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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1075.55862" ID-GBIF-Dataset="cdab907d-0496-4a01-bb09-bdec5f18f314" ID-PMC="PMC8671705" ID-Pensoft-Pub="1313-2970-1075-1" ID-Pensoft-UUID="0F0B3E039F8B56C698A2BFE13EE1C7E6" ID-PubMed="35046750" ID-ZooBank="9A0267042C384B2C9221534780145848" ModsDocID="1313-2970-1075-1" checkinTime="1638937137790" checkinUser="pensoft" docAuthor="Moran, Kevin M. &amp; Skevington, Jeffrey H." docDate="2021" docId="F7344727BD36524F8F12694695D8B0D2" docLanguage="en" docName="ZooKeys 1075: 1-32" docOrigin="ZooKeys 1075" docPubDate="2021-12-07" docSource="http://dx.doi.org/10.3897/zookeys.1075.55862" docTitle="Romaleosyrphus arctophiloides Moran &amp; Skevington 2021, comb. nov." docType="treatment" docVersion="4" id="0F0B3E039F8B56C698A2BFE13EE1C7E6" lastPageNumber="1" masterDocId="0F0B3E039F8B56C698A2BFE13EE1C7E6" masterDocTitle="Taxonomic revision of Romaleosyrphus Bigot (Diptera, Syrphidae), including descriptions of seven new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="1" updateTime="1668151198034" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of Romaleosyrphus Bigot (Diptera, Syrphidae), including descriptions of seven new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moran, Kevin M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9460-4619</mods:nameIdentifier>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada &amp; Carleton University, Department of Biology, 1125 Colonel By Drive, Ottawa Ontario K 1 S 5 B 6, Canada</mods:affiliation>
<mods:nameIdentifier type="email">syrphidae@kevinmoran.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skevington, Jeffrey H.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1445-9870</mods:nameIdentifier>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada &amp; Carleton University, Department of Biology, 1125 Colonel By Drive, Ottawa Ontario K 1 S 5 B 6, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1075</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>32</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1075.55862</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1075.55862</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1075-1</mods:identifier>
<mods:identifier type="ZooBank">9A0267042C384B2C9221534780145848</mods:identifier>
<mods:identifier type="Pensoft-UUID">0F0B3E039F8B56C698A2BFE13EE1C7E6</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="191254599" LSID="urn:lsid:plazi:treatment:F7344727BD36524F8F12694695D8B0D2" httpUri="http://treatment.plazi.org/id/F7344727BD36524F8F12694695D8B0D2" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="F7344727-BD36-524F-8F12-694695D8B0D2" authority="(Giglio-Tos, 1892)" authorityName="Moran &amp; Skevington" authorityYear="2021" baseAuthorityName="Giglio-Tos" baseAuthorityYear="1892" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus arctophiloides" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="arctophiloides" status="comb. nov.">Romaleosyrphus arctophiloides (Giglio-Tos, 1892)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Romaleosyrphus dorsal habitus A: Romaleosyrphus arctophiloides B: Romaleosyrphus vockerothi sp. nov. C: Romaleosyrphus soletluna sp. nov. rufous morph D: Romaleosyrphus soletluna sp. nov. black morph E: Romaleosyrphus villosus F. Romaleosyrphus bigoti sp. nov." figureDoi="10.3897/zookeys.1075.55862.figure4" httpUri="https://binary.pensoft.net/fig/619715" pageId="0" pageNumber="1">Figures 4A</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Romaleosyrphus lateral habitus A Romaleosyrphus arctophiloides B Romaleosyrphus vockerothi sp. nov. C Romaleosyrphus soletluna sp. nov. rufous morph D Romaleosyrphus soletluna sp. nov. black morph E Romaleosyrphus villosus F Romaleosyrphus bigoti sp. nov." figureDoi="10.3897/zookeys.1075.55862.figure6" httpUri="https://binary.pensoft.net/fig/619717" pageId="0" pageNumber="1">, 6A</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Romaleosyrphus 3 rd trochanter tubercle A Romaleosyrphus arctophiloides B Romaleosyrphus villosus." figureDoi="10.3897/zookeys.1075.55862.figure8" httpUri="https://binary.pensoft.net/fig/619719" pageId="0" pageNumber="1">, 8A</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Romaleosyrphus male hind tibia A Romaleosyrphus arctophiloides B Romaleosyrphus bigoti sp. nov. C Romaleosyrphus nephelaeus sp. nov. D Romaleosyrphus soletluna sp. nov. E Romaleosyrphus villosus." figureDoi="10.3897/zookeys.1075.55862.figure9" httpUri="https://binary.pensoft.net/fig/619720" pageId="0" pageNumber="1">, 9A</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Romaleosyrphus male face A Romaleosyrphus arctophiloides B Romaleosyrphus bigoti sp. nov. C Romaleosyrphus nephelaeus sp. nov. D Romaleosyrphus soletluna sp. nov. E Romaleosyrphus villosus F Romaleosyrphus vockerothi sp. nov." figureDoi="10.3897/zookeys.1075.55862.figure10" httpUri="https://binary.pensoft.net/fig/619721" pageId="0" pageNumber="1">, 10A</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Romaleosyrphus male genitalia A Romaleosyrphus arctophiloides B Romaleosyrphus soletluna sp. nov. C Romaleosyrphus nephelaeus sp. nov. D Romaleosyrphus vockerothi sp. nov. E Romaleosyrphus bigoti sp. nov. F Romaleosyrphus villosus." figureDoi="10.3897/zookeys.1075.55862.figure11" httpUri="https://binary.pensoft.net/fig/619722" pageId="0" pageNumber="1">, 11A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Moran &amp; Skevington" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Crioprora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crioprora arctophiloides" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="arctophiloides" status="comb. nov.">Crioprora arctophiloides</taxonomicName>
Giglio-Tos, 1892: 7. -
<bibRefCitation author="Giglio-Tos, E" journalOrPublisher="Universita di Torino" pageId="0" pageNumber="1" refId="B11" refString="Giglio-Tos, E, 1893. Part I. Stratiomyidae - Syrphidae. Torino, 72 pp. [71 pl.]" title="Part I. Stratiomyidae - Syrphidae. Torino, 72 pp. [71 pl.]" year="1893">Giglio-Tos (1893)</bibRefCitation>
: 25. -
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.1681" author="Aldrich, JM" pageId="0" pageNumber="1" refId="B1" refString="Aldrich, JM, 1905. A catalogue of North American Diptera (or two-winged flies) VL - XLVI. Smithsonian Miscellaneous Collections 46 (2): I-680. https://doi.org/10.5962/bhl.title.1681" title="A catalogue of North American Diptera (or two-winged flies) VL - XLVI. Smithsonian Miscellaneous Collections 46 (2): I- 680." url="https://doi.org/10.5962/bhl.title.1681" year="1905">Aldrich (1905)</bibRefCitation>
: 401. Type locality: Mexico, Angang[ueo] [MRSN]
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Moran &amp; Skevington" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Penthesilea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Penthesilea arctophiloides" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="arctophiloides" status="comb. nov.">Penthesilea arctophiloides</taxonomicName>
<normalizedToken originalValue="Kertész">Kertesz</normalizedToken>
, 1910: 286.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Moran &amp; Skevington" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Criorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Criorhina tapeta" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tapeta" status="comb. nov.">Criorhina tapeta</taxonomicName>
Fluke, 1939: 369. -Thompson (1976): 119. Type locality: Mexico City, 10,000 ft. [AMNH]
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Moran &amp; Skevington" authorityYear="2021" baseAuthorityName="Moran &amp; Skevington" baseAuthorityYear="2021" class="Insecta" family="Syrphidae" genus="Criorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Criorhina arctophiloides" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="arctophiloides" status="comb. nov.">Criorhina arctophiloides</taxonomicName>
Thompson, 1976: 118.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1947-09-16" collectingDateMax="1964-06-30" collectingDateMin="1947-09-16" collectorName="W. R. M. Mason, C. Bolivar, Diptera" country="Mexico" elevation="2438" latitude="19.204792" location="Diptera" longLatPrecision="1" longitude="-99.04885" specimenCount="3" specimenCount-male="3" stateProvince="Durango">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Mexico">Mexico</collectingCountry>
</emphasis>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Mexico" name="Durango">Durango</collectingRegion>
</emphasis>
:
<locationDeviation location="Southwest of El Salto">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.2530816" unit="mi" value="14.0">14 miles</quantity>
Southwest of El Salto
</locationDeviation>
,
<geoCoordinate degrees="23.702772" direction="north" orientation="latitude" precision="1" value="23.702772">23.702772</geoCoordinate>
,
<geoCoordinate degrees="105.564053" direction="west" orientation="longitude" precision="1" value="-105.56406">-105.564053</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.438" unit="m" value="2438.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.438" unit="m" value="2438.0">2438m</elevation>
</quantity>
,
<collectingDate value="1964-06-30">30.vi.1964</collectingDate>
,
<collectorName>W.R.M. Mason</collectorName>
, CNC_
<location LSID="urn:lsid:plazi:treatment:F7344727BD36524F8F12694695D8B0D2:189CF3D6CD139631017ADBBF1A10F821" country="Mexico" latitude="19.204792" longLatPrecision="1" longitude="-99.04885" name="Diptera" stateProvince="Durango">Diptera</location>
142464 (
<specimenCount type="male">1♂</specimenCount>
, CNC);
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Mexico" name="Distrito Federal">Mexico City</collectingRegion>
, D.F.
</emphasis>
: San Pedro Atocpan,
<geoCoordinate degrees="19.204792" direction="north" orientation="latitude" precision="1" value="19.204792">19.204792</geoCoordinate>
,
<geoCoordinate degrees="99.048853" direction="west" orientation="longitude" precision="1" value="-99.04885">-99.048853</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600m</elevation>
</quantity>
,
<collectingDate value="1947-09-16">16.ix.1947</collectingDate>
,
<collectorName>C. Bolivar</collectorName>
, CNC_
<collectorName>Diptera</collectorName>
142465 (
<specimenCount type="male">1♂</specimenCount>
, CNC);1910, USNM_ENT1071372 (
<specimenCount type="male">1♂</specimenCount>
, USNM)
</materialsCitation>
;
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Mexico">Mexico</collectingCountry>
</emphasis>
: Edo. de
<materialsCitation collectingDate="1936-07" collectingDateMax="1969-08-23" collectingDateMin="1936-07" collectorName="D. H. Janzen, H. E. Evans, P. D. Hurd, R. R. Dreisbach, G. W. Byers, R. &amp; K. Dreisbach, Skevington, Specimen" country="Mexico" county="Popocatepetel" elevation="3352" latitude="18.924212" location="Lagunas de Zempoala Nat. Park" longLatPrecision="1" longitude="-99.221565" municipality="West Slope" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Morelos">
<collectingCountry name="Mexico">Mexico</collectingCountry>
, km. 73rd to
<normalizedToken originalValue="Popocatépetel">
<collectingCounty>Popocatepetel</collectingCounty>
</normalizedToken>
,
<geoCoordinate degrees="19.075366" direction="north" orientation="latitude" precision="1" value="19.075365">19.075366</geoCoordinate>
,
<geoCoordinate degrees="98.65902" direction="west" orientation="longitude" precision="1" value="-98.65902">-98.65902</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.3519999999999994" unit="m" value="3352.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.3519999999999994" unit="m" value="3352.0">3352m</elevation>
</quantity>
,
<collectingDate value="1961-07-15">15.vii.1961</collectingDate>
,
<collectorName>D.H. Janzen</collectorName>
, EMEC354664 (
<specimenCount type="female">1♀</specimenCount>
, EMEC); Nevado Toluca,
<geoCoordinate degrees="19.110036" direction="north" orientation="latitude" precision="1" value="19.110037">19.110036</geoCoordinate>
,
<geoCoordinate degrees="99.753425" direction="west" orientation="longitude" precision="1" value="-99.753426">-99.753425</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">3200m</elevation>
</quantity>
,
<collectingDate value="1951-07-11">11.vii.1951</collectingDate>
,
<collectorName>H.E. Evans</collectorName>
, Jeff_Skevington_Specimen52560 (
<specimenCount type="male">1♂</specimenCount>
, CNC);
<geoCoordinate degrees="19.110035" direction="north" orientation="latitude" precision="1" value="19.110035">19.110035</geoCoordinate>
,
<geoCoordinate degrees="99.753423" direction="west" orientation="longitude" precision="1" value="-99.753426">-99.753423</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.444" unit="m" value="3444.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.444" unit="m" value="3444.0">3444m</elevation>
</quantity>
,
<collectingDate value="1951-07-11">11.vii.1951</collectingDate>
,
<collectorName>P.D. Hurd</collectorName>
, EMEC354662 (
<specimenCount type="male">1♂</specimenCount>
, EMEC);
<collectingMunicipality>West Slope</collectingMunicipality>
, Cortez Pass,
<geoCoordinate degrees="19.08569" direction="north" orientation="latitude" precision="1" value="19.08569">19.08569</geoCoordinate>
,
<geoCoordinate degrees="98.648296" direction="west" orientation="longitude" precision="1" value="-98.64829">-98.648296</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.743" unit="m" value="2743.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.743" unit="m" value="2743.0">2743m</elevation>
</quantity>
,
<collectingDate value="1954-07-13">13.vii.1954</collectingDate>
,
<collectorName>R.R. Dreisbach</collectorName>
, KMM0919 (
<specimenCount type="male">1♂</specimenCount>
, WIRC);
<geoCoordinate degrees="19.08569" direction="north" orientation="latitude" precision="1" value="19.08569">19.08569</geoCoordinate>
,
<geoCoordinate degrees="98.648296" direction="west" orientation="longitude" precision="1" value="-98.64829">-98.648296</geoCoordinate>
;
<geoCoordinate degrees="19.085692" direction="north" orientation="latitude" precision="1" value="19.085691">19.085692</geoCoordinate>
,
<geoCoordinate degrees="98.648297" direction="west" orientation="longitude" precision="1" value="-98.6483">-98.648297</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.743" unit="m" value="2743.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.743" unit="m" value="2743.0">2743m</elevation>
</quantity>
;~
<collectingDate value="1954-07-13">13.vii.1954</collectingDate>
, CNC_Diptera142466;CNC_Diptera142467 (
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
, CNC); Mexico City,
<geoCoordinate degrees="19.42250" direction="north" orientation="latitude" precision="1" value="19.4225">19.42250</geoCoordinate>
,
<geoCoordinate degrees="99.14389" direction="west" orientation="longitude" precision="1" value="-99.14389">-99.14389</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.048" unit="ft" value="10000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.048" unit="ft" value="10000.0">10000ft</elevation>
</quantity>
,
<collectingDate value="1936-07">vii.1936</collectingDate>
(
<specimenCount type="male">1♂</specimenCount>
HT AMNH);
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Mexico" name="Morelos">Morelos</collectingRegion>
</emphasis>
: #17
<location LSID="urn:lsid:plazi:treatment:F7344727BD36524F8F12694695D8B0D2:BE298FEF1CCD066722947DC3EFD00522" country="Mexico" county="Popocatepetel" latitude="18.924212" longLatPrecision="1" longitude="-99.221565" municipality="West Slope" name="Lagunas de Zempoala Nat. Park" stateProvince="Morelos">Lagunas de Zempoala Nat. Park</location>
,
<geoCoordinate degrees="19.04828" direction="north" orientation="latitude" precision="1" value="19.04828">19.04828</geoCoordinate>
,
<geoCoordinate degrees="99.312179" direction="west" orientation="longitude" precision="1" value="-99.31218">-99.312179</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.8649999999999998" unit="m" value="2865.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.8649999999999998" unit="m" value="2865.0">2865m</elevation>
</quantity>
,
<collectingDate value="1969-08-23">23.viii.1969</collectingDate>
,
<collectorName>G.W. Byers</collectorName>
, KMM0920 (
<specimenCount type="male">1♂</specimenCount>
, SEMC); Cuernavaca,
<geoCoordinate degrees="18.924211" direction="north" orientation="latitude" precision="1" value="18.924212">18.924211</geoCoordinate>
,
<geoCoordinate degrees="99.221567" direction="west" orientation="longitude" precision="1" value="-99.221565">-99.221567</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.133" unit="m" value="2133.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.133" unit="m" value="2133.0">2133m</elevation>
</quantity>
,
<collectingDate value="1961-07-29">29.vii.1961</collectingDate>
,
<collectorName>R. &amp; K. Dreisbach</collectorName>
, J_
<collectorName>Skevington</collectorName>
_
<collectorName>Specimen</collectorName>
50177 (
<specimenCount type="female">1♀</specimenCount>
, ANSP)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Scutellum only partly yellow pilose, black pilose anteriorly and medially. Tergite II-III extensively rufous to yellow pilose. Tergite IV dominantly black pilose, but sometimes with rufous or yellow pile medially or posteriorly. Hind trochanter not tuberculate in male.</paragraph>
<caption doi="10.3897/zookeys.1075.55862.figure4" httpUri="https://binary.pensoft.net/fig/619715" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName authorityName="Bigot" authorityYear="1882" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus</emphasis>
</taxonomicName>
dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
:
<taxonomicName authorityName="Moran &amp; Skevington" authorityYear="2021" baseAuthorityName="Giglio-Tos" baseAuthorityYear="1892" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus arctophiloides" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="arctophiloides">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus arctophiloides</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
:
<taxonomicName authorityName="Moran &amp; Thompson" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus vockerothi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vockerothi">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus vockerothi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
:
<taxonomicName authorityName="Moran &amp; Thompson" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus soletluna" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="soletluna">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus soletluna</emphasis>
</taxonomicName>
sp. nov. rufous morph
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
:
<taxonomicName authorityName="Moran &amp; Thompson" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus soletluna" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="soletluna">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus soletluna</emphasis>
</taxonomicName>
sp. nov. black morph
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
:
<taxonomicName authorityName="Bigot" authorityYear="1882" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus villosus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="villosus">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus villosus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">F.</emphasis>
<taxonomicName authorityName="Moran" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus bigoti" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bigoti">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus bigoti</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">MALE.</emphasis>
Body length: 13.1-14.8 mm. Wing length: 8.6-9.4 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Face shape as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Romaleosyrphus male face A Romaleosyrphus arctophiloides B Romaleosyrphus bigoti sp. nov. C Romaleosyrphus nephelaeus sp. nov. D Romaleosyrphus soletluna sp. nov. E Romaleosyrphus villosus F Romaleosyrphus vockerothi sp. nov." figureDoi="10.3897/zookeys.1075.55862.figure10" httpUri="https://binary.pensoft.net/fig/619721" pageId="0" pageNumber="1">10A</figureCitation>
; face silver or gold pruinose; gena black pilose posteriorly; anterior tentorial pit variable pilose: yellow or black; frons broad, ca. as long as broad at antenna, 2/3 as broad at vertex as at antenna, black pilose and silver-gold pruinose; vertex triangular, longer than broad, black pilose and brown pruinose; postocular setae black; occipital setae variable: yellow or black; antenna reddish orange.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax.</emphasis>
Matte black; postpronotum variable pilose: black or mixed black and yellow; scutum black pilose; scutellum yellow pilose, except black pilose anteromedially; postalar callus variable pilose: yellow, black or mixed black and yellow; proepimeron black pilose; posterior anepisternum yellow pilose; katepisternum yellow pilose posteriorly with broadly separated patches; metasternum variable pilose: black, yellow or mixed black and yellow; anepimeron with anterior portion yellow pilose; lower calypter with long black pile.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs.</emphasis>
Coxae black; femora black except extreme apex of femora; remainder of legs reddish; hind trochanter rounded, not tuberculate as in Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Romaleosyrphus 3 rd trochanter tubercle A Romaleosyrphus arctophiloides B Romaleosyrphus villosus." figureDoi="10.3897/zookeys.1075.55862.figure8" httpUri="https://binary.pensoft.net/fig/619719" pageId="0" pageNumber="1">8A</figureCitation>
; fore and mid-coxae black pilose; hind coxa mixed black and yellow pilose; fore femur black pilose, except occasionally with small mix of yellow pile basally; mid femur fully black pilose or with stretch of yellow pile on posterior side; hind femur black pilose; tibiae and tarsi black pilose; hind tibia as in Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Romaleosyrphus male hind tibia A Romaleosyrphus arctophiloides B Romaleosyrphus bigoti sp. nov. C Romaleosyrphus nephelaeus sp. nov. D Romaleosyrphus soletluna sp. nov. E Romaleosyrphus villosus." figureDoi="10.3897/zookeys.1075.55862.figure9" httpUri="https://binary.pensoft.net/fig/619720" pageId="0" pageNumber="1">9A</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Wing</emphasis>
.
</emphasis>
Microtrichia absent from following areas: broad anterior margin of cell cua.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
.
</emphasis>
Tergites shiny to subshiny black; tergite I with scattered, yellow pile medially, except with short black pile in lateral corners; tergite II with dense yellow pile; tergite III with dense pile which is yellow anteromedially, rufous from anterolateral corners to posteromedial margin and black in posterolateral corners; tergite IV variable, dominantly black pilose, but sometimes with rufous or yellow pile medially or posteriorly; grey pruinosity as follows: tergite I pruinose posteriorly, all of tergite II, tergite III except in posterolateral corners; sternites I-III yellow pilose and not pruinose; sternite IV variable: black or rufous pilose or some mix of the two; pile of postabdomen rufous or yellow.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
.
</emphasis>
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Romaleosyrphus male genitalia A Romaleosyrphus arctophiloides B Romaleosyrphus soletluna sp. nov. C Romaleosyrphus nephelaeus sp. nov. D Romaleosyrphus vockerothi sp. nov. E Romaleosyrphus bigoti sp. nov. F Romaleosyrphus villosus." figureDoi="10.3897/zookeys.1075.55862.figure11" httpUri="https://binary.pensoft.net/fig/619722" pageId="0" pageNumber="1">11A</figureCitation>
) Cercus yellowish brown, broader at apex, covered with long yellow pile; surstylus brown, ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, broadened basally with apical third tapering, directed ventrally and with an acute apex, ventral margin concave, undulated; pile on dorsal surface of surstylus, increasing in length posteriorly; minute spines on ventral surface and apical 3/4th of lateral inner and outer surface.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">FEMALE.</emphasis>
As male, except for usual sexual dimorphism; microtrichia on wing absent in following areas: broad anterior margin of cell cua, medial area of cell bm, anterior margin in cell dm, small region anteriorly in cell m4 near cross-vein m-cu.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Mexico.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">Trans-Mexican Volcanic Belt pine-oak forests ecoregion.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Moran &amp; Skevington" authorityYear="2021" baseAuthorityName="Giglio-Tos" baseAuthorityYear="1892" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus arctophiloides" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="arctophiloides">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus arctophiloides</emphasis>
</taxonomicName>
is the only known member of
<taxonomicName authorityName="Bigot" authorityYear="1882" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus</emphasis>
</taxonomicName>
in which the hind trochanter is not tuberculate in the male. Although males are not known for
<taxonomicName authorityName="Moran" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Romaleosyrphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Romaleosyrphus argosi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="argosi">
<emphasis italics="true" pageId="0" pageNumber="1">Romaleosyrphus argosi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="R. drysus" pageId="0" pageNumber="1" rank="species" species="drysus">
<emphasis italics="true" pageId="0" pageNumber="1">R. drysus</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="R. woodi" pageId="0" pageNumber="1" rank="species" species="woodi">
<emphasis italics="true" pageId="0" pageNumber="1">R. woodi</emphasis>
</taxonomicName>
sp. nov., males of their closest relative in the COI gene tree,
<taxonomicName lsidName="R. nephelaeus" pageId="0" pageNumber="1" rank="species" species="nephelaeus">
<emphasis italics="true" pageId="0" pageNumber="1">R. nephelaeus</emphasis>
</taxonomicName>
sp. nov., possess a tuberculate hind trochanter. It is therefore expected that males of these three species also have a tuberculate hind trochanter.
</paragraph>
<paragraph pageId="0" pageNumber="1">
We suspect that a single specimen
<normalizedToken originalValue="“CNC_Diptera142464”">&quot;CNC_Diptera142464&quot;</normalizedToken>
collected in the Sierra Madre Occidental may represent a distinct species from specimens collected in the Trans-Mexican Volcanic Belt pine-oak forests. Although no genital or discrete morphological differences could be found, the legs of this specimens are fully black pilose while those of all the others have a streak of yellow pile at the base of the fore and mid femora. Unfortunately, while a barcode was obtained for this specimen, no barcode sequences were obtained from specimens from specimens collected in the Trans-Mexican Volcanic Belt pine-oak forests.
</paragraph>
</subSubSection>
</treatment>
</document>