725 lines
112 KiB
XML
725 lines
112 KiB
XML
<document id="5445E36D869E13EF110A1E7651D65700" ID-CLB-Dataset="24097" ID-DOI="10.11646/zootaxa.4731.4.3" ID-GBIF-Dataset="7b208096-3c50-4855-98c8-adbf0bb0902d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3661944" ID-ZooBank="0F693568-B754-4594-BBCB-9B396EBA978B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581407455457" checkinUser="plazi" docAuthor="Pérez-Pech, Wilbert Andrés, Jesús-Navarrate, Alberto De, Demilio, Erica, Anguas-Escalante, Abril & Hansen, Jesper Guldberg" docDate="2020" docId="039BF3253218FF91FF306757408FFDDF" docLanguage="en" docName="zootaxa.4731.4.3.pdf" docOrigin="Zootaxa 4731 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Styraconyx robertoi Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen 2020, sp. nov." docType="treatment" docVersion="5" lastPageNumber="504" masterDocId="FFA28B5D321EFF9DFFA764764468FFAA" masterDocTitle="Marine Tardigrada from the Mexican Caribbean with the description of Styraconyx robertoi sp. nov. (Arthrotardigrada: Styraconyxidae)" masterLastPageNumber="508" masterPageNumber="492" pageNumber="498" updateTime="1698781670076" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AB28F55BD58928824E82C06608276C52">Marine Tardigrada from the Mexican Caribbean with the description of Styraconyx robertoi sp. nov. (Arthrotardigrada: Styraconyxidae)</mods:title>
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<mods:namePart id="3F2E1B41EB522C4FF7DC4B4C9C38A211">Pérez-Pech, Wilbert Andrés</mods:namePart>
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<mods:namePart id="F804D25CE18F6749DC51FDD832F45081">Jesús-Navarrate, Alberto De</mods:namePart>
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<mods:roleTerm id="F051570EAEFB010366135114750D1550">Author</mods:roleTerm>
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<mods:namePart id="30F9550882DBBA26B22E01EF2DFBB155">Demilio, Erica</mods:namePart>
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<mods:namePart id="85BF95483CCD7E7F5E017B3D1970AF39">Anguas-Escalante, Abril</mods:namePart>
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<mods:namePart id="7DBC0E1DEBE19574B99BDBA40C84FC96">Hansen, Jesper Guldberg</mods:namePart>
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<mods:title id="EC6FA5D5E4636F4C600458B216FEB355">Zootaxa</mods:title>
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<mods:part id="55171E485A42CBC69003C49533C16D1D">
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<mods:date id="424CA3AEEC4D25A5DAB2C21E2F1D0726">2020</mods:date>
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<mods:number id="86C2215FB85A8C17FE7802113BC7FD9C">2020-02-10</mods:number>
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<treatment id="039BF3253218FF91FF306757408FFDDF" ID-DOI="http://doi.org/10.5281/zenodo.3664919" ID-GBIF-Taxon="162361610" ID-Zenodo-Dep="3664919" LSID="urn:lsid:plazi:treatment:039BF3253218FF91FF306757408FFDDF" httpUri="http://treatment.plazi.org/id/039BF3253218FF91FF306757408FFDDF" lastPageId="12" lastPageNumber="504" pageId="6" pageNumber="498">
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<paragraph id="8B8D42333218FF9BFF30675745B4FC96" blockId="6.[151,1128,801,864]" box="[151,476,801,828]" pageId="6" pageNumber="498">
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<emphasis id="B9469E213218FF9BFF30675745B4FC96" bold="true" box="[151,476,801,828]" pageId="6" pageNumber="498">
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<taxonomicName id="4C3239B03218FF9BFF3067574513FC96" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[151,379,801,828]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="498" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
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<emphasis id="B9469E213218FF9BFF3067574513FC96" bold="true" box="[151,379,801,828]" italics="true" pageId="6" pageNumber="498">Styraconyx robertoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A275235A3218FF9BFE25675445B4FC96" box="[386,476,802,828]" pageId="6" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8B8D42333218FF9BFF3067334000FCCA" blockId="6.[151,1128,801,864]" box="[151,1128,837,864]" pageId="6" pageNumber="498">
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zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C32811B83218FF9BFF3067FB4777FC66" pageId="6" pageNumber="498" type="materials_examined">
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<paragraph id="8B8D42333218FF9BFF3067FB4777FC66" blockId="6.[151,1436,909,1043]" pageId="6" pageNumber="498">
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<emphasis id="B9469E213218FF9BFF3067FB45EAFC02" bold="true" box="[151,386,909,936]" pageId="6" pageNumber="498">Material examined:</emphasis>
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<specimenCount id="9D3489BA3218FF9BFE2E67FB4649FC0D" box="[393,545,909,935]" pageId="6" pageNumber="498" type="generic">20 specimens</specimenCount>
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(
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:
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<specimenCount id="9D3489BA3218FF9BFDC067FB46A3FC0D" box="[615,715,909,935]" pageId="6" pageNumber="498" type="female">1 female</specimenCount>
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,
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<specimenCount id="9D3489BA3218FF9BFD7167FB474CFC0D" box="[726,804,909,935]" pageId="6" pageNumber="498" type="male">1 male</specimenCount>
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, 2 indeterminable gender;
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<collectionCode id="ED23DAF63218FF9BFBEA67F84013FC02" box="[1101,1147,910,936]" country="France" name="Musee de Lectoure" pageId="6" pageNumber="498">ML</collectionCode>
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: 1 indeterminable gender; XK:
|
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<specimenCount id="9D3489BA3218FF9BFEBD67C745FFFC61" box="[282,407,945,971]" pageId="6" pageNumber="498" type="female">10 females</specimenCount>
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,
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<specimenCount id="9D3489BA3218FF9BFE0567C74593FC61" box="[418,507,945,971]" pageId="6" pageNumber="498" type="male">3 males</specimenCount>
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, 2 indeterminable gender)
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</paragraph>
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<subSubSection id="C32811B83218FF9BFF3067A04551FBB9" pageId="6" pageNumber="498" type="discussion">
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<paragraph id="8B8D42333218FF9BFF3067A04551FBB9" blockId="6.[151,1436,909,1043]" pageId="6" pageNumber="498">
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<emphasis id="B9469E213218FF9BFF3067A04565FC5A" bold="true" box="[151,269,982,1008]" pageId="6" pageNumber="498">Remarks:</emphasis>
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The population occurring at Xcalak is designated as the
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<typeStatus id="5489FC913218FF9BFC3967A047A6FC5A" box="[926,974,982,1008]" pageId="6" pageNumber="498">type</typeStatus>
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population. See Description of the New Species
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</paragraph>
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</subSubSection>
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<subSubSection id="C32811B83218FF9BFF306013458EFB62" pageId="6" pageNumber="498" type="description">
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<subSection id="E2BD5BD83218FF9BFF306013467FFBD5" box="[151,535,1125,1152]" pageId="6" pageNumber="498" type="multiple">
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<paragraph id="8B8D42333218FF9BFF306013467FFBD5" blockId="6.[151,535,1125,1152]" box="[151,535,1125,1152]" pageId="6" pageNumber="498">
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<heading id="D0C5F55F3218FF9BFF306013467FFBD5" bold="true" box="[151,535,1125,1152]" fontSize="11" level="1" pageId="6" pageNumber="498" reason="1">
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<emphasis id="B9469E213218FF9BFF306013467FFBD5" bold="true" box="[151,535,1125,1152]" pageId="6" pageNumber="498">Description of the New Species</emphasis>
|
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</heading>
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</paragraph>
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</subSection>
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<paragraph id="8B8D42333218FF9BFF3060DB458EFB62" blockId="6.[151,486,1197,1224]" box="[151,486,1197,1224]" pageId="6" pageNumber="498">
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<heading id="D0C5F55F3218FF9BFF3060DB458EFB62" bold="true" box="[151,486,1197,1224]" fontSize="11" level="1" pageId="6" pageNumber="498" reason="1">
|
||
<emphasis id="B9469E213218FF9BFF3060DB458EFB62" bold="true" box="[151,486,1197,1224]" pageId="6" pageNumber="498">
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||
<taxonomicName id="4C3239B03218FF9BFF3060DB45EEFB62" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[151,390,1197,1224]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="498" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213218FF9BFF3060DB45EEFB62" bold="true" box="[151,390,1197,1224]" italics="true" pageId="6" pageNumber="498">Styraconyx robertoi</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A275235A3218FF9BFE2A60D8458EFB62" box="[397,486,1198,1224]" pageId="6" pageNumber="498" rank="species">sp. nov</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C32811B83218FF9AFF30608040A4FF19" lastPageId="7" lastPageNumber="499" pageId="6" pageNumber="498" type="materials_examined">
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<paragraph id="8B8D42333218FF9BFF3060804592FA99" blockId="6.[151,1436,1269,1332]" pageId="6" pageNumber="498">
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<emphasis id="B9469E213218FF9BFF3060804522FABA" bold="true" box="[151,330,1269,1296]" pageId="6" pageNumber="498">Type Material:</emphasis>
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<materialsCitation id="3B5A486E3218FF9BFEF76083459EFA99" ID-GBIF-Occurrence="2572158540" location="Morphometric" pageId="6" pageNumber="498" specimenCount="14" specimenCount-female="9" specimenCount-larva="3" specimenCount-male="2" typeStatus="holotype">
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<specimenCount id="9D3489BA3218FF9BFEF7608345CAFAA5" box="[336,418,1269,1295]" pageId="6" pageNumber="498" type="female">Female</specimenCount>
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<typeStatus id="5489FC913218FF9BFE0F60834662FAA5" box="[424,522,1269,1295]" pageId="6" pageNumber="498" type="holotype">holotype</typeStatus>
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and 13 paratypes (
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<specimenCount id="9D3489BA3218FF9BFD7B60834723FAA5" box="[732,843,1269,1295]" pageId="6" pageNumber="498" type="female">8 females</specimenCount>
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,
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<specimenCount id="9D3489BA3218FF9BFCF2608347C5FAA5" box="[853,941,1269,1295]" pageId="6" pageNumber="498" type="male">2 males</specimenCount>
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,
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<specimenCount id="9D3489BA3218FF9BFC1F6083407DFABA" box="[952,1045,1269,1296]" pageId="6" pageNumber="498" type="larva">3 larvae</specimenCount>
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).
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<location id="8EED14E83218FF9BFB80608340A1FAA5" LSID="urn:lsid:plazi:treatment:039BF3253218FF91FF306757408FFDDF:8EED14E83218FF9BFB80608340A1FAA5" box="[1063,1225,1269,1295]" name="Morphometric" pageId="6" pageNumber="498">Morphometric</location>
|
||
values for the type population are given in
|
||
<tableCitation id="C6B077883218FF9BFE06616F459EFA99" box="[417,502,1305,1331]" captionStart="TABLE 2" captionStartId="6.[151,239,1379,1403]" captionTargetBox="[166,1375,1415,2028]" captionTargetPageId="6" captionText="TABLE 2. Measurements (μm) of the type series of Styraconyx robertoi sp. nov. N= number of individuals measured." httpUri="http://table.plazi.org/id/DF4D12BB3218FF9BFF30611541EFFAD1" pageId="6" pageNumber="498" tableUuid="DF4D12BB3218FF9BFF30611541EFFAD1">
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<location id="8EED14E83218FF9BFE06616F45B7FA99" LSID="urn:lsid:plazi:treatment:039BF3253218FF91FF306757408FFDDF:8EED14E83218FF9BFE06616F45B7FA99" box="[417,479,1305,1331]" name="Table" pageId="6" pageNumber="498">Table</location>
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2
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</materialsCitation>
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.
