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<document id="5F61020DD143CDD23E4FCDABBCF3C98B" ID-CLB-Dataset="28347" ID-DOI="10.11646/zootaxa.4567.3.1" ID-GBIF-Dataset="47813581-d217-4a19-8848-a3e9cf2ddaf4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2598945" ID-ZooBank="0AEFAF80-B001-4A18-88AC-5B6A189F6DCD" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553062614686" checkinUser="plazi" docAuthor="Staples, David A." docDate="2019" docId="03895C3329314F25FF01FC07FA8DFDBF" docLanguage="en" docName="zootaxa.4567.3.1.pdf" docOrigin="Zootaxa 4567 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hedgpethia shalei Staples 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="435" masterDocId="FFB0244B29114F07FF96FFB2FF80FFFD" masterDocTitle="Pycnogonids (Arthropoda, Pycnogonida) from the Southwest Indian Ridge" masterLastPageNumber="449" masterPageNumber="401" pageNumber="433" updateTime="1698701998649" updateUser="ExternalLinkService">
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<mods:title id="AF66C0E5B5D56FA04FB8BD47FD89D412">Pycnogonids (Arthropoda, Pycnogonida) from the Southwest Indian Ridge</mods:title>
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<mods:namePart id="4AAB3B1F2012A8AEF8A34ABB7511CCE9">Staples, David A.</mods:namePart>
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<mods:date id="8A89C9CFFB4C0105606F8D59650C216E">2019</mods:date>
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<treatment id="03895C3329314F25FF01FC07FA8DFDBF" ID-DOI="http://doi.org/10.5281/zenodo.5944910" ID-GBIF-Taxon="154435133" ID-Zenodo-Dep="5944910" LSID="urn:lsid:plazi:treatment:03895C3329314F25FF01FC07FA8DFDBF" httpUri="http://treatment.plazi.org/id/03895C3329314F25FF01FC07FA8DFDBF" lastPageId="34" lastPageNumber="435" pageId="32" pageNumber="433">
<subSubSection id="C33ABEAE29314F27FF01FC07FE57FC32" box="[151,471,949,975]" pageId="32" pageNumber="433" type="nomenclature">
<paragraph id="8B9FED2529314F27FF01FC07FE57FC32" blockId="32.[151,471,949,1009]" box="[151,471,949,975]" pageId="32" pageNumber="433">
<heading id="D0D75A4929314F27FF01FC07FE57FC32" bold="true" box="[151,471,949,975]" fontSize="11" level="1" pageId="32" pageNumber="433" reason="1">
<emphasis id="B954313729314F27FF01FC07FE57FC32" bold="true" box="[151,471,949,975]" pageId="32" pageNumber="433">
<taxonomicName id="4C2096A629314F27FF01FC07FEF1FC32" ID-CoL="92f00a9e-234b-41d7-b777-7fe8ca36996b" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[151,369,949,975]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="shalei" status="sp. nov.">
<emphasis id="B954313729314F27FF01FC07FEF1FC32" bold="true" box="[151,369,949,975]" italics="true" pageId="32" pageNumber="433">Hedgpethia shalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678C4C29314F27FEEFFC07FE57FC32" box="[377,471,949,975]" pageId="32" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33ABEAE29314F27FF01FC6AFE48FC0D" box="[151,456,984,1009]" pageId="32" pageNumber="433" type="description">
<paragraph id="8B9FED2529314F27FF01FC6AFE48FC0D" blockId="32.[151,471,949,1009]" box="[151,456,984,1009]" pageId="32" pageNumber="433">
<figureCitation id="131BF1A029314F27FF01FC6AFE81FC0C" box="[151,257,984,1009]" captionStart="FIGURE 16" captionStartId="33.[151,250,1951,1973]" captionTargetBox="[161,1402,311,1813]" captionTargetId="figure@33.[150,1436,301,1929]" captionTargetPageId="33" captionText="FIGURE 16. Hedgpethia shalei sp. nov., holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw." httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="32" pageNumber="433">Figure 16</figureCitation>
AG;
