289 lines
36 KiB
XML
289 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1134.87853" ID-GBIF-Dataset="61572962-d525-4d7b-abe8-1f051fcf166c" ID-PMC="PMC9836628" ID-Pensoft-Pub="1313-2970-1134-129" ID-Pensoft-UUID="0973AEF60C1D54EBA960C4E252F447B0" ID-PubMed="36761113" ID-ZooBank="C79C5E40D9C64E3B816F0201713DBA77" ModsDocID="1313-2970-1134-129" checkinTime="1670590531721" checkinUser="pensoft" docAuthor="Brunke, Adam J." docDate="2022" docId="CE4046D5DAF15D83AC30F5043F1172D7" docLanguage="en" docName="ZooKeys 1134: 129-170" docOrigin="ZooKeys 1134" docPubDate="2022-12-08" docSource="http://dx.doi.org/10.3897/zookeys.1134.87853" docTitle="Iratiquedius Brunke 2022, gen. nov." docType="treatment" docUuid="9DF5C736-97AD-48AD-9BDA-A49203711E06" docUuidSource="ZooBank" docVersion="4" id="0973AEF60C1D54EBA960C4E252F447B0" lastPageNumber="129" masterDocId="0973AEF60C1D54EBA960C4E252F447B0" masterDocTitle="A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages" masterLastPageNumber="170" masterPageNumber="129" pageNumber="129" updateTime="1732489168029" updateUser="ExternalLinkService">
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<mods:title id="57B2E8FFFA7C12D77EA5174F5A52C72C">A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages</mods:title>
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<mods:namePart id="D7953B19C5B8F271539E78534AA03704">Brunke, Adam J.</mods:namePart>
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<mods:affiliation id="1647F53490BA8D5D5C2D35C606179561">Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, 960 Carling Avenue, Ottawa, Ontario, Canada</mods:affiliation>
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<mods:nameIdentifier id="3256235372726092163D6DA3007D0B8F" type="email">adam.j.brunke@gmail.com</mods:nameIdentifier>
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<mods:date id="1A884C31A9B301470B0AB2ED7A0DE02E">2022</mods:date>
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<treatment id="CE4046D5DAF15D83AC30F5043F1172D7" ID-GBIF-Taxon="206092927" LSID="urn:lsid:zoobank.org:act:9DF5C736-97AD-48AD-9BDA-A49203711E06" httpUri="http://treatment.plazi.org/id/CE4046D5DAF15D83AC30F5043F1172D7" lastPageNumber="129" pageId="0" pageNumber="129" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae" scope_tribe="Quediini">
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<subSubSection id="57953F4B8AF90206367463E4FBE41BE7" pageId="0" pageNumber="129" type="nomenclature">
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<paragraph id="969AE966E0C1D0BF388F6546AEFBE5FC" pageId="0" pageNumber="129">
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<taxonomicName id="721F7F1249A0BE9C9E6C8E2CE80B9CDE" LSID="https://zoobank.org/9DF5C736-97AD-48AD-9BDA-A49203711E06" authority="Brunke, 2022" authorityName="Brunke" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus" status="gen. nov.">Iratiquedius</taxonomicName>
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<taxonomicNameLabel id="3977DA06B8F8C67A84340AE60EB23C22" pageId="0" pageNumber="129">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="33984418904A77DA74E76E3906FFEB71" pageId="0" pageNumber="129" type="description">
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<paragraph id="AFABE076A7D15D74D1A68C6E8A4A311A" pageId="0" pageNumber="129">
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<figureCitation id="5780A5DAEE140DE352EE0FE08D822886" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">Figs 2A-F</figureCitation>
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<figureCitation id="84EAC6E6385D0B0C9740CC0B9A01739B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">, 3A, C-F</figureCitation>
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<figureCitation id="3B5D923DDE504F446F5C756121336772" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">, 4A-D</figureCitation>
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<figureCitation id="7742462359FDC3E61A82DCD1C8A78DAE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">, 6A-P</figureCitation>
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<figureCitation id="5AA738E757B9C79A61D52CAFF585D2EB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-K median lobe of aedeagus A-C in ventral view D-K in lateral view H, J internal sac partly everted, showing paired ventral sclerites I, K completely everted L-N underside of paramere showing peg setae A, B, D, E, H, I, L, M Iratiquedius seriatus (Horn) C, F, G, J, K, N I. uncifer sp. nov. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure7" httpUri="https://binary.pensoft.net/fig/779765" pageId="0" pageNumber="129">, 7A-N</figureCitation>
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<figureCitation id="CF7E947E5566AD521CE20424053BBDD6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">, 9A-E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="728012F02F7523C2A951AC1B965EF2C7" pageId="0" pageNumber="129" type="type species">
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<paragraph id="1F0635FA5C67E4EB99AEF31187457BA1" pageId="0" pageNumber="129">Type species.</paragraph>
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<paragraph id="6570AC690B139FB0A486B996B72AC1FA" pageId="0" pageNumber="129">
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<taxonomicName id="93182F3569D698803E1B3CC7C568B116" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
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<emphasis id="0EFA9808F30DA77E0A5BFD4FA5E327C7" italics="true" pageId="0" pageNumber="129">Quedius amabilis</emphasis>
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</taxonomicName>
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Smetana, 1971.
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</paragraph>
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</subSubSection>
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<subSubSection id="0D7D0EB98D14389B721A943316227EF9" pageId="0" pageNumber="129" type="etymology">
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<paragraph id="19AF59884DC3F3DCB95A9481E852801E" pageId="0" pageNumber="129">Etymology.</paragraph>
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<paragraph id="7BF5610A2A5972E40027C29CFEB7E235" pageId="0" pageNumber="129">
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The generic name is a combination of the Latin adjective
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<normalizedToken id="9EB54DB94488830AACBEE0217BDE3EAB" originalValue="‘iratus’">'iratus'</normalizedToken>
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and
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<taxonomicName id="684942D5029801692229375B935BB838" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
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<emphasis id="6660C57888E4F5C228D6A0D4A22F0F60" italics="true" pageId="0" pageNumber="129">Quedius</emphasis>
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</taxonomicName>
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. It refers to the characteristic shape of the eyes, which are strongly convergent anteriad and create a comical, angry appearance.
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</paragraph>
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</subSubSection>
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<subSubSection id="9C28B10CDC456A6EA5FA4906A2A23691" pageId="0" pageNumber="129" type="diagnosis">
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<paragraph id="EDD433C85A8C430B66A0C02734554FF7" pageId="0" pageNumber="129">Diagnosis.</paragraph>
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<paragraph id="39DA54BF5EB830BD95EDEF6467734014" pageId="0" pageNumber="129">
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Within
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<taxonomicName id="9FD3824428625BCD90A13025D1E588BE" lsidName="" pageId="0" pageNumber="129" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
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,
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<taxonomicName id="D8FFF50A3E4288E44CDB6831BF1AB464" authorityName="Brunke" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
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<emphasis id="663866B80C1F9A8FCE7E3271A2BDF738" italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
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</taxonomicName>
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can be distinguished from all other genera of the tribe by the distinctive eyes, which occupy nearly the entire lateral head margin, and are so convergent anteriad that their inner margin forms an obtuse angle with the suprantennal ridge (Fig.
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<figureCitation id="F1DF9268406447275865C90B6B229BA9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3A</figureCitation>
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). The global diagnosis is the same as the regional Nearctic diagnosis.
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</paragraph>
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</subSubSection>
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<subSubSection id="CB17AAC7B7C7FF787D11F25DD256DA07" pageId="0" pageNumber="129" type="description">
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<paragraph id="2CFB8F8E525ABE0106367B1F6D1986C3" pageId="0" pageNumber="129">Description.</paragraph>
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<paragraph id="BE54FA5C972DA9F1BC656798F63FEDFE" pageId="0" pageNumber="129">
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Medium to small rove beetles, with variable coloration (Fig.
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<figureCitation id="D62AD07512C82078F1AF4628D7573080" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2A-F</figureCitation>
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). With the character states of
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<taxonomicName id="12484F2C8A094CF54AD98159F52248D3" lsidName="" pageId="0" pageNumber="129" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
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(see
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<bibRefCitation id="5766E9F6D05547DEEAEC8AFAE3A97318" DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="396 - 421" refId="B4" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46 (2): 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. (2021)</bibRefCitation>
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) and the following: antennomere 3 longer than 2, without dense setation; outer antennomeres (8-10) about as long as wide or shorter; antennae inserted close to inner margin of eye, separated by about the width of the antennal sclerite or less; head with eyes large, strongly convex, bulging from and nearly occupying entire lateral head outline, convergent anteriad and with inner margin forming obtuse angle with suprantennal ridge (Fig.
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<figureCitation id="3A22746BC746BE6BC49F26F2286AB64A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3A</figureCitation>
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); with basal puncture doubled (at least one side), interocular punctures present in some individuals of some species (
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<taxonomicName id="6E3A24601D71EE4065987C4864675D16" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
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<emphasis id="28C62AABFB36E4687CB0A78EB0453DE2" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="17135051CA683A5C68ACEA536550DBBA" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
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<emphasis id="648BF16405D0147D394314CB0F0AD48D" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
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</taxonomicName>
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) or absent, paraocular punctures absent, genal puncture absent (Fig.
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<figureCitation id="EB69DBA7D9F7B953438D627DC2F48984" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3C, D</figureCitation>
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); frons not well-developed anterolaterad of antennal insertions; labrum notched medially, creating two lobes; apical maxillary and labial palpi fusiform and glabrous; infraorbital ridge complete to mandibles; gular sutures converging towards neck and narrowly spaced posteriad; mandibles with dorsal lateral groove absent or rudimentary, right mandible with single proximal tooth, tooth simple (Fig.
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<figureCitation id="A72046E5682C3126E2BA8D812D5851D4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3D</figureCitation>
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) or bifid (Fig.
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<figureCitation id="788CE6BE30FB5A91D6887040F072F4EA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3C</figureCitation>
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); pronotum transverse to elongate, non-explanate, with three punctures in dorsal row, sublateral row at most reaching large lateral puncture, not extended posteriad; with only single large lateral puncture (e.g., Fig.
|
||
<figureCitation id="166D70BC2A4B3FCD31514F1DA57A627A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4A-D</figureCitation>
|
||
); hypomeron strongly inflexed, not visible in lateral view; basisternum with pair of macrosetae (reduced in
|
||
<taxonomicName id="24541554AC8CF2C1A0E9DE71A60CB483" lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis id="8EE41DC48DCF959A7A52F69C30C1C412" italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="56ABCB85D2A91471DF87556763CDB8BA" lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis id="9D0EB0DE4AF0E4B89C9747A1CA772859" italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
) and well-developed longitudinal carina; scutellum impunctate; elytron with subbasal ridge complete and forming scutellar collar, disc without microsculpture between punctures; row of humeral spines present and well-developed; elytral punctation evenly distributed or in serial rows (
|
||
<taxonomicName id="2F45BE20E8A16E04C8C67B7DE417D4C3" lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis id="265FECBD78CAA5E9D31C01753670329C" italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
); foretibia with lateral spines (reduced in
|
||
<taxonomicName id="C48539A23A3B4030D8727CD4B8CF2ABD" lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis id="389072A1641520627ED88B64C3609B7A" italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
, absent in
|
||
<taxonomicName id="6324266B588D33F7A4AEBBC72087DC21" lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis id="8E72E912E3332072CC726F11A9630534" italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
) and apical spurs; metatarsomeres with disc setose; metatibia with at least three spines on outer face; abdominal tergite I with prototergal glands developed as moderately deep impressions, outer margin with row of setae; abdominal tergites not deeply impressed at base but some species with paired median impressions causing a
|
||
<normalizedToken id="4338DE715BF082D20E302070D7B58440" originalValue="‘pinched’">'pinched'</normalizedToken>
|
||
appearance; abdominal sternite III with basal transverse line forming obtuse angle at middle, not produced posteriad; aedeagus with well developed paramere bearing peg setae; at least some species with discrete, paired internal sac sclerites that may be homologous with the ventral paired sclerites described by
|
||
<bibRefCitation id="34CF08B377A46DA58CBA2065980171EE" DOI="https://doi.org/10.1111/cla.12139" author="Brunke, AJ" journalOrPublisher="Cladistics" pageId="0" pageNumber="129" pagination="427 - 451" refId="B2" refString="Brunke, AJ, Chatzimanolis, S, Schillhammer, H, Solodovnikov, A, 2016. Early evolution of the hyperdiverse rove beetle tribe Staphylinini (Coleoptera: Staphylinidae: Staphylininae) and a revision of its higher classification. Cladistics 32 (4): 427 - 451, DOI: https://doi.org/10.1111/cla.12139" title="Early evolution of the hyperdiverse rove beetle tribe Staphylinini (Coleoptera: Staphylinidae: Staphylininae) and a revision of its higher classification." url="https://doi.org/10.1111/cla.12139" volume="32" year="2016">Brunke et al. (2016)</bibRefCitation>
|
||
(e.g.,
|
||
<taxonomicName id="60F917DA2316CAB83CAA195C8033CFD0" lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis id="344E1DD43EA9B3078BB0E51D8BCE4645" italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0A149843C00EE6C0B7FEA7FE48037851" lsidName="I. uncifer" pageId="0" pageNumber="129" rank="species" species="uncifer">
|
||
<emphasis id="F8BD70401346191EC674053B8F76A7DA" italics="true" pageId="0" pageNumber="129">I. uncifer</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="58D49C2D076C3E6A607E1E55C3E1AF72" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-K median lobe of aedeagus A-C in ventral view D-K in lateral view H, J internal sac partly everted, showing paired ventral sclerites I, K completely everted L-N underside of paramere showing peg setae A, B, D, E, H, I, L, M Iratiquedius seriatus (Horn) C, F, G, J, K, N I. uncifer sp. nov. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure7" httpUri="https://binary.pensoft.net/fig/779765" pageId="0" pageNumber="129">7H-K</figureCitation>
|
||
)).
|
||
</paragraph>
|
||
<caption id="525717F2F25CBF4C94D53A0809F04816" doi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129" start="Figure 2" startId="F2">
|
||
<paragraph id="923FB733C1528F082EFF6BF04A44B5F3" pageId="0" pageNumber="129">
|
||
<emphasis id="A9C68DEA33E894E2923D4A6D85552D36" bold="true" pageId="0" pageNumber="129">Figure 2.</emphasis>
|
||
<emphasis id="49B3B646D219DAEE948EBFA111487300" bold="true" pageId="0" pageNumber="129">A-H</emphasis>
|
||
dorsal habitus of
|
||
<emphasis id="977E2612DA0A7EACACFF9F1695C6CEAB" bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||
<taxonomicName id="9AC26B8173F1217EC952952009EE5C15" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||
<emphasis id="35B3BB74119EDFD2DE4A8BA9F6EECF8D" italics="true" pageId="0" pageNumber="129">Iratiquedius amabilis</emphasis>
|
||
</taxonomicName>
|
||
(Smetana)
|
||
<emphasis id="72CD4CF64E7CCD2A8E5FE664F093AA10" bold="true" pageId="0" pageNumber="129">A</emphasis>
|
||
male holotype
|
||
<emphasis id="714083C416AE562C9C48874B7DC557F8" bold="true" pageId="0" pageNumber="129">B</emphasis>
|
||
female non-type
|
||
<emphasis id="23B5FDFB95E2FC0F91562ADEEBE636BE" bold="true" pageId="0" pageNumber="129">C, D</emphasis>
|
||
<taxonomicName id="D9CA3874213DF4C4A97040FF4ECBBEEE" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||
<emphasis id="1FF54A69C504F925A030680CDED1A213" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||
</taxonomicName>
|
||
(Smetana)
|
||
<emphasis id="EBEE33D4E3E8AAC771DB2876529491D2" bold="true" pageId="0" pageNumber="129">C</emphasis>
|
||
male non-type
|
||
<emphasis id="E6CC0E37E55424989B01104FD6BCD1FE" bold="true" pageId="0" pageNumber="129">D</emphasis>
|
||
female holotype
|
||
<emphasis id="C84656262C5578F5408CA1AAE4D901F7" bold="true" pageId="0" pageNumber="129">E</emphasis>
|
||
<taxonomicName id="16307CB96C561BA11A6BAB2DEE23D149" lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis id="5BC669F58C55E09CE2507532C4024A93" italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis id="F712FC1CA8312DFD102C409196898109" bold="true" pageId="0" pageNumber="129">F</emphasis>
|
||
<taxonomicName id="49F71B989124A7CF4BD9DDE9B04279CA" lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis id="38959E8F2246362F2FE3DA93EF246F1E" italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis id="9060BE948497954F7E5434ABE86CDE2A" bold="true" pageId="0" pageNumber="129">G</emphasis>
|
||
<taxonomicName id="022FBB7F263C6DD491D13D8667E080BD" authorityName="Horn" authorityYear="1878" class="Insecta" family="Staphylinidae" genus="Paraquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraquedius puncticeps" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="puncticeps">
|
||
<emphasis id="6DBA1767901B79C84C00AA75EC68B1F1" italics="true" pageId="0" pageNumber="129">Paraquedius puncticeps</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis id="62EEA403DC3315EA83DBE49E55F7FC24" bold="true" pageId="0" pageNumber="129">H</emphasis>
|
||
<taxonomicName id="9236B807C79B551F32144D7170DF19AF" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. marginicollis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="marginicollis">
|
||
<emphasis id="87EC40A32393B957580B0F080A24F63B" italics="true" pageId="0" pageNumber="129">P. marginicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9DAF0F78A5357812612A7D9E5D9E9BE" doi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129" start="Figure 3" startId="F3">
|
||
<paragraph id="AEFCBA726D591E6015EDFFEEDAD8A25F" pageId="0" pageNumber="129">
|
||
<emphasis id="FF7BD17BE143388B66BCA0266454F141" bold="true" pageId="0" pageNumber="129">Figure 3.</emphasis>
|
||
<emphasis id="269627ABD609572A6F12EF676EE224C3" bold="true" pageId="0" pageNumber="129">A-D</emphasis>
|
||
dorsal head
|
||
<emphasis id="BA10529640FD73A588C6E3C7F2C5AE49" bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||
showing confluence of inner eye margin and supra-antennal carina (arrow)
|
||
<emphasis id="0C6FB505D73292C4ECDA27EAAC277DE5" bold="true" pageId="0" pageNumber="129">E, F</emphasis>
|
||
abdominal tergites
|
||
<emphasis id="3E3AC754D7E175222A4CF47378947978" bold="true" pageId="0" pageNumber="129">A, D</emphasis>
|
||
<taxonomicName id="BEBA314D9FD061F535B930A23BAF8B97" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Horn" baseAuthorityYear="1878" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius seriatus" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="seriatus">
|
||
<emphasis id="C097B21D28F5807E0BA8289EA038512D" italics="true" pageId="0" pageNumber="129">Iratiquedius seriatus</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis id="FB559A4B28B2092EB9DB0507998240A4" bold="true" pageId="0" pageNumber="129">B</emphasis>
|
||
<taxonomicName id="A5474942C370315A4D3A7608815E31A9" baseAuthorityName="Casey" baseAuthorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) probus" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="probus" subGenus="Raphirus">Quedius (Raphirus) probus</taxonomicName>
|
||
(Casey)
|
||
<emphasis id="1D9774560F37CF3AD38634E4CFAE23E0" bold="true" pageId="0" pageNumber="129">C, E</emphasis>
|
||
<taxonomicName id="CD08E08B2CC977EE7FCCE301F29EC1D2" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||
<emphasis id="FD1FF2BD3C6FA3B7B9538CCF86F10B4B" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||
</taxonomicName>
|
||
(Smetana)
|
||
<emphasis id="1009907F2B008706345EEEC87330809A" bold="true" pageId="0" pageNumber="129">F</emphasis>
|
||
<taxonomicName id="8F49CD2C3BE95AAEB11EC45F99F2F54B" lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis id="AC01213ED3B1762600CC944265F8FAC2" italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
(Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (
|
||
<emphasis id="72AEAAF4455A79E307821F598D483721" bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis id="FCAEEA87C4BACA73ACCC7C602AAC2E00" bold="true" pageId="0" pageNumber="129">C-F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="637BB0E8B07032F4706DCB7048A598CD" pageId="0" pageNumber="129" type="distribution">
|
||
<paragraph id="08ECC532B4AE6EC1D3B856A527E424CE" pageId="0" pageNumber="129">Distribution.</paragraph>
|
||
<paragraph id="93DD984557F7458C6F61A21FF7DFC691" pageId="0" pageNumber="129">
|
||
<taxonomicName id="AB0A64B41A348AA208C21397E496A0AC" authorityName="Brunke" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="702DC1861210DCDC1570064A9E345BE3" italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
|
||
</taxonomicName>
|
||
is endemic to western North America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F7A67F165CAFA5DC9BA74FF67D2AA043" pageId="0" pageNumber="129" type="bionomics">
|
||
<paragraph id="0ADAA582A6020E20A09BCEDF96C93214" pageId="0" pageNumber="129">Bionomics.</paragraph>
|
||
<paragraph id="1D45103D3E2A70DFF4978AFA6EB68ACD" pageId="0" pageNumber="129">
|
||
Species of
|
||
<taxonomicName id="63C2BAE870892188CB5CCF38723E2AB4" authorityName="Brunke" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D0D23BEB50FB441F86DDC07501BBE405" italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
|
||
</taxonomicName>
|
||
are most often found in wet moss, though
|
||
<taxonomicName id="5DEEAE5EC9CCB469621B15945A95ADCC" lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis id="C5680E039FB1AC0698E849CF02119E0B" italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
is more of a generalist and can be collected from a variety of wet debris along running water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F43D457F31C70EBED5CEDD4CF809D516" pageId="0" pageNumber="129" type="comments">
|
||
<paragraph id="2C471102D1336FCD9178EE5BA8A19171" pageId="0" pageNumber="129">Comments.</paragraph>
|
||
<paragraph id="2530B099C59D1460970031FA109185DD" pageId="0" pageNumber="129">
|
||
The three included species of
|
||
<taxonomicName id="DCF1CFF53072461B56FE27A1AE5D2CAE" authorityName="Brunke" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DE8F332A4FE5B054D7A912DAB61BE57B" italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="C6011D416F53CC21232AEE0F9B9A6C36" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||
<emphasis id="EB5388756A7D057D35B5B8167015D3E7" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7A35874D779B454838F25E1371363AC3" lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis id="5F22FCEE32A23417A85BF6B972011BDA" italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="555449550C19C5F01351FB0FA06CD344" lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis id="76CEF36A4B261E36138DCFD0C9C16018" italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
) were resolved together in 'Clade
|
||
<normalizedToken id="804B2A2ACFCFD2524FC0883DE4A3448E" originalValue="A’">A'</normalizedToken>
|
||
using a phylogenomic dataset (
|
||
<bibRefCitation id="514A39D9938286A7D88A7DEDFAA7FDDA" DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="396 - 421" refId="B4" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46 (2): 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. 2021</bibRefCitation>
|
||
). Although Clade A was recovered by the concatenated but not coalescent analyses, the morphological configuration of the eyes, unique in
|
||
<taxonomicName id="532481A5C6C40C548187E2D8988AB68F" lsidName="" pageId="0" pageNumber="129" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
|
||
, provides strong further evidence for its monophyly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |