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<document id="93BD4CFAACAE04E97399D0625D9DDF3F" ID-CLB-Dataset="25523" ID-DOI="10.11646/zootaxa.4671.2.11" ID-GBIF-Dataset="02c8367f-2b3a-46d1-bf9a-d15c89a3b0be" ID-ISSN="1175-5326" ID-Zenodo-Dep="3442164" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1568799318906" checkinUser="plazi" docAuthor="Yun, Chan-Gyu" docDate="2019" docId="03951F1BC8374D1EFF63AACBFD37FAFA" docLanguage="en" docName="zootaxa.4671.2.11.pdf.imf" docOrigin="Zootaxa 4671 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heyuanninae Yun, 2019, clade. nov" docType="treatment" docVersion="9" lastPageNumber="295" masterDocId="FFAC6763C8374D1EFFF4A842FFB0FF8A" masterDocTitle="Heyuanninae clade nov., a replacement name for the oviraptorid subfamily “ Ingeniinae ” Barsbold, 1981" masterLastPageNumber="296" masterPageNumber="295" pageNumber="295" updateTime="1698755068915" updateUser="ExternalLinkService">
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<mods:title id="63CEDA6427E7CC514C106DF6D8D1BD29">Heyuanninae clade nov., a replacement name for the oviraptorid subfamily “ Ingeniinae ” Barsbold, 1981</mods:title>
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<mods:namePart id="1F6EC3BC43F64831BE6A98FF3FB99B5D">Yun, Chan-Gyu</mods:namePart>
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<treatment id="03951F1BC8374D1EFF63AACBFD37FAFA" ID-DOI="http://doi.org/10.5281/zenodo.5933383" ID-GBIF-Taxon="159608345" ID-Zenodo-Dep="5933383" LSID="urn:lsid:plazi:treatment:03951F1BC8374D1EFF63AACBFD37FAFA" httpUri="http://treatment.plazi.org/id/03951F1BC8374D1EFF63AACBFD37FAFA" lastPageNumber="295" pageId="0" pageNumber="295">
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<paragraph id="8B83AE0DC8374D1EFF63AACBFCDBFCDF" blockId="0.[151,1437,649,1393]" pageId="0" pageNumber="295">
<bibRefCitation id="EFADD3FCC8374D1EFF63AACBFEF8FD2B" author="Barsbold, R." box="[151,328,649,673]" pageId="0" pageNumber="295" pagination="28 - 39" refId="ref534" refString="Barsbold, R. (1981) [Toothless carnivorous dinosaurs of Mongolia]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 15, 28 - 39. [in Russian]" type="journal article" year="1981">Barsbold (1981)</bibRefCitation>
erected
<taxonomicName id="4C3CD58EC8374D1EFE5DAACBFCBEFD2B" authority="Barsbold, 1981" authorityName="Barsbold" authorityYear="1981" box="[425,782,649,673]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFE5DAACBFDEEFD2B" box="[425,606,649,673]" italics="true" pageId="0" pageNumber="295">Ingenia yanshini</emphasis>
<bibRefCitation id="EFADD3FCC8374D1EFD9CAACBFCBEFD2B" author="Barsbold, R." box="[616,782,649,673]" pageId="0" pageNumber="295" pagination="28 - 39" refId="ref534" refString="Barsbold, R. (1981) [Toothless carnivorous dinosaurs of Mongolia]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 15, 28 - 39. [in Russian]" type="journal article" year="1981">Barsbold, 1981</bibRefCitation>
</taxonomicName>
for a nearly complete oviraptorid skeleton from the Upper Cretaceous (Campanian-Maastrichtian) Barun Goyot Formation in
<collectingRegion id="49F860EFC8374D1EFCABAAE9FC7BFD4F" box="[863,971,683,709]" country="Mongolia" name="Omnogovi" pageId="0" pageNumber="295">Ömnögov</collectingRegion>
, southern
<collectingCountry id="F32BEE9DC8374D1EFBC2AAEFFB11FD4F" box="[1078,1185,685,709]" name="Mongolia" pageId="0" pageNumber="295">Mongolia</collectingCountry>
