1924,1950]" pageId="18" pageNumber="126">F</emphasis>
. Right posterior telopod, posterior view.
<emphasis id="B909EAA9FFA4FFC8950BF83BA229F804" bold="true" box="[259,282,1959,1985]" pageId="18" pageNumber="126">G</emphasis>
. Right posteiror telopod, anterior view. Abbreviations: cr-t = crenulated teeth; Cx = coxa; Go = gonopore; St = stigmatic plate. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BC236BBFFA5FFC994B5FE95A0B3FE83" blockId="19.[189,1398,265,327]" pageId="19" pageNumber="127">
25/25. Subanal plate well rounded. Vulvae (
<figureCitation id="13462A3EFFA5FFC996DAFE95A018FEE6" box="[722,811,265,291]" captionStart="Fig" captionStartId="19.[189,232,1916,1942]" captionTargetBox="[360,1211,880,1874]" captionTargetId="figure-186@19.[359,1229,879,1875]" captionTargetPageId="19" captionText="Fig. 11. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (NHM D621696), drawings. A. Second right coxa and prefemur with vulva, posterior view. Abbreviations: Cx = coxa; Op = operculum; Pre = prefemur. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211755" httpUri="https://zenodo.org/record/5211755/files/figure.png" pageId="19" pageNumber="127">Fig. 11</figureCitation>
): with a large pointed operculum (Op). External lateral plate (EP) and inner mesal plate (IP) fused at bottom.
</paragraph>
<paragraph id="8BC236BBFFA5FFC994B5FEF2A7A3FE0B" blockId="19.[189,1399,366,463]" pageId="19" pageNumber="127">
Another female
<typeStatus id="54C68819FFA5FFC9958BFEF2A2DBFE4D" box="[387,488,366,392]" pageId="19" pageNumber="127" type="paratype">paratype</typeStatus>
was used for microcomputed tomography. The resulting volume rendering is shown in
<figureCitation id="13462A3EFFA5FFC99546FE0DA296FE6E" box="[334,421,401,427]" captionStart="Fig" captionStartId="20.[189,232,1739,1765]" captionTargetBox="[189,1393,580,1695]" captionTargetId="figure-116@20.[189,1398,576,1698]" captionTargetPageId="20" captionText="Fig. 12. Zephronia viridisoma Rosenmejer &amp; Wesener sp. nov., paratype, ♀ (ZFMK MYR8787), volume rendering based on micro-computed tomography. Stored as cybertype under https://doi.org/10.5281/zenodo.4548243. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; ts = thoracic shield; Col = collum; Cx = Coxa; Fe = femur; Gn = gnathochilarium; Go = gonopore; IL = incisura lateralis; Lc = locking carina; O = ommatidia; Pl = pleurite; Po = postfemur, Pp = prophragma; Pre = prefemur; Pt = paratergite; st = stigma, Ta = tarsus; Te = tergite; Tib = tibia; Tr = tracheal apodeme; ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5211757" httpUri="https://zenodo.org/record/5211757/files/figure.png" pageId="19" pageNumber="127">Fig. 12</figureCitation>
. While this approach revaled no significant additional details it serves to illustrate morphology of the specimen in an excellent way, much better than any photograph.
</paragraph>
</subSubSection>
</treatment>
</document>
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