<figureCitation id="131BF1A029314F27FEDEFC6AFE48FC0D" box="[328,456,984,1009]" captionStart="PLATE 5" captionStartId="44.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@44.[151,1436,193,1718]" captionTargetPageId="44" captionText="PLATE 5. Colossendeis sp. A, proboscis lateral; B, palp. Colossendeis melancholicus. C, specimen on rock wall. D, specimen with proboscis inserted into Isactinernus quadrilobatus. Hedgpethia filamentus sp. nov., holotype. E, trunk, lateral view; F, trunk, dorsal view. Hedgpethia shalei sp. nov., holotype. G, trunk, lateral view; H, trunk, dorsal view." httpUri="https://zenodo.org/record/2598997/files/figure.png" pageId="32" pageNumber="433">Plate 5G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33ABEAE29314F27FF01FB92FBE2FA80" pageId="32" pageNumber="433" type="materials_examined">
<paragraph id="8B9FED2529314F27FF01FB92FA84FB7C" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">
<emphasis id="B954313729314F27FF01FB92FE01FBC4" bold="true" box="[151,385,1056,1081]" pageId="32" pageNumber="433">Material examined.</emphasis>
<materialsCitation id="3B48E77829314F27FE1CFB92FA84FB7C" ID-GBIF-Occurrence="2013824397" collectingDate="2011-12-02" collectionCode="NHMUK" elevation="1414" location="Southwest Indian Ocean" municipality="Middle of What Seamount" pageId="32" pageNumber="433" specimenCode="NHMUK 2018.41" specimenCount="1" typeStatus="holotype">
<typeStatus id="549B538729314F27FE1CFB92FE75FBC4" box="[394,501,1056,1081]" pageId="32" pageNumber="433" type="holotype">Holotype</typeStatus>
, sex undetermined (
<specimenCode id="DB86455E29314F27FD4CFB92FC2DFBC5" box="[730,941,1056,1080]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="433" type="Museum">NHMUK 2018.41</specimenCode>
),
<location id="8EFFBBFE29314F27FC54FB92FB59FBC5" LSID="urn:lsid:plazi:treatment:03895C3329314F25FF01FC07FA8DFDBF:8EFFBBFE29314F27FC54FB92FB59FBC5" box="[962,1241,1056,1081]" municipality="Middle of What Seamount" name="Southwest Indian Ocean" pageId="32" pageNumber="433">Southwest Indian Ocean</location>
,
<collectingMunicipality id="6BFB775F29314F27FB70FB92FE8AFBA0" pageId="32" pageNumber="433">Middle of What Seamount</collectingMunicipality>
, 37° 56. 38.1´S, 50° 27. 16.9´E, ROV,
<quantity id="4CD840C029314F27FD2BFBF7FC94FBA1" box="[701,788,1093,1117]" metricMagnitude="3" metricUnit="m" metricValue="1.414" pageId="32" pageNumber="433" unit="m" value="1414.0">
<elevation id="000D0A1629314F27FD2BFBF7FC94FBA1" box="[701,788,1093,1117]" metricMagnitude="3" metricUnit="m" metricValue="1.414" pageId="32" pageNumber="433" unit="m" value="1414.0">1414 m</elevation>
</quantity>
, specimen JC066-3522, stn 6.7, parent 2593,
<date id="FF9ECBE529314F27FA82FBF7FF52FB7D" pageId="32" pageNumber="433" value="2011-12-02">
<collectingDate id="EFDA320D29314F27FA82FBF7FF52FB7D" pageId="32" pageNumber="433" value="2011-12-02">2 December 2011</collectingDate>
</date>
. Photographed in aquarium on dead coral in association with yellow and orange
<emphasis id="B954313729314F27FBCEFBDAFA84FB7C" box="[1112,1284,1128,1153]" italics="true" pageId="32" pageNumber="433">Parazoanthids.</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B9FED2529314F27FF51FB3EFE6AFB10" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">
<materialsCitation id="3B48E77829314F27FF51FB3EFE67FB10" ID-GBIF-Occurrence="2013824394" collectingDate="2011-11-20" collectionCode="NHMUK" elevation="732" pageId="32" pageNumber="433" specimenCode="NHMUK 2018.42" specimenCount="1" typeStatus="paratype">
<emphasis id="B954313729314F27FF51FB3EFEBBFB58" bold="true" box="[199,315,1164,1189]" pageId="32" pageNumber="433">
<typeStatus id="549B538729314F27FF51FB3EFEB6FB58" box="[199,310,1164,1189]" pageId="32" pageNumber="433" type="paratype">Paratype</typeStatus>
.
</emphasis>
One specimen (
<specimenCode id="DB86455E29314F27FE6AFB3FFD52FB58" box="[508,722,1165,1189]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="433" type="Museum">NHMUK 2018.42</specimenCode>
), Southwest Indian Ocean, Coral Seamount, 41˚ 22.31'S, 42˚ 54.57'E, ROV,
<quantity id="4CD840C029314F27FED5FB02FE0EFB35" box="[323,398,1200,1224]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="32" pageNumber="433" unit="m" value="732.0">
<elevation id="000D0A1629314F27FED5FB02FE0EFB35" box="[323,398,1200,1224]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="32" pageNumber="433" unit="m" value="732.0">732 m</elevation>
</quantity>
, specimen JC066-873, stn 4.38, parent 1272, mooring site,
<collectingMethod id="5261953229314F27FBA1FB03FBFEFB35" box="[1079,1150,1201,1224]" pageId="32" pageNumber="433">on net</collectingMethod>
containing Mango wood packages,
<date id="FF9ECBE529314F27FE9AFB67FE67FB10" box="[268,487,1236,1261]" pageId="32" pageNumber="433" value="2011-11-20">
<collectingDate id="EFDA320D29314F27FE9AFB67FE67FB10" box="[268,487,1236,1261]" pageId="32" pageNumber="433" value="2011-11-20">20 November 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9FED2529314F27FF51FB4AFBE2FA80" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">
<emphasis id="B954313729314F27FF51FB4AFE03FAEC" bold="true" box="[199,387,1272,1297]" pageId="32" pageNumber="433">Other material.</emphasis>
One specimen (NHMUK 2018.43), Southwest Indian Ocean, Coral Seamount, 41˚ 22.31'S, 42˚ 54.57'E, ROV,
<quantity id="4CD840C029314F27FEE4FAAFFE3DFAC9" box="[370,445,1309,1333]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="32" pageNumber="433" unit="m" value="732.0">732 m</quantity>
, specimen JC066-1298D, parent 1272, stn 4.38, mooring site, on net containing whale bone,
<date id="FF9ECBE529314F27FF4DFAF2FE34FAA5" box="[219,436,1344,1369]" pageId="32" pageNumber="433" value="2011-11-20">20 November 2011</date>
.
<materialsCitation id="3B48E77829314F27FE28FAF2FBDEFA80" ID-GBIF-Occurrence="2013824391" collectingDate="2011-11-20" collectionCode="NHMUK" elevation="687" latitude="-41.357666" pageId="32" pageNumber="433" specimenCode="NHMUK 2018.44" specimenCount="1">
One specimen (
<specimenCode id="DB86455E29314F27FDFDFAF2FCBCFAA5" box="[619,828,1344,1368]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="433" type="Museum">NHMUK 2018.44</specimenCode>
), Southwest Indian Ocean, Coral Seamount,
<geoCoordinate id="EE148BE229314F27FAA6FAF2FF2CFA80" direction="south" orientation="latitude" pageId="32" pageNumber="433" precision="9" value="-41.357666">41° 21.46 'S</geoCoordinate>
, 42˚ 54.54 'E,
<quantity id="4CD840C029314F27FEC6FAD6FE19FA81" box="[336,409,1380,1405]" metricMagnitude="2" metricUnit="m" metricValue="6.87" pageId="32" pageNumber="433" unit="m" value="687.0">
<elevation id="000D0A1629314F27FEC6FAD6FE19FA81" box="[336,409,1380,1405]" metricMagnitude="2" metricUnit="m" metricValue="6.87" pageId="32" pageNumber="433" unit="m" value="687.0">687 m</elevation>
</quantity>
, specimen JC066-3109, stn 4, parent 2542,
<date id="FF9ECBE529314F27FC15FAD7FBDEFA80" box="[899,1118,1380,1405]" pageId="32" pageNumber="433" value="2011-11-20">
<collectingDate id="EFDA320D29314F27FC15FAD7FBDEFA80" box="[899,1118,1380,1405]" pageId="32" pageNumber="433" value="2011-11-20">20 November 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33ABEAE29314F25FF51FA3AFF5DFE73" lastPageId="34" lastPageNumber="435" pageId="32" pageNumber="433" type="description">
<paragraph id="8B9FED2529314F27FF51FA3AFE95F9CC" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">
<emphasis id="B954313729314F27FF51FA3AFED9FA5C" bold="true" box="[199,345,1416,1441]" pageId="32" pageNumber="433">Description.</emphasis>
<typeStatus id="549B538729314F27FEC9FA3AFE47FA5C" box="[351,455,1416,1441]" pageId="32" pageNumber="433" type="holotype">Holotype</typeStatus>
. Leg span about
<quantity id="4CD840C029314F27FD1EFA3AFD5AFA5D" box="[648,730,1416,1441]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="433" unit="mm" value="15.0">15 mm</quantity>
. Trunk (
<figureCitation id="131BF1A029314F27FCAEFA3AFC38FA5D" box="[824,952,1416,1441]" captionStart="FIGURE 16" captionStartId="33.[151,250,1951,1973]" captionTargetBox="[161,1402,311,1813]" captionTargetId="figure@33.[150,1436,301,1929]" captionTargetPageId="33" captionText="FIGURE 16. Hedgpethia shalei sp. nov., holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw." httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="32" pageNumber="433">Fig.16A, B</figureCitation>
.
<figureCitation id="131BF1A029314F27FC52FA3AFBC5FA5D" box="[964,1093,1416,1441]" captionStart="PLATE 5" captionStartId="44.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@44.[151,1436,193,1718]" captionTargetPageId="44" captionText="PLATE 5. Colossendeis sp. A, proboscis lateral; B, palp. Colossendeis melancholicus. C, specimen on rock wall. D, specimen with proboscis inserted into Isactinernus quadrilobatus. Hedgpethia filamentus sp. nov., holotype. E, trunk, lateral view; F, trunk, dorsal view. Hedgpethia shalei sp. nov., holotype. G, trunk, lateral view; H, trunk, dorsal view." httpUri="https://zenodo.org/record/2598997/files/figure.png" pageId="32" pageNumber="433">Plate 5G, H</figureCitation>
) slender, segmented, posterior margin of each segment raised to form a low cowl; lateral processes slightly constricted in mid-region, widest distally, separated by about their own basal width. Ocular tubercle placed on mid-region of cephalon, tall, evenly tapered from base, acuminate, over-reaching base of proboscis; four large eyes of equal size near tubercle base, pigmented.
</paragraph>
<paragraph id="8B9FED2529314F27FF51F98EFB66F984" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">
Proboscis bulbous (
<figureCitation id="131BF1A029314F27FE3BF98EFD98F9A8" box="[429,536,1596,1621]" captionStart="FIGURE 16" captionStartId="33.[151,250,1951,1973]" captionTargetBox="[161,1402,311,1813]" captionTargetId="figure@33.[150,1436,301,1929]" captionTargetPageId="33" captionText="FIGURE 16. Hedgpethia shalei sp. nov., holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw." httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="32" pageNumber="433">Fig. 16E</figureCitation>
), narrowest proximally, expanding evenly to reach maximum width in midregion, slight constriction at about three-quarters length, gently curved downwards, rounded at tip.
</paragraph>
<paragraph id="8B9FED2529314F27FF51F936FC84F93C" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">Abdomen articulated at base, very short, implanted between fourth lateral processes, barely visible dorsally, in lateral view seen extending ventrally beneath the trunk.</paragraph>
<paragraph id="8B9FED2529314F27FF51F97EFD1AF8F5" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">
Palp (
<figureCitation id="131BF1A029314F27FE98F97EFEEDF918" box="[270,365,1740,1765]" captionStart="FIGURE 16" captionStartId="33.[151,250,1951,1973]" captionTargetBox="[161,1402,311,1813]" captionTargetId="figure@33.[150,1436,301,1929]" captionTargetPageId="33" captionText="FIGURE 16. Hedgpethia shalei sp. nov., holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw." httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="32" pageNumber="433">Fig.16D</figureCitation>
) ten-segmented, segment 3 longest, about 1.8 times longer than next longest segment 5, segments 6 to10 with long, fine, ventral setae.
</paragraph>
<paragraph id="8B9FED2529314F27FF51F8A6FE12F8D0" blockId="32.[151,1437,1056,2017]" box="[199,402,1812,1837]" pageId="32" pageNumber="433">Chelifores absent.</paragraph>
<paragraph id="8B9FED2529314F27FF51F88AFD9EF864" blockId="32.[151,1437,1056,2017]" pageId="32" pageNumber="433">
Oviger (
<figureCitation id="131BF1A029314F27FEB1F88AFE01F8AC" box="[295,385,1848,1873]" captionStart="FIGURE 16" captionStartId="33.[151,250,1951,1973]" captionTargetBox="[161,1402,311,1813]" captionTargetId="figure@33.[150,1436,301,1929]" captionTargetPageId="33" captionText="FIGURE 16. Hedgpethia shalei sp. nov., holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw." httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="32" pageNumber="433">Fig.16F</figureCitation>
) ten-segmented, segment 4 longest, claw leaf-like shape (
<figureCitation id="131BF1A029314F27FBB5F88AFB0EF8AC" box="[1059,1166,1848,1873]" captionStart="FIGURE 16" captionStartId="33.[151,250,1951,1973]" captionTargetBox="[161,1402,311,1813]" captionTargetId="figure@33.[150,1436,301,1929]" captionTargetPageId="33" captionText="FIGURE 16. Hedgpethia shalei sp. nov., holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw." httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="32" pageNumber="433">Fig. 16G</figureCitation>
), spines on segments 7 to10 are in several rows but the number of rows and detail of the distal margin of the spines are unclear but appear to be spatulate shaped and smooth.
</paragraph>
<paragraph id="8B9FED2529314F26FF51F816FF71FF05" blockId="32.[151,1437,1056,2017]" lastBlockId="33.[151,1436,151,248]" lastPageId="33" lastPageNumber="434" pageId="32" pageNumber="433">
Third leg (
<figureCitation id="131BF1A029314F27FEDBF816FE2AF840" box="[333,426,1956,1981]" captionStart="FIGURE 16" captionStartId="33.[151,250,1951,1973]" captionTargetBox="[161,1402,311,1813]" captionTargetId="figure@33.[150,1436,301,1929]" captionTargetPageId="33" captionText="FIGURE 16. Hedgpethia shalei sp. nov., holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw." httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="32" pageNumber="433">Fig.16C</figureCitation>
) slender, femur gently curved, widest in mid-region, femur and tibiae with single, prominent, dorsodistal seta about twice segment width, dorsal surfaces more setose generally, tibia 1 longest, femur longer than tibia 2, tarsus slightly more than half length of propodus, 15 to 20 sole spines, spines undifferentiated, terminal claw about one-third length of propodus, auxiliary claws absent. Cement glands and gonopores not evident.
</paragraph>
<caption id="DF5FBDAD29304F26FF01F82DFD72F82E" httpUri="https://zenodo.org/record/2598975/files/figure.png" pageId="33" pageNumber="434" startId="33.[151,250,1951,1973]" targetBox="[161,1402,311,1813]" targetPageId="33">
<paragraph id="8B9FED2529304F26FF01F82DFD72F82E" blockId="33.[151,1435,1951,2003]" pageId="33" pageNumber="434">
<emphasis id="B954313729304F26FF01F82DFEA0F848" bold="true" box="[151,288,1951,1973]" pageId="33" pageNumber="434">FIGURE 16.</emphasis>
<taxonomicName id="4C2096A629304F26FEBEF82DFE61F848" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[296,481,1951,1973]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="33" pageNumber="434" phylum="Arthropoda" rank="species" species="shalei" status="sp. nov.">
<emphasis id="B954313729304F26FEBEF82DFE61F848" box="[296,481,1951,1973]" italics="true" pageId="33" pageNumber="434">Hedgpethia shalei</emphasis>
</taxonomicName>
<emphasis id="B954313729304F26FE7FF82DFDBAF848" bold="true" box="[489,570,1951,1973]" pageId="33" pageNumber="434">
<taxonomicNameLabel id="A2678C4C29304F26FE7FF82DFDBAF848" box="[489,570,1951,1973]" pageId="33" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype. A, trunk, lateral view; B, trunk, dorsal view; C, third leg; D, left palp; E, proboscis, anterior view; F, oviger; G, oviger terminal claw.
</paragraph>
</caption>
<paragraph id="8B9FED2529334F25FF51FF29FF5DFE73" blockId="34.[151,1437,151,578]" pageId="34" pageNumber="435">
Measurements of
<typeStatus id="549B538729334F25FE18FF28FE70FF4E" box="[398,496,154,179]" pageId="34" pageNumber="435" type="holotype">holotype</typeStatus>
(mm
<emphasis id="B954313729334F25FDBAFF28FDBBFF4E" bold="true" box="[556,571,154,179]" pageId="34" pageNumber="435">).</emphasis>
Trunk length (frontal margin of cephalic segment to tip of 4
<superScript id="7C55406D29334F25FB4FFF25FB65FF58" attach="left" box="[1241,1253,151,165]" fontSize="6" pageId="34" pageNumber="435">th</superScript>
lateral process), 1.584; length cephalon, 0.624; width across 2
<superScript id="7C55406D29334F25FD0BFF0FFD2DFF36" attach="left" box="[669,685,189,203]" fontSize="6" pageId="34" pageNumber="435">nd</superScript>
lateral processes, 0.888; proboscis length (dorsal), 1.232; greatest diameter proboscis, 0.600. Third leg: coxa 1, 0.200; coxa 2, 0.240; coxa 3, 0.160; femur, 1.808; tibia 1, 2.192; tibia 2, 1.544; tarsus, 0.328; propodus, 0.600; claw, 0.200. Palp: seg. 1, 0.104; seg. 2, 0.064; seg. 3, 1.264; seg. 4, 0.168; seg. 5, 0.712; seg. 6, 0.104; seg. 7, 0.224; seg. 8, 0.184; seg. 9, 0.184; seg. 10, 0.176. Oviger; seg. 2, 0.160; seg. 3, 0.136; seg. 4, 0.880; seg. 5, 0.208; seg. 6, 0.832; seg. 7, 0.344; seg 8, 0.264; seg. 9, 0.224; seg.10, 0.240; claw, 0.088.
</paragraph>
</subSubSection>
<subSubSection id="C33ABEAE29334F25FF51FE2BFAA3FE2B" pageId="34" pageNumber="435" type="etymology">
<paragraph id="8B9FED2529334F25FF51FE2BFAA3FE2B" blockId="34.[151,1437,151,578]" pageId="34" pageNumber="435">
<emphasis id="B954313729334F25FF51FE2BFECEFE4F" bold="true" box="[199,334,409,434]" pageId="34" pageNumber="435">Etymology.</emphasis>
The specific name acknowledges the contribution made by Dr. David Shale, expedition and natural history photographer for his outstanding work capturing images of the voyage collections fauna.
</paragraph>
</subSubSection>
<subSubSection id="C33ABEAE29334F25FF51FE53FA8DFDBF" pageId="34" pageNumber="435" type="discussion">
<paragraph id="8B9FED2529334F25FF51FE53FA8DFDBF" blockId="34.[151,1437,151,578]" pageId="34" pageNumber="435">
<emphasis id="B954313729334F25FF51FE53FEBBFE07" bold="true" box="[199,315,481,506]" pageId="34" pageNumber="435">Remarks.</emphasis>
This species is most closely related to
<taxonomicName id="4C2096A629334F25FD78FE50FC06FE07" authorityName="Staples" authorityYear="2019" box="[750,902,481,506]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="filamentus">
<emphasis id="B954313729334F25FD78FE50FC06FE07" box="[750,902,481,506]" italics="true" pageId="34" pageNumber="435">H. filamentus</emphasis>
</taxonomicName>
but is noticeably more robust. Further points of difference are in the more rounded proboscis with the basal part being shorter and less defined, the dorsoposterior margins of trunk segments 1 to 3 raised into low cowls and the lateral processes being closer together.
</paragraph>
</subSubSection>
</treatment>
</document>