. Moreover, he erected a subfamily-level clade Ingeniinae
<bibRefCitation id="EFADD3FCC8374D1EFE0FAA93FD2EFD63" author="Barsbold, R." box="[507,670,721,745]" pageId="0" pageNumber="295" pagination="28 - 39" refId="ref534" refString="Barsbold, R. (1981) [Toothless carnivorous dinosaurs of Mongolia]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 15, 28 - 39. [in Russian]" type="journal article" year="1981">Barsbold, 1981</bibRefCitation>
to distinguish
<taxonomicName id="4C3CD58EC8374D1EFCCFAA93FC5DFD63" authorityName="Barsbold" authorityYear="1981" box="[827,1005,721,745]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFCCFAA93FC5DFD63" box="[827,1005,721,745]" italics="true" pageId="0" pageNumber="295">Ingenia yanshini</emphasis>
</taxonomicName>
from other oviraptorids such as
<taxonomicName id="4C3CD58EC8374D1EFABCAA93FD15FC87" authority="Osborn, 1924" authorityName="Osborn" authorityYear="1924" class="Reptilia" family="Oviraptoridae" genus="Oviraptor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="philoceratops">
<emphasis id="B948721FC8374D1EFABCAA93FEE0FC87" italics="true" pageId="0" pageNumber="295">Oviraptor philoceratops</emphasis>
<bibRefCitation id="EFADD3FCC8374D1EFEA2AAB7FE57FC87" author="Osborn, H. F." box="[342,487,757,781]" pageId="0" pageNumber="295" pagination="1 - 12" refId="ref1183" refString="Osborn, H. F. (1924) Three new Theropoda, Protoceratops zone, central Mongolia. American Museum Novitates, 144, 1 - 12." type="journal article" year="1924">Osborn, 1924</bibRefCitation>
(
<bibRefCitation id="EFADD3FCC8374D1EFE01AAB7FD2DFC87" author="Barsbold, R." box="[501,669,757,781]" pageId="0" pageNumber="295" pagination="28 - 39" refId="ref534" refString="Barsbold, R. (1981) [Toothless carnivorous dinosaurs of Mongolia]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 15, 28 - 39. [in Russian]" type="journal article" year="1981">Barsbold, 1981</bibRefCitation>
)
</taxonomicName>
. When Barsbold first erected the subfamily, he considered this subfamily as monotypic (
<bibRefCitation id="EFADD3FCC8374D1EFEAEAB5BFDB0FCBB" author="Barsbold, R." box="[346,512,793,817]" pageId="0" pageNumber="295" pagination="28 - 39" refId="ref534" refString="Barsbold, R. (1981) [Toothless carnivorous dinosaurs of Mongolia]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 15, 28 - 39. [in Russian]" type="journal article" year="1981">Barsbold, 1981</bibRefCitation>
) but now this subfamily comprisesmultiple taxa such as
<taxonomicName id="4C3CD58EC8374D1EFB91AB5BFDB3FCDF" authority="Longrich, Currie, and Dong, 2010" authorityName="Longrich, Currie, and Dong" authorityYear="2010" class="Reptilia" family="Oviraptoridae" genus="Machairasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="leptonychus">
<emphasis id="B948721FC8374D1EFB91AB5BFA2CFCBB" box="[1125,1436,793,817]" italics="true" pageId="0" pageNumber="295">Machairasaurus leptonychus</emphasis>
<bibRefCitation id="EFADD3FCC8374D1EFF63AB7FFDB3FCDF" author="Longrich, N. R. &amp; Currie, P. J. &amp; Dong, Z. - M." box="[151,515,829,853]" pageId="0" pageNumber="295" pagination="945 - 960" refId="ref989" refString="Longrich, N. R., Currie, P. J. &amp; Dong, Z. - M. (2010) A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of Bayan Mandahu, Inner Mongolia. Paleontology, 53, 945 - 960. https: // doi. org / 10.1111 / j. 1475 - 4983.2010.00968. x" type="journal article" year="2010">Longrich, Currie, and Dong, 2010</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3CD58EC8374D1EFDC3AB7FFCD7FCDF" authority="Lu, 2002" authorityName="Lu" authorityYear="2002" box="[567,871,829,853]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="huangi">
