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<document id="DD41792174B254BC9AEA87E8627F9237" ID-DOI="10.11646/zootaxa.4021.2.3" ID-GBIF-Dataset="47fe93e6-3726-46db-a891-01d9ab7b70f3" ID-ISSN="1175-5326" ID-Zenodo-Dep="240666" ID-ZooBank="F12128F6-C542-4FBE-9ACF-414C96D93C6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461274976068" checkinUser="plazi" docAuthor="Guzman, Guillermo L. &amp; Sellanes, Javier" docDate="2015" docId="039B3B0CFFEBFF82FF48F94AE7E899D5" docLanguage="en" docName="zt04021p306.pdf" docOrigin="Zootaxa 4021 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Munidopsis subsquamosa Henderson 1888" docType="treatment" docVersion="9" lastPageNumber="298" masterDocId="FFA24374FFE6FF92FFDFFFBEE4289A38" masterDocTitle="A review of the Munidopsidae Ortmann, 1898 (Decapoda, Galatheoidea) in Chilean waters, including new records for the Southeastern Pacific" masterLastPageNumber="306" masterPageNumber="282" pageNumber="295" updateTime="1698612466786" updateUser="plazi">
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<mods:title id="0CADDA0B9D25A85FBC7BD6E206662C9E">A review of the Munidopsidae Ortmann, 1898 (Decapoda, Galatheoidea) in Chilean waters, including new records for the Southeastern Pacific</mods:title>
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<heading id="D0C53D76FFEBFF9FFF48F94AE6809D36" bold="true" box="[151,680,1780,1806]" fontSize="11" level="1" pageId="13" pageNumber="295" reason="1">
<taxonomicName id="4C32F199FFEBFF9FFF48F94AE6809D36" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[151,680,1780,1806]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="295" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFEBFF9FFF48F94AE6809D36" bold="true" box="[151,680,1780,1806]" pageId="13" pageNumber="295">
<emphasis id="B9465608FFEBFF9FFF48F94AE5E79D36" bold="true" box="[151,463,1780,1806]" italics="true" pageId="13" pageNumber="295">Munidopsis subsquamosa</emphasis>
<bibRefCitation id="EFA3F7EBFFEBFF9FFE0AF94AE6809D36" author="Henderson" box="[469,680,1780,1806]" pageId="13" pageNumber="295" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 27, 1 - 221, pls. 1 - 221." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
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<paragraph id="8B8D8A1AFFEBFF9FFF48F8A8E4C39D17" blockId="13.[151,680,1780,1839]" box="[151,235,1814,1839]" pageId="13" pageNumber="295">
(
<figureCitation id="1309969FFFEBFF9FFF40F8A8E4CB9D17" box="[159,227,1814,1839]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="13" pageNumber="295">Fig. 9</figureCitation>
)
</paragraph>
<paragraph id="8B8D8A1AFFEBFF9FFF48F8E0E55D9D4C" blockId="13.[151,1436,1886,2031]" box="[151,373,1886,1908]" pageId="13" pageNumber="295">Restricted synonymy:</paragraph>
</subSubSection>
<subSubSection id="C328D991FFEBFF9FFF48F8C3E7459DD7" pageId="13" pageNumber="295" type="reference_group">
<paragraph id="8B8D8A1AFFEBFF9FFF48F8C3E7849DAB" blockId="13.[151,1436,1886,2031]" box="[151,940,1917,1939]" pageId="13" pageNumber="295">
<treatmentCitationGroup id="AB22AD34FFEBFF9FFF48F8C3E7849DAB" box="[151,940,1917,1939]" pageId="13" pageNumber="295">
<taxonomicName id="4C32F199FFEBFF9FFF48F8C3E6AD9DAB" authority="Henderson, 1888: 414" authorityName="Henderson" authorityPageNumber="414" authorityYear="1888" box="[151,645,1917,1939]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="295" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFEBFF9FFF48F8C3E5B49DAB" box="[151,412,1917,1939]" italics="true" pageId="13" pageNumber="295">Munidopsis subsquamosa</emphasis>
<treatmentCitation id="0A93AC0BFFEBFF9FFE7DF8C3E6AD9DAB" author="Henderson" box="[418,645,1917,1939]" page="414" pageId="13" pageNumber="295" year="1888">
<bibRefCitation id="EFA3F7EBFFEBFF9FFE7DF8C3E67A9DAB" author="Henderson" box="[418,594,1917,1939]" pageId="13" pageNumber="295" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 27, 1 - 221, pls. 1 - 221." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
: 414
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</taxonomicName>
; 1888: 152, pl. 17, figs. 4, 4a
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</paragraph>
<paragraph id="8B8D8A1AFFEBFF9FFF48F822E6149DE8" blockId="13.[151,1436,1886,2031]" pageId="13" pageNumber="295">
<treatmentCitationGroup id="AB22AD34FFEBFF9FFF48F822E6149DE8" pageId="13" pageNumber="295">
<taxonomicName id="4C32F199FFEBFF9FFF48F822E5B69D8A" box="[151,414,1948,1970]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="295" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFEBFF9FFF48F822E5B69D8A" box="[151,414,1948,1970]" italics="true" pageId="13" pageNumber="295">Munidopsis subsquamosa</emphasis>
</taxonomicName>
<taxonomicName id="4C32F199FFEBFF9FFE7AF823E73F9D8A" authority="Henderson, 1888: 153" authorityName="Henderson" authorityPageNumber="153" authorityYear="1888" box="[421,791,1948,1970]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="295" phylum="Arthropoda" rank="variety" species="subsquamosa" variety="aculeata">
var.
