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<document id="ABEE6734286C62506F41008AB0C2E4C6" ID-DOI="10.1051/acarologia/20152170" ID-ISSN="2107-7207" ID-Zenodo-Dep="5399491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630332251020" checkinUser="marcus" docAuthor="Bertrand, M., Sidorchuk, E. &amp; Hoffeins, C." docDate="2015" docId="0381FC5BC662FFF4AB923FD8FE5BF871" docLanguage="en" docName="Acarologia.2015.55.3.321-336.pdf" docOrigin="Acarologia 55 (3)" docSource="http://dx.doi.org/10.1051/acarologia/20152170" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Eunicolina glaesi Bertrand &amp; Sidorchuk &amp; Hoffeins 2015, n. sp." docType="treatment" docVersion="3" lastPageNumber="329" masterDocId="FFB88423C661FFFDA84F3A79FFAFFFED" masterDocTitle="Before the summer turns to winter: the third labidostommatid genus from Baltic amber has subtropical kin" masterLastPageNumber="336" masterPageNumber="321" pageNumber="323" updateTime="1699231175590" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="222303A0EF4155AC23F7054ADC09B8B1">Before the summer turns to winter: the third labidostommatid genus from Baltic amber has subtropical kin</mods:title>
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<mods:namePart id="102B856446A1E789F8F135223E75232F">Bertrand, M.</mods:namePart>
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<mods:namePart id="7781139B8634D75EE90B48207AFE8D5C">Sidorchuk, E.</mods:namePart>
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<mods:namePart id="0F795B1CC1B0FE0E7687F9A144457496">Hoffeins, C.</mods:namePart>
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<mods:title id="6F9C4DAE88FC5BBC78B2205E3D1EB4CA">Acarologia</mods:title>
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<mods:part id="A15B2A7661BB3CC98C81BB56CDA4A3A7">
<mods:date id="993D9C9F497B5C62770304A87DCCD40B">2015</mods:date>
<mods:detail id="1C26568FBF69C954983FE2AB059D3E6B" type="pubDate">
<mods:number id="4B69DF9FFEC3DFF311CB6BA9A646693C">2015-09-30</mods:number>
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<mods:number id="81D998CD1176C40328199F615818340D">55</mods:number>
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<mods:number id="B7169A66A0018ED63DE345A4CD11FA56">3</mods:number>
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<mods:classification id="770FBB46AC99A55D00C09DAB8D53B330">journal article</mods:classification>
<mods:identifier id="8B7F499EAEA8B79EAAE3A8F6C93A5619" type="DOI">10.1051/acarologia/20152170</mods:identifier>
<mods:identifier id="4E0EDC95A0F14E0008BA44A9A70B6396" type="ISSN">2107-7207</mods:identifier>
<mods:identifier id="EA05B32609524A651A0E38B0F5111CB4" type="Zenodo-Dep">5399491</mods:identifier>
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<treatment id="0381FC5BC662FFF4AB923FD8FE5BF871" LSID="urn:lsid:plazi:treatment:0381FC5BC662FFF4AB923FD8FE5BF871" httpUri="http://treatment.plazi.org/id/0381FC5BC662FFF4AB923FD8FE5BF871" lastPageId="9" lastPageNumber="329" pageId="3" pageNumber="323">
<subSubSection id="C3321EC6C662FFFEAB923FD8FB5DFA57" box="[989,1266,1441,1466]" pageId="3" pageNumber="323" type="nomenclature">
<paragraph id="8B974D4DC662FFFEAB923FD8FB5DFA57" blockId="3.[989,1266,1441,1501]" box="[989,1266,1441,1466]" pageId="3" pageNumber="323">
<heading id="D0DFFA21C662FFFEAB923FD8FB5DFA57" bold="true" box="[989,1266,1441,1466]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="323" reason="2">
<emphasis id="B95C915FC662FFFEAB923FD8FB5DFA57" bold="true" box="[989,1266,1441,1466]" pageId="3" pageNumber="323">
<taxonomicName id="4C2836CEC662FFFEAB923FD8FB06FA54" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[989,1193,1441,1465]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="3" pageNumber="323" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC662FFFEAB923FD8FB06FA54" bold="true" box="[989,1193,1441,1465]" italics="true" pageId="3" pageNumber="323">Eunicolina glaesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F2C24C662FFFEACFF3FDBFB5DFA57" box="[1200,1266,1442,1466]" pageId="3" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3321EC6C662FFF4AC4C3FBDFE5BF871" lastPageId="9" lastPageNumber="329" pageId="3" pageNumber="323" type="description">
<paragraph id="8B974D4DC662FFFEAC4C3FBDFB64FA31" blockId="3.[989,1266,1441,1501]" box="[1027,1227,1476,1501]" pageId="3" pageNumber="323">
<emphasis id="B95C915FC662FFFEAC4C3FBDFB64FA31" bold="true" box="[1027,1227,1476,1501]" pageId="3" pageNumber="323">
(
<figureCitation id="131351C8C662FFFEAC443FBDFBC6FA30" box="[1035,1129,1476,1501]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStart-4="FIGURE" captionStartId-0="3.[159,171,1095,1114]" captionStartId-1="4.[159,171,1716,1735]" captionStartId-2="5.[159,171,1789,1808]" captionStartId-3="6.[159,171,862,881]" captionStartId-4="6.[159,171,1853,1872]" captionTargetBox-0="[257,1346,302,1056]" captionTargetBox-2="[296,1181,368,1706]" captionTargetBox-3="[355,1235,311,834]" captionTargetBox-4="[355,1232,957,1821]" captionTargetId-0="figure-4@3.[242,1346,302,1063]" captionTargetId-2="figure-4@5.[280,1308,350,1758]" captionTargetId-3="figure-4@6.[355,1235,311,834]" captionTargetId-4="figure-43@6.[355,1232,957,1821]" captionTargetPageId-0="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionText-0="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." captionText-1="FIGURE 2: Eunicolina glaesi n. sp.: A – Genu and tibia, first pair of legs, right side, dorsal view; B – First leg, left side, lateral view; C – Ventral view, detail of the proximal articles of leg I, II and acetabulum of fourth leg (the leg IV is not represented here); D – The tridactylous tip of the tarsus II, lateral view. Abbreviations: epI: first epimeral plate; ge: genu; PI, PII, PIII, PIV: Legs 1 to 4; ti: tibia." captionText-2="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." captionText-3="FIGURE 4: Eunicolina glaesi n. sp.: A – The holotype piece, CHHC 588-13 (Hoffeins collection); B – Lateral view of the specimen of Eunicolina glaesi sp.nov.; C – Ventral view of infracapitulum and leg I." captionText-4="FIGURE 5: Eunicolina glaesi n. sp.: A – Oculo-pustular zone. bo.p: trichobthria, la?: supposed insertion of the la seta; lb: seta lb; ly: post-ocular lyriform organ; B – Detail of the PI tarsus showing the bidactylus claw, one of the two tarsal solnidia (ω). F?: possible famulus of the tarsus. Some insertions of setae are marked by circles." figureDoi-0="http://doi.org/10.5281/zenodo.5399493" figureDoi-2="http://doi.org/10.5281/zenodo.5399496" figureDoi-3="http://doi.org/10.5281/zenodo.5399500" figureDoi-4="http://doi.org/10.5281/zenodo.5399502" httpUri-0="https://zenodo.org/record/5399493/files/figure.png" httpUri-2="https://zenodo.org/record/5399496/files/figure.png" httpUri-3="https://zenodo.org/record/5399500/files/figure.png" httpUri-4="https://zenodo.org/record/5399502/files/figure.png" pageId="3" pageNumber="323">Figs 1-5</figureCitation>
,
<figureCitation id="131351C8C662FFFEAC393FBCFB6BFA31" box="[1142,1220,1476,1501]" captionStart="FIGURE" captionStartId="7.[159,171,1248,1267]" captionTargetBox="[281,1309,329,1221]" captionTargetId="figure-4@7.[281,1309,300,1221]" captionTargetPageId="7" captionText="FIGURE 6: Genito-anal area: A – Disposition of genital and anal shields (drawn from the extant species E. travei) Aa, Ab: ventral view, female (a), male (b); Ac: male lateral view. Eunicolina glaesi n. sp.: B-C – extremity of the opisthosoma, lateral view (B) and interpretation (Ca, Cb, Cc)." figureDoi="http://doi.org/10.5281/zenodo.5399504" httpUri="https://zenodo.org/record/5399504/files/figure.png" pageId="3" pageNumber="323">6B, 6C</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B974D4DC662FFFEAB753C7AFAADF871" blockId="3.[826,1429,1539,1948]" pageId="3" pageNumber="323">
<materialsCitation id="3B404710C662FFFEAB753C7AFB51F871" collectionCode="CHHC" collectorName="Baltic &amp; C. &amp; H. W. Hoffeins Collection" county="Museum" location="This" municipality="Naturkunde" pageId="3" pageNumber="323" specimenCode="CHHC 588-13" specimenCount="1" stateProvince="Berlin" typeStatus="holotype">
Type designation and repository — The
<typeStatus id="5493F3EFC662FFFEAD633C7AFA3BF9F6" box="[1324,1428,1539,1563]" pageId="3" pageNumber="323" type="holotype">holotype</typeStatus>
and only described specimen is an inclusion in
<collectorName id="26DD289BC662FFFEAB753C30FCD4F98C" box="[826,891,1609,1633]" pageId="3" pageNumber="323">Baltic</collectorName>
amber from the private
<collectorName id="26DD289BC662FFFEAC893C30FC57F969" pageId="3" pageNumber="323">C. &amp; H.W. Hoffeins Collection</collectorName>
(specimen
<specimenCode id="DB8EE536C662FFFEACC43C15FA97F969" box="[1163,1336,1644,1668]" collectionCode="CHHC" pageId="3" pageNumber="323">CHHC 588-13</specimenCode>
, which will be eventually transferred to the
<collectingCounty id="62F635C1C662FFFEACB13CE9FAC8F94A" box="[1278,1383,1680,1703]" pageId="3" pageNumber="323">Museum</collectingCounty>
für
<collectingMunicipality id="6BF3D737C662FFFEAB753CCBFC61F927" box="[826,974,1714,1738]" pageId="3" pageNumber="323">Naturkunde</collectingMunicipality>
,
<collectingRegion id="49EC83AFC662FFFEABAF3CCBFB84F927" box="[992,1067,1714,1738]" country="Germany" name="Berlin" pageId="3" pageNumber="323">Berlin</collectingRegion>
).
<location id="8EF71B96C662FFFEAC013CCBFB2EF927" LSID="urn:lsid:plazi:treatment:0381FC5BC662FFF4AB923FD8FE5BF871:8EF71B96C662FFFEAC013CCBFB2EF927" box="[1102,1153,1714,1738]" county="Museum" municipality="Naturkunde" name="This" pageId="3" pageNumber="323" stateProvince="Berlin">This</location>
specimen is an adult, having the entire dorsal shield (nymphal stases of labidostommatids are less chitinized, with the dorsal shield divided into paired lateral and dorsolateral sclerotized zones), and, having the rounded genital ring (
<figureCitation id="131351C8C662FFFEAB9B3D18FBB0F894" box="[980,1055,1889,1913]" captionStart="FIGURE" captionStartId="7.[159,171,1248,1267]" captionTargetBox="[281,1309,329,1221]" captionTargetId="figure-4@7.[281,1309,300,1221]" captionTargetPageId="7" captionText="FIGURE 6: Genito-anal area: A – Disposition of genital and anal shields (drawn from the extant species E. travei) Aa, Ab: ventral view, female (a), male (b); Ac: male lateral view. Eunicolina glaesi n. sp.: B-C – extremity of the opisthosoma, lateral view (B) and interpretation (Ca, Cb, Cc)." figureDoi="http://doi.org/10.5281/zenodo.5399504" httpUri="https://zenodo.org/record/5399504/files/figure.png" pageId="3" pageNumber="323">Fig. 6</figureCitation>
), it is a male (labidostommatid females have an oval ano-genital ring)
</materialsCitation>
.
</paragraph>
<caption id="DF571DC5C665FFF9A8D03CCDFA2CF913" pageId="4" pageNumber="324" startId="4.[159,171,1716,1735]">
<paragraph id="8B974D4DC665FFF9A8D03CCDFA2CF913" blockId="4.[159,1429,1715,1790]" pageId="4" pageNumber="324">
FIGURE 2:
<taxonomicName id="4C2836CEC665FFF9A9473CCAFE32F92B" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[264,413,1715,1734]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="4" pageNumber="324" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC665FFF9A9473CCAFE32F92B" box="[264,413,1715,1734]" italics="true" pageId="4" pageNumber="324">Eunicolina glaesi</emphasis>
</taxonomicName>
<emphasis id="B95C915FC665FFF9A9EB3CCDFE73F92A" bold="true" box="[420,476,1716,1735]" pageId="4" pageNumber="324">
<taxonomicNameLabel id="A26F2C24C665FFF9A9EB3CCDFE73F92A" box="[420,476,1716,1735]" pageId="4" pageNumber="324" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A – Genu and tibia, first pair of legs, right side, dorsal view; B – First leg, left side, lateral view; C – Ventral view, detail of the proximal articles of leg I, II and acetabulum of fourth leg (the leg IV is not represented here); D – The tridactylous tip of the tarsus II, lateral view. Abbreviations:
<emphasis id="B95C915FC665FFF9AABD3C95FCA3F913" box="[754,780,1772,1790]" italics="true" pageId="4" pageNumber="324">epI</emphasis>
: first epimeral plate;
<emphasis id="B95C915FC665FFF9AB9C3C95FC49F913" box="[979,998,1772,1790]" italics="true" pageId="4" pageNumber="324">ge</emphasis>
: genu;
<emphasis id="B95C915FC665FFF9AC633C95FBEFF913" box="[1068,1088,1772,1790]" italics="true" pageId="4" pageNumber="324">PI</emphasis>
,
<emphasis id="B95C915FC665FFF9AC053C95FB3DF913" box="[1098,1170,1772,1790]" italics="true" pageId="4" pageNumber="324">PII, PIII</emphasis>
,
<emphasis id="B95C915FC665FFF9ACD23C95FB6FF913" box="[1181,1216,1772,1790]" italics="true" pageId="4" pageNumber="324">PIV</emphasis>
: Legs 1 to 4;
<emphasis id="B95C915FC665FFF9AD753C95FAE8F913" box="[1338,1351,1772,1790]" italics="true" pageId="4" pageNumber="324">ti</emphasis>
: tibia.
</paragraph>
</caption>
<caption id="DF571DC5C664FFF8A8D03C84FDCAF88E" ID-DOI="http://doi.org/10.5281/zenodo.5399496" ID-Zenodo-Dep="5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="5" pageNumber="325" startId="5.[159,171,1789,1808]" targetBox="[296,1181,368,1706]" targetPageId="5">
<paragraph id="8B974D4DC664FFF8A8D03C84FDCAF88E" blockId="5.[159,1429,1789,1892]" pageId="5" pageNumber="325">
FIGURE 3:.
<taxonomicName id="4C2836CEC664FFF8A9543C84FE1FF8FD" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[283,432,1789,1808]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="5" pageNumber="325" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC664FFF8A9543C84FE1FF8FD" box="[283,432,1789,1808]" italics="true" pageId="5" pageNumber="325">Eunicolina glaesi</emphasis>
</taxonomicName>
<emphasis id="B95C915FC664FFF8A9F73C84FE5DF8FD" bold="true" box="[440,498,1789,1808]" pageId="5" pageNumber="325">
<taxonomicNameLabel id="A26F2C24C664FFF8A9F73C84FE5DF8FD" box="[440,498,1789,1808]" pageId="5" pageNumber="325" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum;
<emphasis id="B95C915FC664FFF8ACA23D60FAA0F8C1" box="[1261,1295,1817,1836]" italics="true" pageId="5" pageNumber="325">bo.a</emphasis>
,
<emphasis id="B95C915FC664FFF8AD553D60FA92F8C1" box="[1306,1341,1817,1836]" italics="true" pageId="5" pageNumber="325">bo.p</emphasis>
: anterior and posterior bothridia;
<emphasis id="B95C915FC664FFF8A9E83D4CFE69F8A5" box="[423,454,1845,1864]" italics="true" pageId="5" pageNumber="325">CH</emphasis>
: Chelicera;
<emphasis id="B95C915FC664FFF8AA7A3D4CFDFDF8A5" box="[565,594,1845,1864]" italics="true" pageId="5" pageNumber="325">cha</emphasis>
,
<emphasis id="B95C915FC664FFF8AA123D4CFDD5F8A5" box="[605,634,1845,1864]" italics="true" pageId="5" pageNumber="325">chb</emphasis>
cheliceral setae;
<emphasis id="B95C915FC664FFF8AB563D4CFC83F8AA" box="[793,812,1845,1863]" italics="true" pageId="5" pageNumber="325">gr</emphasis>
,
<emphasis id="B95C915FC664FFF8AB783D4CFCE9F8A5" box="[823,838,1845,1864]" italics="true" pageId="5" pageNumber="325">la</emphasis>
: ocular and lateral setae of the dorsal shield;
<emphasis id="B95C915FC664FFF8ACBC3D4CFAA8F8AA" box="[1267,1287,1845,1863]" italics="true" pageId="5" pageNumber="325">PI</emphasis>
,
<emphasis id="B95C915FC664FFF8AD5D3D4CFA82F8AA" box="[1298,1325,1845,1863]" italics="true" pageId="5" pageNumber="325">PII</emphasis>
: legs I and II;
<emphasis id="B95C915FC664FFF8A8953D28FF5DF88E" box="[218,242,1873,1891]" italics="true" pageId="5" pageNumber="325">Pp</emphasis>
: palp.
<emphasis id="B95C915FC664FFF8A97C3D29FEEEF88E" box="[307,321,1872,1891]" italics="true" pageId="5" pageNumber="325">ω</emphasis>
1: tarsal solenidion of the palp.
</paragraph>
</caption>
<caption id="DF571DC5C667FFFBA8D03927FCE6FC60" ID-DOI="http://doi.org/10.5281/zenodo.5399500" ID-Zenodo-Dep="5399500" httpUri="https://zenodo.org/record/5399500/files/figure.png" pageId="6" pageNumber="326" startId="6.[159,171,862,881]" targetBox="[355,1235,311,834]" targetPageId="6">
<paragraph id="8B974D4DC667FFFBA8D03927FCE6FC60" blockId="6.[159,1429,862,909]" pageId="6" pageNumber="326">
FIGURE 4:
<taxonomicName id="4C2836CEC667FFFBA9443927FE0FFC9C" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[267,416,862,881]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="6" pageNumber="326" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC667FFFBA9443927FE0FFC9C" box="[267,416,862,881]" italics="true" pageId="6" pageNumber="326">Eunicolina glaesi</emphasis>
</taxonomicName>
<emphasis id="B95C915FC667FFFBA9E73927FE4CFC9C" bold="true" box="[424,483,862,881]" pageId="6" pageNumber="326">
<taxonomicNameLabel id="A26F2C24C667FFFBA9E73927FE4CFC9C" box="[424,483,862,881]" pageId="6" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A – The holotype piece, CHHC 588-13 (Hoffeins collection); B – Lateral view of the specimen of
<taxonomicName id="4C2836CEC667FFFBA88E3903FEFBFC60" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[193,340,890,909]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="6" pageNumber="326" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC667FFFBA88E3903FEFBFC60" box="[193,340,890,909]" italics="true" pageId="6" pageNumber="326">Eunicolina glaesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F2C24C667FFFBA9163903FE31FC60" box="[345,414,890,909]" pageId="6" pageNumber="326" rank="species">sp.nov.</taxonomicNameLabel>
; C – Ventral view of infracapitulum and leg I.
</paragraph>
</caption>
<caption id="DF571DC5C667FFFBA8D03D44FCE7F86A" ID-DOI="http://doi.org/10.5281/zenodo.5399502" ID-Zenodo-Dep="5399502" httpUri="https://zenodo.org/record/5399502/files/figure.png" pageId="6" pageNumber="326" startId="6.[159,171,1853,1872]" targetBox="[355,1232,957,1821]" targetPageId="6">
<paragraph id="8B974D4DC667FFFBA8D03D44FCE7F86A" blockId="6.[159,1429,1852,1928]" pageId="6" pageNumber="326">
FIGURE 5:
<taxonomicName id="4C2836CEC667FFFBA9443D44FE0EF8BD" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[267,417,1853,1872]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="6" pageNumber="326" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC667FFFBA9443D44FE0EF8BD" box="[267,417,1853,1872]" italics="true" pageId="6" pageNumber="326">Eunicolina glaesi</emphasis>
</taxonomicName>
<emphasis id="B95C915FC667FFFBA9E73D44FE4CF8BD" bold="true" box="[424,483,1853,1872]" pageId="6" pageNumber="326">
<taxonomicNameLabel id="A26F2C24C667FFFBA9E73D44FE4CF8BD" box="[424,483,1853,1872]" pageId="6" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A – Oculo-pustular zone.
<emphasis id="B95C915FC667FFFBAABA3D44FCB7F8BD" box="[757,792,1853,1872]" italics="true" pageId="6" pageNumber="326">bo.p</emphasis>
: trichobthria,
<emphasis id="B95C915FC667FFFBABEA3D44FC1BF8BD" box="[933,948,1853,1872]" italics="true" pageId="6" pageNumber="326">la</emphasis>
?: supposed insertion of the
<emphasis id="B95C915FC667FFFBAC843D44FB75F8BD" box="[1227,1242,1853,1872]" italics="true" pageId="6" pageNumber="326">la</emphasis>
seta;
<emphasis id="B95C915FC667FFFBAD5A3D44FA8AF8BD" box="[1301,1317,1853,1872]" italics="true" pageId="6" pageNumber="326">lb</emphasis>
: seta
<emphasis id="B95C915FC667FFFBAD2E3D44FADEF8BD" box="[1377,1393,1853,1872]" italics="true" pageId="6" pageNumber="326">lb</emphasis>
;
<emphasis id="B95C915FC667FFFBAD303D44FA3FF8BD" box="[1407,1424,1853,1872]" italics="true" pageId="6" pageNumber="326">ly</emphasis>
: post-ocular lyriform organ; B – Detail of the
<emphasis id="B95C915FC667FFFBAA263D20FDD2F886" box="[617,637,1881,1899]" italics="true" pageId="6" pageNumber="326">PI</emphasis>
tarsus showing the bidactylus claw, one of the two tarsal solnidia (
<emphasis id="B95C915FC667FFFBACB03D21FAA2F886" box="[1279,1293,1880,1899]" italics="true" pageId="6" pageNumber="326">ω</emphasis>
).
<emphasis id="B95C915FC667FFFBAD6B3D20FA95F881" box="[1316,1338,1881,1900]" italics="true" pageId="6" pageNumber="326">F?</emphasis>
: possible famulus of the tarsus. Some insertions of setae are marked by circles.
</paragraph>
</caption>
<caption id="DF571DC5C666FFFAA8D03E99FEDBFAC7" ID-DOI="http://doi.org/10.5281/zenodo.5399504" ID-Zenodo-Dep="5399504" httpUri="https://zenodo.org/record/5399504/files/figure.png" pageId="7" pageNumber="327" startId="7.[159,171,1248,1267]" targetBox="[281,1309,329,1221]" targetPageId="7">
<paragraph id="8B974D4DC666FFFAA8D03E99FEDBFAC7" blockId="7.[159,1429,1247,1322]" pageId="7" pageNumber="327">
FIGURE 6: Genito-anal area: A – Disposition of genital and anal shields (drawn from the extant species
<taxonomicName id="4C2836CEC666FFFAAC3C3E99FB11FB1F" authorityName="Coineau" authorityYear="1964" box="[1139,1214,1248,1266]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="travei">
<emphasis id="B95C915FC666FFFAAC3C3E99FB11FB1F" box="[1139,1214,1248,1266]" italics="true" pageId="7" pageNumber="327">E. travei</emphasis>
</taxonomicName>
) Aa, Ab: ventral view, female (a), male (b); Ac: male lateral view.
<taxonomicName id="4C2836CEC666FFFAAA1C3E82FD48FAE3" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[595,743,1275,1294]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC666FFFAAA1C3E82FD48FAE3" box="[595,743,1275,1294]" italics="true" pageId="7" pageNumber="327">Eunicolina glaesi</emphasis>
</taxonomicName>
<emphasis id="B95C915FC666FFFAAAA33E82FC8EFAE3" bold="true" box="[748,801,1275,1294]" pageId="7" pageNumber="327">
<taxonomicNameLabel id="A26F2C24C666FFFAAAA33E82FC8EFAE3" box="[748,801,1275,1294]" pageId="7" pageNumber="327" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: B-C – extremity of the opisthosoma, lateral view (B) and interpretation (Ca, Cb, Cc).
</paragraph>
</caption>
<paragraph id="8B974D4DC666FFFAA88A3F12FE70FA24" blockId="7.[159,762,1387,1481]" pageId="7" pageNumber="327">Description — The habitus is that of an animal with a rounded body, robust, well-sclerotized, yellow to brown in coloration.</paragraph>
<paragraph id="8B974D4DC666FFFAA88A3F95FF56F980" blockId="7.[159,762,1516,1645]" pageId="7" pageNumber="327">
Dimensions and proportions — Body length (
<emphasis id="B95C915FC666FFFAAAA73F95FD55F9EE" box="[744,762,1516,1539]" italics="true" pageId="7" pageNumber="327">c.</emphasis>
580 – 600 µm) is about 1.7 times its maximum width (
<emphasis id="B95C915FC666FFFAA8E73C4AFF1CF9A7" box="[168,179,1587,1610]" italics="true" pageId="7" pageNumber="327">c</emphasis>
. 390 – 400 µm), about as wide as high (
<figureCitation id="131351C8C666FFFAAAD83C4BFD5BF9A7" box="[663,756,1586,1610]" captionStart="FIGURE" captionStartId="3.[159,171,1095,1114]" captionTargetBox="[257,1346,302,1056]" captionTargetId="figure-4@3.[242,1346,302,1063]" captionTargetPageId="3" captionText="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." figureDoi="http://doi.org/10.5281/zenodo.5399493" httpUri="https://zenodo.org/record/5399493/files/figure.png" pageId="7" pageNumber="327">Figs 1A</figureCitation>
,
<figureCitation id="131351C8C666FFFAA8D03C2CFF43F980" box="[159,236,1621,1645]" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="7" pageNumber="327">3B, 3C</figureCitation>
).
</paragraph>
<paragraph id="8B974D4DC666FFFAA88A3CF6FAC8FA4B" blockId="7.[159,762,1679,1949]" lastBlockId="7.[826,1429,1387,1948]" pageId="7" pageNumber="327">
Cuticle ornamentation and cerotegument — The cuticle is ornamented with more or less rounded alveoli: the dorsal shield is covered with a regular polygonal pattern laterally, whereas partitions are more rounded on the central zone of the dorsum, and on the ventral shields. The borders of alveoli are crossed with more or less large costules (
<figureCitation id="131351C8C666FFFAAA823D18FF6FF870" captionStart="FIGURE" captionStartId="3.[159,171,1095,1114]" captionTargetBox="[257,1346,302,1056]" captionTargetId="figure-4@3.[242,1346,302,1063]" captionTargetPageId="3" captionText="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." figureDoi="http://doi.org/10.5281/zenodo.5399493" httpUri="https://zenodo.org/record/5399493/files/figure.png" pageId="7" pageNumber="327">Fig. 1C</figureCitation>
). As in other representatives of the genus, the polygonal pattern extends onto the articles of the legs and on the chelicerae, but not on the palps.
</paragraph>
<paragraph id="8B974D4DC666FFF5AB2F3FC4FD48FD8A" blockId="7.[826,1429,1387,1948]" lastBlockId="8.[159,762,311,1949]" lastPageId="8" lastPageNumber="328" pageId="7" pageNumber="327">
Dorsal shield — Chaetotaxy as for the genus (
<figureCitation id="131351C8C666FFFAAB0D3F99FC0EFA15" box="[834,929,1504,1528]" captionStart="FIGURE" captionStartId="3.[159,171,1095,1114]" captionTargetBox="[257,1346,302,1056]" captionTargetId="figure-4@3.[242,1346,302,1063]" captionTargetPageId="3" captionText="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." figureDoi="http://doi.org/10.5281/zenodo.5399493" httpUri="https://zenodo.org/record/5399493/files/figure.png" pageId="7" pageNumber="327">Figs 1A</figureCitation>
,
<figureCitation id="131351C8C666FFFAABFE3F99FC7AFA15" box="[945,981,1504,1528]" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="7" pageNumber="327">3A</figureCitation>
), the dorsal setae are long, (
<emphasis id="B95C915FC666FFFAAD7D3F99FA21FA1A" box="[1330,1422,1504,1528]" italics="true" pageId="7" pageNumber="327">la – le c</emphasis>
. 75 – 100 µm,
<emphasis id="B95C915FC666FFFAAB933C7AFBEFF9F6" box="[988,1088,1539,1563]" italics="true" pageId="7" pageNumber="327">da – de c</emphasis>
. 75 – 80 µm), anterior setae
<emphasis id="B95C915FC666FFFAAB753C5EFCFCF9D3" box="[826,851,1575,1598]" italics="true" pageId="7" pageNumber="327">ga</emphasis>
,
<emphasis id="B95C915FC666FFFAAB2D3C5EFCD5F9D3" box="[866,890,1575,1598]" italics="true" pageId="7" pageNumber="327">ge</emphasis>
, and
<emphasis id="B95C915FC666FFFAABF23C5EFC70F9D3" box="[957,991,1575,1598]" italics="true" pageId="7" pageNumber="327">gm</emphasis>
less visible or broken, their presence revealed only by the mark of insertion. The setae
<emphasis id="B95C915FC666FFFAAD323C33FA3AF98C" box="[1405,1429,1610,1633]" italics="true" pageId="7" pageNumber="327">gr</emphasis>
are short (&lt;40 µm). The trichobothria are similar to those of
<taxonomicName id="4C2836CEC666FFFAABEF3CE9FB00F94A" authority="Coineau, 1964" authorityName="Coineau" authorityYear="1964" box="[928,1199,1679,1703]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="travei">
<emphasis id="B95C915FC666FFFAABEF3CE9FC52F94A" box="[928,1021,1680,1703]" italics="true" pageId="7" pageNumber="327">E. travei</emphasis>
<bibRefCitation id="EFB930BCC666FFFAAC4A3CF6FB00F94A" author="Coineau Y." box="[1029,1199,1679,1703]" pageId="7" pageNumber="327" pagination="153 - 175" refId="ref8442" refString="Coineau Y. 1964 - Une nouvelle espLce francaise de Labidostomidae (Acariens, Prostigmates) Eunicolina travei sp. n. - Vie et Milieu, 15: 153 - 175." type="journal article" year="1964">Coineau, 1964</bibRefCitation>
</taxonomicName>
, with 3-4 branches. With more than 100 µm in length, the bothridial setae are longer than the lateral and the dorsal setae (
<figureCitation id="131351C8C666FFFAAB0D3C81FC6CF8FD" box="[834,963,1784,1808]" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="7" pageNumber="327">Figs 3A, B</figureCitation>
). The frontal edge of the dorsal shield bears two cornua. Close to twenty pustules were counted on the lateral fields of the dorsal shield. On each side, the field of the pustules begins at the level of the second coxal plate. In the usual position of the lateral eyes, close to the (
<emphasis id="B95C915FC669FFF5AA0C3B41FDF4FEA2" box="[579,603,312,335]" italics="true" pageId="8" pageNumber="328">gr</emphasis>
) setae, a convex structure (
<figureCitation id="131351C8C669FFF5A9013B23FE72FE9F" box="[334,477,346,370]" captionStart="FIGURE" captionStartId="3.[159,171,1095,1114]" captionTargetBox="[257,1346,302,1056]" captionTargetId="figure-4@3.[242,1346,302,1063]" captionTargetPageId="3" captionText="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." figureDoi="http://doi.org/10.5281/zenodo.5399493" httpUri="https://zenodo.org/record/5399493/files/figure.png" pageId="8" pageNumber="328">Figs 1A, 1B</figureCitation>
, star) could be either a pustule or the lateral eye (the usual striation of the lens surface, different from the smooth surface of the pustules, is not visible). No median anterior eye was discernible. The lateral lyrifissure, poorly preserved, may be located between the most anterior pustules. The density of the pustules is maximal behind the supposed lateral eye (
<figureCitation id="131351C8C669FFF5AA6C3836FD05FD8A" box="[547,682,591,615]" captionStart="FIGURE" captionStartId="3.[159,171,1095,1114]" captionTargetBox="[257,1346,302,1056]" captionTargetId="figure-4@3.[242,1346,302,1063]" captionTargetPageId="3" captionText="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." figureDoi="http://doi.org/10.5281/zenodo.5399493" httpUri="https://zenodo.org/record/5399493/files/figure.png" pageId="8" pageNumber="328">Figs 1A, 1B</figureCitation>
,
<figureCitation id="131351C8C669FFF5AAFA3836FD75FD8A" box="[693,730,591,615]" captionStart="FIGURE" captionStartId="6.[159,171,1853,1872]" captionTargetBox="[355,1232,957,1821]" captionTargetId="figure-43@6.[355,1232,957,1821]" captionTargetPageId="6" captionText="FIGURE 5: Eunicolina glaesi n. sp.: A – Oculo-pustular zone. bo.p: trichobthria, la?: supposed insertion of the la seta; lb: seta lb; ly: post-ocular lyriform organ; B – Detail of the PI tarsus showing the bidactylus claw, one of the two tarsal solnidia (ω). F?: possible famulus of the tarsus. Some insertions of setae are marked by circles." figureDoi="http://doi.org/10.5281/zenodo.5399502" httpUri="https://zenodo.org/record/5399502/files/figure.png" pageId="8" pageNumber="328">5A</figureCitation>
).
</paragraph>
<paragraph id="8B974D4DC669FFF5A88A3807FDF8F9EE" blockId="8.[159,762,311,1949]" pageId="8" pageNumber="328">
Ventral view — The posterior end of the ventral shield is damaged, hence the anal sclerites were not seen and the genital ones were only partially observable. The infracapitulum (162 µm long, 150 µm wide) is ornamented by alveoli, the pair of setae
<emphasis id="B95C915FC669FFF5AA963972FD55FCCE" box="[729,762,779,803]" italics="true" pageId="8" pageNumber="328">mb</emphasis>
is well visible, plus at least one pair of additional posterior setae (
<emphasis id="B95C915FC669FFF5A92C3928FE2AFC85" box="[355,389,849,872]" italics="true" pageId="8" pageNumber="328">
<collectingCountry id="F33F0DDDC669FFF5A92C3928FE2AFC85" box="[355,389,849,872]" name="American Samoa" pageId="8" pageNumber="328">AS</collectingCountry>
</emphasis>
on
<figureCitation id="131351C8C669FFF5A9FE3928FDA5FC84" box="[433,522,849,873]" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="8" pageNumber="328">Fig. 3C</figureCitation>
). The lateral lips are narrower than the anterior margin of the infracapitulum. The palps (the right palp was well observable in ventral view) are characteristic for the genus (1-1-3-4+
<emphasis id="B95C915FC669FFF5A94839A4FEB7FC18" box="[263,280,989,1013]" italics="true" pageId="8" pageNumber="328">ω</emphasis>
), with the palpal solenidion identifiable and the long apical seta (eupathidium?) (45 – 60 µm long) (
<figureCitation id="131351C8C669FFF5A8A03E5AFEE3FBD6" box="[239,332,1059,1083]" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="8" pageNumber="328">Figs 3C</figureCitation>
&amp;
<figureCitation id="131351C8C669FFF5A9393E5AFE36FBD6" box="[374,409,1059,1083]" captionStart="FIGURE" captionStartId="6.[159,171,862,881]" captionTargetBox="[355,1235,311,834]" captionTargetId="figure-4@6.[355,1235,311,834]" captionTargetPageId="6" captionText="FIGURE 4: Eunicolina glaesi n. sp.: A – The holotype piece, CHHC 588-13 (Hoffeins collection); B – Lateral view of the specimen of Eunicolina glaesi sp.nov.; C – Ventral view of infracapitulum and leg I." figureDoi="http://doi.org/10.5281/zenodo.5399500" httpUri="https://zenodo.org/record/5399500/files/figure.png" pageId="8" pageNumber="328">4C</figureCitation>
). The chelicerae are partially hidden by the first pair of legs and the palps, and cannot be observed in lateral view. The chelicerae are robust (
<emphasis id="B95C915FC669FFF5A9653EF4FE93FB49" box="[298,316,1165,1188]" italics="true" pageId="8" pageNumber="328">c.</emphasis>
250 µm long) and are strongly sclerotized, the proximal part covered with alveolar ornamentation. Examination in alternate dorsal and ventral views permitted identification of the anterior seta (
<emphasis id="B95C915FC669FFF5A95E3F60FE9AFADC" box="[273,309,1305,1329]" italics="true" pageId="8" pageNumber="328">chb</emphasis>
) of the fixed digit and the long posterior seta
<emphasis id="B95C915FC669FFF5A94B3F45FE87FAB9" box="[260,296,1340,1364]" italics="true" pageId="8" pageNumber="328">cha</emphasis>
, whose insertion was not seen (
<figureCitation id="131351C8C669FFF5AAD63F45FF6EFA9A" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="8" pageNumber="328">Figs 3A, 3C</figureCitation>
). The coxisternal area (
<figureCitation id="131351C8C669FFF5A9863F26FDB1FA9A" box="[457,542,1375,1399]" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="8" pageNumber="328">Fig. 3C</figureCitation>
) is armoured, sclerotized and ornamented with strong alveoli (
<figureCitation id="131351C8C669FFF5AA823FFBFF6EFA50" captionStart="FIGURE" captionStartId="5.[159,171,1789,1808]" captionTargetBox="[296,1181,368,1706]" captionTargetId="figure-4@5.[280,1308,350,1758]" captionTargetPageId="5" captionText="FIGURE 3:. Eunicolina glaesi n. sp.: A – Dorsal shield, anterior part; B – Detail of the posterior trichobothrium; C – Ventral view of epimeral plates and detail of ornamentation. Abbreviations: AS: additional posterior seate of the infracapitulum; bo.a, bo.p: anterior and posterior bothridia; CH: Chelicera; cha, chb cheliceral setae; gr, la: ocular and lateral setae of the dorsal shield; PI, PII: legs I and II; Pp: palp. ω1: tarsal solenidion of the palp." figureDoi="http://doi.org/10.5281/zenodo.5399496" httpUri="https://zenodo.org/record/5399496/files/figure.png" pageId="8" pageNumber="328">Fig. 3C</figureCitation>
), the epimeral setae are poorly preserved, and only some simple setae were identified on the anterior fringe of the first epimeral plates.
</paragraph>
<paragraph id="8B974D4DC669FFF5A88A3C63FD55F8E9" blockId="8.[159,762,311,1949]" pageId="8" pageNumber="328">
At the level of the fourth epimera, on each side, the integument forms a large cavity, a pit, that receives the basal parts of the legs IV (&quot;
<emphasis id="B95C915FC669FFF5AA193C18FF73F976" italics="true" pageId="8" pageNumber="328">puits tØgumentaire&quot;</emphasis>
in
<bibRefCitation id="EFB930BCC669FFF5A94D3CFAFE0BF976" author="Coineau Y." box="[258,420,1667,1691]" pageId="8" pageNumber="328" pagination="153 - 175" refId="ref8442" refString="Coineau Y. 1964 - Une nouvelle espLce francaise de Labidostomidae (Acariens, Prostigmates) Eunicolina travei sp. n. - Vie et Milieu, 15: 153 - 175." type="journal article" year="1964">Coineau 1964</bibRefCitation>
).The cuticle of these hollows is covered with dense villose integument similar to what has been observed in congeneric species (
<emphasis id="B95C915FC669FFF5AA9D3CB3FD5DF90C" box="[722,754,1738,1761]" italics="true" pageId="8" pageNumber="328">PT</emphasis>
,
<figureCitation id="131351C8C669FFF5A8D03C95FF56F8E9" box="[159,249,1772,1796]" captionStart="FIGURE" captionStartId="3.[159,171,1095,1114]" captionTargetBox="[257,1346,302,1056]" captionTargetId="figure-4@3.[242,1346,302,1063]" captionTargetPageId="3" captionText="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." figureDoi="http://doi.org/10.5281/zenodo.5399493" httpUri="https://zenodo.org/record/5399493/files/figure.png" pageId="8" pageNumber="328">Figs 1D</figureCitation>
,
<figureCitation id="131351C8C669FFF5A94B3C95FE8AF8E9" box="[260,293,1772,1796]" captionStart="FIGURE" captionStartId="4.[159,171,1716,1735]" captionText="FIGURE 2: Eunicolina glaesi n. sp.: A – Genu and tibia, first pair of legs, right side, dorsal view; B – First leg, left side, lateral view; C – Ventral view, detail of the proximal articles of leg I, II and acetabulum of fourth leg (the leg IV is not represented here); D – The tridactylous tip of the tarsus II, lateral view. Abbreviations: epI: first epimeral plate; ge: genu; PI, PII, PIII, PIV: Legs 1 to 4; ti: tibia." pageId="8" pageNumber="328">2C</figureCitation>
) (
<emphasis id="B95C915FC669FFF5A9733C95FEF9F8E9" box="[316,342,1772,1796]" italics="true" pageId="8" pageNumber="328">cf.</emphasis>
<bibRefCitation id="EFB930BCC669FFF5A9123C95FE51F8E9" author="Coineau Y." box="[349,510,1772,1796]" pageId="8" pageNumber="328" pagination="153 - 175" refId="ref8442" refString="Coineau Y. 1964 - Une nouvelle espLce francaise de Labidostomidae (Acariens, Prostigmates) Eunicolina travei sp. n. - Vie et Milieu, 15: 153 - 175." type="journal article" year="1964">Coineau 1964</bibRefCitation>
;
<bibRefCitation id="EFB930BCC669FFF5AA473C95FD41F8E9" author="Feider Z. &amp; Vasiliu N." box="[520,750,1772,1796]" pageId="8" pageNumber="328" pagination="643 - 662" refId="ref8683" refString="Feider Z., Vasiliu N. 1968 - Un nouveau genre de la famille Nicoletiellidae Canestrini 1891 et description de l'espLce Grandjeanellina nova (Sellnick) 1931 - Trav. Mus. Hist. natur. &quot; Grigore Antipa &quot;. VII &quot; Le centenaire Grigore Antipa, 1867 - 1967), Bucarest: 643 - 662." type="book chapter" year="1968">Feider, Vasiliu 1968</bibRefCitation>
).
</paragraph>
<paragraph id="8B974D4DC669FFF5A88A3D65FB35FCB0" blockId="8.[159,762,311,1949]" lastBlockId="8.[826,1429,311,1949]" pageId="8" pageNumber="328">
Legs (
<figureCitation id="131351C8C669FFF5A9463D62FECCF8DE" box="[265,355,1819,1843]" captionStart="FIGURE" captionStartId="3.[159,171,1095,1114]" captionTargetBox="[257,1346,302,1056]" captionTargetId="figure-4@3.[242,1346,302,1063]" captionTargetPageId="3" captionText="FIGURE 1: Eunicolina glaesi n. sp. — A. Lateral view, right side. B. Oculo-pustular zone, details. C. Detail of the cuticle, pattern in the lateral zone. D. Enlarged view of the acetabula IV, showing the villosity in pit PT (&quot;puits tØgumentaire&quot; in Coineau 1964). Abbreviations: BO.A, BO.P: anterior and posterior trichobothria; la, lb: lateral setae of the dorsal shield; da, db, dc: dorsal setae of the dorsal shield; EPI IV: fourth epimeral plate; PI, PIV: first and fourth legs. Star: possible lateral eye." figureDoi="http://doi.org/10.5281/zenodo.5399493" httpUri="https://zenodo.org/record/5399493/files/figure.png" pageId="8" pageNumber="328">Figs 1A</figureCitation>
,
<figureCitation id="131351C8C669FFF5A9213D62FE10F8DE" box="[366,447,1819,1843]" captionStart="FIGURE" captionStartId="4.[159,171,1716,1735]" captionText="FIGURE 2: Eunicolina glaesi n. sp.: A – Genu and tibia, first pair of legs, right side, dorsal view; B – First leg, left side, lateral view; C – Ventral view, detail of the proximal articles of leg I, II and acetabulum of fourth leg (the leg IV is not represented here); D – The tridactylous tip of the tarsus II, lateral view. Abbreviations: epI: first epimeral plate; ge: genu; PI, PII, PIII, PIV: Legs 1 to 4; ti: tibia." pageId="8" pageNumber="328">2A, 2D</figureCitation>
) — The legs are robust and sclerotized, the polygonal pattern ornamentation of the body is continued on the articles and discernible on femora and genua on the first pair of legs. The articles of the first pair of legs are thick, and not elongated as in the other genera of labidostommatids. However, asymmetry between right and left legs of the first pair attests that either the fossilisation process or optical properties of amber induced some deformation, the legs of the right side keeping more &quot;natural&quot; proportions. The third leg is missing on the left side. The alveolate ornamentation of the legs is similar to that of the shields, but locally it is resolved in cristules as on the edges of the genu and tibia of leg I (
<figureCitation id="131351C8C669FFF5AB9A38ECFB9EFD43" box="[981,1073,661,686]" captionStart="FIGURE" captionStartId="4.[159,171,1716,1735]" captionText="FIGURE 2: Eunicolina glaesi n. sp.: A – Genu and tibia, first pair of legs, right side, dorsal view; B – First leg, left side, lateral view; C – Ventral view, detail of the proximal articles of leg I, II and acetabulum of fourth leg (the leg IV is not represented here); D – The tridactylous tip of the tarsus II, lateral view. Abbreviations: epI: first epimeral plate; ge: genu; PI, PII, PIII, PIV: Legs 1 to 4; ti: tibia." pageId="8" pageNumber="328">Fig. 2A</figureCitation>
). The genu, is shorter than the telofemur and the separation of the femora in three articles is functionally reduced to basi- and [mesotelo] femora, due to &quot;ankylosis&quot; of the distal article of the femora (
<figureCitation id="131351C8C669FFF5ABA9395BFBC3FCD7" box="[998,1132,802,826]" captionStart="FIGURE" captionStartId="4.[159,171,1716,1735]" captionText="FIGURE 2: Eunicolina glaesi n. sp.: A – Genu and tibia, first pair of legs, right side, dorsal view; B – First leg, left side, lateral view; C – Ventral view, detail of the proximal articles of leg I, II and acetabulum of fourth leg (the leg IV is not represented here); D – The tridactylous tip of the tarsus II, lateral view. Abbreviations: epI: first epimeral plate; ge: genu; PI, PII, PIII, PIV: Legs 1 to 4; ti: tibia." pageId="8" pageNumber="328">Figs 2A, 2B</figureCitation>
) already described in the extant species (
<bibRefCitation id="EFB930BCC669FFF5ABA4393CFB21FCB0" author="Coineau Y." box="[1003,1166,837,861]" pageId="8" pageNumber="328" pagination="153 - 175" refId="ref8442" refString="Coineau Y. 1964 - Une nouvelle espLce francaise de Labidostomidae (Acariens, Prostigmates) Eunicolina travei sp. n. - Vie et Milieu, 15: 153 - 175." type="journal article" year="1964">Coineau 1964</bibRefCitation>
).
</paragraph>
<paragraph id="8B974D4DC669FFF5AB2F390DFC40FBB3" blockId="8.[826,1429,311,1949]" pageId="8" pageNumber="328">
Tarsi as for the genus, tarsi II, III, IV are tridactylous, and tarsi of legs I bidactylous. The tarsi I are provided with long dorsal setae, and on the left leg, one of the two usual recumbent solenidia (
<emphasis id="B95C915FC669FFF5AD2F39A4FADEFC18" box="[1376,1393,989,1013]" italics="true" pageId="8" pageNumber="328">ω</emphasis>
) is clearly visible, and a spine which, by both the position and its shape, could be identified as the famulus (
<figureCitation id="131351C8C669FFF5AB233E3FFC7AFBB3" box="[876,981,1094,1118]" captionStart="FIGURE" captionStartId="6.[159,171,1853,1872]" captionTargetBox="[355,1232,957,1821]" captionTargetId="figure-43@6.[355,1232,957,1821]" captionTargetPageId="6" captionText="FIGURE 5: Eunicolina glaesi n. sp.: A – Oculo-pustular zone. bo.p: trichobthria, la?: supposed insertion of the la seta; lb: seta lb; ly: post-ocular lyriform organ; B – Detail of the PI tarsus showing the bidactylus claw, one of the two tarsal solnidia (ω). F?: possible famulus of the tarsus. Some insertions of setae are marked by circles." figureDoi="http://doi.org/10.5281/zenodo.5399502" httpUri="https://zenodo.org/record/5399502/files/figure.png" pageId="8" pageNumber="328">
Fig.5B,
<emphasis id="B95C915FC669FFF5AB8C3E3EFC7AFBB3" box="[963,981,1095,1118]" italics="true" pageId="8" pageNumber="328">F</emphasis>
</figureCitation>
?).
</paragraph>
<paragraph id="8B974D4DC669FFF5AB2F3E0CFC4DF953" blockId="8.[826,1429,311,1949]" pageId="8" pageNumber="328">
Differential diagnosis — By the characters described here above, there is no doubt that this specimen belongs to the genus
<emphasis id="B95C915FC669FFF5ACC23EC2FAA6FB3E" box="[1165,1289,1211,1235]" italics="true" pageId="8" pageNumber="328">
<taxonomicName id="4C2836CEC669FFF5ACC23EC2FAA9FB3E" authorityName="Berlese" authorityYear="1911" box="[1165,1286,1211,1235]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">Eunicolina</taxonomicName>
.
</emphasis>
The fossil species is distinct from the extant members of the genus by the supposed absence of frontal eye, reduced lateral eyes and by the lower number of pustules than in modern species. It clearly differs from
<emphasis id="B95C915FC669FFF5AB753F12FC20FA6F" box="[826,911,1387,1410]" italics="true" pageId="8" pageNumber="328">E. nova</emphasis>
(
<bibRefCitation id="EFB930BCC669FFF5ABEE3F12FBF8FA6E" author="Sellnick M." box="[929,1111,1387,1411]" pageId="8" pageNumber="328" pagination="693 - 776" refId="ref9906" refString="Sellnick M. 1931 b - Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes von Max Beier, Wien. XVI. - Acari Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, 140 (9 - 10): 693 - 776." type="journal article" year="1931">Sellnick, 1931b</bibRefCitation>
), which has 20 to 35 pustules, by absence of neotrichy; from
<taxonomicName id="4C2836CEC669FFF5ACB83FF7FAF9FA48" authorityName="Coineau" authorityYear="1964" box="[1271,1366,1422,1445]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="travei">
<emphasis id="B95C915FC669FFF5ACB83FF7FAF9FA48" box="[1271,1366,1422,1445]" italics="true" pageId="8" pageNumber="328">E. travei</emphasis>
</taxonomicName>
– by the absence of modified integument near the posterior trichobothria. On the ventral surface of the infracapitulum it clearly differs from both
<taxonomicName id="4C2836CEC669FFF5AD783F81FA3AF9E2" authorityName="Coineau" authorityYear="1964" box="[1335,1429,1528,1551]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="travei">
<emphasis id="B95C915FC669FFF5AD783F81FA3AF9E2" box="[1335,1429,1528,1551]" italics="true" pageId="8" pageNumber="328">E. travei</emphasis>
</taxonomicName>
and
<emphasis id="B95C915FC669FFF5AB213C62FC69F9DF" box="[878,966,1563,1586]" italics="true" pageId="8" pageNumber="328">E. nova,</emphasis>
these latter species having respectively (
<emphasis id="B95C915FC669FFF5AB0C3C47FCCCF9B8" box="[835,867,1598,1621]" italics="true" pageId="8" pageNumber="328">ma</emphasis>
) and (
<emphasis id="B95C915FC669FFF5ABE13C44FC60F9B8" box="[942,975,1597,1621]" italics="true" pageId="8" pageNumber="328">mb</emphasis>
) and 4-5 or 7 pairs of infracapitular setae, while
<taxonomicName id="4C2836CEC669FFF5ABFE3C18FBA4F995" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[945,1035,1632,1656]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="glaesi">
<emphasis id="B95C915FC669FFF5ABFE3C18FBA4F995" box="[945,1035,1632,1656]" italics="true" pageId="8" pageNumber="328">E. glaesi</emphasis>
</taxonomicName>
shows two pairs of infracapitular setae: (
<emphasis id="B95C915FC669FFF5ABDE3CFAFC14F976" box="[913,955,1667,1691]" italics="true" pageId="8" pageNumber="328">mb)</emphasis>
and one other pair with discernible insertion marks.
</paragraph>
<paragraph id="8B974D4DC669FFF4AB2F3CACFF4BFA9D" blockId="8.[826,1429,311,1949]" lastBlockId="9.[159,762,1017,1392]" lastPageId="9" lastPageNumber="329" pageId="8" pageNumber="328">
Remark — The pustules of
<taxonomicName id="4C2836CEC669FFF5ACE53CACFAB0F900" authorityName="Berlese" authorityYear="1911" box="[1194,1311,1749,1773]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B95C915FC669FFF5ACE53CACFAB0F900" box="[1194,1311,1749,1773]" italics="true" pageId="8" pageNumber="328">Eunicolina</emphasis>
</taxonomicName>
mites are distributed &quot;on each side from the level of the leg II and extending uninterrupted caudad and absent from a short space of the posterior end of the dorsal shield&quot; (
<bibRefCitation id="EFB930BCC669FFF5AB883D18FB29F894" author="Greenberg B." box="[967,1158,1889,1913]" pageId="8" pageNumber="328" pagination="201" refId="ref9008" refString="Greenberg B. 1952 - New Labidostommidae with keys to the New World species (Acarina) - J. New York Entomol. Soc., 60 (4): 201." type="journal article" year="1952">Greenberg 1952</bibRefCitation>
), but each species differs by their number (
<tableCitation id="C6AA78F6C669FFF5AC093DFDFB33F870" box="[1094,1180,1924,1949]" captionStart="TABLE" captionStartId="9.[159,172,308,327]" captionText="TABLE 1: The tubercles and distributions of the Eunicolina species (data from Berlese 1911; Sellnick 1931b; Feider et Vasiliu 1968; Vistorin, 1980; Coineau 1964; Bertrand, 1988)" pageId="8" pageNumber="328">Table 1</tableCitation>
).
<bibRefCitation id="EFB930BCC669FFF5ACF13DFDFACCF870" author="Berlese A." box="[1214,1379,1924,1949]" pageId="8" pageNumber="328" pagination="429 - 435" refId="ref7902" refString="Berlese A. 1911 - Acarorum species novae quindecim - Redia, 7 [2]: 429 - 435." type="journal article" year="1911">Berlese (1911)</bibRefCitation>
has noted the arrangement of the pustules in two &quot;series&quot; &quot;
<emphasis id="B95C915FC668FFF4A8A83E65FF5EFB97" italics="true" pageId="9" pageNumber="329">tuberculi seriei marginalis 6 numero; marginalibus serie usque ad marginalis posticum producta, ad 25 numero</emphasis>
&quot;,
<emphasis id="B95C915FC668FFF4A9483E1AFE8FFB97" box="[263,288,1123,1146]" italics="true" pageId="9" pageNumber="329">i.e</emphasis>
a total of 31 pustules, in
<emphasis id="B95C915FC668FFF4AA713E1AFF72FB70" italics="true" pageId="9" pageNumber="329">
<taxonomicName id="4C2836CEC668FFF4AA713E1AFD77FB97" authorityName="Berlese" authorityYear="1911" box="[574,728,1122,1146]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="tuberculata">E. tuberculata</taxonomicName>
.
<taxonomicName id="4C2836CEC668FFF4AAAC3E1AFF72FB70" authorityName="Coineau" authorityYear="1964" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="travei">E. travei</taxonomicName>
</emphasis>
presents the largest neotaxy, with more than 40 pustules on each side, in
<emphasis id="B95C915FC668FFF4A9BC3ED0FDE8FB2D" box="[499,583,1193,1216]" italics="true" pageId="9" pageNumber="329">E. nova</emphasis>
the number of pustules varies from 20 to 35 on each side (
<bibRefCitation id="EFB930BCC668FFF4AAD33EB5FF4AFAEA" author="Sellnick M." pageId="9" pageNumber="329" pagination="693 - 776" refId="ref9906" refString="Sellnick M. 1931 b - Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes von Max Beier, Wien. XVI. - Acari Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, 140 (9 - 10): 693 - 776." type="journal article" year="1931">Sellnick 1931b</bibRefCitation>
;
<bibRefCitation id="EFB930BCC668FFF4A8BF3E96FDA9FAEA" author="Feider Z. &amp; Vasiliu N." box="[240,518,1263,1287]" pageId="9" pageNumber="329" pagination="643 - 662" refId="ref8683" refString="Feider Z., Vasiliu N. 1968 - Un nouveau genre de la famille Nicoletiellidae Canestrini 1891 et description de l'espLce Grandjeanellina nova (Sellnick) 1931 - Trav. Mus. Hist. natur. &quot; Grigore Antipa &quot;. VII &quot; Le centenaire Grigore Antipa, 1867 - 1967), Bucarest: 643 - 662." type="book chapter" year="1968">Feider and Vasiliu 1968</bibRefCitation>
). The fourth species, the American
<taxonomicName id="4C2836CEC668FFF4A9093F6BFD2EFAC7" authority="Greenberg, 1952" authorityName="Greenberg" authorityYear="1952" box="[326,641,1298,1322]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="porifera">
<emphasis id="B95C915FC668FFF4A9093F6BFE17FAC7" box="[326,440,1298,1322]" italics="true" pageId="9" pageNumber="329">E. porifera</emphasis>
<bibRefCitation id="EFB930BCC668FFF4A9F03F6BFD2EFAC7" author="Greenberg B." box="[447,641,1298,1322]" pageId="9" pageNumber="329" pagination="201" refId="ref9008" refString="Greenberg B. 1952 - New Labidostommidae with keys to the New World species (Acarina) - J. New York Entomol. Soc., 60 (4): 201." type="journal article" year="1952">Greenberg, 1952</bibRefCitation>
</taxonomicName>
, from
<collectingRegion id="49EC83AFC668FFF4AA823F6BFF73FAA0" country="United States of America" name="Virginia" pageId="9" pageNumber="329">Virginia</collectingRegion>
, averages 30 pustules on each side (
<bibRefCitation id="EFB930BCC668FFF4AA323F4CFF77FA9D" author="Greenberg B." pageId="9" pageNumber="329" pagination="201" refId="ref9008" refString="Greenberg B. 1952 - New Labidostommidae with keys to the New World species (Acarina) - J. New York Entomol. Soc., 60 (4): 201." type="journal article" year="1952">Greenberg 1952</bibRefCitation>
).
</paragraph>
<caption id="DF571DC5C668FFF4A8D03B4DFDA1FE8F" pageId="9" pageNumber="329" startId="9.[159,172,308,327]">
<paragraph id="8B974D4DC668FFF4A8D03B4DFDA1FE8F" blockId="9.[159,1429,307,354]" pageId="9" pageNumber="329">
TABLE 1: The tubercles and distributions of the
<taxonomicName id="4C2836CEC668FFF4AA163B4AFD51FEAB" box="[601,766,307,326]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95C915FC668FFF4AA163B4AFD18FEAB" box="[601,695,307,326]" italics="true" pageId="9" pageNumber="329">Eunicolina</emphasis>
species
</taxonomicName>
(data from
<bibRefCitation id="EFB930BCC668FFF4AB253B4AFC71FEAB" author="Berlese A." box="[874,990,307,326]" pageId="9" pageNumber="329" pagination="429 - 435" refId="ref7902" refString="Berlese A. 1911 - Acarorum species novae quindecim - Redia, 7 [2]: 429 - 435." type="journal article" year="1911">Berlese 1911</bibRefCitation>
;
<bibRefCitation id="EFB930BCC668FFF4ABA93B4AFBC2FEAB" author="Sellnick M." box="[998,1133,307,326]" pageId="9" pageNumber="329" pagination="693 - 776" refId="ref9906" refString="Sellnick M. 1931 b - Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes von Max Beier, Wien. XVI. - Acari Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, 140 (9 - 10): 693 - 776." type="journal article" year="1931">Sellnick 1931b</bibRefCitation>
;
<bibRefCitation id="EFB930BCC668FFF4AC3A3B4AFA93FEAB" author="Feider Z. &amp; Vasiliu N." box="[1141,1340,307,327]" pageId="9" pageNumber="329" pagination="643 - 662" refId="ref8683" refString="Feider Z., Vasiliu N. 1968 - Un nouveau genre de la famille Nicoletiellidae Canestrini 1891 et description de l'espLce Grandjeanellina nova (Sellnick) 1931 - Trav. Mus. Hist. natur. &quot; Grigore Antipa &quot;. VII &quot; Le centenaire Grigore Antipa, 1867 - 1967), Bucarest: 643 - 662." type="book chapter" year="1968">Feider et Vasiliu 1968</bibRefCitation>
;
<bibRefCitation id="EFB930BCC668FFF4AD0B3B4AFF41FE8F" author="Vistorin H. E." pageId="9" pageNumber="329" pagination="367 - 383" refId="ref10293" refString="Vistorin H. E. 1980 - Morphologische Untersuchungen an europaischen Nicoletielliden (Acari, Trombidiformes). - Acarologia, 21 (3): 367 - 383." type="journal article" year="1980">Vistorin, 1980</bibRefCitation>
;
<bibRefCitation id="EFB930BCC668FFF4A8B93B36FED8FE8F" author="Coineau Y." box="[246,375,335,354]" pageId="9" pageNumber="329" pagination="153 - 175" refId="ref8442" refString="Coineau Y. 1964 - Une nouvelle espLce francaise de Labidostomidae (Acariens, Prostigmates) Eunicolina travei sp. n. - Vie et Milieu, 15: 153 - 175." type="journal article" year="1964">Coineau 1964</bibRefCitation>
;
<bibRefCitation id="EFB930BCC668FFF4A9303B36FDA6FE8F" author="Bertrand M." box="[383,521,335,354]" pageId="9" pageNumber="329" pagination="1 - 345" refId="ref7994" refString="Bertrand M. 1988 - Les Labidostomidae Oudemans, 1904 (Acariens, Actinedida) - ThLse d'Etat, UniversitO des Sciences et Techniques du Languedoc, Montpellier, France. 1 - 345." type="book chapter" year="1988">Bertrand, 1988</bibRefCitation>
)
</paragraph>
</caption>
<paragraph id="8B974D4DC668FFF4A8AC3BEAFB24FC69" pageId="9" pageNumber="329">
<table id="F928BFEDC6680002A8E03BEAFA29FC44" box="[175,1414,403,937]" gridcols="4" gridrows="7" pageId="9" pageNumber="329">
<tr id="35184F0FC6680002A8E03BEAFA29FE46" box="[175,1414,403,427]" gridrow="0" pageId="9" pageNumber="329">
<th id="76C92673C6680002A8E03BEAFEE0FE46" box="[175,335,403,427]" gridcol="0" gridrow="0" pageId="9" pageNumber="329">Species</th>
<th id="76C92673C6680002A93C3BEAFD7DFE46" box="[371,722,403,427]" gridcol="1" gridrow="0" pageId="9" pageNumber="329">Number of pustules</th>
<th id="76C92673C6680002AA903BEAFBA0FE46" box="[735,1039,403,427]" gridcol="2" gridrow="0" pageId="9" pageNumber="329">Distribution</th>
<th id="76C92673C6680002AC513BEAFA29FE46" box="[1054,1414,403,427]" gridcol="3" gridrow="0" pageId="9" pageNumber="329">Habitat</th>
</tr>
<tr id="35184F0FC6680002A8E03BC1FA29FE1A" box="[175,1414,440,503]" gridrow="1" pageId="9" pageNumber="329">
<th id="76C92673C6680002A8E03BC1FEE0FE1A" box="[175,335,440,503]" gridcol="0" gridrow="1" pageId="9" pageNumber="329">
<taxonomicName id="4C2836CEC668FFF4A8E03BC0FEEFFE3D" authorityName="Berlese" authorityYear="1911" box="[175,320,440,464]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B95C915FC668FFF4A8E03BC0FEEFFE3D" box="[175,320,440,464]" italics="true" pageId="9" pageNumber="329">E. tuberculata</emphasis>
</taxonomicName>
</th>
<td id="76C92673C6680002A93C3BC1FD7DFE1A" box="[371,722,440,503]" gridcol="1" gridrow="1" pageId="9" pageNumber="329">
<emphasis id="B95C915FC668FFF4A93C3BC0FE26FE3D" box="[371,393,441,464]" italics="true" pageId="9" pageNumber="329">ca</emphasis>
28–32. 5–6 anterior, 25 posterior to the eye.
</td>
<td id="76C92673C6680002AA903BC1FBA0FE1A" box="[735,1039,440,503]" gridcol="2" gridrow="1" pageId="9" pageNumber="329">North Italy, Adriatic coast.</td>
<td id="76C92673C6680002AC513BC1FA29FE1A" box="[1054,1414,440,503]" gridcol="3" gridrow="1" pageId="9" pageNumber="329">fields, natural vegetal cover, shrubs</td>
</tr>
<tr id="35184F0FC6680002A8E03866FA29FDDA" box="[175,1414,543,567]" gridrow="2" pageId="9" pageNumber="329">
<th id="76C92673C6680002A8E03866FEE0FDDA" box="[175,335,543,567]" gridcol="0" gridrow="2" pageId="9" pageNumber="329">
<taxonomicName id="4C2836CEC668FFF4A8E03866FEB5FDDA" authorityName="Greenberg" authorityYear="1952" box="[175,282,543,567]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="porifera">
<emphasis id="B95C915FC668FFF4A8E03866FEB5FDDA" box="[175,282,543,567]" italics="true" pageId="9" pageNumber="329">E. porifera</emphasis>
</taxonomicName>
</th>
<td id="76C92673C6680002A93C3866FD7DFDDA" box="[371,722,543,567]" gridcol="1" gridrow="2" pageId="9" pageNumber="329">30. None anterior to the eye.</td>
<td id="76C92673C6680002AA903866FBA0FDDA" box="[735,1039,543,567]" gridcol="2" gridrow="2" pageId="9" pageNumber="329">Virginia (USA)</td>
<td id="76C92673C6680002AC513866FA29FDDA" box="[1054,1414,543,567]" gridcol="3" gridrow="2" pageId="9" pageNumber="329">leaf mold</td>
</tr>
<tr id="35184F0FC6680002A8E03819FA29FD73" box="[175,1414,608,670]" gridrow="3" pageId="9" pageNumber="329">
<th id="76C92673C6680002A8E03819FEE0FD73" box="[175,335,608,670]" gridcol="0" gridrow="3" pageId="9" pageNumber="329">
<emphasis id="B95C915FC668FFF4A8E03819FF52FD9A" box="[175,253,608,631]" italics="true" pageId="9" pageNumber="329">E. nova</emphasis>
</th>
<td id="76C92673C6680002A93C3819FD7DFD73" box="[371,722,608,670]" gridcol="1" gridrow="3" pageId="9" pageNumber="329">
20–35. First four are anterior to the seta
<emphasis id="B95C915FC668FFF4A99F38FFFE48FD70" box="[464,487,646,669]" italics="true" pageId="9" pageNumber="329">gr</emphasis>
</td>
<td id="76C92673C6680002AA903819FBA0FD73" box="[735,1039,608,670]" gridcol="2" gridrow="3" pageId="9" pageNumber="329">East Mediterranean basin, Greece, Romania.</td>
<td id="76C92673C6680002AC513819FA29FD73" box="[1054,1414,608,670]" gridcol="3" gridrow="3" pageId="9" pageNumber="329">leaf litter, sclerophyllous Mediterranean forest</td>
</tr>
<tr id="35184F0FC6680002A8E038BDFA29FD31" box="[175,1414,708,732]" gridrow="4" pageId="9" pageNumber="329">
<th id="76C92673C6680002A8E038BDFEE0FD31" box="[175,335,708,732]" gridcol="0" gridrow="4" pageId="9" pageNumber="329">
<taxonomicName id="4C2836CEC668FFF4A8E038BCFEA8FD31" authorityName="Coineau" authorityYear="1964" box="[175,263,709,732]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="travei">
<emphasis id="B95C915FC668FFF4A8E038BCFEA8FD31" box="[175,263,709,732]" italics="true" pageId="9" pageNumber="329">E. travei</emphasis>
</taxonomicName>
</th>
<td id="76C92673C6680002A93C38BDFD7DFD31" box="[371,722,708,732]" gridcol="1" gridrow="4" pageId="9" pageNumber="329">More than 40. More than 7</td>
<td id="76C92673C6680002AA9038BDFBA0FD31" box="[735,1039,708,732]" gridcol="2" gridrow="4" pageId="9" pageNumber="329">Mediterranean France</td>
<td id="76C92673C6680002AC5138BDFA29FD31" box="[1054,1414,708,732]" gridcol="3" gridrow="4" pageId="9" pageNumber="329">under tree cover, under</td>
</tr>
<tr id="35184F0FC6680002A8E03893FA29FCA3" box="[175,1414,746,846]" gridrow="5" pageId="9" pageNumber="329" rowspan-0="1" rowspan-2="1">
<td id="76C92673C6680002A93C3893FD7DFCA3" box="[371,722,746,846]" gridcol="1" gridrow="5" pageId="9" pageNumber="329">anterior to the lateral eye.</td>
<td id="76C92673C6680002AC513893FA29FCA3" box="[1054,1414,746,846]" gridcol="3" gridrow="5" pageId="9" pageNumber="329">Mediterranean and temperate climatic conditions, upper layer of litter</td>
</tr>
<tr id="35184F0FC6680002A8E03915FA29FC44" box="[175,1414,876,937]" gridrow="6" pageId="9" pageNumber="329">
<th id="76C92673C6680002A8E03915FEE0FC44" box="[175,335,876,937]" gridcol="0" gridrow="6" pageId="9" pageNumber="329">
<taxonomicName id="4C2836CEC668FFF4A8E03915FEA9FC69" authority="Bertrand &amp; Sidorchuk &amp; Hoffeins, 2015" authorityName="Bertrand &amp; Sidorchuk &amp; Hoffeins" authorityYear="2015" box="[175,262,876,900]" class="Arachnida" family="Labidostommatidae" genus="Eunicolina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="glaesi" status="sp. nov.">
<emphasis id="B95C915FC668FFF4A8E03915FEA9FC69" box="[175,262,876,900]" italics="true" pageId="9" pageNumber="329">E. glaesi</emphasis>
</taxonomicName>
<emphasis id="B95C915FC668FFF4A95E3915FEE0FC69" bold="true" box="[273,335,876,900]" pageId="9" pageNumber="329">
<taxonomicNameLabel id="A26F2C24C668FFF4A95E3915FEE0FC69" box="[273,335,876,900]" pageId="9" pageNumber="329" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76C92673C6680002A93C3915FD7DFC44" box="[371,722,876,937]" gridcol="1" gridrow="6" pageId="9" pageNumber="329">
18–20. None anterior to the seta
<emphasis id="B95C915FC668FFF4A9E939EBFE12FC44" box="[422,445,914,937]" italics="true" pageId="9" pageNumber="329">gr</emphasis>
</td>
<td id="76C92673C6680002AA903915FBA0FC44" box="[735,1039,876,937]" gridcol="2" gridrow="6" pageId="9" pageNumber="329">Fossil, in Baltic amber</td>
<td id="76C92673C6680002AC513915FA29FC44" box="[1054,1414,876,937]" gridcol="3" gridrow="6" pageId="9" pageNumber="329">unknown</td>
</tr>
</table>
</paragraph>
<paragraph id="8B974D4DC668FFF4A88A3F99FE5BF871" blockId="9.[159,762,1504,1948]" pageId="9" pageNumber="329">
Etymology — from &quot;
<emphasis id="B95C915FC668FFF4A9813F99FD88FA15" box="[462,551,1504,1528]" italics="true" pageId="9" pageNumber="329">glaesum</emphasis>
&quot;, term, used by Pliny the Elder (77) for amber
<emphasis id="B95C915FC668FFF4AA4B3C7AFDECF94B" italics="true" pageId="9" pageNumber="329">
&quot;.../... certum est gigni in insulis septentrionalis oceani et ab germanis appellari
<emphasis id="B95C915FC668FFF4A8853C31FE84F98D" bold="true" box="[202,299,1608,1632]" italics="true" pageId="9" pageNumber="329">glaesum</emphasis>
, itaque et ab nostris ob id unam insularum glaesariam appellatam, .../... nascitur autem defluente medulla pinei generis arboribus .../...&quot;.
</emphasis>
Translation: It is well established that the amber is a product of islands in the Northern Ocean, that the Germans named &quot;glaesum&quot;, and that, as a result, one of these islands was nicknamed &quot;Glaesaria&quot; by our (legion), .../... (Amber is) formed of a liquid seeping from the interior of a species of pine .../... [adapted from Woolley’s translation (1969)].
</paragraph>
</subSubSection>
</treatment>
</document>
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