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</paragraph>
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<caption id="DF4D12BB3218FF9BFF30611541EFFAD1" ID-Table-UUID="DF4D12BB3218FF9BFF30611541EFFAD1" box="[151,1415,1379,1404]" httpUri="http://table.plazi.org/id/DF4D12BB3218FF9BFF30611541EFFAD1" pageId="6" pageNumber="498" targetBox="[166,1375,1415,2028]" targetIsTable="true" targetPageId="6">
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<paragraph id="8B8D42333218FF9BFF30611541EFFAD1" blockId="6.[151,1415,1379,1404]" box="[151,1415,1379,1404]" pageId="6" pageNumber="498">
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<emphasis id="B9469E213218FF9BFF306115456BFAD1" bold="true" box="[151,259,1379,1403]" pageId="6" pageNumber="498">TABLE 2</emphasis>
|
||
. Measurements (μm) of the type series of
|
||
<taxonomicName id="4C3239B03218FF9BFD6F611547F3FAD1" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[712,923,1379,1403]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="498" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
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<emphasis id="B9469E213218FF9BFD6F611547F3FAD1" box="[712,923,1379,1403]" italics="true" pageId="6" pageNumber="498">Styraconyx robertoi</emphasis>
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</taxonomicName>
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<emphasis id="B9469E213218FF9BFC056115479FFAD1" bold="true" box="[930,1015,1379,1403]" pageId="6" pageNumber="498">
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<taxonomicNameLabel id="A275235A3218FF9BFC056115479FFAD1" box="[930,1015,1379,1403]" pageId="6" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
N= number of individuals measured.
|
||
</paragraph>
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||
</caption>
|
||
<paragraph id="8B8D42333218FF9BFF0161F1412EF846" pageId="6" pageNumber="498">
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<table id="F932B09332180062FF0161F14137F846" box="[166,1375,1415,2028]" gridcols="9" gridrows="16" pageId="6" pageNumber="498">
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<tr id="3502407132180062FF0161F14137FA0A" box="[166,1375,1415,1440]" gridrow="0" pageId="6" pageNumber="498" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
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<th id="76D3290D32180062FF0161F1450FFA0A" box="[166,359,1415,1440]" gridcol="0" gridrow="0" pageId="6" pageNumber="498">Character</th>
|
||
<th id="76D3290D32180062FE6E61F14644FA0A" box="[457,556,1415,1440]" gridcol="1" gridrow="0" pageId="6" pageNumber="498">Holotype</th>
|
||
<th id="76D3290D32180062FDCA61F14696FA0A" box="[621,766,1415,1440]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="6" pageNumber="498">
|
||
<emphasis id="B9469E213218FF9BFDCA61F146B1FA35" bold="true" box="[621,729,1415,1439]" pageId="6" pageNumber="498">Paratypes</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="3502407132180062FF0161D14137FA4A" box="[166,1375,1447,1504]" gridrow="1" pageId="6" pageNumber="498" rowspan-0="1">
|
||
<td id="76D3290D32180062FE6E61D14644FA4A" box="[457,556,1447,1504]" gridcol="1" gridrow="1" pageId="6" pageNumber="498">Female</td>
|
||
<td id="76D3290D32180062FDCA61D14696FA4A" box="[621,766,1447,1504]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="6" pageNumber="498">Females N Range</td>
|
||
<td id="76D3290D32180062FC9C61D147A2FA4A" box="[827,970,1447,1504]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="6" pageNumber="498">Males N Range</td>
|
||
<td id="76D3290D32180062FBAE61D140E0FA4A" box="[1033,1160,1447,1504]" gridcol="6" gridrow="1" pageId="6" pageNumber="498">Male Larva</td>
|
||
<td id="76D3290D32180062FB7361D14137FA4A" box="[1236,1375,1447,1504]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="6" pageNumber="498">
|
||
<emphasis id="B9469E213218FF9BFB7361D14134FA15" bold="true" box="[1236,1372,1447,1472]" pageId="6" pageNumber="498">Other Larva</emphasis>
|
||
N Range
|
||
</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF01619E4137F9AB" box="[166,1375,1512,1537]" gridrow="2" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF01619E450FF9AB" box="[166,359,1512,1537]" gridcol="0" gridrow="2" pageId="6" pageNumber="498">Body length</th>
|
||
<td id="76D3290D32180062FE6E619E4644F9AB" box="[457,556,1512,1537]" gridcol="1" gridrow="2" pageId="6" pageNumber="498">139</td>
|
||
<td id="76D3290D32180062FDCA619E4612F9AB" box="[621,634,1512,1537]" gridcol="2" gridrow="2" pageId="6" pageNumber="498">8</td>
|
||
<td id="76D3290D32180062FD02619E4696F9AB" box="[677,766,1512,1537]" gridcol="3" gridrow="2" pageId="6" pageNumber="498">128–150</td>
|
||
<td id="76D3290D32180062FC9C619E4720F9AB" box="[827,840,1512,1537]" gridcol="4" gridrow="2" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD4619E47A2F9AB" box="[883,970,1512,1537]" gridcol="5" gridrow="2" pageId="6" pageNumber="498">112–154</td>
|
||
<td id="76D3290D32180062FBAE619E40E0F9AB" box="[1033,1160,1512,1537]" gridcol="6" gridrow="2" pageId="6" pageNumber="498">112</td>
|
||
<td id="76D3290D32180062FB73619E4089F9AB" box="[1236,1249,1512,1537]" gridcol="7" gridrow="2" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB4619E4137F9AB" box="[1299,1375,1512,1537]" gridcol="8" gridrow="2" pageId="6" pageNumber="498">95–114</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF01627B4137F98C" box="[166,1375,1549,1574]" gridrow="3" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF01627B450FF98C" box="[166,359,1549,1574]" gridcol="0" gridrow="3" pageId="6" pageNumber="498">Body width</th>
|
||
<td id="76D3290D32180062FE6E627B4644F98C" box="[457,556,1549,1574]" gridcol="1" gridrow="3" pageId="6" pageNumber="498">46</td>
|
||
<td id="76D3290D32180062FDCA627B4612F98C" box="[621,634,1549,1574]" gridcol="2" gridrow="3" pageId="6" pageNumber="498">8</td>
|
||
<td id="76D3290D32180062FD02627B4696F98C" box="[677,766,1549,1574]" gridcol="3" gridrow="3" pageId="6" pageNumber="498">47–69</td>
|
||
<td id="76D3290D32180062FC9C627B4720F98C" box="[827,840,1549,1574]" gridcol="4" gridrow="3" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD4627B47A2F98C" box="[883,970,1549,1574]" gridcol="5" gridrow="3" pageId="6" pageNumber="498">36–51</td>
|
||
<td id="76D3290D32180062FBAE627B40E0F98C" box="[1033,1160,1549,1574]" gridcol="6" gridrow="3" pageId="6" pageNumber="498">40</td>
|
||
<td id="76D3290D32180062FB73627B4089F98C" box="[1236,1249,1549,1574]" gridcol="7" gridrow="3" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB4627B4137F98C" box="[1299,1375,1549,1574]" gridcol="8" gridrow="3" pageId="6" pageNumber="498">40–43</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0162454137F9E6" box="[166,1375,1587,1612]" gridrow="4" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF016245450FF9E6" box="[166,359,1587,1612]" gridcol="0" gridrow="4" pageId="6" pageNumber="498">Caudal width</th>
|
||
<td id="76D3290D32180062FE6E62454644F9E6" box="[457,556,1587,1612]" gridcol="1" gridrow="4" pageId="6" pageNumber="498">36</td>
|
||
<td id="76D3290D32180062FDCA62454612F9E6" box="[621,634,1587,1612]" gridcol="2" gridrow="4" pageId="6" pageNumber="498">8</td>
|
||
<td id="76D3290D32180062FD0262454696F9E6" box="[677,766,1587,1612]" gridcol="3" gridrow="4" pageId="6" pageNumber="498">33–47</td>
|
||
<td id="76D3290D32180062FC9C62454720F9E6" box="[827,840,1587,1612]" gridcol="4" gridrow="4" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD4624547A2F9E6" box="[883,970,1587,1612]" gridcol="5" gridrow="4" pageId="6" pageNumber="498">26–32</td>
|
||
<td id="76D3290D32180062FBAE624540E0F9E6" box="[1033,1160,1587,1612]" gridcol="6" gridrow="4" pageId="6" pageNumber="498">32</td>
|
||
<td id="76D3290D32180062FB7362454089F9E6" box="[1236,1249,1587,1612]" gridcol="7" gridrow="4" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB462454137F9E6" box="[1299,1375,1587,1612]" gridcol="8" gridrow="4" pageId="6" pageNumber="498">30–32</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF01622F4137F9D8" box="[166,1375,1625,1650]" gridrow="5" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF01622F450FF9D8" box="[166,359,1625,1650]" gridcol="0" gridrow="5" pageId="6" pageNumber="498">Median cirrus</th>
|
||
<td id="76D3290D32180062FE6E622F4644F9D8" box="[457,556,1625,1650]" gridcol="1" gridrow="5" pageId="6" pageNumber="498">9</td>
|
||
<td id="76D3290D32180062FDCA622F4612F9D8" box="[621,634,1625,1650]" gridcol="2" gridrow="5" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FD02622F4696F9D8" box="[677,766,1625,1650]" gridcol="3" gridrow="5" pageId="6" pageNumber="498">9–13</td>
|
||
<td id="76D3290D32180062FC9C622F4720F9D8" box="[827,840,1625,1650]" gridcol="4" gridrow="5" pageId="6" pageNumber="498">1</td>
|
||
<td id="76D3290D32180062FCD4622F47A2F9D8" box="[883,970,1625,1650]" gridcol="5" gridrow="5" pageId="6" pageNumber="498">8</td>
|
||
<td id="76D3290D32180062FBAE622F40E0F9D8" box="[1033,1160,1625,1650]" gridcol="6" gridrow="5" pageId="6" pageNumber="498">8</td>
|
||
<td id="76D3290D32180062FB73622F4089F9D8" box="[1236,1249,1625,1650]" gridcol="7" gridrow="5" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB4622F4137F9D8" box="[1299,1375,1625,1650]" gridcol="8" gridrow="5" pageId="6" pageNumber="498">7–8</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0162094137F932" box="[166,1375,1663,1688]" gridrow="6" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF016209450FF932" box="[166,359,1663,1688]" gridcol="0" gridrow="6" pageId="6" pageNumber="498">Internal cirrus</th>
|
||
<td id="76D3290D32180062FE6E62094644F932" box="[457,556,1663,1688]" gridcol="1" gridrow="6" pageId="6" pageNumber="498">13</td>
|
||
<td id="76D3290D32180062FDCA62094612F932" box="[621,634,1663,1688]" gridcol="2" gridrow="6" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FD0262094696F932" box="[677,766,1663,1688]" gridcol="3" gridrow="6" pageId="6" pageNumber="498">10–13</td>
|
||
<td id="76D3290D32180062FC9C62094720F932" box="[827,840,1663,1688]" gridcol="4" gridrow="6" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD4620947A2F932" box="[883,970,1663,1688]" gridcol="5" gridrow="6" pageId="6" pageNumber="498">11</td>
|
||
<td id="76D3290D32180062FBAE620940E0F932" box="[1033,1160,1663,1688]" gridcol="6" gridrow="6" pageId="6" pageNumber="498">10</td>
|
||
<td id="76D3290D32180062FB7362094089F932" box="[1236,1249,1663,1688]" gridcol="7" gridrow="6" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB462094137F932" box="[1299,1375,1663,1688]" gridcol="8" gridrow="6" pageId="6" pageNumber="498">10</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0162D34137F914" box="[166,1375,1701,1726]" gridrow="7" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF0162D3450FF914" box="[166,359,1701,1726]" gridcol="0" gridrow="7" pageId="6" pageNumber="498">External cirrus</th>
|
||
<td id="76D3290D32180062FE6E62D34644F914" box="[457,556,1701,1726]" gridcol="1" gridrow="7" pageId="6" pageNumber="498">12</td>
|
||
<td id="76D3290D32180062FDCA62D34612F914" box="[621,634,1701,1726]" gridcol="2" gridrow="7" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FD0262D34696F914" box="[677,766,1701,1726]" gridcol="3" gridrow="7" pageId="6" pageNumber="498">8–13</td>
|
||
<td id="76D3290D32180062FC9C62D34720F914" box="[827,840,1701,1726]" gridcol="4" gridrow="7" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD462D347A2F914" box="[883,970,1701,1726]" gridcol="5" gridrow="7" pageId="6" pageNumber="498">12–13</td>
|
||
<td id="76D3290D32180062FBAE62D340E0F914" box="[1033,1160,1701,1726]" gridcol="6" gridrow="7" pageId="6" pageNumber="498">8</td>
|
||
<td id="76D3290D32180062FB7362D34089F914" box="[1236,1249,1701,1726]" gridcol="7" gridrow="7" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB462D34137F914" box="[1299,1375,1701,1726]" gridcol="8" gridrow="7" pageId="6" pageNumber="498">8–9</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0162BC4137F949" box="[166,1375,1738,1763]" gridrow="8" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF0162BC450FF949" box="[166,359,1738,1763]" gridcol="0" gridrow="8" pageId="6" pageNumber="498">Lateral cirrus</th>
|
||
<td id="76D3290D32180062FE6E62BC4644F949" box="[457,556,1738,1763]" gridcol="1" gridrow="8" pageId="6" pageNumber="498">28</td>
|
||
<td id="76D3290D32180062FDCA62BC4612F949" box="[621,634,1738,1763]" gridcol="2" gridrow="8" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FD0262BC4696F949" box="[677,766,1738,1763]" gridcol="3" gridrow="8" pageId="6" pageNumber="498">23–29</td>
|
||
<td id="76D3290D32180062FC9C62BC4720F949" box="[827,840,1738,1763]" gridcol="4" gridrow="8" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD462BC47A2F949" box="[883,970,1738,1763]" gridcol="5" gridrow="8" pageId="6" pageNumber="498">22–26</td>
|
||
<td id="76D3290D32180062FBAE62BC40E0F949" box="[1033,1160,1738,1763]" gridcol="6" gridrow="8" pageId="6" pageNumber="498">18</td>
|
||
<td id="76D3290D32180062FB7362BC4089F949" box="[1236,1249,1738,1763]" gridcol="7" gridrow="8" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB462BC4137F949" box="[1299,1375,1738,1763]" gridcol="8" gridrow="8" pageId="6" pageNumber="498">18–20</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0162864137F8A3" box="[166,1375,1776,1801]" gridrow="9" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF016286450FF8A3" box="[166,359,1776,1801]" gridcol="0" gridrow="9" pageId="6" pageNumber="498">Primary clava</th>
|
||
<td id="76D3290D32180062FE6E62864644F8A3" box="[457,556,1776,1801]" gridcol="1" gridrow="9" pageId="6" pageNumber="498">10</td>
|
||
<td id="76D3290D32180062FDCA62864612F8A3" box="[621,634,1776,1801]" gridcol="2" gridrow="9" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FD0262864696F8A3" box="[677,766,1776,1801]" gridcol="3" gridrow="9" pageId="6" pageNumber="498">9–12</td>
|
||
<td id="76D3290D32180062FC9C62864720F8A3" box="[827,840,1776,1801]" gridcol="4" gridrow="9" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD4628647A2F8A3" box="[883,970,1776,1801]" gridcol="5" gridrow="9" pageId="6" pageNumber="498">8–10</td>
|
||
<td id="76D3290D32180062FBAE628640E0F8A3" box="[1033,1160,1776,1801]" gridcol="6" gridrow="9" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FB7362864089F8A3" box="[1236,1249,1776,1801]" gridcol="7" gridrow="9" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB462864137F8A3" box="[1299,1375,1776,1801]" gridcol="8" gridrow="9" pageId="6" pageNumber="498">7–8</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0163604137F885" box="[166,1375,1814,1839]" gridrow="10" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF016360450FF885" box="[166,359,1814,1839]" gridcol="0" gridrow="10" pageId="6" pageNumber="498">Secondary clava</th>
|
||
<td id="76D3290D32180062FE6E63604644F885" box="[457,556,1814,1839]" gridcol="1" gridrow="10" pageId="6" pageNumber="498">12</td>
|
||
<td id="76D3290D32180062FDCA63604612F885" box="[621,634,1814,1839]" gridcol="2" gridrow="10" pageId="6" pageNumber="498">5</td>
|
||
<td id="76D3290D32180062FD0263604696F885" box="[677,766,1814,1839]" gridcol="3" gridrow="10" pageId="6" pageNumber="498">12</td>
|
||
<td id="76D3290D32180062FC9C63604720F885" box="[827,840,1814,1839]" gridcol="4" gridrow="10" pageId="6" pageNumber="498">0</td>
|
||
<td id="76D3290D32180062FCD4636047A2F885" box="[883,970,1814,1839]" gridcol="5" gridrow="10" pageId="6" pageNumber="498">-</td>
|
||
<td id="76D3290D32180062FBAE636040E0F885" box="[1033,1160,1814,1839]" gridcol="6" gridrow="10" pageId="6" pageNumber="498">10</td>
|
||
<td id="76D3290D32180062FB7363604089F885" box="[1236,1249,1814,1839]" gridcol="7" gridrow="10" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB463604137F885" box="[1299,1375,1814,1839]" gridcol="8" gridrow="10" pageId="6" pageNumber="498">10–12</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF01634A4137F8FF" box="[166,1375,1852,1877]" gridrow="11" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF01634A450FF8FF" box="[166,359,1852,1877]" gridcol="0" gridrow="11" pageId="6" pageNumber="498">Cirrus E</th>
|
||
<td id="76D3290D32180062FE6E634A4644F8FF" box="[457,556,1852,1877]" gridcol="1" gridrow="11" pageId="6" pageNumber="498">26</td>
|
||
<td id="76D3290D32180062FDCA634A4612F8FF" box="[621,634,1852,1877]" gridcol="2" gridrow="11" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FD02634A4696F8FF" box="[677,766,1852,1877]" gridcol="3" gridrow="11" pageId="6" pageNumber="498">21–28</td>
|
||
<td id="76D3290D32180062FC9C634A4720F8FF" box="[827,840,1852,1877]" gridcol="4" gridrow="11" pageId="6" pageNumber="498">1</td>
|
||
<td id="76D3290D32180062FCD4634A47A2F8FF" box="[883,970,1852,1877]" gridcol="5" gridrow="11" pageId="6" pageNumber="498">23</td>
|
||
<td id="76D3290D32180062FBAE634A40E0F8FF" box="[1033,1160,1852,1877]" gridcol="6" gridrow="11" pageId="6" pageNumber="498">22</td>
|
||
<td id="76D3290D32180062FB73634A4089F8FF" box="[1236,1249,1852,1877]" gridcol="7" gridrow="11" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB4634A4137F8FF" box="[1299,1375,1852,1877]" gridcol="8" gridrow="11" pageId="6" pageNumber="498">22</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0163144137F8D1" box="[166,1375,1890,1915]" gridrow="12" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF016314450FF8D1" box="[166,359,1890,1915]" gridcol="0" gridrow="12" pageId="6" pageNumber="498">Sense organ leg I</th>
|
||
<td id="76D3290D32180062FE6E63144644F8D1" box="[457,556,1890,1915]" gridcol="1" gridrow="12" pageId="6" pageNumber="498">9</td>
|
||
<td id="76D3290D32180062FDCA63144612F8D1" box="[621,634,1890,1915]" gridcol="2" gridrow="12" pageId="6" pageNumber="498">6</td>
|
||
<td id="76D3290D32180062FD0263144696F8D1" box="[677,766,1890,1915]" gridcol="3" gridrow="12" pageId="6" pageNumber="498">8–11</td>
|
||
<td id="76D3290D32180062FC9C63144720F8D1" box="[827,840,1890,1915]" gridcol="4" gridrow="12" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD4631447A2F8D1" box="[883,970,1890,1915]" gridcol="5" gridrow="12" pageId="6" pageNumber="498">8–9</td>
|
||
<td id="76D3290D32180062FBAE631440E0F8D1" box="[1033,1160,1890,1915]" gridcol="6" gridrow="12" pageId="6" pageNumber="498">10</td>
|
||
<td id="76D3290D32180062FB7363144089F8D1" box="[1236,1249,1890,1915]" gridcol="7" gridrow="12" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB463144137F8D1" box="[1299,1375,1890,1915]" gridcol="8" gridrow="12" pageId="6" pageNumber="498">7–10</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0163F14137F80A" box="[166,1375,1927,1952]" gridrow="13" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF0163F1450FF80A" box="[166,359,1927,1952]" gridcol="0" gridrow="13" pageId="6" pageNumber="498">Sense organ leg II</th>
|
||
<td id="76D3290D32180062FE6E63F14644F80A" box="[457,556,1927,1952]" gridcol="1" gridrow="13" pageId="6" pageNumber="498">-</td>
|
||
<td id="76D3290D32180062FDCA63F14612F80A" box="[621,634,1927,1952]" gridcol="2" gridrow="13" pageId="6" pageNumber="498">3</td>
|
||
<td id="76D3290D32180062FD0263F14696F80A" box="[677,766,1927,1952]" gridcol="3" gridrow="13" pageId="6" pageNumber="498">7–11</td>
|
||
<td id="76D3290D32180062FC9C63F14720F80A" box="[827,840,1927,1952]" gridcol="4" gridrow="13" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD463F147A2F80A" box="[883,970,1927,1952]" gridcol="5" gridrow="13" pageId="6" pageNumber="498">7–10</td>
|
||
<td id="76D3290D32180062FBAE63F140E0F80A" box="[1033,1160,1927,1952]" gridcol="6" gridrow="13" pageId="6" pageNumber="498">7</td>
|
||
<td id="76D3290D32180062FB7363F14089F80A" box="[1236,1249,1927,1952]" gridcol="7" gridrow="13" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB463F14137F80A" box="[1299,1375,1927,1952]" gridcol="8" gridrow="13" pageId="6" pageNumber="498">6–7</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0163DB4137F86C" box="[166,1375,1965,1990]" gridrow="14" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF0163DB450FF86C" box="[166,359,1965,1990]" gridcol="0" gridrow="14" pageId="6" pageNumber="498">Sense organ leg III</th>