<emphasis id="B9465608FFEBFF9FFE0DF822E6019D8A" box="[466,553,1948,1970]" italics="true" pageId="13" pageNumber="295">aculeata</emphasis>
<treatmentCitation id="0A93AC0BFFEBFF9FFDEDF822E73F9D8A" author="Henderson" box="[562,791,1948,1970]" page="153" pageId="13" pageNumber="295" year="1888">
<bibRefCitation id="EFA3F7EBFFEBFF9FFDEDF822E6CB9D8A" author="Henderson" box="[562,739,1948,1970]" pageId="13" pageNumber="295" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 27, 1 - 221, pls. 1 - 221." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
: 153
</treatmentCitation>
</taxonomicName>
, pl. 16: figs. 1, la.—
<treatmentCitation id="0A93AC0BFFEBFF9FFC32F822E09C9D8A" author="Gordon" box="[1005,1204,1948,1970]" page="244" pageId="13" pageNumber="295" year="1955">
<bibRefCitation id="EFA3F7EBFFEBFF9FFC32F822E0A89D8A" author="Gordon" box="[1005,1152,1948,1970]" pageId="13" pageNumber="295" refString="Gordon, I. (1955) Crustacea Decapoda. Reports of the Swedish Deep-Sea Expedition, 1947 - 1948, 2, 237 - 245." type="journal article" year="1955">Gordon, 1955</bibRefCitation>
: 244
</treatmentCitation>
, figs. lC, 1D, 2B, 2B', 3B, 3C.—
<treatmentCitation id="0A93AC0BFFEBFF9FFEF5F804E5D79DE8" author="Wicksten" box="[298,511,1978,2000]" page="136" pageId="13" pageNumber="295" year="1989">
<bibRefCitation id="EFA3F7EBFFEBFF9FFEF5F804E5F89DE8" author="Wicksten" box="[298,464,1978,2000]" pageId="13" pageNumber="295" refString="Wicksten, M. K. (1989) Ranges of offshore decapod crustacean in the eastern Pacific Ocean. Transactions of the san Diego Society of Natural History, 21 (19), 291 - 316." type="journal article" year="1989">Wicksten, 1989</bibRefCitation>
: 136
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8D8A1AFFEBFF9FFF48F867E7459DD7" blockId="13.[151,1436,1886,2031]" box="[151,877,2009,2031]" pageId="13" pageNumber="295">
<treatmentCitationGroup id="AB22AD34FFEBFF9FFF48F867E7459DD7" box="[151,877,2009,2031]" pageId="13" pageNumber="295">
<taxonomicName id="4C32F199FFEBFF9FFF48F867E62E9DD7" authority="Haig, 1955: 41" authorityName="Haig" authorityPageNumber="41" authorityYear="1955" box="[151,518,2009,2031]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="295" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B9465608FFEBFF9FFF48F867E5429DD7" box="[151,362,2009,2031]" italics="true" pageId="13" pageNumber="295">Munidopsis aculeata</emphasis>
<treatmentCitation id="0A93AC0BFFEBFF9FFEAFF867E62E9DD7" author="Haig" box="[368,518,2009,2031]" page="41" pageId="13" pageNumber="295" year="1955">
<bibRefCitation id="EFA3F7EBFFEBFF9FFEAFF867E5CC9DD7" author="Haig" box="[368,484,2009,2031]" pageId="13" pageNumber="295" refString="Haig, J. (1955) The Crustacea Anomura of Chile. Reports of the Lund University Chile Expedition 1948 - 49 No. 20. Lunds Universitets Arsskrift Avd. 2, 51 (12), 1 - 68, figs 1 - 12." type="journal article" year="1955">Haig, 1955</bibRefCitation>
: 41
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="0A93AC0BFFEBFF9FFDFAF867E6C99DD7" author="Retamal" box="[549,737,2009,2031]" page="23" pageId="13" pageNumber="295" year="1981">
<bibRefCitation id="EFA3F7EBFFEBFF9FFDFAF867E6969DD7" author="Retamal" box="[549,702,2009,2031]" pageId="13" pageNumber="295" refString="Retamal, M. A. (1981) Catalogo ilustrado de los crustaceos decapodos de Chile, Gayana, 44, 1 - 110." type="journal article" year="1981">Retamal, 1981</bibRefCitation>
