466 lines
57 KiB
XML
466 lines
57 KiB
XML
<document id="F15F5C1A3213EBD3DF3291D35C82CEAE" ID-DOI="10.11646/zootaxa.3718.5.4" ID-GBIF-Dataset="322d05d4-fbc0-4ef9-b50e-7392eda1ff0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="216599" ID-ZooBank="2C30C73E-3154-4AD1-A3FF-CA6F7210194A" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460687034545" checkinUser="plazi" docAuthor="Yin, Zi-Wei & Li, Li-Zhen" docDate="2013" docId="C6722A3AEC4CFFD5FF3DFE10F1F1FE27" docLanguage="en" docName="zt03718p482.pdf" docOrigin="Zootaxa 3718 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Smetanabatrus Yin & Li, 2013, new species" docType="treatment" docVersion="8" lastPageNumber="480" masterDocId="3A4B5242EC4DFFD6FFAAFFC3F46AFFC8" masterDocTitle="Smetanabatrus kinabalu (Staphylinidae: Pselaphinae: Batrisitae), a new genus and new species from Sabah, Borneo" masterLastPageNumber="482" masterPageNumber="477" pageNumber="478" updateTime="1698351348321" updateUser="plazi">
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<mods:mods id="E99AE63E577EC840940AA8B24B7751DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D9879F3FBEA0D573CED847720BCE946A">
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<mods:title id="F562C5DAA2E66D1453BE94830224E593">Smetanabatrus kinabalu (Staphylinidae: Pselaphinae: Batrisitae), a new genus and new species from Sabah, Borneo</mods:title>
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<mods:name id="29AEB146EAE135B1075B9EE8526E4F8F" type="personal">
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<mods:role id="1B00F5E851E2A22CC7759E398B18C48F">
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<mods:roleTerm id="991D978A5F56AD42B826950701CB1BEF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3E96A9870F2C7A6AC977FC5F60FB8C15">Yin, Zi-Wei</mods:namePart>
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</mods:name>
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<mods:name id="157C8BB15FE39DB40C80913C0A4FA9A2" type="personal">
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<mods:role id="E0F548FD341DAC726A8ED7147F8498F4">
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<mods:roleTerm id="1AB87B2916D560B673EB64DCF8680B19">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CABD7CE87A800A05473A9C2BA9F1D3BB">Li, Li-Zhen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1EA8A250FCCB3DBD0581FC5807AE61A7">text</mods:typeOfResource>
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<mods:titleInfo id="656074313A6736D6A3320356790F11A0">
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<mods:title id="517A0F5E945F6EBD4A8A1BE968BF6F21">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D3383FF720D505CDC20DFCA8191766E9">
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<mods:date id="9FFC757B9ADA0FAD8E3F1B67B15AD973">2013</mods:date>
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<mods:detail id="918B44F6162CE28FFC678F2D2348D31E" type="volume">
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<mods:number id="27B9A38EE8BC92CCC54FA8F950401B8B">3718</mods:number>
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</mods:detail>
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<mods:detail id="9B25265C929DEBD4D3090896EF8A2E2E" type="issue">
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<mods:number id="A7C847A17126FB3FB331EB9EB647EC00">5</mods:number>
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<mods:classification id="78AD1A311213DD8284D908697D79BC8E">journal article</mods:classification>
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<mods:identifier id="9E2B4650E41CE4FCF7B3B032E1E9DE8D" type="DOI">10.11646/zootaxa.3718.5.4</mods:identifier>
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<mods:identifier id="C47D97A068E5DD2F1975D4DF0BCB3A03" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A8B8EDE5D5E823DECB01CFCE0A2B0FE2" type="Zenodo-Dep">216599</mods:identifier>
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<treatment id="C6722A3AEC4CFFD5FF3DFE10F1F1FE27" ID-DOI="http://doi.org/10.5281/zenodo.6150399" ID-GBIF-Taxon="119580779" ID-Zenodo-Dep="6150399" LSID="urn:lsid:plazi:treatment:C6722A3AEC4CFFD5FF3DFE10F1F1FE27" httpUri="http://treatment.plazi.org/id/C6722A3AEC4CFFD5FF3DFE10F1F1FE27" lastPageId="3" lastPageNumber="480" pageId="1" pageNumber="478">
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<subSubSection id="06C1C8A7EC4CFFD7FF3DFE10F732FDF8" pageId="1" pageNumber="478" type="nomenclature">
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<paragraph id="4E649B2CEC4CFFD7FF3DFE10F603FE26" blockId="1.[151,617,467,494]" box="[151,617,467,494]" pageId="1" pageNumber="478">
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<heading id="152C2C40EC4CFFD7FF3DFE10F603FE26" bold="true" box="[151,617,467,494]" fontSize="11" level="1" pageId="1" pageNumber="478" reason="1">
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<emphasis id="7CAF473EEC4CFFD7FF3DFE10F603FE26" bold="true" box="[151,617,467,494]" pageId="1" pageNumber="478">
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<taxonomicName id="89DBE0AFEC4CFFD7FF3DFE10F527FE25" ID-CoL="8XPWV" box="[151,333,467,493]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="7CAF473EEC4CFFD7FF3DFE10F527FE25" bold="true" box="[151,333,467,493]" italics="true" pageId="1" pageNumber="478">Smetanabatrus</emphasis>
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</taxonomicName>
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Yin and Li, new genus
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4E649B2CEC4CFFD7FF3DFDD8F732FDF8" blockId="1.[151,856,538,560]" box="[151,856,538,560]" pageId="1" pageNumber="478">
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<typeStatus id="9160258EEC4CFFD7FF3DFDD8F4A3FDF8" box="[151,201,539,560]" pageId="1" pageNumber="478">Type</typeStatus>
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species:
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<taxonomicName id="89DBE0AFEC4CFFD7FE8CFDD8F676FDF8" box="[294,540,539,560]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="478" phylum="Arthropoda" rank="species" species="kinabalu" status="sp. nov.">
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<emphasis id="7CAF473EEC4CFFD7FE8CFDD8F676FDF8" box="[294,540,539,560]" italics="true" pageId="1" pageNumber="478">Smetanabatrus kinabalu</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="679CFA45EC4CFFD7FD83FDD8F6C9FDF8" box="[553,675,538,560]" pageId="1" pageNumber="478" rank="species">new species</taxonomicNameLabel>
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(here designated)
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</paragraph>
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</subSubSection>
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<subSubSection id="06C1C8A7EC4CFFD7FF3DFD9EF130FD76" pageId="1" pageNumber="478" type="diagnosis">
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<paragraph id="4E649B2CEC4CFFD7FF3DFD9EF130FD76" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">
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<emphasis id="7CAF473EEC4CFFD7FF3DFD9EF57BFDBE" bold="true" box="[151,273,605,630]" pageId="1" pageNumber="478">Diagnosis.</emphasis>
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Head trapezoidal; lacking frontal rostrum, antennal tubercles distinct, antennomeres elongate. Pronotal disc barely convex, with median longitudinal sulcus, antebasal spines present, lacking spines on lateral margins. Elytra with three basal foveae, discal striae present. Tergite IV (first visible tergite) lacking marginal carinae.
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</paragraph>
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</subSubSection>
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<subSubSection id="06C1C8A7EC4CFFD7FF6DFD0AF0D7F83A" pageId="1" pageNumber="478" type="description">
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<paragraph id="4E649B2CEC4CFFD7FF6DFD0AF5EAFC72" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">
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<emphasis id="7CAF473EEC4CFFD7FF6DFD0AF533FD2A" bold="true" box="[199,345,713,738]" pageId="1" pageNumber="478">Description.</emphasis>
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Length
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<quantity id="892336C9EC4CFFD7FE14FD09F602FD29" box="[446,616,714,738]" metricMagnitude="-3" metricUnit="m" metricValue="2.82" metricValueMax="2.9" metricValueMin="2.74" pageId="1" pageNumber="478" unit="mm" value="2.82" valueMax="2.9" valueMin="2.74">2.74–2.90 mm</quantity>
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. Head trapezoidal; lacking frontal rostrum and frontal fovea, antennal tubercles distinct, area between them broadly depressed; vertexal foveae nude, located below level of posterior margins of eyes, lacking sulcus connecting foveae, median sulcus present; with 11 antennomeres, clubs indistinct; ocular-mandibular carinae present; eyes round, ocular canthi conspicuously spinose; maxillary palpi with palpomeres II elongate, basally pedunculate, III nearly quadrate, IV with mesal margins greatly lobed; single large gular fovea present, lacking gular carina or sulcus, gula greatly extended to form elongate projections at anterolateral margin.
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</paragraph>
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<paragraph id="4E649B2CEC4CFFD7FF6DFC05F1F7FB81" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">Pronotum with distinct median longitudinal sulcus, followed by elongate median pit, with short median carina at base; lateral longitudinal impressions shallow; with lateral antebasal foveae, antebasal spines prominent, thin laterodorsal carinae extending anteriorly from antebasal spines; inner- and outer pair of basolateral foveae present; lateral margins lacking spines. Paranotal carinae extending from base to midlength; lateral procoxal foveae present.</paragraph>
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<paragraph id="4E649B2CEC4CFFD7FF6DFB96F613FB7E" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">Each elytron with three basal foveae, discal striae extending past elytral midlength; lacking subbasal foveae; sutural striae complete; subhumeral foveae absent, marginal striae extending through basal two-thirds of flank; apicolateral margins shallowly emarginate.</paragraph>
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<paragraph id="4E649B2CEC4CFFD7FF6DFB02F0F6FAEA" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">Mesoventrite with forked lateral foveae, anterior forks as wide as median ones, median foveae widely separated, opening in shared transverse cavity; lateral mesocoxal foveae present; lateral metaventral foveae moved medially, close; metacoxae moderately separated; posterior margin with narrow median slit.</paragraph>
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<paragraph id="4E649B2CEC4CFFD7FF6DFAEEF142FA32" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">Abdominal characters largely overlapped in male by sexual modifications. Tergite IV (first visible tergite) longest, mediobasal foveae at inner margins of basal sulcus, two pairs of basolateral foveae in lateral sulci; lacking dorsal and marginal carinae; tergites V–VI short, subequal in length, VII longer than VI, V–VII lacking mediobasal sulcus and foveae, with basolateral foveae. Sternite IV (second visible sternite) longest, about twice length of V in midlength, mediobasal foveae at inner margins of basolateral sulci, with basolateral foveae and marginal sockets; V–VII subequal in length, each with basolateral foveae. Second and third tarsomeres subequal in length.</paragraph>
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<paragraph id="4E649B2CEC4CFFD7FF6DF9C6F79CF98A" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">Males with metatibiae, tergites IV–VI, and sternite V modified, profemora spinose. Aedeagal median lobe elongate; dorsal lobe slender; with large, semi-sclerotized ventral membrane.</paragraph>
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<paragraph id="4E649B2CEC4CFFD7FF6DF98EF0D7F83A" blockId="1.[151,1437,605,2034]" pageId="1" pageNumber="478">
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<emphasis id="7CAF473EEC4CFFD7FF6DF98EF5DDF9AE" bold="true" box="[199,439,1613,1638]" pageId="1" pageNumber="478">Comparative notes.</emphasis>
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The prominent antennal tubercles, the presence of prominent antebasal spines and laterodorsal carinae of the pronotum, the lack of a pronotal antebasal sulcus, and the presence of three basal elytral foveae places
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<taxonomicName id="89DBE0AFEC4CFFD7FE9FF956F5B7F965" box="[309,477,1685,1709]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="7CAF473EEC4CFFD7FE9FF956F5B7F965" box="[309,477,1685,1709]" italics="true" pageId="1" pageNumber="478">Smetanabatrus</emphasis>
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</taxonomicName>
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near a group of genera centered on
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<taxonomicName id="89DBE0AFEC4CFFD7FCC4F956F0D6F966" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[878,1212,1685,1710]" class="Insecta" family="Staphylinidae" genus="Coryphomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="478" phylum="Arthropoda" rank="genus">
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<emphasis id="7CAF473EEC4CFFD7FCC4F956F07DF965" box="[878,1047,1685,1709]" italics="true" pageId="1" pageNumber="478">Coryphomodes</emphasis>
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Jeannel, 1960
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</taxonomicName>
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.
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<taxonomicName id="89DBE0AFEC4CFFD7FB6CF956F104F965" box="[1222,1390,1685,1709]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="7CAF473EEC4CFFD7FB6CF956F104F965" box="[1222,1390,1685,1709]" italics="true" pageId="1" pageNumber="478">Smetanabatrus</emphasis>
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</taxonomicName>
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can be separated from
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<taxonomicName id="89DBE0AFEC4CFFD7FECCF97AF67AF919" box="[358,528,1721,1745]" class="Insecta" family="Staphylinidae" genus="Coryphomodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="478" phylum="Arthropoda" rank="genus">
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<emphasis id="7CAF473EEC4CFFD7FECCF97AF67AF919" box="[358,528,1721,1745]" italics="true" pageId="1" pageNumber="478">Coryphomodes</emphasis>
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</taxonomicName>
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by the distinctly transverse head and pronotum, much stouter body form, and the lack of dorsal carinae on abdominal tergite IV. The pronotal lateral margins lacking spines in
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<taxonomicName id="89DBE0AFEC4CFFD7FB16F91EF10EF93D" box="[1212,1380,1757,1781]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="7CAF473EEC4CFFD7FB16F91EF10EF93D" box="[1212,1380,1757,1781]" italics="true" pageId="1" pageNumber="478">Smetanabatrus</emphasis>
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</taxonomicName>
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, and the base of tergite IV with mediobasal foveae at the lateral margins of a short basal sulcus readily separates the new genus from
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<taxonomicName id="89DBE0AFEC4CFFD7FE8AF8E5F63AF8F6" authority="Jeannel, 1957" authorityName="Jeannel" authorityYear="1957" box="[288,592,1829,1854]" class="Insecta" family="Staphylinidae" genus="Ancistromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="478" phylum="Arthropoda" rank="genus">
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<emphasis id="7CAF473EEC4CFFD7FE8AF8E5F5C1F8F5" box="[288,427,1830,1853]" italics="true" pageId="1" pageNumber="478">Ancistromus</emphasis>
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Jeannel, 1957
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</taxonomicName>
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(pronotal marginal spines present) and
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<taxonomicName id="89DBE0AFEC4CFFD7FBBCF8E5F15DF8F6" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" box="[1046,1335,1829,1854]" class="Insecta" family="Staphylinidae" genus="Batristerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="478" phylum="Arthropoda" rank="genus">
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<emphasis id="7CAF473EEC4CFFD7FBBCF8E5F0F9F8F5" box="[1046,1171,1830,1853]" italics="true" pageId="1" pageNumber="478">Batristerus</emphasis>
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Jeannel, 1949
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</taxonomicName>
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(lacking basal sulcus and foveae on abdominal tergite IV).
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<taxonomicName id="89DBE0AFEC4CFFD7FCACF88AF7C4F8A9" box="[774,942,1865,1889]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="7CAF473EEC4CFFD7FCACF88AF7C4F8A9" box="[774,942,1865,1889]" italics="true" pageId="1" pageNumber="478">Smetanabatrus</emphasis>
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</taxonomicName>
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also shares many external features with
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<taxonomicName id="89DBE0AFEC4CFFD7FF3DF8AEF6EAF84E" authority="Lobl & Kurbatov, 2001" authorityName="Lobl & Kurbatov" authorityYear="2001" box="[151,640,1901,1926]" class="Insecta" family="Staphylinidae" genus="Coryphomobatrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="478" phylum="Arthropoda" rank="genus">
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<emphasis id="7CAF473EEC4CFFD7FF3DF8AEF509F84D" box="[151,355,1901,1925]" italics="true" pageId="1" pageNumber="478">Coryphomobatrus</emphasis>
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Löbl & Kurbatov, 2001
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</taxonomicName>
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from
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<collectingCountry id="36CCDBBCEC4CFFD7FD66F8AEF72BF84E" box="[716,833,1901,1926]" name="Sri Lanka" pageId="1" pageNumber="478">Sri Lanka</collectingCountry>
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, but can be separated by the lack of the discal and marginal carinae of abdominal tergite IV. Together with all of the above,
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<taxonomicName id="89DBE0AFEC4CFFD7FC7DF852F015F861" box="[983,1151,1937,1961]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="7CAF473EEC4CFFD7FC7DF852F015F861" box="[983,1151,1937,1961]" italics="true" pageId="1" pageNumber="478">Smetanabatrus</emphasis>
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</taxonomicName>
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has a distinctive form of the maxillary palpi, with palpomeres IV being greatly protruding at the posteromesal margins in both sexes, and has abdominal tergites IV–VI and sternite V exhibiting conspicuous modifications in the male.
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</paragraph>
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</subSubSection>
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<subSubSection id="06C1C8A7EC4FFFD4FF6DFF54F492FE40" pageId="2" pageNumber="479" type="etymology">
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<paragraph id="4E649B2CEC4FFFD4FF6DFF54F6EEFF30" blockId="2.[151,1436,151,248]" pageId="2" pageNumber="479">
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<emphasis id="7CAF473EEC4FFFD4FF6DFF54F524FF78" bold="true" box="[199,334,151,176]" pageId="2" pageNumber="479">Etymology.</emphasis>
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The genus is named after Aleš Smetana (Ottawa,
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<collectingCountry id="36CCDBBCEC4FFFD4FC0EFF54F790FF78" box="[932,1018,151,176]" name="Canada" pageId="2" pageNumber="479">Canada</collectingCountry>
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) who collected the
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<typeStatus id="9160258EEC4FFFD4FB4CFF5BF17CFF78" box="[1254,1302,152,176]" pageId="2" pageNumber="479">type</typeStatus>
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series, and combined with ‘
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<emphasis id="7CAF473EEC4FFFD4FEE7FF7FF5FFFF1C" box="[333,405,188,212]" italics="true" pageId="2" pageNumber="479">batrus</emphasis>
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’, an arbitrary abbreviation of
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<taxonomicName id="89DBE0AFEC4FFFD4FD49FF7EF789FF1D" authority="Aube. Gender" authorityName="Aube. Gender" box="[739,995,188,213]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="genus">
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<emphasis id="7CAF473EEC4FFFD4FD49FF7EF72AFF1C" box="[739,832,189,212]" italics="true" pageId="2" pageNumber="479">Batrisus</emphasis>
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Aubé. Gender
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</taxonomicName>
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is masculine. The species epithet refers to the
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<typeStatus id="9160258EEC4FFFD4FF77FF23F567FF30" box="[221,269,224,248]" pageId="2" pageNumber="479">type</typeStatus>
|
||
locality,
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FEDEFF1CF5B1FF30" box="[372,475,223,248]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="kinabalu">Kinabalu</taxonomicName>
|
||
National Park.
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DFE88F492FE40" blockId="2.[151,721,331,392]" pageId="2" pageNumber="479">
|
||
<emphasis id="7CAF473EEC4FFFD4FF3DFE88F6BBFEAD" bold="true" box="[151,721,331,357]" pageId="2" pageNumber="479">
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FF3DFE88F58EFEAD" authority="Yin" authorityName="Yin" box="[151,484,331,357]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="kinabalu">
|
||
<emphasis id="7CAF473EEC4FFFD4FF3DFE88F5D9FEAC" bold="true" box="[151,435,331,356]" italics="true" pageId="2" pageNumber="479">Smetanabatrus kinabalu</emphasis>
|
||
Yin
|
||
</taxonomicName>
|
||
and Li, new species
|
||
</emphasis>
|
||
<figureCitation id="D6E087A9EC4FFFD4FF3DFEACF492FE40" box="[151,248,367,392]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1937,1959]" captionStartId-1="4.[151,250,1896,1918]" captionTargetBox-0="[162,1429,556,1910]" captionTargetBox-1="[163,1434,201,1856]" captionTargetId-0="figure@3.[151,1436,547,1915]" captionTargetId-1="figure@4.[151,1436,193,1875]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Dorsal habitus of Smetanabatrus kinabalu. A. Male. B. Female. Scales (mm): 0.5 mm." captionText-1="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri-0="https://zenodo.org/record/216600/files/figure.png" httpUri-1="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Figs 1–2</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06C1C8A7EC4FFFD4FF3DFE74F7DFFDF4" pageId="2" pageNumber="479" type="materials_examined">
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DFE74F7DFFDF4" blockId="2.[151,1437,439,1509]" pageId="2" pageNumber="479">
|
||
<emphasis id="7CAF473EEC4FFFD4FF3DFE74F52AFE18" bold="true" box="[151,320,439,464]" pageId="2" pageNumber="479">
|
||
<typeStatus id="9160258EEC4FFFD4FF3DFE74F4B8FE18" box="[151,210,439,464]" pageId="2" pageNumber="479">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
(2 ♂♂, 1 ♀).
|
||
<emphasis id="7CAF473EEC4FFFD4FE40FE74F68AFE18" bold="true" box="[490,736,439,464]" pageId="2" pageNumber="479">
|
||
<typeStatus id="9160258EEC4FFFD4FE40FE74F631FE18" box="[490,603,439,464]" pageId="2" pageNumber="479" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="36CCDBBCEC4FFFD4FDC0FE74F6B1FE18" box="[618,731,439,464]" name="Malaysia" pageId="2" pageNumber="479">Malaysia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
♂, labeled ‘BORNEO, SABAH, Mt.
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FB3CFE74F097FE18" box="[1174,1277,439,464]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="kinabalu">Kinabalu</taxonomicName>
|
||
Nat. Pk., HQ
|
||
<quantity id="892336C9EC4FFFD4FF3DFE1FF55DFE3C" box="[151,311,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.61" metricValueMax="1.66" metricValueMin="1.56" pageId="2" pageNumber="479" unit="m" value="1610.0" valueMax="1660.0" valueMin="1560.0">1560–1660 m</quantity>
|
||
,
|
||
<date id="3A65BDECEC4FFFD4FEEDFE1FF5C9FE3D" box="[327,419,476,501]" pageId="2" pageNumber="479" value="1987-04-24">24.iv.87</date>
|
||
, A. Smetana /
|
||
<typeStatus id="9160258EEC4FFFD4FDFFFE1FF683FE3C" box="[597,745,476,500]" pageId="2" pageNumber="479" type="holotype">HOLOTYPE</typeStatus>
|
||
[red],
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FC91FE1FF025FE3C" box="[827,1103,476,500]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="kinabalu" status="sp. nov.">
|
||
<emphasis id="7CAF473EEC4FFFD4FC91FE1FF025FE3C" box="[827,1103,476,500]" italics="true" pageId="2" pageNumber="479">Smetanabatrus kinabalu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="679CFA45EC4FFFD4FBF0FE1EF0F1FE3C" box="[1114,1179,477,500]" pageId="2" pageNumber="479" rank="species">sp. n.</taxonomicNameLabel>
|
||
, Yin & Li det., 2013, MHNG’.
|
||
<emphasis id="7CAF473EEC4FFFD4FEACFE3CF66CFDD0" bold="true" box="[262,518,511,536]" pageId="2" pageNumber="479">
|
||
<typeStatus id="9160258EEC4FFFD4FEACFE3CF5E8FDD0" box="[262,386,511,536]" pageId="2" pageNumber="479" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectingCountry id="36CCDBBCEC4FFFD4FE3AFE3CF66BFDD0" box="[400,513,511,536]" name="Malaysia" pageId="2" pageNumber="479">Malaysia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="58DD50A5EC4FFFD4FDBAFDC3F654FDDF" box="[528,574,512,536]" pageId="2" pageNumber="479" type="generic">1 ♂</specimenCount>
|
||
, 1 ♀, same label data as
|
||
<typeStatus id="9160258EEC4FFFD4FCCEFE3CF7A3FDD0" box="[868,969,511,536]" pageId="2" pageNumber="479" type="holotype">holotype</typeStatus>
|
||
, each bears a
|
||
<typeStatus id="9160258EEC4FFFD4FBC7FDC3F0F7FDD0" box="[1133,1181,512,536]" pageId="2" pageNumber="479">type</typeStatus>
|
||
label as ‘
|
||
<typeStatus id="9160258EEC4FFFD4FAA1FDC3F1F6FDD0" box="[1291,1436,512,536]" pageId="2" pageNumber="479" type="paratype">PARATYPE</typeStatus>
|
||
[yellow],
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FEAEFDE7F67FFDF4" box="[260,533,548,572]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="kinabalu" status="sp. nov.">
|
||
<emphasis id="7CAF473EEC4FFFD4FEAEFDE7F67FFDF4" box="[260,533,548,572]" italics="true" pageId="2" pageNumber="479">Smetanabatrus kinabalu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="679CFA45EC4FFFD4FDB6FDE6F633FDF4" box="[540,601,549,572]" pageId="2" pageNumber="479" rank="species">sp. n.</taxonomicNameLabel>
|
||
, Yin & Li det., 2013, MHNG’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06C1C8A7EC4FFFD4FF6DFD84F53DFA08" pageId="2" pageNumber="479" type="description">
|
||
<paragraph id="4E649B2CEC4FFFD4FF6DFD84F5BCFD4D" blockId="2.[151,1437,439,1509]" pageId="2" pageNumber="479">
|
||
<emphasis id="7CAF473EEC4FFFD4FF6DFD84F533FDA8" bold="true" box="[199,345,583,608]" pageId="2" pageNumber="479">Description.</emphasis>
|
||
Male (
|
||
<figureCitation id="D6E087A9EC4FFFD4FE01FD84F586FDA8" box="[427,492,583,608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1937,1959]" captionTargetBox="[162,1429,556,1910]" captionTargetId="figure@3.[151,1436,547,1915]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of Smetanabatrus kinabalu. A. Male. B. Female. Scales (mm): 0.5 mm." httpUri="https://zenodo.org/record/216600/files/figure.png" pageId="2" pageNumber="479">Fig. 1</figureCitation>
|
||
A). BL
|
||
<quantity id="892336C9EC4FFFD4FDE2FD84F69AFD97" box="[584,752,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="2.925" metricValueMax="2.95" metricValueMin="2.9" pageId="2" pageNumber="479" unit="mm" value="2.925" valueMax="2.95" valueMin="2.9">2.90–2.95 mm</quantity>
|
||
. Body reddish brown, tarsi lighter in color; most part of the dorsal surface densely hairy.
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4FFFD4FF6DFD4CF706FB68" blockId="2.[151,1437,439,1509]" pageId="2" pageNumber="479">
|
||
Head distinctly transverse, HL
|
||
<quantity id="892336C9EC4FFFD4FD86FD4CF6BEFD6F" box="[556,724,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="5.8" metricValueMin="5.699999999999999" pageId="2" pageNumber="479" unit="mm" value="0.575" valueMax="0.58" valueMin="0.57">0.57–0.58 mm</quantity>
|
||
, HW
|
||
<quantity id="892336C9EC4FFFD4FCB3FD53F7ABFD6F" box="[793,961,656,680]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.3" metricValueMin="7.199999999999999" pageId="2" pageNumber="479" unit="mm" value="0.725" valueMax="0.73" valueMin="0.72">0.72–0.73 mm</quantity>
|
||
; surface roughly punctate; vertex slightly convex, foveae below level of posterior margins of eyes, lacking sulcus connecting foveae, median sulcus present from base toward level of antennal tubercles; eyes relatively small, each composed of about 35 facets; maxillary palpi (
|
||
<figureCitation id="D6E087A9EC4FFFD4FF74FD3FF54BFCDC" box="[222,289,764,789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
A) nearly securiform; antennomeres each elongate, clubs indistinct. Pronotum nearly cordiform, wider than long, PL
|
||
<quantity id="892336C9EC4FFFD4FEECFCDCF59AFCFF" box="[326,496,799,824]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" metricValueMax="6.2" metricValueMin="6.0" pageId="2" pageNumber="479" unit="mm" value="0.61" valueMax="0.62" valueMin="0.6">0.60–0.62 mm</quantity>
|
||
, PW
|
||
<quantity id="892336C9EC4FFFD4FD92FCE3F689FCFF" box="[568,739,800,824]" metricMagnitude="-4" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="7.199999999999999" metricValueMin="7.1" pageId="2" pageNumber="479" unit="mm" value="0.715" valueMax="0.72" valueMin="0.71">0.71–0.72 mm</quantity>
|
||
; with rough punctation; lateral margins rounded, gradually narrowed from middle toward base. Elytra slightly wider than long, EL
|
||
<quantity id="892336C9EC4FFFD4FC13FC87F034FC94" box="[953,1118,836,861]" metricMagnitude="-4" metricUnit="m" metricValue="9.549999999999999" metricValueMax="9.6" metricValueMin="9.5" pageId="2" pageNumber="479" unit="mm" value="0.955" valueMax="0.96" valueMin="0.95">0.95–0.96 mm</quantity>
|
||
, EW
|
||
<quantity id="892336C9EC4FFFD4FB31FC87F12AFC94" box="[1179,1344,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="1.095" metricValueMax="1.11" metricValueMin="1.08" pageId="2" pageNumber="479" unit="mm" value="1.095" valueMax="1.11" valueMin="1.08">1.08–1.11 mm</quantity>
|
||
; surface finely punctate; with three large, deep basal foveae. Metathoracic wings fully developed. Profemora (
|
||
<figureCitation id="D6E087A9EC4FFFD4FAAFFCA4F122FC48" box="[1285,1352,871,896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
B) with large ventral spine at middle; metatibiae (
|
||
<figureCitation id="D6E087A9EC4FFFD4FDC5FC4FF6D8FC6C" box="[623,690,908,933]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
C) angularly expanded laterally near apices. Abdomen wider than long, AL
|
||
<quantity id="892336C9EC4FFFD4FEA7FC73F5DDFC0F" box="[269,439,944,968]" metricMagnitude="-4" metricUnit="m" metricValue="7.85" metricValueMax="7.9" metricValueMin="7.8" pageId="2" pageNumber="479" unit="mm" value="0.785" valueMax="0.79" valueMin="0.78">0.78–0.79 mm</quantity>
|
||
, AW
|
||
<quantity id="892336C9EC4FFFD4FE55FC6CF6C0FC0F" box="[511,682,943,968]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" metricValueMax="9.7" metricValueMin="9.5" pageId="2" pageNumber="479" unit="mm" value="0.96" valueMax="0.97" valueMin="0.95">0.95–0.97 mm</quantity>
|
||
; tergite IV (
|
||
<figureCitation id="D6E087A9EC4FFFD4FC95FC6CF7EDFC00" box="[831,903,943,968]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
D) with large median cavity, apicolateral and posteromedial margins of cavity with tufts of setae, posterolateral margins greatly projecting posteriorly, apices of projection pointed, each densely setose; tergite V conspicuously convex medially, with setose patch at middle; tergite VI with posterior margin greatly and angularly protruding posteriorly, tergite VIII (
|
||
<figureCitation id="D6E087A9EC4FFFD4FB2CFBDFF0A3FBFC" box="[1158,1225,1052,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
F) about as long as wide; sternite V (
|
||
<figureCitation id="D6E087A9EC4FFFD4FEF1FBFCF5F4FB90" box="[347,414,1087,1112]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
E) with flat erect median projection, sternite VIII (
|
||
<figureCitation id="D6E087A9EC4FFFD4FC70FBFCF077FB90" box="[986,1053,1087,1112]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
G) transverse. Aedeagus (Figs H– K) asymmetric, length
|
||
<quantity id="892336C9EC4FFFD4FE30FBA7F668FBB4" box="[410,514,1124,1148]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="2" pageNumber="479" unit="mm" value="0.47">0.47 mm</quantity>
|
||
; basal bulb with large, transverse foramen; median lobe elongate, flattened dorsoventrally, slightly curved leftwards; ventral membrane lamellate.
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4FFFD4FF6DFB6FF778FA9C" blockId="2.[151,1437,439,1509]" pageId="2" pageNumber="479">
|
||
Female (
|
||
<figureCitation id="D6E087A9EC4FFFD4FE81FB6FF504FB0C" box="[299,366,1196,1221]" captionStart="FIGURE 1" captionStartId="3.[151,250,1937,1959]" captionTargetBox="[162,1429,556,1910]" captionTargetId="figure@3.[151,1436,547,1915]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of Smetanabatrus kinabalu. A. Male. B. Female. Scales (mm): 0.5 mm." httpUri="https://zenodo.org/record/216600/files/figure.png" pageId="2" pageNumber="479">Fig. 1</figureCitation>
|
||
B). Similar to male in general. Antennae and elytra shorter; each eye composed of about 25 facets; metathoracic wings greatly reduced; profemora and metatibiae simple; abdomen lacking modifications. Tergite (
|
||
<figureCitation id="D6E087A9EC4FFFD4FF5EFB37F55DFAC4" box="[244,311,1268,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
L) and sternite (
|
||
<figureCitation id="D6E087A9EC4FFFD4FE43FB37F641FAC4" box="[489,555,1268,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
M) VIII each transverse. Genital complex (
|
||
<figureCitation id="D6E087A9EC4FFFD4FBA4FB37F03AFAC4" box="[1038,1104,1268,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1896,1918]" captionTargetBox="[163,1434,201,1856]" captionTargetId="figure@4.[151,1436,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Diagnostic features of Smetanabatrus kinabalu (A – K. Male. L – N. Female). A. Maxillary palpus. B. profemur. C. apical portion of metatibia. D. abdomen, in dorsal view. E. same, in lateral view. F., L. tergite VIII. G., M. Sternite VIII. H. Apical portion of aedeagus. I. Aedeagus, in dorsal view. J. Same, in lateral view. K. Same, in ventral view. N. Female genital complex. Scales (mm): A, H – K, N = 0.1; B – G, L, M = 0.2." httpUri="https://zenodo.org/record/216601/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
|
||
N)
|
||
<quantity id="892336C9EC4FFFD4FBDFFB37F0B2FAC4" box="[1141,1240,1268,1292]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="479" unit="mm" value="0.33">0.33 mm</quantity>
|
||
wide, transverse, with pair of lateral sclerites. Measurements: BL
|
||
<quantity id="892336C9EC4FFFD4FD19FADBF772FAE7" box="[691,792,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="2.74" pageId="2" pageNumber="479" unit="mm" value="2.74">2.74 mm</quantity>
|
||
, HL
|
||
<quantity id="892336C9EC4FFFD4FCFAFAD4F7DCFAE7" box="[848,950,1303,1328]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="2" pageNumber="479" unit="mm" value="0.59">0.59 mm</quantity>
|
||
, HW
|
||
<quantity id="892336C9EC4FFFD4FC5DFAD4F037FAE7" box="[1015,1117,1303,1328]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="479" unit="mm" value="0.76">0.76 mm</quantity>
|
||
, PL
|
||
<quantity id="892336C9EC4FFFD4FB25FAD4F09FFAE7" box="[1167,1269,1303,1328]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="479" unit="mm" value="0.63">0.63 mm</quantity>
|
||
, PW
|
||
<quantity id="892336C9EC4FFFD4FA98FADBF1FDFAE7" box="[1330,1431,1304,1328]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="479" unit="mm" value="0.74">0.74 mm</quantity>
|
||
, EL
|
||
<quantity id="892336C9EC4FFFD4FF6AFAFFF54CFA9C" box="[192,294,1340,1365]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="2" pageNumber="479" unit="mm" value="0.86">0.86 mm</quantity>
|
||
, EW
|
||
<quantity id="892336C9EC4FFFD4FECEFAFFF5A0FA9C" box="[356,458,1340,1364]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="2" pageNumber="479" unit="mm" value="1.02">1.02 mm</quantity>
|
||
, AL
|
||
<quantity id="892336C9EC4FFFD4FDA8FAFFF602FA9C" box="[514,616,1340,1365]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="479" unit="mm" value="0.66">0.66 mm</quantity>
|
||
, AW
|
||
<quantity id="892336C9EC4FFFD4FD0DFAFFF764FA9C" box="[679,782,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="2" pageNumber="479" unit="mm" value="0.97">0.97 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4FFFD4FF6DFA9CF53DFA08" blockId="2.[151,1437,439,1509]" pageId="2" pageNumber="479">
|
||
<emphasis id="7CAF473EEC4FFFD4FF6DFA9CF5D8FAB0" bold="true" box="[199,434,1375,1400]" pageId="2" pageNumber="479">Comparative notes.</emphasis>
|
||
The large body size, combined with the spinose profemora, enlarged apical portions of the metatibiae, and the heavy abdominal modifications of the male readily separates
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FBDAFA47F09BFA54" box="[1136,1265,1412,1436]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="kinabalu">
|
||
<emphasis id="7CAF473EEC4FFFD4FBDAFA47F014FA53" box="[1136,1150,1412,1435]" italics="true" pageId="2" pageNumber="479">S</emphasis>
|
||
.
|
||
<emphasis id="7CAF473EEC4FFFD4FB25FA47F09BFA54" box="[1167,1265,1412,1436]" italics="true" pageId="2" pageNumber="479">kinabalu</emphasis>
|
||
</taxonomicName>
|
||
from all other batrisine species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06C1C8A7EC4FFFD4FF6DFA0FF60AFA2D" box="[199,608,1484,1509]" pageId="2" pageNumber="479" type="distribution">
|
||
<paragraph id="4E649B2CEC4FFFD4FF6DFA0FF60AFA2D" blockId="2.[151,1437,439,1509]" box="[199,608,1484,1509]" pageId="2" pageNumber="479">
|
||
<emphasis id="7CAF473EEC4FFFD4FF6DFA0FF50AFA2D" bold="true" box="[199,352,1484,1509]" pageId="2" pageNumber="479">Distribution.</emphasis>
|
||
East
|
||
<collectingCountry id="36CCDBBCEC4FFFD4FE34FA0FF663FA2D" box="[414,521,1484,1509]" name="Malaysia" pageId="2" pageNumber="479">Malaysia</collectingCountry>
|
||
: Sabah.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06C1C8A7EC4FFFD5FF3DF9F4F1F1FE27" lastPageId="3" lastPageNumber="480" pageId="2" pageNumber="479" type="key">
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DF9F4F626F998" blockId="2.[151,588,1591,1616]" box="[151,588,1591,1616]" pageId="2" pageNumber="479">
|
||
<emphasis id="7CAF473EEC4FFFD4FF3DF9F4F626F998" bold="true" box="[151,588,1591,1616]" pageId="2" pageNumber="479">
|
||
Key to
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FF46F9F4F53EF998" ID-CoL="8VR76" box="[236,340,1591,1616]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
genera from Borneo
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DF943F69EF8E0" blockId="2.[151,1436,1663,1832]" pageId="2" pageNumber="479">
|
||
A number of Bornean species described by Raffray, Reitter and Schaufuss are listed under
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FB0AF943F101F950" ID-CoL="8VRHR" authority="Reitter" authorityName="Reitter" box="[1184,1387,1663,1688]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4FFFD4FB0AF943F172F950" box="[1184,1304,1664,1688]" italics="true" pageId="2" pageNumber="479">Batrisodes</emphasis>
|
||
Reitter
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FF3DF966F593F974" ID-CoL="8VVHR" authority="Aube (Nomura 2008)" authorityName="Aube (Nomura" authorityYear="2008" box="[151,505,1700,1725]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4FFFD4FF3DF966F49FF974" box="[151,245,1701,1724]" italics="true" pageId="2" pageNumber="479">Batrisus</emphasis>
|
||
Aubé (Nomura 2008)
|
||
</taxonomicName>
|
||
. Since both genera are largely Holarctic or Palaearctic in distribution (Besuchet 1979, 1981), and are not likely to reach the tropical region of
|
||
<collectingCountry id="36CCDBBCEC4FFFD4FCC6F90AF7B6F928" box="[876,988,1735,1760]" name="China" pageId="2" pageNumber="479">East Asia</collectingCountry>
|
||
, these species were probably based on misidentifications. Correct placements of these species require further examination of the
|
||
<typeStatus id="9160258EEC4FFFD4FB2FF92EF0AEF8CD" box="[1157,1220,1773,1797]" pageId="2" pageNumber="479">types</typeStatus>
|
||
. For this reason we exclude
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FF5FF8D3F507F8E0" ID-CoL="8VRHR" box="[245,365,1808,1832]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4FFFD4FF5FF8D3F507F8E0" box="[245,365,1808,1832]" italics="true" pageId="2" pageNumber="479">Batrisodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FE09F8D3F66AF8EF" ID-CoL="8VVHR" box="[419,512,1808,1831]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4FFFD4FE09F8D3F66AF8EF" box="[419,512,1808,1831]" italics="true" pageId="2" pageNumber="479">Batrisus</emphasis>
|
||
</taxonomicName>
|
||
from the present key.
|
||
</paragraph>
|
||
<key id="3A4ACC1DEC4FFFD5FF3DF894F1F1FE27" lastPageId="3" lastPageNumber="480" pageId="2" pageNumber="479">
|
||
<keyStep id="F52F8389EC4FFFD4FF3DF894F1F1F842" pageId="2" pageNumber="479">
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DF894F1F1F8A5" blockId="2.[151,1436,1879,2018]" box="[151,1435,1879,1901]" pageId="2" pageNumber="479">
|
||
<keyLead id="F52A3819EC4FFFD4FF3DF894F1F1F8A5" box="[151,1435,1879,1901]" pageId="2" pageNumber="479">1 Each elytron with four basal foveae, or fovea lacking........................................................ 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DF8B6F1F1F842" blockId="2.[151,1436,1879,2018]" box="[151,1435,1908,1930]" pageId="2" pageNumber="479">
|
||
<keyLead id="F52A3819EC4FFFD4FF3DF8B6F1F1F842" box="[151,1435,1908,1930]" pageId="2" pageNumber="479">- Each elytron with two to three basal foveae................................................................. 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F52F8389EC4FFFD4FF3DF851F1F6F80D" pageId="2" pageNumber="479">
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DF851F1F6F860" blockId="2.[151,1436,1879,2018]" box="[151,1436,1938,1960]" pageId="2" pageNumber="479">
|
||
<keyLead id="F52A3819EC4FFFD4FF3DF851F1F6F860" box="[151,1436,1938,1960]" pageId="2" pageNumber="479">
|
||
2 Elytra each with four basal foveae; surface of body coarsely punctate.............................
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FB4BF851F1F6F860" ID-CoL="8W3BW" authority="Westwood" authorityName="Westwood" box="[1249,1436,1938,1960]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4FFFD4FB4BF851F15BF860" box="[1249,1329,1938,1960]" italics="true" pageId="2" pageNumber="479">Sathytes</emphasis>
|
||
Westwood
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DF86CF1F6F80D" blockId="2.[151,1436,1879,2018]" box="[151,1436,1967,1989]" pageId="2" pageNumber="479">
|
||
<keyLead id="F52A3819EC4FFFD4FF3DF86CF1F6F80D" box="[151,1436,1967,1989]" pageId="2" pageNumber="479">
|
||
- Elytra lacking basal fovea; surface of body smooth or finely punctate.......................
|
||
<taxonomicName id="89DBE0AFEC4FFFD4FB0BF873F1F6F80D" ID-CoL="8XMWB" authority="Newton & Chandler" authorityName="Newton & Chandler" box="[1185,1436,1967,1989]" class="Insecta" family="Staphylinidae" genus="Mnia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="479" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4FFFD4FB0BF873F0BEF80D" box="[1185,1236,1968,1989]" italics="true" pageId="2" pageNumber="479">Mnia</emphasis>
|
||
Newton & Chandler
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F52F8389EC4FFFD5FF3DF80FF1F1FF65" lastPageId="3" lastPageNumber="480" pageId="2" pageNumber="479">
|
||
<paragraph id="4E649B2CEC4FFFD4FF3DF80FF1F1F82A" blockId="2.[151,1436,1879,2018]" box="[151,1435,1996,2018]" pageId="2" pageNumber="479">
|
||
<keyLead id="F52A3819EC4FFFD4FF3DF80FF1F1F82A" box="[151,1435,1996,2018]" pageId="2" pageNumber="479">3 Elytra each with two basal foveae........................................................................ 4</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFF54F1F1FF65" blockId="3.[151,1436,151,495]" box="[151,1435,151,173]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFF54F1F1FF65" box="[151,1435,151,173]" pageId="3" pageNumber="480">- Elytra each with three basal foveae........................................................................ 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F52F8389EC4EFFD5FF3DFF77F1F1FF2F" pageId="3" pageNumber="480">
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFF77F1F6FF02" blockId="3.[151,1436,151,495]" box="[151,1436,180,202]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFF77F1F6FF02" box="[151,1436,180,202]" pageId="3" pageNumber="480">
|
||
4 Postgenae at least three times as long as diameter of eyes..........................................
|
||
<taxonomicName id="89DBE0AFEC4EFFD5FB55FF77F1F6FF02" ID-CoL="8XPWQ" authority="Lobl" authorityName="Lobl" box="[1279,1436,180,202]" class="Insecta" family="Staphylinidae" genus="Siteromina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4EFFD5FB55FF77F102FF02" box="[1279,1384,180,202]" italics="true" pageId="3" pageNumber="480">Siteromina</emphasis>
|
||
Löbl
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFF11F1F1FF2F" blockId="3.[151,1436,151,495]" box="[151,1435,209,231]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFF11F1F1FF2F" box="[151,1435,209,231]" pageId="3" pageNumber="480">- Postgenae short, less than twice length of diameter of eyes..................................................... 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F52F8389EC4EFFD5FF3DFF2CF1F1FEF7" pageId="3" pageNumber="480">
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFF2CF1F1FEEA" blockId="3.[151,1436,151,495]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFF2CF1F1FEEA" pageId="3" pageNumber="480">
|
||
5 Anterolateral margins of scapes more or less protruding laterally; maxillary palpi conspicuously elongate, geniculate................................................................................................
|
||
<taxonomicName id="89DBE0AFEC4EFFD5FAAEFECFF1F1FEEA" ID-CoL="8VWJG" authority="Raffray" authorityName="Raffray" box="[1284,1435,268,290]" class="Insecta" family="Staphylinidae" genus="Cratna" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4EFFD5FAAEFECFF126FEEA" box="[1284,1356,268,290]" italics="true" pageId="3" pageNumber="480">Cratna</emphasis>
|
||
Raffray
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFEE9F1F1FEF7" blockId="3.[151,1436,151,495]" box="[151,1435,297,319]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFEE9F1F1FEF7" box="[151,1435,297,319]" pageId="3" pageNumber="480">- Anterolateral margins of scapes flat; maxillary palpi short, never geniculate....................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F52F8389EC4EFFD5FF3DFE84F1F1FEB2" pageId="3" pageNumber="480">
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFE84F1F6FE95" blockId="3.[151,1436,151,495]" box="[151,1436,327,349]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFE84F1F6FE95" box="[151,1436,327,349]" pageId="3" pageNumber="480">
|
||
6 Head and pronotum roughly punctate; lacking median longitudinal sulcus on pronotum..............
|
||
<taxonomicName id="89DBE0AFEC4EFFD5FB7FFE8BF1F6FE95" ID-CoL="8XK3Q" authority="Schaufuss" authorityName="Schaufuss" box="[1237,1436,327,349]" class="Insecta" family="Staphylinidae" genus="Borneana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4EFFD5FB7FFE8BF15EFE95" box="[1237,1332,328,349]" italics="true" pageId="3" pageNumber="480">Borneana</emphasis>
|
||
Schaufuss
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFEA6F1F1FEB2" blockId="3.[151,1436,151,495]" box="[151,1435,356,378]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFEA6F1F1FEB2" box="[151,1435,356,378]" pageId="3" pageNumber="480">- Head and pronotum finely punctate; pronotum with distinct median longitudinal sulcus.............................. 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F52F8389EC4EFFD5FF3DFE41F1F1FE7D" pageId="3" pageNumber="480">
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFE41F1F1FE5F" blockId="3.[151,1436,151,495]" box="[151,1435,385,407]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFE41F1F1FE5F" box="[151,1435,385,407]" pageId="3" pageNumber="480">
|
||
7 Posterior border of pronotal transverse antebasal sulcus forming evenly curved ridge................
|
||
<taxonomicName id="89DBE0AFEC4EFFD5FB65FE41F1F1FE5F" ID-CoL="8VVHK" authority="Raffray" authorityName="Raffray" box="[1231,1435,385,407]" class="Insecta" family="Staphylinidae" genus="Batrisocenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4EFFD5FB65FE41F126FE5F" box="[1231,1356,386,407]" italics="true" pageId="3" pageNumber="480">Batrisocenus</emphasis>
|
||
Raffray
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFE5CF1F1FE7D" blockId="3.[151,1436,151,495]" box="[151,1435,415,437]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFE5CF1F1FE7D" box="[151,1435,415,437]" pageId="3" pageNumber="480">
|
||
- Posterior border of pronotal transverse antebasal sulcus not raised................................
|
||
<taxonomicName id="89DBE0AFEC4EFFD5FB4FFE5CF1F1FE7D" ID-CoL="8XL9Q" authority="Raffray" authorityName="Raffray" box="[1253,1435,415,437]" class="Insecta" family="Staphylinidae" genus="Eubatrisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4EFFD5FB4FFE5CF127FE7D" box="[1253,1357,415,437]" italics="true" pageId="3" pageNumber="480">Eubatrisus</emphasis>
|
||
Raffray
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F52F8389EC4EFFD5FF3DFE7FF1F1FE27" pageId="3" pageNumber="480">
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFE7FF1F6FE1A" blockId="3.[151,1436,151,495]" box="[151,1436,444,466]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFE7FF1F6FE1A" box="[151,1436,444,466]" pageId="3" pageNumber="480">
|
||
8 Pronotal lateral margins lacking spines; abdominal tergite IV lacking marginal carinae..............
|
||
<taxonomicName id="89DBE0AFEC4EFFD5FB68FE7FF138FE1A" ID-CoL="8XPWV" box="[1218,1362,444,466]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="7CAF473EEC4EFFD5FB68FE7FF138FE1A" box="[1218,1362,444,466]" italics="true" pageId="3" pageNumber="480">Smetanabatrus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="679CFA45EC4EFFD5FAF2FE7FF1F6FE1A" box="[1368,1436,444,466]" pageId="3" pageNumber="480" rank="genus">
|
||
<emphasis id="7CAF473EEC4EFFD5FAF2FE7FF1F6FE1A" bold="true" box="[1368,1436,444,466]" pageId="3" pageNumber="480">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4E649B2CEC4EFFD5FF3DFE19F1F1FE27" blockId="3.[151,1436,151,495]" box="[151,1435,473,495]" pageId="3" pageNumber="480">
|
||
<keyLead id="F52A3819EC4EFFD5FF3DFE19F1F1FE27" box="[151,1435,473,495]" pageId="3" pageNumber="480">
|
||
- Pronotal marginal spines present; inner and outer pairs of marginal carinae on tergite IV complete....
|
||
<taxonomicName id="89DBE0AFEC4EFFD5FB6CFE1AF1F1FE27" ID-CoL="8VVHN" authority="Reitter" authorityName="Reitter" box="[1222,1435,473,495]" class="Insecta" family="Staphylinidae" genus="Batrisoschema" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7CAF473EEC4EFFD5FB6CFE1AF13EFE27" box="[1222,1364,473,495]" italics="true" pageId="3" pageNumber="480">Batrisoschema</emphasis>
|
||
Reitter
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |