treatments-xml/data/CE/32/CB/CE32CBABFD0F54F1A4632E4460DC8FB5.xml

<document id="E1DBB5E1E1A96DBA752215CC0B6BA45D" ID-CLB-Dataset="299501" ID-DOI="10.3897/zse.100.120244" ID-GBIF-Dataset="f155e856-0a0c-4ed9-82e6-288d514654ca" ID-ZooBank="C025A8A6-116F-4BAF-94AA-6276BC84D2C8" ID-publisher-id="120244" URI-arpha="B4215F95-B9BA-5441-8E04-39CB230B9B03" URI-zoobank="http://zoobank.org/C025A8A6-116F-4BAF-94AA-6276BC84D2C8" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1720023924482" checkinUser="pensoft" docAuthor="Tsuyuki, Aoi, Reyes, Jhoe, Oya, Yuki, Wakeman, Kevin C., Leander, Brian S. &amp; Van Steenkiste, Niels W. L." docDate="2024" docId="CE32CBABFD0F54F1A4632E4460DC8FB5" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 877-895" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/120244/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Reinhardorhynchus sagamianus Tsuyuki, Reyes, Oya, &amp; Van Steenkiste 2024, sp. nov." docType="treatment" docUuid="B2534AFD-3461-4329-A62F-3C6AC8467FB2" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="895" masterDocId="B4215F95B9BA54418E0439CB230B9B03" masterDocTitle="Marine microturbellarians from Japan, with descriptions of two new species of Reinhardorhynchus (Platyhelminthes, Rhabdocoela, Koinocystididae)" masterLastPageNumber="895" masterPageNumber="877" pageNumber="877" updateTime="1720036997304" updateUser="ExternalLinkService">
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<mods:title id="893B4840B28B362757C8A83F60AF4753">Marine microturbellarians from Japan, with descriptions of two new species of Reinhardorhynchus (Platyhelminthes, Rhabdocoela, Koinocystididae)</mods:title>
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<mods:namePart id="1A34C79198D322EAF48F5E9511E39A06">Tsuyuki, Aoi</mods:namePart>
<mods:nameIdentifier id="62984E1296778C65B751966D2C43E6EA" type="ORCID">0000-0002-6001-0679</mods:nameIdentifier>
<mods:affiliation id="F1F1E8FD9262CFF1B9D1ED07E8276F0C">Faculty of Science, Hokkaido University, Sapporo, Hokkaido, 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="A434E4313422D7AB6558568AB5EAB24C">Reyes, Jhoe</mods:namePart>
<mods:nameIdentifier id="468EE15ED637BEAC5E6CBE56B53AF6F3" type="ORCID">0000-0002-4753-5577</mods:nameIdentifier>
<mods:affiliation id="9949E5A3AFACA56FC40700B10BEE1A9A">Creative Research Institution, Hokkaido University, Sapporo, Hokkaido, 001 - 0021, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="BC99ECD65E56295FB586C129A1DD10F5">Oya, Yuki</mods:namePart>
<mods:nameIdentifier id="837808E8605BC84448CC59E6772ADE23" type="ORCID">0000-0001-6862-6868</mods:nameIdentifier>
<mods:affiliation id="0DE769661B2C2329C18E0BA5B9CDB3EF">Facultad de Ciencias de la Vida y de la Salud, Universidad Científica del Sur, Lima, Peru</mods:affiliation>
</mods:name>
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<mods:namePart id="01BB42E2330903DA1B91CF6B339390E0">Wakeman, Kevin C.</mods:namePart>
<mods:nameIdentifier id="3F8115219C19AF52BA624240B5DADE89" type="ORCID">0000-0003-3501-7702</mods:nameIdentifier>
<mods:affiliation id="7E86D03C36664BD73BF17935DD328A5E">College of Arts and Sciences, J. F. Oberlin University, 3758 Tokiwa, Machida, Tokyo, 194 - 0294, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="822072AB54EA394893B14276BAEE5C1C">Leander, Brian S.</mods:namePart>