: 23
</treatmentCitation>
(list), fig. 98.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF4DDA92FFE8FF9CFF48F8F4E5029DA5" httpUri="https://zenodo.org/record/240674/files/figure.png" pageId="14" pageNumber="296" targetBox="[163,1419,205,1830]" targetPageId="14">
<paragraph id="8B8D8A1AFFE8FF9CFF48F8F4E5029DA5" blockId="14.[151,1436,1866,1949]" pageId="14" pageNumber="296">
<emphasis id="B9465608FFE8FF9CFF48F8F4E53C9D67" bold="true" box="[151,276,1866,1888]" pageId="14" pageNumber="296">FIGURE 8.</emphasis>
<taxonomicName id="4C32F199FFE8FF9CFEC2F8F4E6AB9D58" authority="Faxon, 1893" authorityName="Faxon" authorityYear="1893" box="[285,643,1866,1888]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="296" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B9465608FFE8FF9CFEC2F8F4E5D09D58" box="[285,504,1866,1888]" italics="true" pageId="14" pageNumber="296">Munidopsis quadrata</emphasis>
Faxon, 1893
</taxonomicName>
, male cl 9.30 mm, MUAP(CD)-04111: a, lateral view of carapace; b, lateral view of abdomen; c, right cheliped; d, third right pereiopod; e, detail of carpus of right cheliped; f, merus of right cheliped. Scale = 1 mm.
</paragraph>
</caption>
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<paragraph id="8B8D8A1AFFE9FF9DFF48F921E17F9CCA" blockId="15.[151,1436,1695,1778]" pageId="15" pageNumber="297">
<emphasis id="B9465608FFE9FF9DFF48F921E53B9C8C" bold="true" box="[151,275,1695,1717]" pageId="15" pageNumber="297">FIGURE 9.</emphasis>
<taxonomicName id="4C32F199FFE9FF9DFEC5F921E6FF9C8D" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[282,727,1695,1717]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="297" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFE9FF9DFEC5F921E6379C8D" box="[282,543,1695,1717]" italics="true" pageId="15" pageNumber="297">Munidopsis subsquamosa</emphasis>
Henderson, 1888
</taxonomicName>
, ovigerous female cl 18.28 mm, MUAP(CD)-0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm.
</paragraph>
</caption>
<subSubSection id="C328D991FFE9FF82FF48F8A4E1B49B24" lastPageId="16" lastPageNumber="298" pageId="15" pageNumber="297" type="materials_examined">
<paragraph id="8B8D8A1AFFE9FF9DFF48F8A4E5C69D6E" blockId="15.[151,1436,1818,2023]" pageId="15" pageNumber="297">
<emphasis id="B9465608FFE9FF9DFF48F8A4E5079D0B" bold="true" box="[151,303,1818,1843]" pageId="15" pageNumber="297">
<typeStatus id="548934B8FFE9FF9DFF48F8A4E4FA9D0B" box="[151,210,1818,1843]" pageId="15" pageNumber="297">Type</typeStatus>
locality
</emphasis>
. Stn 237 expedition “Challenger” off the Japanese coast, SE of Nojimazaki, Boso Peninsula,
<collectingCountry id="F325CA8AFFE9FF9DFA83F8A5E1B49D0A" box="[1372,1436,1819,1842]" name="Japan" pageId="15" pageNumber="297">Japan</collectingCountry>
(
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,
<geoCoordinate id="EE06ECDDFFE9FF9DFED0F881E5569D6F" box="[271,382,1855,1879]" direction="east" orientation="longitude" pageId="15" pageNumber="297" precision="925" value="140.53334">140°32E</geoCoordinate>
),
<quantity id="4CCA27FFFFE9FF9DFE4DF881E5C29D6E" box="[402,490,1855,1879]" metricMagnitude="3" metricUnit="m" metricValue="3.431" pageId="15" pageNumber="297" unit="m" value="3431.0">3431 m</quantity>
.
</paragraph>
<paragraph id="8B8D8A1AFFE9FF9DFF18F8DCE0909DDF" blockId="15.[151,1436,1818,2023]" pageId="15" pageNumber="297">
<emphasis id="B9465608FFE9FF9DFF18F8DCE5929D43" bold="true" box="[199,442,1890,1915]" pageId="15" pageNumber="297">Distributional range</emphasis>
.
<collectingCountry id="F325CA8AFFE9FF9DFE16F8DDE6259D42" box="[457,525,1891,1914]" name="Japan" pageId="15" pageNumber="297">Japan</collectingCountry>
: Boso Peninsula, Izu Shoto (
<bibRefCitation id="EFA3F7EBFFE9FF9DFC90F8DCE0249D43" author="Henderson" box="[847,1036,1890,1915]" pageId="15" pageNumber="297" refString="Henderson, J. R. (1885) Diagnoses of new species of Galatheidae collected during the &quot; Challenger &quot; expedition. Annals and Magazine of Natural History, 16 (5), 407 - 421." type="journal article" year="1885">Henderson 1885</bibRefCitation>
;
<bibRefCitation id="EFA3F7EBFFE9FF9DFBC7F8DCE0BD9D43" author="Baba" box="[1048,1173,1890,1915]" pageId="15" pageNumber="297" refString="Baba, K. (1982) Deep-sea galatheidean Crustacea (Decapoda, Anomura) taken by the RIV Soyo-Maru in Japanese waters. 11. Galatheidae. Bulletin of the National Science Museum, Series A (Zoology), 8, 103 - 118, pls. 1, 2. [Tokyo]" type="journal article" year="1982">Baba 1982</bibRefCitation>
).
<collectingCountry id="F325CA8AFFE9FF9DFB76F8DCE13C9D43" box="[1193,1300,1890,1915]" name="Australia" pageId="15" pageNumber="297">Australia</collectingCountry>
: New South
<collectingCountry id="F325CA8AFFE9FF9DFF48F838E4F39DA7" box="[151,219,1926,1951]" name="United Kingdom" pageId="15" pageNumber="297">Wales</collectingCountry>
(
<bibRefCitation id="EFA3F7EBFFE9FF9DFF35F838E5DB9DA7" author="Ahyong" box="[234,499,1926,1951]" pageId="15" pageNumber="297" refString="Ahyong, S. T. &amp; Poore, G. C. B. (2004) Deep-water Galatheidae (Crustacea: Decapoda: Anomura) from southern and eastern Australia. Zootaxa, 472, 1 - 76" type="journal article" year="2004">Ahyong &amp; Poore 2004</bibRefCitation>
). Marion and Crozet Islands and west of Patagonia,
<collectingCountry id="F325CA8AFFE9FF9DFB98F838E0AC9DA7" box="[1095,1156,1926,1951]" name="Chile" pageId="15" pageNumber="297">Chile</collectingCountry>
(
<bibRefCitation id="EFA3F7EBFFE9FF9DFB4CF838E5BC9DFA" author="Henderson" pageId="15" pageNumber="297" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 27, 1 - 221, pls. 1 - 221." type="journal article" year="1888">
Henderson 1888, as
<taxonomicName id="4C32F199FFE9FF9DFAA2F839E5BC9DFA" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="297" phylum="Arthropoda" rank="subSpecies" species="subsquamosa" subSpecies="aculeata">
<emphasis id="B9465608FFE9FF9DFAA2F839E5BC9DFA" italics="true" pageId="15" pageNumber="297">M. subsquamosa aculeata</emphasis>
</taxonomicName>
</bibRefCitation>
).
<collectingCountry id="F325CA8AFFE9FF9DFE74F814E6179DFB" box="[427,575,1962,1987]" name="South Africa" pageId="15" pageNumber="297">South Africa</collectingCountry>
: West Cape Point (
<bibRefCitation id="EFA3F7EBFFE9FF9DFCD0F814E07E9DFA" author="Kensley" box="[783,1110,1962,1987]" pageId="15" pageNumber="297" refString="Kensley, B. F. (1968) Deep sea decapod Crustacea from west of Cape Point, South Africa. Annals of the South African Museum, 50 (12), 283 - 323." type="journal article" year="1968">
Kensley 1968 as
<taxonomicName id="4C32F199FFE9FF9DFC10F815E07E9DFA" box="[975,1110,1962,1986]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="297" phylum="Arthropoda" rank="species" species="barnardi">
<emphasis id="B9465608FFE9FF9DFC10F815E07E9DFA" box="[975,1110,1962,1986]" italics="true" pageId="15" pageNumber="297">M. barnardi</emphasis>
</taxonomicName>
</bibRefCitation>
) between Marion and Crozet islands. Coasts of the South American Pacific, Gulf of
<collectingCountry id="F325CA8AFFE9FF9DFD21F871E7729DDE" box="[766,858,1999,2022]" name="Panama" pageId="15" pageNumber="297">Panama</collectingCountry>
, off Chiloe,
<collectingCountry id="F325CA8AFFE9FF9DFC3BF870E0099DDF" box="[996,1057,1998,2023]" name="Chile" pageId="15" pageNumber="297">Chile</collectingCountry>
(
<bibRefCitation id="EFA3F7EBFFE9FF9DFBEFF870E0839DDF" author="Haig" box="[1072,1195,1998,2023]" pageId="15" pageNumber="297" refString="Haig, J. (1955) The Crustacea Anomura of Chile. Reports of the Lund University Chile Expedition 1948 - 49 No. 20. Lunds Universitets Arsskrift Avd. 2, 51 (12), 1 - 68, figs 1 - 12." type="journal article" year="1955">Haig 1955</bibRefCitation>
).
</paragraph>
<paragraph id="8B8D8A1AFFF6FF82FF18FF29E6629A97" blockId="16.[151,1436,151,1005]" box="[199,586,151,176]" pageId="16" pageNumber="298">
<emphasis id="B9465608FFF6FF82FF18FF29E58F9A88" bold="true" box="[199,423,151,176]" pageId="16" pageNumber="298">Bathymetric range</emphasis>
.
<quantity id="4CCA27FFFFF6FF82FE6AFF29E66E9A97" box="[437,582,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.148" metricValueMax="3.431" metricValueMin="0.865" pageId="16" pageNumber="298" unit="m" value="2148.0" valueMax="3431.0" valueMin="865.0">8653431 m</quantity>
.
</paragraph>
<paragraph id="8B8D8A1AFFF6FF82FF18FF02E1B49B24" blockId="16.[151,1436,151,1005]" pageId="16" pageNumber="298">
<emphasis id="B9465608FFF6FF82FF18FF02E57A9AED" bold="true" box="[199,338,188,213]" pageId="16" pageNumber="298">New record</emphasis>
.
<collectionCode id="ED2312DFFFF6FF82FE80FF02E59D9AED" box="[351,437,188,213]" pageId="16" pageNumber="298">MUAP</collectionCode>
(CD)-0468, ovigerous female, cl
<quantity id="4CCA27FFFFF6FF82FCFBFF02E7BF9AEC" box="[804,919,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.828" pageId="16" pageNumber="298" unit="mm" value="18.28">18.28 mm</quantity>
, off Caldera (~27°S; about
<quantity id="4CCA27FFFFF6FF82FB14FF02E10B9AEC" box="[1227,1315,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="298" unit="m" value="1000.0">1000 m</quantity>
),
<date id="FF8CACDAFFF6FF82FAE8FF02E4FA9AC0" pageId="16" pageNumber="298" value="2004-02">February 2004</date>
, taken in Patagonian Toothfish (
<taxonomicName id="4C32F199FFF6FF82FD8BFF5EE7449AC0" box="[596,876,224,248]" class="Actinopterygii" family="Nototheniidae" genus="Dissostichus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="298" phylum="Chordata" rank="species" species="eleginoides">
<emphasis id="B9465608FFF6FF82FD8BFF5EE7449AC0" box="[596,876,224,248]" italics="true" pageId="16" pageNumber="298">Dissostichus eleginoides</emphasis>
</taxonomicName>
) long-line at about
<quantity id="4CCA27FFFFF6FF82FB83FF5EE0919ACF" box="[1116,1209,224,248]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="298" unit="m" value="1000.0">1000 m</quantity>
;
<collectionCode id="ED2312DFFFF6FF82FB16FF61E1379AC0" box="[1225,1311,223,248]" pageId="16" pageNumber="298">MUAP</collectionCode>
(CD)-0469, ovigerous female cl
<quantity id="4CCA27FFFFF6FF82FEA6FEBAE5C49B24" box="[377,492,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.7559999999999998" pageId="16" pageNumber="298" unit="mm" value="17.56">17.56 mm</quantity>
, off Concepción “ONR” expedition AGT 0 7 (
<geoCoordinate id="EE06ECDDFFF6FF82FC33FEBAE0469B25" box="[1004,1134,260,285]" direction="south" orientation="latitude" pageId="16" pageNumber="298" precision="9" value="-32.36067">32°21.64S</geoCoordinate>
,
<geoCoordinate id="EE06ECDDFFF6FF82FBA6FEBAE12B9B25" box="[1145,1283,260,285]" direction="west" orientation="longitude" pageId="16" pageNumber="298" precision="9" value="-73.726166">73°43.57W</geoCoordinate>
,
<quantity id="4CCA27FFFFF6FF82FAD1FEBAE1B89B24" box="[1294,1424,260,285]" metricMagnitude="2" metricUnit="m" metricValue="8.955" metricValueMax="9.26" metricValueMin="8.65" pageId="16" pageNumber="298" unit="m" value="895.5" valueMax="926.0" valueMin="865.0">865926 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C328D991FFF6FF82FF18FE99E08198BD" pageId="16" pageNumber="298" type="description">
<paragraph id="8B8D8A1AFFF6FF82FF18FE99E08198BD" blockId="16.[151,1436,151,1005]" pageId="16" pageNumber="298">
<emphasis id="B9465608FFF6FF82FF18FE99E5799B78" bold="true" box="[199,337,295,320]" pageId="16" pageNumber="298">Description</emphasis>
. Rostrum acute, short, almost half carapace length; with medial keel. Carapace almost pentagonal, densely covered by scale-like tubercles ending in minute spine with row of single setae; few spines in cardiac region and along margins (see Henderson 1895); antennal spine acute, short, not reaching the cornea; carapace surface flat, without well delimited regions; cervical groove poorly developed (
<figureCitation id="1309969FFFF6FF82FBF0FE2AE05E9B95" box="[1071,1142,404,429]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">Fig. 9</figureCitation>
b). Eyes with long ocular spine (
<figureCitation id="1309969FFFF6FF82FF3BFE09E5039BE8" box="[228,299,439,464]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">Fig. 9</figureCitation>
a). Abdominal segments unarmed, with 1 transverse groove on second to fourth somites (
<figureCitation id="1309969FFFF6FF82FAE5FE09E1A99BE8" box="[1338,1409,439,464]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">Fig. 9</figureCitation>
c). Chelipeds short, almost same length as carapace; palm as long as dactyl, both unarmed (
<figureCitation id="1309969FFFF6FF82FBA7FE62E0949BCD" box="[1144,1212,476,501]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">Fig. 9</figureCitation>
d), surface with low tubercles ending in row of setae; carpus and merus armed with strong spines on the distal margin, merus with mesial row of spines (
<figureCitation id="1309969FFFF6FF82FE54FD9AE5E69804" box="[395,462,548,573]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">Fig. 9</figureCitation>
e). Walking legs as long as two-thirds total length carapace, armed on the dorsal margin of carpus and merus; propodus and dactyl unarmed dorsally (
<figureCitation id="1309969FFFF6FF82FC96FDF9E7A59858" box="[841,909,583,608]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">Fig. 9</figureCitation>
f); dactyl with ventral row of spines, each with seta on anterior margin (
<figureCitation id="1309969FFFF6FF82FE76FDD2E5C498BC" box="[425,492,620,645]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">
Fig.
<quantity id="4CCA27FFFFF6FF82FE02FDD2E5C498BC" box="[477,492,620,644]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="16" pageNumber="298" unit="g" value="9.0">9</quantity>
</figureCitation>
g); distoventral end of propodus with mobile spine (
<figureCitation id="1309969FFFF6FF82FBEEFDD2E05C98BD" box="[1073,1140,620,645]" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">Fig. 9</figureCitation>
f, h).
</paragraph>
</subSubSection>
<subSubSection id="C328D991FFF6FF82FF18FD31E7B49965" pageId="16" pageNumber="298" type="discussion">
<paragraph id="8B8D8A1AFFF6FF82FF18FD31E7B49965" blockId="16.[151,1436,151,1005]" pageId="16" pageNumber="298">
<emphasis id="B9465608FFF6FF82FF18FD31E51C9890" bold="true" box="[199,308,655,680]" pageId="16" pageNumber="298">Remarks</emphasis>
. The main differences between the
<typeStatus id="548934B8FFF6FF82FD19FD2EE7359890" box="[710,797,656,680]" pageId="16" pageNumber="298" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C32F199FFF6FF82FC9BFD2FE09D989F" authority="Faxon" authorityName="Faxon" box="[836,1205,656,680]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="subSpecies" species="subsquamosa" subSpecies="aculeata">
<emphasis id="B9465608FFF6FF82FC9BFD2FE0409890" box="[836,1128,656,680]" italics="true" pageId="16" pageNumber="298">M. subsquamosa aculeata</emphasis>
Faxon
</taxonomicName>
and the
<typeStatus id="548934B8FFF6FF82FAC9FD31E1559890" box="[1302,1405,655,680]" pageId="16" pageNumber="298" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C32F199FFF6FF82FF48FD0BE5FA98F5" authority="Henderson" authorityName="Henderson" box="[151,466,692,717]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFF6FF82FF48FD0BE57B98F4" box="[151,339,692,716]" italics="true" pageId="16" pageNumber="298">M. subsquamosa</emphasis>
Henderson
</taxonomicName>
compared to the specimens analyzed here are in the straighter dactyls of the P24 (
<figureCitation id="1309969FFFF6FF82FAB0FD0AE48E98C8" captionStart="FIGURE 9" captionStartId="15.[151,250,1695,1717]" captionTargetBox="[160,1435,196,1657]" captionTargetId="figure@15.[151,1436,193,1674]" captionTargetPageId="15" captionText="FIGURE 9. Munidopsis subsquamosa Henderson, 1888, ovigerous female cl 18.28 mm, MUAP (CD) - 0468: a, anterior region of carapace; b, lateral view of carapace; c, lateral view of abdomen; d, left cheliped; e, carpus and merus of left cheliped; f, third left pereiopod; g, dactyl of third pereiopod; h, detail of ventrodistal spine of propodus of third pereiopod. Scale = 1 mm." httpUri="https://zenodo.org/record/240675/files/figure.png" pageId="16" pageNumber="298">
Fig.
<quantity id="4CCA27FFFFF6FF82FF48FD66E48E98C8" box="[151,166,728,752]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="16" pageNumber="298" unit="g" value="9.0">9</quantity>
</figureCitation>
g). Regarding the differences noted by
<bibRefCitation id="EFA3F7EBFFF6FF82FDBEFD69E6C598C8" author="Baba" box="[609,749,727,752]" pageId="16" pageNumber="298" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-Pacific, with a list of species. Galathea Report, 20, 1 - 317." type="journal article" year="2005">Baba (2005)</bibRefCitation>
in the more spinous form of the
<typeStatus id="548934B8FFF6FF82FBBAFD66E09498C8" box="[1125,1212,728,752]" pageId="16" pageNumber="298" type="syntype">syntype</typeStatus>
specimens than the
<typeStatus id="548934B8FFF6FF82FF48FD42E4D6992D" box="[151,254,764,789]" pageId="16" pageNumber="298" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C32F199FFF6FF82FEF9FD43E5CC992C" box="[294,484,764,788]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFF6FF82FEF9FD43E5CC992C" box="[294,484,764,788]" italics="true" pageId="16" pageNumber="298">M. subsquamosa</emphasis>
</taxonomicName>
, the specimens observed here are scarcely spinous and the rostrum is upturned. By this, our specimens are more similar to
<taxonomicName id="4C32F199FFF6FF82FD93FC9FE7469900" box="[588,878,800,824]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="subSpecies" species="subsquamosa" subSpecies="aculeata">
<emphasis id="B9465608FFF6FF82FD93FC9FE7469900" box="[588,878,800,824]" italics="true" pageId="16" pageNumber="298">M. subsquamosa aculeata</emphasis>
</taxonomicName>
. Nevertheless, the differences are not sufficient to separate then into two subspecies, as also concluded by
<bibRefCitation id="EFA3F7EBFFF6FF82FCD5FCFAE7B09965" author="Baba" box="[778,920,836,861]" pageId="16" pageNumber="298" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-Pacific, with a list of species. Galathea Report, 20, 1 - 317." type="journal article" year="2005">Baba (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328D991FFF6FF82FF18FCD6E7E899D5" pageId="16" pageNumber="298" type="reference_group">
<paragraph id="8B8D8A1AFFF6FF82FF18FCD6E7E899D5" blockId="16.[151,1436,151,1005]" pageId="16" pageNumber="298">
<taxonomicName id="4C32F199FFF6FF82FF18FCD6E5C299B8" box="[199,490,872,896]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFF6FF82FF18FCD6E5C299B8" box="[199,490,872,896]" italics="true" pageId="16" pageNumber="298">Munidopsis subsquamosa</emphasis>
</taxonomicName>
has been cited many times in Chilean waters as
<taxonomicName id="4C32F199FFF6FF82FBC3FCD7E15899B8" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[1052,1392,871,896]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B9465608FFF6FF82FBC3FCD7E08C99B8" box="[1052,1188,872,896]" italics="true" pageId="16" pageNumber="298">M. aculeata</emphasis>
<bibRefCitation id="EFA3F7EBFFF6FF82FB73FCD9E15899B8" author="Henderson" box="[1196,1392,871,896]" pageId="16" pageNumber="298" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 27, 1 - 221, pls. 1 - 221." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C32F199FFF6FF82FAA2FCD7E506999C" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="B9465608FFF6FF82FAA2FCD7E506999C" italics="true" pageId="16" pageNumber="298">M. subsquamosa</emphasis>
</taxonomicName>
<taxonomicName id="4C32F199FFF6FF82FEE6FC33E689999C" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[313,673,908,933]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="variety" species="subsquamosa" variety="aculeata">
var.
<emphasis id="B9465608FFF6FF82FEB1FC32E5E6999C" box="[366,462,908,932]" italics="true" pageId="16" pageNumber="298">aculeata</emphasis>
<bibRefCitation id="EFA3F7EBFFF6FF82FE05FC32E689999C" author="Henderson" box="[474,673,908,933]" pageId="16" pageNumber="298" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 27, 1 - 221, pls. 1 - 221." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
</taxonomicName>
, or
<taxonomicName id="4C32F199FFF6FF82FD0CFC33E0B4999D" authority="Faxon, 1895" authorityName="Faxon" authorityYear="1895" box="[723,1180,908,933]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="298" phylum="Arthropoda" rank="subSpecies" species="subsquamosa" subSpecies="aculeata">
<emphasis id="B9465608FFF6FF82FD0CFC33E028999C" box="[723,1024,908,932]" italics="true" pageId="16" pageNumber="298">M. subsquamosa aculeata</emphasis>
<bibRefCitation id="EFA3F7EBFFF6FF82FBD3FC33E0B4999D" author="Faxon" box="[1036,1180,908,933]" pageId="16" pageNumber="298" refString="Faxon, W. (1895) Reports on an exploration off the west coasts of Mexico, Central and South America, and off the Galapagos Islands, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; during 1891, Lieut. - Commander Z. L. Tanner, U. S. N. commanding. XV. The stalk-eyed Crustacea. Memoirs of the Museum of Comparative Zoology at Harvard College, 18, 1 - 292." type="journal article" year="1895">Faxon, 1895</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFA3F7EBFFF6FF82FB6FFC32E106999D" author="Haig" box="[1200,1326,908,933]" pageId="16" pageNumber="298" refString="Haig, J. (1955) The Crustacea Anomura of Chile. Reports of the Lund University Chile Expedition 1948 - 49 No. 20. Lunds Universitets Arsskrift Avd. 2, 51 (12), 1 - 68, figs 1 - 12." type="journal article" year="1955">Haig 1955</bibRefCitation>
;
<bibRefCitation id="EFA3F7EBFFF6FF82FAE0FC32E4F999F0" author="Retamal" pageId="16" pageNumber="298" refString="Retamal, M. A. (1981) Catalogo ilustrado de los crustaceos decapodos de Chile, Gayana, 44, 1 - 110." type="journal article" year="1981">Retamal 1981</bibRefCitation>
,
<bibRefCitation id="EFA3F7EBFFF6FF82FF01FC0EE53199F0" author="Retamal" box="[222,281,944,968]" pageId="16" pageNumber="298" refString="Retamal, M. A. (2000) Decapodos de Chile. ETI-Universidad de Concepcion. Springer-Verlag, Berlin. [CD-ROM]" type="book" year="2000">2000</bibRefCitation>
;
<bibRefCitation id="EFA3F7EBFFF6FF82FEFAFC11E66199F0" author="Retamal" box="[293,585,943,968]" pageId="16" pageNumber="298" refString="Retamal, M. A. &amp; Moyano, H. I. (2011) Zoogeografia de los crustaceos decapodos chilenos marinos y dulceacuicolas. Latin American Journal of Aquatic Research, 38 (3), 302 - 328." type="journal article" year="2011">Retamal &amp; Moyano 2011</bibRefCitation>
). Despite these citations, specimens of this species were known from
<collectingCountry id="F325CA8AFFF6FF82FABFFC11E1B499F0" box="[1376,1436,943,968]" name="Chile" pageId="16" pageNumber="298">Chile</collectingCountry>
only from the original description, so this is a new record of the species.
</paragraph>
</subSubSection>
</treatment>
</document>