<emphasis id="B948721FC8374D1EFDC3AB7FFD4CFCDF" box="[567,764,829,853]" italics="true" pageId="0" pageNumber="295">Heyuannia huangi</emphasis>
<bibRefCitation id="EFADD3FCC8374D1EFCF7AB7FFCD7FCDF" author="Lu, J." box="[771,871,829,853]" pageId="0" pageNumber="295" pagination="871 - 875" refId="ref1056" refString="Lu, J. (2002) A new oviraptorosaurid (Theropoda: Oviraptorosauria) from the Late Cretaceous of southern China. Journal of Vertebrate Paleontology, 22, 871 - 875. https: // doi. org / 10.1671 / 0272 - 4634 (2002) 022 [0871: ANOTOF] 2.0. CO; 2" type="journal article" year="2002">Lü, 2002</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C326FD86C8374D1EFF33AB23FA35FC4B" pageId="0" pageNumber="295" type="discussion">
<paragraph id="8B83AE0DC8374D1EFF33AB23FA35FC4B" blockId="0.[151,1437,649,1393]" pageId="0" pageNumber="295">
However, the genus name
<taxonomicName id="4C3CD58EC8374D1EFE2BAB23FD80FCF2" authorityName="Gerlach" authorityYear="1957" box="[479,560,865,888]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="genus">
<emphasis id="B948721FC8374D1EFE2BAB23FD80FCF2" box="[479,560,865,888]" italics="true" pageId="0" pageNumber="295">Ingenia</emphasis>
</taxonomicName>
is a junior homonym of the tripyloidid nematode genus
<taxonomicName id="4C3CD58EC8374D1EFB76AB23FADFFCF2" authority="Gerlach, 1957" authorityName="Gerlach" authorityYear="1957" box="[1154,1391,864,889]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="genus">
<emphasis id="B948721FC8374D1EFB76AB23FB63FCF2" box="[1154,1235,865,888]" italics="true" pageId="0" pageNumber="295">Ingenia</emphasis>
<bibRefCitation id="EFADD3FCC8374D1EFB2DAB23FADFFCF2" author="Gerlach, S. A." box="[1241,1391,864,889]" pageId="0" pageNumber="295" pagination="411 - 459" refId="ref737" refString="Gerlach, S. A. (1957) Die Nematodenfauna des Sandstrandes an der Kuste von Mittelbrasilien (Brasilianische MeeresNematoden IV). Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, Berlin, 33, 411 - 459. https: // doi. org / 10.1002 / mmnz. 19570330206" type="journal article" year="1957">Gerlach, 1957</bibRefCitation>
</taxonomicName>
and according to the Article 23.1 of the International Code of Zoological Nomenclature (
<bibRefCitation id="EFADD3FCC8374D1EFBD4ABC7FB16FC17" author="ICZN" box="[1056,1190,901,925]" pageId="0" pageNumber="295" refId="ref885" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, c / o The Natural History Museum, London, 126 pp." type="book" year="1999">ICZN, 1999</bibRefCitation>
),
<bibRefCitation id="EFADD3FCC8374D1EFB4DABC7FAEBFC16" author="Gerlach, S. A." box="[1209,1371,900,925]" pageId="0" pageNumber="295" pagination="411 - 459" refId="ref737" refString="Gerlach, S. A. (1957) Die Nematodenfauna des Sandstrandes an der Kuste von Mittelbrasilien (Brasilianische MeeresNematoden IV). Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, Berlin, 33, 411 - 459. https: // doi. org / 10.1002 / mmnz. 19570330206" type="journal article" year="1957">Gerlachs 1957</bibRefCitation>
name has priority. Therefore, the usage of the subfamily name Ingeniinae is problematic as it is based on a junior homonym.
</paragraph>
</subSubSection>
<paragraph id="8B83AE0DC8374D1EFF33AB8FFC7FFB6B" blockId="0.[151,1437,649,1393]" pageId="0" pageNumber="295">
<bibRefCitation id="EFADD3FCC8374D1EFF33AB8FFEF6FC6F" author="Paul, G. S." box="[199,326,973,997]" pageId="0" pageNumber="295" refId="ref1288" refString="Paul, G. S. (1988) Predatory Dinosaurs of the World: a Complete Illustrated Guide. Simon and Schuster Co., New York, 464 pp." type="book" year="1988">Paul (1988)</bibRefCitation>
suggested a new combination
<taxonomicName id="4C3CD58EC8374D1EFD67AB8FFCD0FC6F" baseAuthorityName="Barsbold" baseAuthorityYear="1981" box="[659,864,973,997]" class="Reptilia" family="Oviraptoridae" genus="Oviraptor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFD67AB8FFCD0FC6F" box="[659,864,973,997]" italics="true" pageId="0" pageNumber="295">Oviraptor yanshini</emphasis>
</taxonomicName>
for “
<taxonomicName id="4C3CD58EC8374D1EFC6FAB8FFC5CFC6E" authorityName="Gerlach" authorityYear="1957" box="[923,1004,973,996]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="genus">
<emphasis id="B948721FC8374D1EFC6FAB8FFC5CFC6E" box="[923,1004,973,996]" italics="true" pageId="0" pageNumber="295">Ingenia</emphasis>
</taxonomicName>
<taxonomicName id="4C3CD58EC8374D1EFBF4AB8FFBEAFC6F" baseAuthorityName="Barsbold" baseAuthorityYear="1981" box="[1024,1114,973,997]" class="Reptilia" family="Oviraptoridae" genus="Oviraptor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFBF4AB8FFBEAFC6F" box="[1024,1114,973,997]" italics="true" pageId="0" pageNumber="295">yanshini</emphasis>
</taxonomicName>
but this classification has not been supported by any subsequent authors (
<bibRefCitation id="EFADD3FCC8374D1EFD96ABB3FD5CFB83" author="Easter, J." box="[610,748,1009,1033]" pageId="0" pageNumber="295" pagination="184 - 190" refId="ref565" refString="Easter, J. (2013) A new name for the oviraptorid dinosaur &quot; Ingenia &quot; yanshini (Barsbold, 1981; preoccupied by Gerlach, 1957). Zootaxa, 3737 (2), 184 - 190." type="journal article" year="2013">Easter, 2013</bibRefCitation>
).
<bibRefCitation id="EFADD3FCC8374D1EFD09ABB3FC3CFB83" author="Easter, J." box="[765,908,1009,1033]" pageId="0" pageNumber="295" pagination="184 - 190" refId="ref565" refString="Easter, J. (2013) A new name for the oviraptorid dinosaur &quot; Ingenia &quot; yanshini (Barsbold, 1981; preoccupied by Gerlach, 1957). Zootaxa, 3737 (2), 184 - 190." type="journal article" year="2013">Easter (2013)</bibRefCitation>
proposed a new genus name
<taxonomicName id="4C3CD58EC8374D1EFB34ABB3FF7CFBA7" authority="Easter, 2013" authorityName="Easter" authorityYear="2013" class="Reptilia" family="Oviraptoridae" genus="Ajancingenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="genus">
<emphasis id="B948721FC8374D1EFB34ABB3FAFEFB82" box="[1216,1358,1009,1032]" italics="true" pageId="0" pageNumber="295">Ajancingenia</emphasis>
<bibRefCitation id="EFADD3FCC8374D1EFAA0ABB3FF7CFBA7" author="Easter, J." pageId="0" pageNumber="295" pagination="184 - 190" refId="ref565" refString="Easter, J. (2013) A new name for the oviraptorid dinosaur &quot; Ingenia &quot; yanshini (Barsbold, 1981; preoccupied by Gerlach, 1957). Zootaxa, 3737 (2), 184 - 190." type="journal article" year="2013">Easter, 2013</bibRefCitation>
</taxonomicName>
for “
<emphasis id="B948721FC8374D1EFEF6AC57FE72FBA7" box="[258,450,1045,1069]" italics="true" pageId="0" pageNumber="295">
<taxonomicName id="4C3CD58EC8374D1EFEF6AC57FEEBFBA6" authorityName="Gerlach" authorityYear="1957" box="[258,347,1045,1068]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="genus">Ingenia</taxonomicName>
<taxonomicName id="4C3CD58EC8374D1EFE9CAC57FE72FBA7" baseAuthorityName="Barsbold" baseAuthorityYear="1981" box="[360,450,1045,1069]" class="Reptilia" family="Oviraptoridae" genus="Oviraptor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="yanshini">yanshini</taxonomicName>
</emphasis>
, but subsequent phylogenetic analyses have found “
<taxonomicName id="4C3CD58EC8374D1EFC1DAC57FB8AFBA6" authorityName="Gerlach" authorityYear="1957" box="[1001,1082,1045,1068]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="genus">
<emphasis id="B948721FC8374D1EFC1DAC57FB8AFBA6" box="[1001,1082,1045,1068]" italics="true" pageId="0" pageNumber="295">Ingenia</emphasis>
</taxonomicName>
<taxonomicName id="4C3CD58EC8374D1EFBB8AC57FB16FBA7" baseAuthorityName="Barsbold" baseAuthorityYear="1981" box="[1100,1190,1045,1069]" class="Reptilia" family="Oviraptoridae" genus="Oviraptor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFBB8AC57FB16FBA7" box="[1100,1190,1045,1069]" italics="true" pageId="0" pageNumber="295">yanshini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3CD58EC8374D1EFB2CAC57FA2CFBA7" authorityName="Lu" authorityYear="2002" box="[1240,1436,1045,1069]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="huangi">
<emphasis id="B948721FC8374D1EFB2CAC57FA2CFBA7" box="[1240,1436,1045,1069]" italics="true" pageId="0" pageNumber="295">Heyuannia huangi</emphasis>
</taxonomicName>
as sister taxa (e.g.,
<bibRefCitation id="EFADD3FCC8374D1EFE9EAC7BFDE6FBDB" author="Lamanna, M. C. &amp; Sues, H. - D. &amp; Schachner, E. R. &amp; Lyson, T. R." box="[362,598,1081,1105]" pageId="0" pageNumber="295" pagination="92022" refId="ref918" refString="Lamanna, M. C., Sues, H. - D., Schachner, E. R. &amp; Lyson, T. R. (2014) A New Large-Bodied Oviraptorosaurian Theropod Dinosaur from the Latest Cretaceous of Western North America. PLoS ONE, 9 (3), e 92022. https: // doi. org / 10.1371 / journal. pone. 0092022" type="journal article" year="2014">
Lamanna
<emphasis id="B948721FC8374D1EFE22AC7BFDBEFBDB" box="[470,526,1081,1105]" italics="true" pageId="0" pageNumber="295">et al.</emphasis>
, 2014
</bibRefCitation>
;
<bibRefCitation id="EFADD3FCC8374D1EFD96AC7BFCCCFBDA" author="Funston, G. F. &amp; Currie, P. J." box="[610,892,1080,1105]" pageId="0" pageNumber="295" pagination="1160910" refId="ref606" refString="Funston, G. F. &amp; Currie, P. J. (2016) A new caenagnathid (Dinosauria: Oviraptorosauria) from the Horseshoe Canyon Formation of Alberta, Canada, and a reevaluation of the relationships of Caenagnathidae. Journal of Vertebrate Paleontology, 36 (4), e 1160910. https: // doi. org / 10.1080 / 02724634.2016.1160910" type="journal article" year="2016">Funston and Currie, 2016</bibRefCitation>
;
<bibRefCitation id="EFADD3FCC8374D1EFC7CAC7BFB9EFBDA" author="Lu, J. &amp; Chen, R. &amp; Brusatte, S. L. &amp; Zhu, Y. &amp; Shen, C." box="[904,1070,1080,1105]" pageId="0" pageNumber="295" pagination="35780" refId="ref1116" refString="Lu, J., Chen, R., Brusatte, S. L., Zhu, Y. &amp; Shen, C. (2016) A Late Cretaceous diversification of Asian oviraptorid dinosaurs; evidence from a new species preserved in an unusual posture. Scientific Reports, 6, 35780. https: // doi. org / 10.1038 / srep 35780" type="journal article" year="2016">
<emphasis id="B948721FC8374D1EFC5BAC7BFC56FBDB" box="[943,998,1081,1105]" italics="true" pageId="0" pageNumber="295">et al.</emphasis>
, 2016
</bibRefCitation>
;
<bibRefCitation id="EFADD3FCC8374D1EFBCEAC7BFAA9FBDB" author="Funston, G. F. &amp; Mendonca, S. E. &amp; Currie, P. J. &amp; Barsbold, R." box="[1082,1305,1081,1105]" pageId="0" pageNumber="295" pagination="101 - 120" refId="ref672" refString="Funston, G. F., Mendonca, S. E., Currie, P. J. &amp; Barsbold, R. (2018) Oviraptorosaur anatomy, diversity and ecology in the Nemegt Basin. Palaeogeography, Palaeoclimatology, Palaeoecology, 494, 101 - 120. https: // doi. org / 10.1016 / j. palaeo. 2017.10.023" type="journal article" year="2018">
Funston
<emphasis id="B948721FC8374D1EFB6DAC7BFB61FBDB" box="[1177,1233,1081,1105]" italics="true" pageId="0" pageNumber="295">et al.</emphasis>
, 2018
</bibRefCitation>
). Therefore, most recent studies have considered
<taxonomicName id="4C3CD58EC8374D1EFDE3AC1FFD15FBFE" authorityName="Easter" authorityYear="2013" box="[535,677,1117,1140]" class="Reptilia" family="Oviraptoridae" genus="Ajancingenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="genus">
<emphasis id="B948721FC8374D1EFDE3AC1FFD15FBFE" box="[535,677,1117,1140]" italics="true" pageId="0" pageNumber="295">Ajancingenia</emphasis>
</taxonomicName>
as junior synonym of
<taxonomicName id="4C3CD58EC8374D1EFC7AAC1FFBB3FBFE" authorityName="Lu" authorityYear="2003" box="[910,1027,1117,1140]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="genus">
<emphasis id="B948721FC8374D1EFC7AAC1FFBB3FBFE" box="[910,1027,1117,1140]" italics="true" pageId="0" pageNumber="295">Heyuannia</emphasis>
</taxonomicName>
and used new combination
<taxonomicName id="4C3CD58EC8374D1EFAD3AC1FFF41FB13" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFAD3AC1FFF41FB13" italics="true" pageId="0" pageNumber="295">Heyuannia yanshini</emphasis>
</taxonomicName>
for “
<taxonomicName id="4C3CD58EC8374D1EFEDFACC3FECCFB12" authorityName="Gerlach" authorityYear="1957" box="[299,380,1153,1176]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="genus">
<emphasis id="B948721FC8374D1EFEDFACC3FECCFB12" box="[299,380,1153,1176]" italics="true" pageId="0" pageNumber="295">Ingenia</emphasis>
</taxonomicName>
<taxonomicName id="4C3CD58EC8374D1EFE64ACC3FE5AFB13" box="[400,490,1153,1177]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFE64ACC3FE5AFB13" box="[400,490,1153,1177]" italics="true" pageId="0" pageNumber="295">yanshini</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="EFADD3FCC8374D1EFDDBACC3FCB9FB13" author="Funston, G. F. &amp; Mendonca, S. E. &amp; Currie, P. J. &amp; Barsbold, R." box="[559,777,1153,1177]" pageId="0" pageNumber="295" pagination="101 - 120" refId="ref672" refString="Funston, G. F., Mendonca, S. E., Currie, P. J. &amp; Barsbold, R. (2018) Oviraptorosaur anatomy, diversity and ecology in the Nemegt Basin. Palaeogeography, Palaeoclimatology, Palaeoecology, 494, 101 - 120. https: // doi. org / 10.1016 / j. palaeo. 2017.10.023" type="journal article" year="2018">
Funston
<emphasis id="B948721FC8374D1EFD78ACC3FD73FB13" box="[652,707,1153,1177]" italics="true" pageId="0" pageNumber="295">et al.</emphasis>
, 2018
</bibRefCitation>
;
<bibRefCitation id="EFADD3FCC8374D1EFCE0ACC3FC6BFB13" author="Herne, M. C. &amp; Tait, A. M. &amp; Weisbecker, V. &amp; Hall, M. &amp; Nair, J. P. &amp; Cleeland, M. &amp; Salisbury, S. W." box="[788,987,1153,1177]" pageId="0" pageNumber="295" pagination="4113" refId="ref798" refString="Herne, M. C., Tait, A. M., Weisbecker, V., Hall, M., Nair, J. P., Cleeland, M. &amp; Salisbury, S. W. (2018) A new small-bodied ornithopod (Dinosauria, Ornithischia) from a deep, high-energy Early Cretaceous river of the Australian-Antarctic rift system. PeerJ, 5, e 4113. https: // doi. org / 10.7717 / peerj. 4113" type="journal article" year="2018">
Herne
<emphasis id="B948721FC8374D1EFCA9ACC3FC23FB13" box="[861,915,1153,1177]" italics="true" pageId="0" pageNumber="295">et al.</emphasis>
, 2018
</bibRefCitation>
). Thus,
<taxonomicName id="4C3CD58EC8374D1EFBC4ACC3FB15FB12" authorityName="Lu" authorityYear="2003" box="[1072,1189,1153,1176]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="genus">
<emphasis id="B948721FC8374D1EFBC4ACC3FB15FB12" box="[1072,1189,1153,1176]" italics="true" pageId="0" pageNumber="295">Heyuannia</emphasis>
</taxonomicName>
is the proper synonym available as a replacement name of the junior homonym. Therefore, the author proposes a new replacement subfamily name
<subSubSection id="C326FD86C8374D1EFF2CAC8BFE5BFB6B" box="[216,491,1225,1249]" pageId="0" pageNumber="295" type="nomenclature">
<taxonomicName id="4C3CD58EC8374D1EFF2CAC8BFED5FB6B" authorityName="Chan-Gyu" authorityYear="2019" box="[216,357,1225,1249]" class="Reptilia" family="Oviraptoridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="subFamily" status="clade. nov" subFamily="Heyuanninae">Heyuanninae</taxonomicName>
<taxonomicNameLabel id="A27BCF64C8374D1EFE9FAC8BFE6BFB6A" box="[363,475,1225,1249]" pageId="0" pageNumber="295">clade. nov</taxonomicNameLabel>
. (
</subSubSection>
<subSubSection id="C326FD86C8374D1EFE1FAC8BFC7FFB6B" box="[491,975,1225,1249]" pageId="0" pageNumber="295" type="etymology">
from
<taxonomicName id="4C3CD58EC8374D1EFDDCAC8BFD2DFB6A" authorityName="Lu" authorityYear="2003" box="[552,669,1225,1248]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="genus">
<emphasis id="B948721FC8374D1EFDDCAC8BFD2DFB6A" box="[552,669,1225,1248]" italics="true" pageId="0" pageNumber="295">Heyuannia</emphasis>
</taxonomicName>
and inae) for “Ingeniinae”.
</subSubSection>
</paragraph>
<subSubSection id="C326FD86C8374D1EFF33ACAFFD37FAFA" pageId="0" pageNumber="295" type="discussion">
<paragraph id="8B83AE0DC8374D1EFF33ACAFFD37FAFA" blockId="0.[151,1437,649,1393]" pageId="0" pageNumber="295">
<bibRefCitation id="EFADD3FCC8374D1EFF33ACAFFE0CFA8F" author="Osmolska, H. &amp; Currie, P. J. &amp; Barsbold, R." box="[199,444,1261,1285]" pageId="0" pageNumber="295" pagination="165 - 183" refId="ref1212" refString="Osmolska, H., Currie, P. J. &amp; Barsbold, R. (2004) Oviraptorosauria. In: Weishampel, D. B., Dodson, P. &amp; Osmolska, H. (Eds.), The Dinosauria. 2 nd Edition. University of California Press, Berkeley, pp. 165 - 183. https: // doi. org / 10.1525 / california / 9780520242098.003.0010" type="book chapter" year="2004">
Osmolska
<emphasis id="B948721FC8374D1EFECDACAFFEDFFA8F" box="[313,367,1261,1285]" italics="true" pageId="0" pageNumber="295">et al.</emphasis>
(2004)
</bibRefCitation>
defined “Ingeniinae” as
<taxonomicName id="4C3CD58EC8374D1EFD33ACAFFC06FA8F" authorityName="Barsbold" authorityYear="1986" box="[711,950,1261,1285]" class="Reptilia" family="Oviraptoridae" genus="Conchoraptor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B948721FC8374D1EFD33ACAFFC06FA8F" box="[711,950,1261,1285]" italics="true" pageId="0" pageNumber="295">Conchoraptor gracilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3CD58EC8374D1EFC30ACAFFBC6FA8F" authorityName="Barsbold" authorityYear="1981" box="[964,1142,1261,1285]" class="Adenophorea" family="Tripyloididae" genus="Ingenia" kingdom="Animalia" order="Araeolaimida" pageId="0" pageNumber="295" phylum="Nematoda" rank="species" species="yanshini">
<emphasis id="B948721FC8374D1EFC30ACAFFBC6FA8F" box="[964,1142,1261,1285]" italics="true" pageId="0" pageNumber="295">Ingenia yanshini</emphasis>
</taxonomicName>
, their most recent common ancestor, and all descendants. Here the author revises this definition for Heyuanninae, using
<taxonomicName id="4C3CD58EC8374D1EFB78AD53FAE4FAA3" authorityName="Lu" authorityYear="2002" box="[1164,1364,1297,1321]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="huangi">
<emphasis id="B948721FC8374D1EFB78AD53FAE4FAA3" box="[1164,1364,1297,1321]" italics="true" pageId="0" pageNumber="295">Heyuannia huangi</emphasis>
</taxonomicName>
as the specifier. Heyuanninae is a node-based subfamily-level clade that can be defined as the least inclusive clade containing
<taxonomicName id="4C3CD58EC8374D1EFF63AD1BFEEDFAFB" authorityName="Lu" authorityYear="2002" box="[151,349,1369,1393]" class="Reptilia" family="Oviraptoridae" genus="Heyuannia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="huangi">
<emphasis id="B948721FC8374D1EFF63AD1BFEEDFAFB" box="[151,349,1369,1393]" italics="true" pageId="0" pageNumber="295">Heyuannia huangi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3CD58EC8374D1EFE65AD1BFD30FAFB" authorityName="Barsbold" authorityYear="1986" box="[401,640,1369,1393]" class="Reptilia" family="Oviraptoridae" genus="Conchoraptor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="295" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B948721FC8374D1EFE65AD1BFD30FAFB" box="[401,640,1369,1393]" italics="true" pageId="0" pageNumber="295">Conchoraptor gracilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>