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<mods:affiliation id="AFAA1AFCA3A193A7871C983C8EB43737">Institute for the Advancement of High Education, Hokkaido University, Sapporo, Hokkaido, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="4D85DC0904D692A464B2103929F6A13B">Van Steenkiste, Niels W. L.</mods:namePart>
<mods:nameIdentifier id="599C09B769FE39ABA51F81902E7D3E24" type="ORCID">0000-0002-2676-7862</mods:nameIdentifier>
<mods:affiliation id="AEE2ACB4190FC9D8DF1C643EB4EF2204">Institute for the Advancement of High Education, Hokkaido University, Sapporo, Hokkaido, Japan &amp; Graduate School of Science, Hokkaido University, Sapporo, Hokkaido, 080 - 0810, Japan</mods:affiliation>
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<mods:date id="3E8130310A1787A6C2A145F439CF0B55">2024</mods:date>
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<treatment id="CE32CBABFD0F54F1A4632E4460DC8FB5" ID-DOI="http://doi.org/10.5281/zenodo.12636675" ID-GBIF-Taxon="234422976" ID-Zenodo-Dep="12636675" ID-arpha="CE32CBAB-FD0F-54F1-A463-2E4460DC8FB5" ID-zoobank="https://zoobank.org/B2534AFD-3461-4329-A62F-3C6AC8467FB2" LSID="urn:lsid:zoobank.org:act:B2534AFD-3461-4329-A62F-3C6AC8467FB2" LSID-ZBK="urn:lsid:zoobank.org:act:B2534AFD-3461-4329-A62F-3C6AC8467FB2" httpUri="http://treatment.plazi.org/id/CE32CBABFD0F54F1A4632E4460DC8FB5">
<subSubSection id="82FF691F864A322E25FC4871C6367F64" type="nomenclature">
<paragraph id="2AE03761834AF095559CF3DAD5CEB595">
<taxonomicName id="E83447975F060D3CC16857BFBF2E17CC" authority="Tsuyuki, Reyes, Oya, &amp; Van Steenkiste" authorityName="Tsuyuki, Reyes, Oya, &amp; Van Steenkiste" authorityYear="2024" family="Koinocystididae" genus="Reinhardorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" phylum="Platyhelminthes" rank="species" species="sagamianus" status="sp. nov.">
<emphasis id="83201D0137F287BE2E2525AA488D4E12" italics="true">Reinhardorhynchus sagamianus</emphasis>
Tsuyuki, Reyes, Oya, &amp; Van Steenkiste
</taxonomicName>
<taxonomicNameLabel id="B8EFB0C1B5A8DDDB1C4B0CF03A68A949" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="478FEE7E94E674CE84EE2A76F1C075F0" type="description">
<paragraph id="E9C982DDCBB20D54E0C6218F3A204E16">
<figureCitation id="E7BB73BD8FC8FAFC388A5DCDD6467800" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUDBG" type="material">
<paragraph id="5092F003E8794B6E56836920133F56A1">
<heading id="EFD879A7C1FF3976392F5F8F25874D6D" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="8031601BF2967B5E7500E0CBB25384CB">
<materialsCitation id="6BBBC1207FA3C56AAC39CE53029F8606" ID-GBIF-Occurrence="4902918305" collectedFrom="sandy substrates" collectingDate="2023-04-21" collectionCode="NSMT" collectorName="Aoi Tsuyuki &amp; Yuki Oya" country="Japan" latitude="35.266193" location="Sangashita beach" longLatPrecision="1" longitude="139.57211" municipality="Hayama" specimenCode="Pl 6460" specimenCount="1" stateProvince="Kanagawa Prefecture" typeStatus="holotype">
<emphasis id="92EFEDB32CE5BA151DE6A7E921D128F2" bold="true">
<emphasis id="87490738B55EE8F2BAA316ABBFCB7800" bold="true" italics="true">
<typeStatus id="D5EDAA264E86E40251014FD7E5FD36F1" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectingCountry id="8EBD79AD5598060CBA3BC11C8227C201" name="Japan">Japan</collectingCountry>
• 1
</emphasis>
; 
<collectingRegion id="228315F63831609127976F616E23FC3F" country="Japan" name="Kanagawa">Kanagawa Prefecture</collectingRegion>
, 
<collectingMunicipality id="F763212855FF48393EB924B777D771BD">Hayama</collectingMunicipality>
, 
<location id="C2312EC5F1084EB294A8A0AE715B68ED" LSID="urn:lsid:plazi:treatment:CE32CBABFD0F54F1A4632E4460DC8FB5:C2312EC5F1084EB294A8A0AE715B68ED" country="Japan" latitude="35.266193" longLatPrecision="1" longitude="139.57211" municipality="Hayama" name="Sangashita beach" stateProvince="Kanagawa Prefecture">Sangashita beach</location>
; (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECEBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[139.572111,35.266194]}">
<geoCoordinate id="C49A2FD6C0BDF4ED3AE93807A57EF118" degrees="35" direction="north" minutes="15" orientation="latitude" precision="1" seconds="58.3" value="35.266193">35 ° 15 ' 58.3 &quot; N</geoCoordinate>
, 
<geoCoordinate id="6DDFA3A06CCCC769652C059892C57257" degrees="139" direction="east" minutes="34" orientation="longitude" precision="1" seconds="19.6" value="139.57211">139 ° 34 ' 19.6 &quot; E</geoCoordinate>
</named-content>
); 
<date id="E95719AF91358F393D691FCDD46DDFA8" value="2023-04-21">
<collectingDate id="97021828312E3429E664478085C06EA1" value="2023-04-21">21 April 2023</collectingDate>
</date>
; 
<collectedFrom id="DB80ACA1D8DEA0FE67073A3E197359C6">sandy substrates</collectedFrom>
; 
<collectorName id="250549B72248FC8AA7B707D5F5082DBD">Aoi Tsuyuki</collectorName>
and 
<collectorName id="471561E08D82CEA7BAB898688B9B1ADD">Yuki Oya</collectorName>
leg.; one individual worm in a single slide; [Holotype: 
<abbrev id="ABBRID0EHEBG" xlink_title="National Museum of Nature and Science, Tokyo">
<collectionCode id="DA357763F63EE6D32BA83B7F3EB73ED8" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" type="">NSMT</collectionCode>
</abbrev>
- 
<specimenCode id="19EBD6F8D8BF30CAE119C8326B40BF90">Pl 6460</specimenCode>
].
</materialsCitation>
</paragraph>
<paragraph id="D84B3E90E801B1A6AA34ED8A4593B36F">
<materialsCitation id="6BBB43E899B9407A6C734E8D9FD00D6F" ID-GBIF-Occurrence="4902918304" collectedFrom="sandy substrates" collectingDate="2023-08-30" collectionCode="NSMT" collectorName="Yuki Oya" country="Japan" latitude="35.266193" location="Japan" longLatPrecision="1" longitude="139.57211" municipality="Hayama" specimenCode="DNA 56985" specimenCount="1" stateProvince="Kanagawa" typeStatus="paratype">
<emphasis id="08A6C5DA269880B45B080BBCAFD55FD8" bold="true">
<emphasis id="2D8070F4ED8CB62FE279B76C5C10DB84" bold="true" italics="true">
<typeStatus id="EDD5C2D77C312A0DB1E7C28DBCBB6532" type="paratype">Paratype</typeStatus>
</emphasis>
: 
<collectingCountry id="9F20BB9D015F403DA6FF8C4F021EF916" name="Japan">Japan</collectingCountry>
• 1
</emphasis>
; locality same as for holotype; 
<date id="BB8B8C6265F9E7F31CF519338EEF6E40" value="2023-08-30">
<collectingDate id="E7E4E11D17007217E78A109B8DFECB05" value="2023-08-30">30 Aug 2023</collectingDate>
</date>
; 
<collectorName id="D78E2073A43753C01279F782A0B58A01">Yuki Oya</collectorName>
leg.; genomic DNA extract from one individual stored at - 20 ° C; GenBank: 
<ext-link id="6FBC97CA7842FF92616AE6B545A100CE" ext-link-type="gen" xlink_href="LC807767" xlink_type="simple">
<accessionNumber id="538EF21F9E16786A5468FA2A251475A2" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/LC807767" type="GenBank">LC 807767</accessionNumber>
</ext-link>
(18 S rDNA; 1,654 bp), 
<ext-link id="498B4E7DC94618D62F41C7B5338A0D96" ext-link-type="gen" xlink_href="LC807770" xlink_type="simple">
<accessionNumber id="B3873FF1A4F16BF29F4C4F35622C2A49" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/LC807770" type="GenBank">LC 807770</accessionNumber>
</ext-link>
(28 S rDNA; 1,667 bp); [Paratype: 
<abbrev id="ABBRID0E3EBG" xlink_title="National Museum of Nature and Science, Tokyo">
<collectionCode id="3249F2A45861570EBE36C77766AF0617" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" type="">NSMT</collectionCode>
</abbrev>
- 
<specimenCode id="A8D9E5A39271F31087C5C22FDA69CF66" collectionCode="NSMT" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" type="">DNA 56985</specimenCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAFBG" type="type locality">
<paragraph id="EFAD8920EB9777E8172D342EE1B64519">
<heading id="90C691CB53F0FC65653662D74D7C0E42" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="FCA165F7922A9276C5648DEB32D2C131">
<collectingCountry id="FC0C21BF45F9C778A1BBE182D4EE6844" name="Japan">Japan</collectingCountry>
, 
<collectingRegion id="9739C129A167E25CBE1A465A14F37F87" country="Japan" name="Kanagawa">Kanagawa Prefecture</collectingRegion>
, Hayama, Sangashita Beach (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJFBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[139.572111,35.266194]}">
<geoCoordinate id="44E18D9F474A4E3DB151B7241941019C" degrees="35" direction="north" minutes="15" orientation="latitude" precision="1" seconds="58.3" value="35.266193">35 ° 15 ' 58.3 &quot; N</geoCoordinate>
, 
<geoCoordinate id="D9B69AE801B7662742DE482D99BE370C" degrees="139" direction="east" minutes="34" orientation="longitude" precision="1" seconds="19.6" value="139.57211">139 ° 34 ' 19.6 &quot; E</geoCoordinate>
</named-content>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOFBG" type="diagnosis">
<paragraph id="326F4F4D4547BAC620AEA09A8718DC00">
<heading id="2ED7A488E004457F4D4748326572EDDC" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="ABA9664269742E215CD2E3128DC23BDF">
Species of 
<taxonomicName id="C13D821EBD01B6C83760BFCE556D2C5F" authorityName="Diez, Monnens, &amp; Artois" authorityYear="2021" family="Koinocystididae" genus="Reinhardorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" phylum="Platyhelminthes" rank="genus">
<emphasis id="C7FF8F40E66715CD9614D0341B6F9DD9" italics="true">Reinhardorhynchus</emphasis>
</taxonomicName>
with a copulatory organ encompassing an armed cirrus, an unarmed accessory cirrus, and two distal hooks. Bipartite armed cirrus consisting of two sacs lined with a continuous ± 295.2 - μm-long sclerotized belt of overlapping scale-like spines. Larger sac with more spaced-out, triangular, ± 20.1 μm-long spines on the proximal end of the belt. Spines gradually decrease in size distally as the belt runs towards and folds into the smaller sac, increasing in size (± 6.1 to ± 22.2 μm long) towards the proximal tip of the smaller sac, and decreasing in size again from the proximal to the distal tip of the smaller sac. Unarmed accessory cirrus as an elongated sac. The larger distal hook is slightly curved, ± 111.5 μm long and ± 43.5 μm wide at its base; its base is provided with a slightly curved, ± 42.3 μm-long projection with a blunt distal tip forming a ~ 90 ° angle with the axis of the hook. The smaller hook is funnel-shaped, ± 58.9 μm long and ± 46.6 μm wide at its base.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2FBG" type="description">
<paragraph id="965CAF70E10A347C5F8E8474FED4BF54">
<heading id="AFE270C153A16E2E2BB120D6EC1AAF0C" reason="title">Description.</heading>
</paragraph>
<paragraph id="5EE68555480E9BD67909A176FDCDE536">
<emphasis id="F425728CBF4867D1FE8E92BC61EF37D2" bold="true" italics="true">General morphology.</emphasis>
Live mature specimens are 1500–1800 µm long (n = 2), with two eyes (Fig. 
<figureCitation id="5D5987FBC8F6522710430114EA7E9CB7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 A, B</figureCitation>
). The proboscis is 297–304 μm (n = 2) long in swimming animals and is characteristic for koinocystidids (
<bibRefCitation id="BFDE4808F740D026C88460BE3A3980CB" DOI="10.1111/j.1463-6409.1972.tb00673.x" author="Brunet" firstAuthor="Brunet" issue="3" journalOrPublisher="Zoologica Scripta" pagination="157-174" refId="B16" refString="Brunet M (1972) Koinocystididae de la région de Marseille (Turbellaria, Kalyptorhynchia). Zoologica Scripta 1 (3): 157–174. https://doi.org/10.1111/j.1463-6409.1972.tb00673.x" title="Koinocystididae de la région de Marseille (Turbellaria, Kalyptorhynchia)." volume="1" year="1972">Brunet 1972</bibRefCitation>
; 
<bibRefCitation id="C123E6F828FEFCF5A8461C4E5DEC8083" DOI="10.1111/j.1463-6409.1980.tb00666.x" author="Karling" firstAuthor="Karling" issue="1-4" journalOrPublisher="Zoologica Scripta" pagination="241-269" refId="B42" refString="Karling TG (1980) Revision of Koinocystididae (Turbellaria). Zoologica Scripta 9 (1–4): 241–269. https://doi.org/10.1111/j.1463-6409.1980.tb00666.x" title="Revision of Koinocystididae (Turbellaria)." volume="9" year="1980">Karling 1980</bibRefCitation>
; 
<bibRefCitation id="4E9687BC23025C5D3605BD840B8330D1" DOI="10.11646/zootaxa.4948.4.1" author="Diez" etAl="et al." firstAuthor="Diez" issue="4" journalOrPublisher="Zootaxa" pagination="451-500" refId="B21" refString="Diez YL, Monnens M, Aguirre RI, Yurduseven R, Jouk P, Van Steenkiste NWL, Leander BS, Schockaert E, Reygel P, Smeets K, Artois T (2021) Taxonomy and phylogeny of Koinocystididae (Platyhelminthes, Kalyptorhynchia), with the description of three new genera and twelve new species. Zootaxa 4948 (4): 451–500. https://doi.org/10.11646/zootaxa.4948.4.1" title="Taxonomy and phylogeny of Koinocystididae (Platyhelminthes, Kalyptorhynchia), with the description of three new genera and twelve new species." volume="4948" year="2021">Diez et al. 2021</bibRefCitation>
). The pharynx (ph, Fig. 
<figureCitation id="8B78EB61D99B0170C3E1E9F458AC3DE3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 A, B</figureCitation>
) is positioned near the body’s midpoint and has an approximate diameter of 231–258 μm (n = 2) in the live specimens. The oesophagus is visible as a clear zone surrounded by oesophageal glands behind the pharynx. It empties into the intestine, which is situated in the posterior portion of the body. The male and female reproductive systems are mainly located in the third posterior region of the body.
</paragraph>
<caption id="1B38559169814943362DB6FC7E889030" ID-DOI="10.3897/zse.100.120244.figure3" ID-arpha="DACD3884-6536-54C7-A1A9-B340B1B76778" httpUri="https://binary.pensoft.net/fig/1084561" startId="F3">
<paragraph id="DE0095E5F951BD59A23C830BB386928C">
<label id="02835496E6664B761898718339FC0C89">Figure 3.</label>
</paragraph>
<paragraph id="3C7C507FFD3D5BFD6F1EB037B0EBA0EF">
<taxonomicName id="8149430147DE9FCE72BCF389A82E91FC" authorityName="Tsuyuki &amp; Reyes &amp; Oya &amp; Wakeman &amp; Leander &amp; Van Steenkiste" authorityYear="2024" family="Koinocystididae" genus="Reinhardorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" phylum="Platyhelminthes" rank="species" species="sagamianus" status="sp. nov.">
<emphasis id="DE901A66387A772A6FC58905112F6230" italics="true">Reinhardorhynchus sagamianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56DDA3AA61A99E33142E13A976D075F6" rank="species">n. sp.</taxonomicNameLabel>
, 
<abbrev id="ABBRID0ENHBG" xlink_title="National Museum of Nature and Science, Tokyo">NSMT</abbrev>
- DNA 56985 (paratype) (
<emphasis id="1BA4E7CB7788276A3F87D7039DBD621C" bold="true">A</emphasis>
) and 
<abbrev id="ABBRID0ETHBG" xlink_title="National Museum of Nature and Science, Tokyo">NSMT</abbrev>
- Pl 6460 (holotype) (
<emphasis id="4E8784629EE352086693EEC40E2106C1" bold="true">B</emphasis>
– 
<emphasis id="074E6CBBEB976E6228B744F1CA2D57D3" bold="true">H</emphasis>
). 
<emphasis id="EB0BC1A3ACAE88D9CC1B4AE23F377658" bold="true">A, B.</emphasis>
Micrograph and drawing of live animals. 
<emphasis id="F0819C7A625C8A7B627D454FC4CE1AF0" bold="true">C.</emphasis>
Detail of the male copulatory organ in a live animal. 
<emphasis id="D585D02ADBB2C9AD3B01E5E1FBE02DC7" bold="true">D, E.</emphasis>
Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. 
<emphasis id="A58EE597D3D46F0C2761D1259578CA41" bold="true">F – H.</emphasis>
Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New.
</paragraph>
</caption>
<paragraph id="30A12A1676D5DCF6CF3C715E2CA3F5D3">
<emphasis id="09F8B1A8FE448CFD9CDECC166155E712" bold="true" italics="true">Male reproductive system.</emphasis>
Paired testes anterior to the pharynx are on each side of the body (t, Fig. 
<figureCitation id="187F15AA16A12280C288766DAFB3DD2C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 A, B</figureCitation>
). A pair of sac-like seminal vesicles (sv, Fig. 
<figureCitation id="300C152C20FAE1A3852E1A49C5739265" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B – D</figureCitation>
) fuse distally before entering the copulatory organ and forming an ejaculatory duct (ed; Fig. 
<figureCitation id="1694D04FD8E7929231B14492AACEF522" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 C, D</figureCitation>
). The piriform copulatory organ is 323–376 μm (n = 2) long and encompasses a proximal prostate vesicle, a bipartite armed cirrus, and an unarmed accessory cirrus. Distally, it bears two large hooks of varying shapes (h 1, h 2, Fig. 
<figureCitation id="0126ACD2A3A2F71F914C8F766A65A6DF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 D</figureCitation>
). The prostate vesicle consists of one type of intracapsular prostate gland (pg 1; Fig. 
<figureCitation id="071074212A5D5369DCF693CC101F0971" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B – D</figureCitation>
) opening into the transition zone between the ejaculatory duct and armed cirrus through filiform ducts (Fig. 
<figureCitation id="990A41B0BDF5FA864FFB12E0FE697B30" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B – D</figureCitation>
; Suppl. material 
<xref id="6BAEBDEB0818E091EB400A6BE360F6C1" ref-type="supplementary-material" rid="S1">1</xref>
). A second type of extracapsular prostate gland (pg 2; Fig. 
<figureCitation id="8F96010E2CAF7525A4ACE0DDA7A98FA8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B – D</figureCitation>
) enters the copulatory organ proximally. The armed cirrus bears small spines on its entire surface (cia, Fig. 
<figureCitation id="0BF608BAFD0174788D8D46D8ED33A774" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B – D</figureCitation>
; Suppl. materials 
<xref id="69CAD17BA42D5FFEF4DFAC97AA5D42F0" ref-type="supplementary-material" rid="S1">1</xref>
, 
<xref id="DD1965902FB73CAF6F95B9470AB3E294" ref-type="supplementary-material" rid="S2">2</xref>
) and has two sacs of differing sizes, both of which are equipped with an interconnecting belt of overlapping, scale-like spines (Fig. 
<figureCitation id="23FAEF7F63B5EC1258FCD5F45AFCC34A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B – F</figureCitation>
; Suppl. material 
<xref id="67562806E15B8BA0F0F8281EE2F7982C" ref-type="supplementary-material" rid="S1">1</xref>
). The spines of the proximal part of the spiny belt start in the larger sac. Here, the scale-like spines are triangular, more spaced out, and 20.1 μm long (cps, Fig. 
<figureCitation id="BB8A41D4B6BADA796D5ADABA1ADF4943" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B – F</figureCitation>
). Gradually, these spines become less triangular, more overlapping, and decrease in size (8.8 μm long) until they reach the transition to the smaller sac, where they fold backwards and continue into the smaller sac (cms, Fig. 
<figureCitation id="5AC54443968B2432236A382FD8251659" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 C, E, F</figureCitation>
; Suppl. materials 
<xref id="F71A1B55F6419A6B541C147F5ACB4097" ref-type="supplementary-material" rid="S1">1</xref>
– 
<xref id="CA0D8BE66BD4F4BB94DF9ECD503C0639" ref-type="supplementary-material" rid="S3">3</xref>
). From this fold, the spines gradually increase in size (from 6.1 to 22.2 μm long) towards the proximal tip of the smaller sac, where they fold again to continue along the distal side of this sac while gradually decreasing in size again (cds, Fig. 
<figureCitation id="1AB761DF9C80A7CB07C5A273B52A86DC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 C, E, F</figureCitation>
). The total length of this spiny belt is 295.2 μm. The unarmed accessory cirrus (ciu, Fig. 
<figureCitation id="CA763ACB3835231BDC85F73B4B88D9F2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 C, D</figureCitation>
; Suppl. material 
<xref id="EF771F71D40833BE3512D230C659BC17" ref-type="supplementary-material" rid="S1">1</xref>
) runs alongside the armed cirrus from the proximal end of the copulatory organ to the distal end of the armed cirrus. It appears as an elongated sac that narrows proximally. It is possible that this narrow proximal part of the unarmed accessory cirrus connects to the bundles of extracapsular prostate glands (pg 2, Fig. 
<figureCitation id="172BCBF12F68FA15E06BCF2CFC0CC21F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 C</figureCitation>
) visible around the copulatory organ and seminal vesicles, but this connection could not be established with certainty. The larger distal hook is 111.5 μm long and 43.5 μm wide at its base (h 1, Fig. 
<figureCitation id="26A5B6F8947CC674761F2D16A3FC41D6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 G, H</figureCitation>
; Suppl. material 
<xref id="749C75D0B8994A426985C3F41CB11C22" ref-type="supplementary-material" rid="S1">1</xref>
). One side of the base features a sturdy, slightly curved projection with a blunt distal tip. The projection itself is 42.3 μm long and forms an ~ 90 ° angle with the axis of the hook. The smaller distal hook is 58.9 μm long and has a 46.6 μm-wide base (h 2, Fig. 
<figureCitation id="D15DBFFD54C14C77920A9C5F4E2B6B58" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 G, H</figureCitation>
).
</paragraph>
<paragraph id="66B2A781EBA6C77F0130862F69BD0A82">
<emphasis id="A431603E0DD58E657E1F49C7ABA05EA7" bold="true" italics="true">Female reproductive system.</emphasis>
The vitellaria (vi; Fig. 
<figureCitation id="488BDF60F54A4BA10B122544140A7AAC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 A</figureCitation>
) extends from the rear end of the pharynx to the posterior body end. Paired ovaries with oocytes arranged in a single line are situated anterior to the copulatory organ (ov, Fig. 
<figureCitation id="54B7CBA8E6F4AEDCA8628DAEBF0921A7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B, C</figureCitation>
). The bursa (bu, Fig. 
<figureCitation id="32DAAC668459E1FD1A9937C4BB970C39" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B</figureCitation>
) opens into the female atrium through a muscular bursal stalk (bs, Fig. 
<figureCitation id="BE833714F2E478589015CCC6831E1351" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B</figureCitation>
). The female atrium also receives the female duct (fd, Fig. 
<figureCitation id="2A7D480761FF3E593791CA3B6B724540" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B</figureCitation>
), which consists of a narrow distal duct in which bundles of female glands (fg, Fig. 
<figureCitation id="871AACD11771C1FB1B6A0283DBE06D3F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reinhardorhynchus sagamianus n. sp., NSMT - DNA 56985 (paratype) (A) and NSMT - Pl 6460 (holotype) (B – H). A, B. Micrograph and drawing of live animals. C. Detail of the male copulatory organ in a live animal. D, E. Drawing and micrograph of the belt of overlapping spines in the armed cirrus of the male copulatory organ in a whole mounted specimen fixed in Entellan New. F – H. Drawings and micrograph of the distal hooks associated with the male copulatory organ in a whole mounted specimen fixed in Entellan New." figureDoi="10.3897/zse.100.120244.figure3" httpUri="https://binary.pensoft.net/fig/1084561">3 B</figureCitation>
) discharge, and a wide, muscular proximal part that receives the oviducts.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOMBG" type="etymology">
<paragraph id="159B248674F19E5577D24367EE1E3204">
<heading id="B3FF74178AD27C4F85430591801B7D48" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="49EC7796DD060457B5C87D8CE5D7D54E">
The species epithet 
<emphasis id="F6EE1857DFAB967B3AD386B5E09362CA" italics="true">sagamianus</emphasis>
refers to the type locality, which is located in Sagami Bay.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5MBG" type="distribution">
<paragraph id="634066FB27A8545F84225736027F7675">
<heading id="A4284B310F79C84CCF6AFC4808458B56" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EE1F86C05475DDF6A5E37F081FA4CD36">
Kanawaga, Sangashita Beach, 
<collectingCountry id="1F1B791EDE31F27D1E40CF1F3D7B8E13" name="Japan">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>