|
||
<td id="76D3290D32180062FE6E63DB4644F86C" box="[457,556,1965,1990]" gridcol="1" gridrow="14" pageId="6" pageNumber="498">-</td>
|
||
<td id="76D3290D32180062FDCA63DB4612F86C" box="[621,634,1965,1990]" gridcol="2" gridrow="14" pageId="6" pageNumber="498">1</td>
|
||
<td id="76D3290D32180062FD0263DB4696F86C" box="[677,766,1965,1990]" gridcol="3" gridrow="14" pageId="6" pageNumber="498">10</td>
|
||
<td id="76D3290D32180062FC9C63DB4720F86C" box="[827,840,1965,1990]" gridcol="4" gridrow="14" pageId="6" pageNumber="498">1</td>
|
||
<td id="76D3290D32180062FCD463DB47A2F86C" box="[883,970,1965,1990]" gridcol="5" gridrow="14" pageId="6" pageNumber="498">9</td>
|
||
<td id="76D3290D32180062FBAE63DB40E0F86C" box="[1033,1160,1965,1990]" gridcol="6" gridrow="14" pageId="6" pageNumber="498">9</td>
|
||
<td id="76D3290D32180062FB7363DB4089F86C" box="[1236,1249,1965,1990]" gridcol="7" gridrow="14" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB463DB4137F86C" box="[1299,1375,1965,1990]" gridcol="8" gridrow="14" pageId="6" pageNumber="498">7–9</td>
|
||
</tr>
|
||
<tr id="3502407132180062FF0163A54137F846" box="[166,1375,2003,2028]" gridrow="15" pageId="6" pageNumber="498">
|
||
<th id="76D3290D32180062FF0163A5450FF846" box="[166,359,2003,2028]" gridcol="0" gridrow="15" pageId="6" pageNumber="498">Sense organ leg IV</th>
|
||
<td id="76D3290D32180062FE6E63A54644F846" box="[457,556,2003,2028]" gridcol="1" gridrow="15" pageId="6" pageNumber="498">13</td>
|
||
<td id="76D3290D32180062FDCA63A54612F846" box="[621,634,2003,2028]" gridcol="2" gridrow="15" pageId="6" pageNumber="498">6</td>
|
||
<td id="76D3290D32180062FD0263A54696F846" box="[677,766,2003,2028]" gridcol="3" gridrow="15" pageId="6" pageNumber="498">11–12</td>
|
||
<td id="76D3290D32180062FC9C63A54720F846" box="[827,840,2003,2028]" gridcol="4" gridrow="15" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FCD463A547A2F846" box="[883,970,2003,2028]" gridcol="5" gridrow="15" pageId="6" pageNumber="498">10–11</td>
|
||
<td id="76D3290D32180062FBAE63A540E0F846" box="[1033,1160,2003,2028]" gridcol="6" gridrow="15" pageId="6" pageNumber="498">8</td>
|
||
<td id="76D3290D32180062FB7363A54089F846" box="[1236,1249,2003,2028]" gridcol="7" gridrow="15" pageId="6" pageNumber="498">2</td>
|
||
<td id="76D3290D32180062FAB463A54137F846" box="[1299,1375,2003,2028]" gridcol="8" gridrow="15" pageId="6" pageNumber="498">8–10</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B8D42333219FF9AFF6064EF40A4FF19" blockId="7.[151,1437,152,2024]" box="[199,1228,152,179]" pageId="7" pageNumber="499">
|
||
<emphasis id="B9469E213219FF9AFF6064EF451CFF19" bold="true" box="[199,372,152,179]" pageId="7" pageNumber="499">Type Locality:</emphasis>
|
||
reef lagoon, 0.2–1.0 m depth, subtidal zone, Xcalak,
|
||
<collectingRegion id="49F68CD13219FF9AFC6B64EF4002FF19" box="[972,1130,153,179]" country="Mexico" name="Quintana Roo" pageId="7" pageNumber="499">Quintana Roo</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32502A33219FF9AFBD164EF40A4FF19" box="[1142,1228,153,179]" name="Mexico" pageId="7" pageNumber="499">Mexico</collectingCountry>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32811B83219FF9AFF6064CA453EFE59" pageId="7" pageNumber="499" type="diagnosis">
|
||
<paragraph id="8B8D42333219FF9AFF6064CA453EFE59" blockId="7.[151,1437,152,2024]" pageId="7" pageNumber="499">
|
||
<emphasis id="B9469E213219FF9AFF6064CA452CFF7D" bold="true" box="[199,324,188,215]" pageId="7" pageNumber="499">Diagnosis:</emphasis>
|
||
<taxonomicName id="4C3239B03219FF9AFEED64CB45AEFF7D" authorityName="Thulin" authorityYear="1942" box="[330,454,189,215]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="7" pageNumber="499" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B9469E213219FF9AFEED64CB45AEFF7D" box="[330,454,189,215]" italics="true" pageId="7" pageNumber="499">Styraconyx</emphasis>
|
||
</taxonomicName>
|
||
with subterminal mouth cone, pigmented eyespots present. Asymmetric primary clavae and lateral cirri with common pedestal and surrounded basally by common membrane. Lateral cirri not segmented and with asymmetrically expanded base. Lenticular secondary clavae. Cephalic cirri composed of scapus and tubular, flagellar part only, cirrophore and protruding flagellum absent. Peduncles present only on external digits and reaching claw base, internal digits with proximal pads. In males, internal digits with peduncles present only on leg pair IV. Three-pointed claws with strong primary hook, thinner secondary and accessory hook. Claw sheath not covering tip of claw hooks. Sense organs of first leg two-parted, unsegmented spines on second and third leg. Asymmetric papilla with broad base and terminal spine on fourth leg. Dorsal cuticle with grid-like sculpture and fine punctuation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32811B83219FF96FF60658A41E5F9DA" lastPageId="11" lastPageNumber="503" pageId="7" pageNumber="499" type="description">
|
||
<paragraph id="8B8D42333219FF9AFF60658A46F4FC98" blockId="7.[151,1437,152,2024]" pageId="7" pageNumber="499">
|
||
<emphasis id="B9469E213219FF9AFF60658A45AFFDBC" bold="true" box="[199,455,507,534]" pageId="7" pageNumber="499">
|
||
<typeStatus id="5489FC913219FF9AFF60658A455CFDBC" box="[199,308,508,534]" pageId="7" pageNumber="499" type="holotype">Holotype</typeStatus>
|
||
description.
|
||
</emphasis>
|
||
Female (
|
||
<figureCitation id="13095EB63219FF9AFD89658A4615FDBC" box="[558,637,508,534]" captionStart="FIGURE 2" captionStartId="8.[151,250,1939,1964]" captionTargetBox="[152,1435,409,1918]" captionTargetId="figure@8.[151,1435,409,1919]" captionTargetPageId="8" captionText="FIGURE 2. DIC-micrographs of Styraconyx robertoi sp. nov., holotypic female. A. Overview. B. Grid-like sculpture of the dorsal cuticle. C. Gonopore and anus." figureDoi="http://doi.org/10.5281/zenodo.3661950" httpUri="https://zenodo.org/record/3661950/files/figure.png" pageId="7" pageNumber="499">Figs. 2</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFD2F658A46AFFDBC" box="[648,711,508,534]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="7" pageNumber="499">3B,D</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFD75658A4688FDBC" box="[722,736,508,534]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">4</figureCitation>
|
||
A–C; 7) with roughly cylindrical body form (139 μm in length) broadest between leg pairs II and III (46 μm) and distinctly narrower (36 μm) at the point of the insertion of cirri E. Eye spots (
|
||
<figureCitation id="13095EB63219FF9AFE94663545C6FDF7" box="[307,430,579,605]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="7" pageNumber="499">Figs. 3A,E</figureCitation>
|
||
) visible as congregations of brown pigment granules on each side of the pharyngeal bulb slightly posterior to the pedestals bearing the primary clavae. Pillars highly conspicuous in the dorsal cuticle and to a lesser degree in the ventral cuticle. Twenty-two ribs traverse the dorsal surface at right angles to eight equally spaced longitudinal ribs forming a distinct grid-like pattern (
|
||
<figureCitation id="13095EB63219FF9AFC9166DB47DCFD62" box="[822,948,685,712]" captionStart="FIGURE 2" captionStartId="8.[151,250,1939,1964]" captionTargetBox="[152,1435,409,1918]" captionTargetId="figure@8.[151,1435,409,1919]" captionTargetPageId="8" captionText="FIGURE 2. DIC-micrographs of Styraconyx robertoi sp. nov., holotypic female. A. Overview. B. Grid-like sculpture of the dorsal cuticle. C. Gonopore and anus." figureDoi="http://doi.org/10.5281/zenodo.3661950" httpUri="https://zenodo.org/record/3661950/files/figure.png" pageId="7" pageNumber="499">Figs. 2A,B</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFC1966D847A6FD62" box="[958,974,686,712]" captionStart="FIGURE 7" captionStartId="13.[151,250,2007,2032]" captionTargetBox="[194,1379,179,1971]" captionTargetId="figure@13.[194,1393,166,1988]" captionTargetPageId="13" captionText="FIGURE 7. Habitus drawing of Styraconyx robertoi sp. nov., female holotype." figureDoi="http://doi.org/10.5281/zenodo.3661960" httpUri="https://zenodo.org/record/3661960/files/figure.png" pageId="7" pageNumber="499">7</figureCitation>
|
||
), beginning just posterior to the pedestals bearing the primary clavae and extending to a position slightly anterior to the cirri E. Distinct dorso-caudal region without grid-like sculpture. Instead, two longitudinal ribs occur which originate on either side of the median line and curve slightly toward the lateral sides (
|
||
<figureCitation id="13095EB63219FF9AFDED676E46E6FC98" box="[586,654,792,818]" captionStart="FIGURE 7" captionStartId="13.[151,250,2007,2032]" captionTargetBox="[194,1379,179,1971]" captionTargetId="figure@13.[194,1393,166,1988]" captionTargetPageId="13" captionText="FIGURE 7. Habitus drawing of Styraconyx robertoi sp. nov., female holotype." figureDoi="http://doi.org/10.5281/zenodo.3661960" httpUri="https://zenodo.org/record/3661960/files/figure.png" pageId="7" pageNumber="499">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D42333219FF9AFF60674D4490FC6A" blockId="7.[151,1437,152,2024]" pageId="7" pageNumber="499">
|
||
The subterminal mouth cone is short and ends in a smooth, annular dome with a central mouth opening. The buccal tube (16 μm x 2 μm) has a pair of lateral apophyses on each side (
|
||
<figureCitation id="13095EB63219FF9AFC756729404FFCD3" box="[978,1063,863,889]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="7" pageNumber="499">Fig. 3E</figureCitation>
|
||
) and stylet supports with a distal swelling. Stylets are long (20 μm) with strong furcae. Pharyngeal bulb (15 μm) with three placoids of equal length (11 μm).
|
||
</paragraph>
|
||
<paragraph id="8B8D42333219FF9AFF6067BF412DFAEC" blockId="7.[151,1437,152,2024]" pageId="7" pageNumber="499">
|
||
Primary clavae, secondary clavae, and a full set of cephalic cirri are present. The primary clava (10 μm) is highly asymmetric and the form appears different depending on the angle in which it is observed (
|
||
<figureCitation id="13095EB63219FF9AFB44679B4136FBAD" box="[1251,1374,1005,1031]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4D,F</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFACD679A41F8FBAD" box="[1386,1424,1004,1031]" captionStart="FIGURE 5" captionStartId="11.[151,250,1239,1264]" captionTargetBox="[151,1435,167,1213]" captionTargetId="figure@11.[151,1436,166,1214]" captionTargetPageId="11" captionText="FIGURE 5. SEM-micrographs of Styraconyx robertoi sp. nov. A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle.Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows)." figureDoi="http://doi.org/10.5281/zenodo.3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="7" pageNumber="499">5A</figureCitation>
|
||
). Primary clava and lateral cirrus inserted on a common pedestal and surrounded basally by a short, common membrane. Primary clavae with van der Land’s body present at the base. The lateral cirrus (28 μm) is not segmented and has an asymmetrically expanded base, which is fused to the proximal end of the primary clava (
|
||
<figureCitation id="13095EB63219FF9AFB7A6021412BFBDB" box="[1245,1347,1111,1137]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4D</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFAF7602041E7FBDB" box="[1360,1423,1110,1137]" captionStart="FIGURE 5" captionStartId="11.[151,250,1239,1264]" captionTargetBox="[151,1435,167,1213]" captionTargetId="figure@11.[151,1436,166,1214]" captionTargetPageId="11" captionText="FIGURE 5. SEM-micrographs of Styraconyx robertoi sp. nov. A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle.Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows)." figureDoi="http://doi.org/10.5281/zenodo.3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="7" pageNumber="499">5A,B</figureCitation>
|
||
). Ventral lenticular secondary clava (12 μm) inserted antero-laterally to the external cirrus close to the anterior margin of the head (
|
||
<figureCitation id="13095EB63219FF9AFE8760E845E9FB12" box="[288,385,1182,1208]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="7" pageNumber="499">Figs. 3E</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFE2B60E845ADFB12" box="[396,453,1182,1208]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">4E,F</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFE7760E94588FB13" box="[464,480,1183,1209]" captionStart="FIGURE 7" captionStartId="13.[151,250,2007,2032]" captionTargetBox="[194,1379,179,1971]" captionTargetId="figure@13.[194,1393,166,1988]" captionTargetPageId="13" captionText="FIGURE 7. Habitus drawing of Styraconyx robertoi sp. nov., female holotype." figureDoi="http://doi.org/10.5281/zenodo.3661960" httpUri="https://zenodo.org/record/3661960/files/figure.png" pageId="7" pageNumber="499">7</figureCitation>
|
||
). The median cirrus (9 μm) is placed dorsally (
|
||
<figureCitation id="13095EB63219FF9AFC4060E84022FB12" box="[999,1098,1182,1208]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4A</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFBF260E84013FB13" box="[1109,1147,1182,1209]" captionStart="FIGURE 6" captionStartId="12.[151,250,1661,1686]" captionTargetBox="[152,1435,642,1641]" captionTargetId="figure@12.[151,1436,642,1641]" captionTargetPageId="12" captionText="FIGURE 6. SEM-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus. B. Internal cirrus. C. External cirrus. D. Sense organ of leg I. E. Sense organ of leg II. F. Sense organ of leg III." figureDoi="http://doi.org/10.5281/zenodo.3661958" httpUri="https://zenodo.org/record/3661958/files/figure.png" pageId="7" pageNumber="499">6A</figureCitation>
|
||
), well back from the ante- rior margin of the head. Internal cirri (13 μm) are dorsal (
|
||
<figureCitation id="13095EB63219FF9AFCB060B74712FB76" box="[791,890,1217,1244]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4B</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFC2160B747CFFB76" box="[902,935,1217,1244]" captionStart="FIGURE 6" captionStartId="12.[151,250,1661,1686]" captionTargetBox="[152,1435,642,1641]" captionTargetId="figure@12.[151,1436,642,1641]" captionTargetPageId="12" captionText="FIGURE 6. SEM-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus. B. Internal cirrus. C. External cirrus. D. Sense organ of leg I. E. Sense organ of leg II. F. Sense organ of leg III." figureDoi="http://doi.org/10.5281/zenodo.3661958" httpUri="https://zenodo.org/record/3661958/files/figure.png" pageId="7" pageNumber="499">6B</figureCitation>
|
||
) and more medial than the ventrally inserted external cirri (12 μm) (
|
||
<figureCitation id="13095EB63219FF9AFE346093459CFB55" box="[403,500,1253,1279]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4C</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFE586093464BFAAA" box="[511,547,1253,1280]" captionStart="FIGURE 6" captionStartId="12.[151,250,1661,1686]" captionTargetBox="[152,1435,642,1641]" captionTargetId="figure@12.[151,1436,642,1641]" captionTargetPageId="12" captionText="FIGURE 6. SEM-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus. B. Internal cirrus. C. External cirrus. D. Sense organ of leg I. E. Sense organ of leg II. F. Sense organ of leg III." figureDoi="http://doi.org/10.5281/zenodo.3661958" httpUri="https://zenodo.org/record/3661958/files/figure.png" pageId="7" pageNumber="499">6C</figureCitation>
|
||
). Except for the lateral cirri, all cephalic cirri (
|
||
<figureCitation id="13095EB63219FF9AFB876093400CFB55" box="[1056,1124,1253,1279]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs.4</figureCitation>
|
||
A–C; 6A–C) are two-parted (which can be difficult to recognize in LM) and each composed of a conical scapus and a tapering, flagellar part with a terminal pore. The flagellum is strictly internal and evident in all cephalic cirri except for the lateral cirri.
|
||
</paragraph>
|
||
<paragraph id="8B8D42333219FF9AFF6061394110F9CB" blockId="7.[151,1437,152,2024]" pageId="7" pageNumber="499">
|
||
Sense organs present on all legs and inserted coxally. Sense organs of leg pair I (9 μm) two-parted and similar to the cephalic cirri having a conical scapus and a tapering, flagellar part with a terminal pore (
|
||
<figureCitation id="13095EB63219FF9AFB6A6105415BFA27" box="[1229,1331,1395,1421]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4G</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFAE76105410DFA24" box="[1344,1381,1395,1422]" captionStart="FIGURE 6" captionStartId="12.[151,250,1661,1686]" captionTargetBox="[152,1435,642,1641]" captionTargetId="figure@12.[151,1436,642,1641]" captionTargetPageId="12" captionText="FIGURE 6. SEM-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus. B. Internal cirrus. C. External cirrus. D. Sense organ of leg I. E. Sense organ of leg II. F. Sense organ of leg III." figureDoi="http://doi.org/10.5281/zenodo.3661958" httpUri="https://zenodo.org/record/3661958/files/figure.png" pageId="7" pageNumber="499">6D</figureCitation>
|
||
). On leg pairs II and III the sense organs are simple, without any recognizable segmentation, and resemble the shape of curved spines (
|
||
<figureCitation id="13095EB63219FF9AFE9961CC45DEFA7F" box="[318,438,1466,1493]" captionStart="FIGURE 6" captionStartId="12.[151,250,1661,1686]" captionTargetBox="[152,1435,642,1641]" captionTargetId="figure@12.[151,1436,642,1641]" captionTargetPageId="12" captionText="FIGURE 6. SEM-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus. B. Internal cirrus. C. External cirrus. D. Sense organ of leg I. E. Sense organ of leg II. F. Sense organ of leg III." figureDoi="http://doi.org/10.5281/zenodo.3661958" httpUri="https://zenodo.org/record/3661958/files/figure.png" pageId="7" pageNumber="499">Figs. 6E,F</figureCitation>
|
||
). Sense organ of leg pair IV (
|
||
<figureCitation id="13095EB63219FF9AFD5D61CC4704FA7E" box="[762,876,1466,1492]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4H,I</figureCitation>
|
||
) consists of a smooth papilla (7 μm) with terminal spine (6 μm), which has a terminal pore. The papilla is asymmetrical, having the postero-lateral portion less curved than the antero-lateral portion. It is inserted with a broad base on a short pedestal and appears hemispherical rather than spherical. Dorso-lateral cirrus E (
|
||
<figureCitation id="13095EB63219FF9AFDEB625246A9F994" box="[588,705,1572,1598]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4H,I</figureCitation>
|
||
) inserted on a prominent pedestal. It consists of a short (3 μm), oversized cirrophore (
|
||
<figureCitation id="13095EB63219FF9AFE2A6231458FF9C8" box="[397,487,1607,1634]" captionStart="FIGURE 4" captionStartId="10.[151,250,1708,1733]" captionTargetBox="[151,1435,182,1681]" captionTargetId="figure@10.[151,1435,181,1682]" captionTargetPageId="10" captionText="FIGURE 4. DIC-micrographs of Styraconyx robertoi sp. nov. A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="7" pageNumber="499">Figs. 4I</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFE54623E466AF9C8" box="[499,514,1608,1634]" captionStart="FIGURE 7" captionStartId="13.[151,250,2007,2032]" captionTargetBox="[194,1379,179,1971]" captionTargetId="figure@13.[194,1393,166,1988]" captionTargetPageId="13" captionText="FIGURE 7. Habitus drawing of Styraconyx robertoi sp. nov., female holotype." figureDoi="http://doi.org/10.5281/zenodo.3661960" httpUri="https://zenodo.org/record/3661960/files/figure.png" pageId="7" pageNumber="499">7</figureCitation>
|
||
) and a long, tapering spine (23 μm) without evident accordion-pleated portion.
|
||
</paragraph>
|
||
<paragraph id="8B8D42333219FF9AFF60621D456FF8D7" blockId="7.[151,1437,152,2024]" pageId="7" pageNumber="499">
|
||
Each leg with evident coxa, femur, tibia and tarsus. Tarsus of all legs bearing four digits of different length with external digits being shorter than internal digits. Peduncles (
|
||
<figureCitation id="13095EB63219FF9AFC9262F847FDF903" box="[821,917,1678,1705]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="7" pageNumber="499">Figs. 3B</figureCitation>
|
||
) present only inside the external digits and are of the same
|
||
<typeStatus id="5489FC913219FF9AFEBC62C54523F967" box="[283,331,1715,1741]" pageId="7" pageNumber="499">type</typeStatus>
|
||
as
|
||
<taxonomicName id="4C3239B03219FF9AFEC962C4477BF966" authority="(Pollock, 1983)" baseAuthorityName="Pollock" baseAuthorityYear="1983" box="[366,787,1714,1740]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="7" pageNumber="499" phylum="Tardigrada" rank="species" species="craticulus">
|
||
<emphasis id="B9469E213219FF9AFEC962C44608F966" box="[366,608,1714,1740]" italics="true" pageId="7" pageNumber="499">Styraconyx craticulus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA33FC23219FF9AFDCA62C44762F966" author="Pollock, L. W." box="[621,778,1714,1740]" pageId="7" pageNumber="499" pagination="109 - 117" refId="ref8477" refString="Pollock, L. W. (1983) A closer look at some marine Heterotardigrada. II: The morphology and taxonomy of Bathyechiniscus with a description of B. craticulus n. sp. from the Caribbean. The Bulletin of Marine Science, 33, 109 - 117." type="journal article" year="1983">Pollock, 1983</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
reaching the claw bases (see
|
||
<bibRefCitation id="EFA33FC23219FF9AFBF162C441F8F966" author="Kristensen, R. M. & Higgins, R. P." box="[1110,1424,1714,1740]" pageId="7" pageNumber="499" pagination="1 - 40" refId="ref7808" refString="Kristensen, R. M. & Higgins, R. P. (1984) Revision of Styraconyx (Tardigrada: Halechiniscidae) with description of two new species from Disko Bay, West Greenland. Smithsonian Contributions to Zoology, 391, 1 - 40. https: // doi. org / 10.5479 / si. 00810282.391" type="journal article" year="1984">Kristensen & Higgins, 1984</bibRefCitation>
|
||
). They have a strongly curved basal portion and an enlarged medial portion with two small lateral points (
|
||
<figureCitation id="13095EB63219FF9AFA9362A341E7F95A" box="[1332,1423,1749,1776]" captionStart="FIGURE 8" captionStartId="14.[151,250,982,1007]" captionTargetBox="[151,1436,184,938]" captionTargetId="figure@14.[151,1436,184,938]" captionTargetPageId="14" captionText="FIGURE 8. Drawings of leg IV, Styraconyx robertoi sp. nov. A. Female. B. Male" figureDoi="http://doi.org/10.5281/zenodo.3661962" httpUri="https://zenodo.org/record/3661962/files/figure.png" pageId="7" pageNumber="499">Fig. 8A</figureCitation>
|
||
). The internal digits have thin, proximal pads. All digits bearing three-pointed claws (
|
||
<figureCitation id="13095EB63219FF9AFBE8628F40A3F8B9" box="[1103,1227,1785,1811]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="7" pageNumber="499">Figs. 3D,F</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFB7F628E4091F8B9" box="[1240,1273,1784,1811]" captionStart="FIGURE 5" captionStartId="11.[151,250,1239,1264]" captionTargetBox="[151,1435,167,1213]" captionTargetId="figure@11.[151,1436,166,1214]" captionTargetPageId="11" captionText="FIGURE 5. SEM-micrographs of Styraconyx robertoi sp. nov. A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle.Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows)." figureDoi="http://doi.org/10.5281/zenodo.3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="7" pageNumber="499">5C</figureCitation>
|
||
) with primary hook, secondary and accessory hook. The primary hook is the strongest. The secondary hook is thinner but slightly longer than primary hook, and accessory hook is the shortest. The tips of all three hooks are free of the claw sheath (
|
||
<figureCitation id="13095EB63219FF9AFF0763154491F8D7" box="[160,249,1891,1917]" captionStart="FIGURE 8" captionStartId="14.[151,250,982,1007]" captionTargetBox="[151,1436,184,938]" captionTargetId="figure@14.[151,1436,184,938]" captionTargetPageId="14" captionText="FIGURE 8. Drawings of leg IV, Styraconyx robertoi sp. nov. A. Female. B. Male" figureDoi="http://doi.org/10.5281/zenodo.3661962" httpUri="https://zenodo.org/record/3661962/files/figure.png" pageId="7" pageNumber="499">Fig. 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D42333219FF9AFF6063F14635F842" blockId="7.[151,1437,152,2024]" pageId="7" pageNumber="499">
|
||
Seminal receptacles are not evident. The gonopore (
|
||
<figureCitation id="13095EB63219FF9AFCB163F14713F80B" box="[790,891,1927,1953]" captionStart="FIGURE 2" captionStartId="8.[151,250,1939,1964]" captionTargetBox="[152,1435,409,1918]" captionTargetId="figure@8.[151,1435,409,1919]" captionTargetPageId="8" captionText="FIGURE 2. DIC-micrographs of Styraconyx robertoi sp. nov., holotypic female. A. Overview. B. Grid-like sculpture of the dorsal cuticle. C. Gonopore and anus." figureDoi="http://doi.org/10.5281/zenodo.3661950" httpUri="https://zenodo.org/record/3661950/files/figure.png" pageId="7" pageNumber="499">Figs. 2C</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFC2063F047C3F80B" box="[903,939,1926,1953]" captionStart="FIGURE 5" captionStartId="11.[151,250,1239,1264]" captionTargetBox="[151,1435,167,1213]" captionTargetId="figure@11.[151,1436,166,1214]" captionTargetPageId="11" captionText="FIGURE 5. SEM-micrographs of Styraconyx robertoi sp. nov. A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle.Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows)." figureDoi="http://doi.org/10.5281/zenodo.3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="7" pageNumber="499">5D</figureCitation>
|
||
) is relatively large (6 μm) and consists of a rosette of six cells situated close to the anus. The anus is an undulating slit and composed of two larger lateral lobes and a small anterior lobe (
|
||
<figureCitation id="13095EB63219FF9AFE1C63B84677F842" box="[443,543,1998,2024]" captionStart="FIGURE 2" captionStartId="8.[151,250,1939,1964]" captionTargetBox="[152,1435,409,1918]" captionTargetId="figure@8.[151,1435,409,1919]" captionTargetPageId="8" captionText="FIGURE 2. DIC-micrographs of Styraconyx robertoi sp. nov., holotypic female. A. Overview. B. Grid-like sculpture of the dorsal cuticle. C. Gonopore and anus." figureDoi="http://doi.org/10.5281/zenodo.3661950" httpUri="https://zenodo.org/record/3661950/files/figure.png" pageId="7" pageNumber="499">Figs. 2C</figureCitation>
|
||
;
|
||
<figureCitation id="13095EB63219FF9AFD8D63BB4627F842" box="[554,591,1997,2024]" captionStart="FIGURE 5" captionStartId="11.[151,250,1239,1264]" captionTargetBox="[151,1435,167,1213]" captionTargetId="figure@11.[151,1436,166,1214]" captionTargetPageId="11" captionText="FIGURE 5. SEM-micrographs of Styraconyx robertoi sp. nov. A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle.Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows)." figureDoi="http://doi.org/10.5281/zenodo.3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="7" pageNumber="499">5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D42333216FF95FF6064EF456DFEE9" blockId="8.[151,1437,152,395]" pageId="8" pageNumber="500">
|
||
<emphasis id="B9469E213216FF95FF6064EF457EFF19" bold="true" box="[199,278,153,179]" pageId="8" pageNumber="500">Males.</emphasis>
|
||
Only
|
||
<specimenCount id="9D3489BA3216FF95FEF964EF45DDFF19" box="[350,437,153,179]" pageId="8" pageNumber="500" type="male">2 males</specimenCount>
|
||
were found. The smaller one measures 112 μm in body length, the larger measures 154 μm. Both males display a peculiar combination of digit characters in having only two peduncles on each foot of leg pairs I–III (similar to the females) and four peduncles on each foot of leg pair IV. On leg pair IV the peduncles of internal digits are rod-shaped and extend from the base of the digits, through the proximal pads, to the claw bases (
|
||
<figureCitation id="13095EB63216FF95FF07655F449FFEE9" box="[160,247,297,323]" captionStart="FIGURE 8" captionStartId="14.[151,250,982,1007]" captionTargetBox="[151,1436,184,938]" captionTargetId="figure@14.[151,1436,184,938]" captionTargetPageId="14" captionText="FIGURE 8. Drawings of leg IV, Styraconyx robertoi sp. nov. A. Female. B. Male" figureDoi="http://doi.org/10.5281/zenodo.3661962" httpUri="https://zenodo.org/record/3661962/files/figure.png" pageId="8" pageNumber="500">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D42333216FF95FF60653B4696FE21" blockId="8.[151,1437,152,395]" pageId="8" pageNumber="500">The testes are not evident. The male gonopore is a simple, oval opening situated close to the anus. No other differences between males and females were observed.</paragraph>
|
||
<caption id="DF4D12BB3216FF95FF3063E5467DF865" ID-DOI="http://doi.org/10.5281/zenodo.3661950" ID-Zenodo-Dep="3661950" httpUri="https://zenodo.org/record/3661950/files/figure.png" pageId="8" pageNumber="500" startId="8.[151,250,1939,1964]" targetBox="[152,1435,409,1918]" targetPageId="8">
|
||
<paragraph id="8B8D42333216FF95FF3063E5467DF865" blockId="8.[151,1436,1939,2000]" pageId="8" pageNumber="500">
|
||
<emphasis id="B9469E213216FF95FF3063E5457DF806" bold="true" box="[151,277,1939,1964]" pageId="8" pageNumber="500">FIGURE 2.</emphasis>
|
||
DIC-micrographs of
|
||
<taxonomicName id="4C3239B03216FF95FE5163E246A8F806" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[502,704,1940,1964]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="500" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213216FF95FE5163E246A8F806" box="[502,704,1940,1964]" italics="true" pageId="8" pageNumber="500">Styraconyx robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213216FF95FD6E63E24773F806" bold="true" box="[713,795,1940,1964]" pageId="8" pageNumber="500">
|
||
<taxonomicNameLabel id="A275235A3216FF95FD6E63E24773F806" box="[713,795,1940,1964]" pageId="8" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotypic female. A. Overview. B. Grid-like sculpture of the dorsal cuticle. C. Gonopore and anus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D12BB3217FF94FF30620E449EF88B" ID-DOI="http://doi.org/10.5281/zenodo.3661952" ID-Zenodo-Dep="3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="9" pageNumber="501" startId="9.[151,250,1656,1681]" targetBox="[151,1435,175,1636]" targetPageId="9">
|
||
<paragraph id="8B8D42333217FF94FF30620E449EF88B" blockId="9.[151,1437,1656,1825]" pageId="9" pageNumber="501">
|
||
<emphasis id="B9469E213217FF94FF30620E457BF93B" bold="true" box="[151,275,1656,1681]" pageId="9" pageNumber="501">FIGURE 3.</emphasis>
|
||
DIC-micrographs of
|
||
<taxonomicName id="4C3239B03217FF94FE4A620F46DEF93B" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[493,694,1657,1681]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="501" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213217FF94FE4A620F46DEF93B" box="[493,694,1657,1681]" italics="true" pageId="9" pageNumber="501">Styraconyx robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213217FF94FD1B620F4765F93B" bold="true" box="[700,781,1657,1681]" pageId="9" pageNumber="501">
|
||
<taxonomicNameLabel id="A275235A3217FF94FD1B620F4765F93B" box="[700,781,1657,1681]" pageId="9" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes. Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D42333217FF94FF60633941E2F853" blockId="9.[151,1436,1870,2042]" pageId="9" pageNumber="501">
|
||
<emphasis id="B9469E213217FF94FF606339454CF8C3" bold="true" box="[199,292,1871,1897]" pageId="9" pageNumber="501">Larvae.</emphasis>
|
||
Three larvae were found, all with four claws on each foot and with body lengths of 95 μm, 112 μm and 114 μm. The specimens display larval characters in the shape and size of the primary clava and in the overall body form (
|
||
<figureCitation id="13095EB63217FF94FF7963E1455FF81B" box="[222,311,1943,1969]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="9" pageNumber="501">Fig. 3C</figureCitation>
|
||
). The primary clavae are asymmetric but shorter (6–8 μm) and less slender than in adults. The head is not well-differentiated from the trunk and the frontal margin appears blunt, whereas in adults it is protruded (
|
||
<figureCitation id="13095EB63217FF94FAC863CD44D5F853" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="9" pageNumber="501">Fig. 3A</figureCitation>
|
||
). One of the specimens has four peduncles on the digits of leg pair IV and so is considered to be a male larva.
|
||
</paragraph>
|
||
<caption id="DF4D12BB3214FF97FF3062DA46E3F8FF" ID-DOI="http://doi.org/10.5281/zenodo.3661954" ID-Zenodo-Dep="3661954" httpUri="https://zenodo.org/record/3661954/files/figure.png" pageId="10" pageNumber="502" startId="10.[151,250,1708,1733]" targetBox="[151,1435,182,1681]" targetPageId="10">
|
||
<paragraph id="8B8D42333214FF97FF3062DA46E3F8FF" blockId="10.[151,1437,1708,1877]" pageId="10" pageNumber="502">
|
||
<emphasis id="B9469E213214FF97FF3062DA457BF96F" bold="true" box="[151,275,1708,1733]" pageId="10" pageNumber="502">FIGURE 4.</emphasis>
|
||
DIC-micrographs of
|
||
<taxonomicName id="4C3239B03214FF97FE4962DB46DFF96F" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[494,695,1709,1733]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="502" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213214FF97FE4962DB46DFF96F" box="[494,695,1709,1733]" italics="true" pageId="10" pageNumber="502">Styraconyx robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213214FF97FD1A62DB4766F96F" bold="true" box="[701,782,1709,1733]" pageId="10" pageNumber="502">
|
||
<taxonomicNameLabel id="A275235A3214FF97FD1A62DB4766F96F" box="[701,782,1709,1733]" pageId="10" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Median cirrus, holotypic female. B. Internal cirrus, holotypic female. C. External cirrus, holotypic female. D. Primary clava and lateral cirrus with expanded base (arrow), female paratype. E. Ventro-lateral view of secondary clava (arrow), female paratype. F. Dorso-lateral view of secondary clava (arrow), female paratype. G. Sense organ of leg I, female paratype. H. Cirrus E and leg IV sense organ, female paratype. I. Cirrus E and leg IV sense organ from another angle, female paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D12BB3215FF96FF3060A14661FA92" ID-DOI="http://doi.org/10.5281/zenodo.3661956" ID-Zenodo-Dep="3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="11" pageNumber="503" startId="11.[151,250,1239,1264]" targetBox="[151,1435,167,1213]" targetPageId="11">
|
||
<paragraph id="8B8D42333215FF96FF3060A14661FA92" blockId="11.[151,1437,1239,1336]" pageId="11" pageNumber="503">
|
||
<emphasis id="B9469E213215FF96FF3060A1457BFB5A" bold="true" box="[151,275,1239,1264]" pageId="11" pageNumber="503">FIGURE 5.</emphasis>
|
||
SEM-micrographs of
|
||
<taxonomicName id="4C3239B03215FF96FE5F60AE46AAFB5A" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[504,706,1240,1264]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213215FF96FE5F60AE46AAFB5A" box="[504,706,1240,1264]" italics="true" pageId="11" pageNumber="503">Styraconyx robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213215FF96FD6E60AE4772FB5A" bold="true" box="[713,794,1240,1264]" pageId="11" pageNumber="503">
|
||
<taxonomicNameLabel id="A275235A3215FF96FD6E60AE4772FB5A" box="[713,794,1240,1264]" pageId="11" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle. Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D42333215FF96FF60611541E5F9DA" blockId="11.[151,1437,1379,2030]" pageId="11" pageNumber="503">
|
||
<emphasis id="B9469E213215FF96FF6061154647FAD4" bold="true" box="[199,559,1379,1406]" pageId="11" pageNumber="503">Scanning electron microscopy.</emphasis>
|
||
The SEM observations of a single specimen confirmed most of the diagnostic features of this species and aided to a better understanding of their true nature. In LM, the claws, and especially their hooks, can look very different depending on their orientation in the preparation (
|
||
<figureCitation id="13095EB63215FF96FB8761DF40F3FA69" box="[1056,1179,1449,1475]" captionStart="FIGURE 3" captionStartId="9.[151,250,1656,1681]" captionTargetBox="[151,1435,175,1636]" captionTargetId="figure@9.[151,1436,174,1637]" captionTargetPageId="9" captionText="FIGURE 3. DIC-micrographs of Styraconyx robertoi sp. nov. A. Overview, female paratype with evident eyes and large ovary occupying more than half the body. B. Peduncles of external digits on leg IV, holotypic female. C. Overview, 4-clawed larva. D. Claws of digits on leg IV, holotypic female. E. Female paratype showing buccal apparatus, secondary clava and eyes.Apophyses (apo), eyespot (eye), placoid (pl), secondary clava (sc), stylet support (ss), stylet (st). F. Claws and peduncles on leg III, female paratype." figureDoi="http://doi.org/10.5281/zenodo.3661952" httpUri="https://zenodo.org/record/3661952/files/figure.png" pageId="11" pageNumber="503">Figs. 3D,F</figureCitation>
|
||
), whereas the three-dimensional SEM image leaves no mistake of interpretation. This applies also to the sense organs where segmentation and terminal pores can be difficult to recognise by LM, but clearly evident with SEM. In this case where both the primary clava and the base of the lateral cirrus are asymmetric, their true shape is only recognizable with SEM (
|
||
<figureCitation id="13095EB63215FF96FAC3626744BEF9E4" captionStart="FIGURE 5" captionStartId="11.[151,250,1239,1264]" captionTargetBox="[151,1435,167,1213]" captionTargetId="figure@11.[151,1436,166,1214]" captionTargetPageId="11" captionText="FIGURE 5. SEM-micrographs of Styraconyx robertoi sp. nov. A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle.Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows)." figureDoi="http://doi.org/10.5281/zenodo.3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="11" pageNumber="503">Figs. 5A,B</figureCitation>
|
||
). Not visible in LM but revealed by SEM is a pair of large pores (
|
||
<figureCitation id="13095EB63215FF96FC3A6245479AF9E4" box="[925,1010,1587,1614]" captionStart="FIGURE 5" captionStartId="11.[151,250,1239,1264]" captionTargetBox="[151,1435,167,1213]" captionTargetId="figure@11.[151,1436,166,1214]" captionTargetPageId="11" captionText="FIGURE 5. SEM-micrographs of Styraconyx robertoi sp. nov. A. Primary clava and lateral cirrus with fused base (arrow). B. Grid-like sculpture of the dorsal cuticle.Arrow indicates the asymmetric base of lateral cirrus. C. Digits and claws of leg III. D. Gonopore, anus, and pores (arrows)." figureDoi="http://doi.org/10.5281/zenodo.3661956" httpUri="https://zenodo.org/record/3661956/files/figure.png" pageId="11" pageNumber="503">Fig. 5D</figureCitation>
|
||
) posterior to the gonopore. While their function is unknown, the position close to the gonopore suggests that they are openings of the seminal receptacles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32811B83215FF91FF60620E40D6FE62" lastPageId="12" lastPageNumber="504" pageId="11" pageNumber="503" type="diagnosis">
|
||
<paragraph id="8B8D42333215FF91FF60620E40D6FE62" blockId="11.[151,1437,1379,2030]" lastBlockId="12.[151,1436,153,630]" lastPageId="12" lastPageNumber="504" pageId="11" pageNumber="503">
|
||
<emphasis id="B9469E213215FF96FF60620E45A5F939" bold="true" box="[199,461,1656,1683]" pageId="11" pageNumber="503">Differential diagnosis.</emphasis>
|
||
The 14 previously described taxa (13 species and 1 subspecies) of the genus
|
||
<taxonomicName id="4C3239B03215FF96FA87620F41F4F939" authorityName="Thulin" authorityYear="1942" box="[1312,1436,1657,1683]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B9469E213215FF96FA87620F41F4F939" box="[1312,1436,1657,1683]" italics="true" pageId="11" pageNumber="503">Styraconyx</emphasis>
|
||
</taxonomicName>
|
||
can generally be divided into two major groups (
|
||
<bibRefCitation id="EFA33FC23215FF96FD1F62ED4798F91C" author="Kristensen, R. M. & Higgins, R. P." box="[696,1008,1691,1718]" pageId="11" pageNumber="503" pagination="1 - 40" refId="ref7808" refString="Kristensen, R. M. & Higgins, R. P. (1984) Revision of Styraconyx (Tardigrada: Halechiniscidae) with description of two new species from Disko Bay, West Greenland. Smithsonian Contributions to Zoology, 391, 1 - 40. https: // doi. org / 10.5479 / si. 00810282.391" type="journal article" year="1984">Kristensen & Higgins 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA33FC23215FF96FC5B62ED412FF91C" author="D'Addabbo Gallo, M. & Morone De Lucia, M. R. & de Zio Grimaldi, S." box="[1020,1351,1691,1718]" pageId="11" pageNumber="503" pagination="17 - 33" refId="ref7316" refString="D'Addabbo Gallo, M., Morone De Lucia, M. R. & de Zio Grimaldi, S. (1989) Two new species of the genus Styraconyx (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine, 30, 17 - 33." type="journal article" year="1989">
|
||
D’Addabbo Gallo
|
||
<emphasis id="B9469E213215FF96FB6A62EA416DF91F" box="[1229,1285,1691,1717]" italics="true" pageId="11" pageNumber="503">et al.</emphasis>
|
||
1989
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA33FC23215FF96FAF462ED454BF972" author="Chang, C. Y. & Rho, H. S." pageId="11" pageNumber="503" pagination="323 - 331" refId="ref7209" refString="Chang, C. Y. & Rho, H. S. (1998) Three new tardigrade species associated with barnacles from the Thai coast of Andaman Sea. Korean Journal of Biological Sciences, 2, 323 - 331. https: // doi. org / 10.1080 / 12265071.1998.9647426" type="journal article" year="1998">Chang & Rho 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA33FC23215FF96FE8962C84596F972" author="Bartels, P. J. & Fontoura, P. & Nelson, D. R." box="[302,510,1725,1753]" pageId="11" pageNumber="503" pagination="389 - 402" refId="ref6997" refString="Bartels, P. J., Fontoura, P. & Nelson, D. R. (2015) New records of marine tardigrades from Moorea, French Polynesia, with the description of Styraconyx turbinarium sp. nov. (Arthrotardigrada, Halechiniscidae). Zootaxa, 3955 (3), 389 - 402. https: // doi. org / 10.11646 / zootaxa. 3955.3.6" type="journal article" year="2015">
|
||
Bartels
|
||
<emphasis id="B9469E213215FF96FE2362C945DCF972" box="[388,436,1726,1752]" italics="true" pageId="11" pageNumber="503">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
); the
|
||
<taxonomicName id="4C3239B03215FF96FD9E62C946C8F972" authorityName="Kristensen" authorityYear="1977" box="[569,672,1726,1753]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="hallasi">
|
||
<emphasis id="B9469E213215FF96FD9E62C946C8F972" box="[569,672,1726,1753]" italics="true" pageId="11" pageNumber="503">S. hallasi</emphasis>
|
||
</taxonomicName>
|
||
group and the
|
||
<taxonomicName id="4C3239B03215FF96FCE262C947D5F972" authorityName="Thulin" authorityYear="1942" box="[837,957,1727,1753]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="sargassi">
|
||
<emphasis id="B9469E213215FF96FCE262C947D5F972" box="[837,957,1727,1753]" italics="true" pageId="11" pageNumber="503">S. sargassi</emphasis>
|
||
</taxonomicName>
|
||
group. Species in the
|
||
<taxonomicName id="4C3239B03215FF96FB1562C94171F972" authorityName="Kristensen" authorityYear="1977" box="[1202,1305,1726,1753]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="hallasi">
|
||
<emphasis id="B9469E213215FF96FB1562C94171F972" box="[1202,1305,1726,1753]" italics="true" pageId="11" pageNumber="503">S. hallasi</emphasis>
|
||
</taxonomicName>
|
||
group have peduncles on two digits (the external) of each leg and claws with reduced accessory hooks, while species in the
|
||
<taxonomicName id="4C3239B03215FF96FA206297449BF8B7" authorityName="Thulin" authorityYear="1942" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="sargassi">
|
||
<emphasis id="B9469E213215FF96FA206297449BF8B7" italics="true" pageId="11" pageNumber="503">S. sargassi</emphasis>
|
||
</taxonomicName>
|
||
group (including the
|
||
<typeStatus id="5489FC913215FF96FE406372467FF8B4" box="[487,535,1796,1822]" pageId="11" pageNumber="503">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C3239B03215FF96FDD0637247C0F8B4" authority="Thulin, 1942" authorityName="Thulin" authorityYear="1942" box="[631,936,1795,1822]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="haplocerus">
|
||
<emphasis id="B9469E213215FF96FDD063724767F8B7" box="[631,783,1795,1822]" italics="true" pageId="11" pageNumber="503">S. haplocerus</emphasis>
|
||
<bibRefCitation id="EFA33FC23215FF96FCB1637547C0F8B4" author="Thulin, G." box="[790,936,1795,1822]" pageId="11" pageNumber="503" pagination="1 - 10" refId="ref9327" refString="Thulin, G. (1942) Ein neuer mariner Tardigrad. Meddelanden fran Goteborgs Musei Zoologiska Avdeling, 99, 1 - 10." type="journal article" year="1942">Thulin, 1942</bibRefCitation>
|
||
</taxonomicName>
|
||
) have peduncles on all four digits and claws with three hooks of nearly equal size. While females of
|
||
<taxonomicName id="4C3239B03215FF96FCB06351479FF8EA" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[791,1015,1830,1857]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213215FF96FCB06351479FF8EA" box="[791,1015,1830,1857]" italics="true" pageId="11" pageNumber="503">Styraconyx robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213215FF96FC5863504032F8EA" bold="true" box="[1023,1114,1830,1856]" pageId="11" pageNumber="503">
|
||
<taxonomicNameLabel id="A275235A3215FF96FC5863504032F8EA" box="[1023,1114,1830,1856]" pageId="11" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
should be placed within the
|
||
<taxonomicName id="4C3239B03215FF96FF30633F4568F8C9" authorityName="Kristensen" authorityYear="1977" box="[151,256,1865,1891]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="hallasi">
|
||
<emphasis id="B9469E213215FF96FF30633F4568F8C9" box="[151,256,1865,1891]" italics="true" pageId="11" pageNumber="503">S. hallasi</emphasis>
|
||
</taxonomicName>
|
||
group due to the presence of peduncles only on the external digits of legs I–IV and claws with reduced accessory hooks, the unusual digit configuration of the males,
|
||
<emphasis id="B9469E213215FF96FCE8631A4719F82F" box="[847,881,1900,1925]" italics="true" pageId="11" pageNumber="503">i.e.</emphasis>
|
||
peduncles only on the external digits of legs I–III and peduncles on all four digits of leg IV, complicates the assignment of the new species to either of the two groups. Within the
|
||
<taxonomicName id="4C3239B03215FF96FEB363C74514F861" authorityName="Kristensen" authorityYear="1977" box="[276,380,1969,1995]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="hallasi">
|
||
<emphasis id="B9469E213215FF96FEB363C74514F861" box="[276,380,1969,1995]" italics="true" pageId="11" pageNumber="503">S. hallasi</emphasis>
|
||
</taxonomicName>
|
||
group only
|
||
<emphasis id="B9469E213215FF96FDA263C74712F861" box="[517,890,1969,1995]" italics="true" pageId="11" pageNumber="503">
|
||
<taxonomicName id="4C3239B03215FF96FDA263C7476AF861" authorityName="D'Addabbo Gallo, de Zio Grimaldi, Morone De Lucia & Troccoli" authorityYear="1992" box="[517,770,1969,1995]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" sensu="lato" species="kristenseni">Styraconyx kristenseni</taxonomicName>
|
||
<taxonomicNameLabel id="A275235A3215FF96FCAE63C44712F861" box="[777,890,1969,1995]" pageId="11" pageNumber="503" sensu="lato">sensu lato</taxonomicNameLabel>
|
||
</emphasis>
|
||
<bibRefCitation id="EFA33FC23215FF96FC2663C740EEF861" author="Renaud-Mornant, J." box="[897,1158,1969,1995]" pageId="11" pageNumber="503" pagination="799 - 813" refId="ref8668" refString="Renaud-Mornant, J. (1981) Tardigrades marins (Arthrotardigrada) du Pacifique Sud. Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e, Section A, 3, 799 - 813." type="journal article" year="1981">Renaud-Mornant, 1981</bibRefCitation>
|
||
has asymmetric primary clavae that are similar to those of the new species. However,
|
||
<taxonomicName id="4C3239B03215FF96FCE263A247B2F847" authorityName="D'Addabbo Gallo, de Zio Grimaldi, Morone De Lucia & Troccoli" authorityYear="1992" box="[837,986,2003,2030]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="kristenseni">
|
||
<emphasis id="B9469E213215FF96FCE263A247B2F847" box="[837,986,2003,2030]" italics="true" pageId="11" pageNumber="503">S. kristenseni</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3239B03215FF96FBC863A2408EF847" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[1135,1254,2003,2030]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="503" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213215FF96FBC863A2408EF847" box="[1135,1254,2003,2030]" italics="true" pageId="11" pageNumber="503">S. robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213215FF96FB4A63A2412EF844" bold="true" box="[1261,1350,2004,2030]" pageId="11" pageNumber="503">
|
||
<taxonomicNameLabel id="A275235A3215FF96FB4A63A2412EF844" box="[1261,1350,2004,2030]" pageId="11" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in hav- ing equal sized claw hooks, slender papilla on leg IV with evident cirrophore, and by lacking secondary clavae and the characteristic grid-like dorsal sculpture. Interestingly, the grid-like sculpture of the dorsal cuticle of
|
||
<taxonomicName id="4C3239B03212FF91FA8264CA41F4FF7C" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[1317,1436,188,214]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213212FF91FA8264CA41F4FF7C" box="[1317,1436,188,214]" italics="true" pageId="12" pageNumber="504">S. robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213212FF91FF3064A8449AFF52" bold="true" box="[151,242,222,248]" pageId="12" pageNumber="504">
|
||
<taxonomicNameLabel id="A275235A3212FF91FF3064A8449AFF52" box="[151,242,222,248]" pageId="12" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
bears an immediate resemblance to that of
|
||
<taxonomicName id="4C3239B03212FF91FD4464A94719FF52" baseAuthorityName="Pollock" baseAuthorityYear="1983" box="[739,881,222,249]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="craticulus">
|
||
<emphasis id="B9469E213212FF91FD4464A94699FF53" box="[739,753,223,249]" italics="true" pageId="12" pageNumber="504">S</emphasis>
|
||
.
|
||
<emphasis id="B9469E213212FF91FCA664A84719FF52" box="[769,881,222,248]" italics="true" pageId="12" pageNumber="504">craticulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3239B03212FF91FC0D64A94136FF52" authority="Chang & Rho, 1998" authorityName="Chang & Rho" authorityYear="1998" box="[938,1374,222,249]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="craticuliformis">
|
||
<emphasis id="B9469E213212FF91FC0D64A947D0FF53" box="[938,952,223,249]" italics="true" pageId="12" pageNumber="504">S</emphasis>
|
||
.
|
||
<emphasis id="B9469E213212FF91FC6064A84005FF52" box="[967,1133,222,248]" italics="true" pageId="12" pageNumber="504">craticuliformis</emphasis>
|
||
<bibRefCitation id="EFA33FC23212FF91FBD164A84136FF52" author="Chang, C. Y. & Rho, H. S." box="[1142,1374,222,248]" pageId="12" pageNumber="504" pagination="323 - 331" refId="ref7209" refString="Chang, C. Y. & Rho, H. S. (1998) Three new tardigrade species associated with barnacles from the Thai coast of Andaman Sea. Korean Journal of Biological Sciences, 2, 323 - 331. https: // doi. org / 10.1080 / 12265071.1998.9647426" type="journal article" year="1998">Chang & Rho, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
, both of which belong to the
|
||
<taxonomicName id="4C3239B03212FF91FE0565774674FEB1" authorityName="Thulin" authorityYear="1942" box="[418,540,257,283]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="sargassi">
|
||
<emphasis id="B9469E213212FF91FE0565774674FEB1" box="[418,540,257,283]" italics="true" pageId="12" pageNumber="504">S. sargassi</emphasis>
|
||
</taxonomicName>
|
||
group. This sculpture is the main decisive character differentiating these three species from all other
|
||
<taxonomicName id="4C3239B03212FF91FE3265524679FE94" authorityName="Thulin" authorityYear="1942" box="[405,529,292,318]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B9469E213212FF91FE3265524679FE94" box="[405,529,292,318]" italics="true" pageId="12" pageNumber="504">Styraconyx</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C3239B03212FF91FD866552468CFE94" authorityName="Chang & Rho" authorityYear="1998" box="[545,740,292,318]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="craticuliformis">
|
||
<emphasis id="B9469E213212FF91FD866552468CFE94" box="[545,740,292,318]" italics="true" pageId="12" pageNumber="504">S. craticuliformis</emphasis>
|
||
</taxonomicName>
|
||
and the new species both differ from
|
||
<taxonomicName id="4C3239B03212FF91FB316552414CFE94" baseAuthorityName="Pollock" baseAuthorityYear="1983" box="[1174,1316,292,318]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="craticulus">
|
||
<emphasis id="B9469E213212FF91FB31655240CCFE94" box="[1174,1188,292,318]" italics="true" pageId="12" pageNumber="504">S</emphasis>
|
||
.
|
||
<emphasis id="B9469E213212FF91FB136552414CFE94" box="[1204,1316,292,318]" italics="true" pageId="12" pageNumber="504">craticulus</emphasis>
|
||
</taxonomicName>
|
||
by having reduced accessory hooks, a thin common membrane at the base of the primary clavae and lateral cirri, asymmetric primary clavae, and a smooth papilla on leg IV. In summary,
|
||
<taxonomicName id="4C3239B03212FF91FCD5651F4039FE29" authority="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen, 2020" authorityName="Pérez-Pech & Jesús-Navarrate & Demilio & Anguas-Escalante & Hansen" authorityYear="2020" box="[882,1105,361,387]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="robertoi" status="sp. nov.">
|
||
<emphasis id="B9469E213212FF91FCD5651F4039FE29" box="[882,1105,361,387]" italics="true" pageId="12" pageNumber="504">Styraconyx robertoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9469E213212FF91FBFE651F40DBFE29" bold="true" box="[1113,1203,361,387]" pageId="12" pageNumber="504">
|
||
<taxonomicNameLabel id="A275235A3212FF91FBFE651F40DBFE29" box="[1113,1203,361,387]" pageId="12" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="4C3239B03212FF91FA20651F4555FE0C" authorityName="Chang & Rho" authorityYear="1998" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="craticuliformis">
|
||
<emphasis id="B9469E213212FF91FA20651F41FDFE29" box="[1415,1429,361,387]" italics="true" pageId="12" pageNumber="504">S</emphasis>
|
||
.
|
||
<emphasis id="B9469E213212FF91FF3065FA4555FE0C" box="[151,317,396,422]" italics="true" pageId="12" pageNumber="504">craticuliformis</emphasis>
|
||
</taxonomicName>
|
||
but can be readily distinguished from it by the number of peduncles, the presence of distinct secondary clavae, and the shape of the papilla on leg IV, which in
|
||
<taxonomicName id="4C3239B03212FF91FC8965D94798FE62" authorityName="Chang & Rho" authorityYear="1998" box="[814,1008,430,457]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="504" phylum="Tardigrada" rank="species" species="craticuliformis">
|
||
<emphasis id="B9469E213212FF91FC8965D94754FE63" box="[814,828,431,457]" italics="true" pageId="12" pageNumber="504">S</emphasis>
|
||
.
|
||
<emphasis id="B9469E213212FF91FCED65D84798FE62" box="[842,1008,430,456]" italics="true" pageId="12" pageNumber="504">craticuliformis</emphasis>
|
||
</taxonomicName>
|
||
is more spherical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32811B83212FF91FF6065A74142FD9A" pageId="12" pageNumber="504" type="materials_examined">
|
||
<paragraph id="8B8D42333212FF91FF6065A74142FD9A" blockId="12.[151,1436,153,630]" pageId="12" pageNumber="504">
|
||
<emphasis id="B9469E213212FF91FF6065A745C1FE41" bold="true" box="[199,425,464,491]" pageId="12" pageNumber="504">Type Repositories:</emphasis>
|
||
<materialsCitation id="3B5A486E3212FF91FE1465A74142FD9A" ID-GBIF-Occurrence="2572158539" collectionCode="NHMD, ECOSUR" country="Mexico" location="Quintana Roo" pageId="12" pageNumber="504" specimenCount="7" specimenCount-female="2" specimenCount-larva="3" specimenCount-male="2" stateProvince="Quintana Roo" typeStatus="holotype">
|
||
<typeStatus id="5489FC913212FF91FE1465A74673FE41" box="[435,539,465,491]" pageId="12" pageNumber="504" type="holotype">Holotype</typeStatus>
|
||
(NHMD-633434) and seven
|
||
<typeStatus id="5489FC913212FF91FCD665A747B4FE41" box="[881,988,465,491]" pageId="12" pageNumber="504" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D3489BA3212FF91FC4965A74036FE41" box="[1006,1118,465,491]" pageId="12" pageNumber="504" type="female">2 females</specimenCount>
|
||
NHMD-633435–633436,
|
||
<specimenCount id="9D3489BA3212FF91FA2965A744B0FDA7" pageId="12" pageNumber="504" type="male">2 males</specimenCount>
|
||
NHMD-633437–633438,
|
||
<specimenCount id="9D3489BA3212FF91FDA565824633FDA7" box="[514,603,499,526]" pageId="12" pageNumber="504" type="larva">3 larvae</specimenCount>
|
||
NHMD-633439–633441) are deposited in the Tardigrada collection of the
|
||
<collectionCode id="ED23DAF63212FF91FF3066614487FD9B" box="[151,239,535,561]" pageId="12" pageNumber="504">NHMD</collectionCode>
|
||
, Copenhagen. Six
|
||
<typeStatus id="5489FC913212FF91FE6666614644FD9B" box="[449,556,535,561]" pageId="12" pageNumber="504" type="paratype">paratypes</typeStatus>
|
||
are deposited in the collection at
|
||
<collectionCode id="ED23DAF63212FF91FC016660407FFD9A" box="[934,1047,534,560]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="12" pageNumber="504" type="Museum">ECOSUR</collectionCode>
|
||
,
|
||
<collectingRegion id="49F68CD13212FF91FB85666040A9FD9B" box="[1058,1217,534,561]" country="Mexico" name="Quintana Roo" pageId="12" pageNumber="504">Quintana Roo</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32502A33212FF91FB6B6660414DFD9A" box="[1228,1317,534,560]" name="Mexico" pageId="12" pageNumber="504">Mexico</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32811B83212FF91FF60664F408FFDDF" pageId="12" pageNumber="504" type="etymology">
|
||
<paragraph id="8B8D42333212FF91FF60664F408FFDDF" blockId="12.[151,1436,153,630]" pageId="12" pageNumber="504">
|
||
<emphasis id="B9469E213212FF91FF60664F453AFDF9" bold="true" box="[199,338,569,595]" pageId="12" pageNumber="504">Etymology:</emphasis>
|
||
The species is dedicated to our valued friend, Dr. Roberto Guidetti of the University of Modena and Reggio Emilia,
|
||
<collectingCountry id="F32502A33212FF91FED0662D45C2FDDF" box="[375,426,603,629]" name="Italy" pageId="12" pageNumber="504">Italy</collectingCountry>
|
||
in recognition of his remarkable contributions to the study of tardigrades.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |