treatments-xml/data/C4/FC/A2/C4FCA297B86056A5A534B1F02D40B928.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="C4FCA297B86056A5A534B1F02D40B928" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila elevata" docType="treatment" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1147</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>177</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928" httpUri="http://treatment.plazi.org/id/C4FCA297B86056A5A534B1F02D40B928" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="66QN6" LSID="C4FCA297-B860-56A5-A534-B1F02D40B928" authority="(Thompson &amp; Upatham, 1997)" baseAuthorityName="Thompson &amp; Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevata">Angustopila elevata (Thompson &amp; Upatham, 1997)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Angustopila elevata (Thompson &amp; Upatham, 1997) (specimen from the type locality, HNHM 100184). Apertural (A), ventral (B), lateral (C) and apical (D) sides of the shell; granular sculpture on the protoconch (E), ventral (F) and frontal (G) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure12" httpUri="https://binary.pensoft.net/fig/808821" pageId="0" pageNumber="1">Figs 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Angustopila elevata (Thompson &amp; Upatham, 1997) A Perpant Cave (Myanmar) B Chiang Dao C Tha Song Yang D Panara E 2019 / 123 F 2019 / 119 G 2020 / 18 H 2020 / 26 I Cong Troi Cave J 2020 / 52 K 2020 / 48 L A. subelevata holotype M La. 2." figureDoi="10.3897/zookeys.1147.93824.figure13" httpUri="https://binary.pensoft.net/fig/808822" pageId="0" pageNumber="1">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="F. G. Thompson &amp; Upatham" authorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Systenostoma elevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Bryozoa" rank="species" species="elevata">Systenostoma elevata</taxonomicName>
Thompson &amp; Upatham, 1997: 232-233, figs 39-43.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Thompson &amp; Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevata">Angustopila elevata</taxonomicName>
- 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.410.7488" author="Jochum, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="23 - 40" refId="B30" refString="Jochum, A, Slapnik, R, Kampschulte, M, Martels, G, Heneka, M, Pall-Gergely, B, 2014. A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae). ZooKeys 410: 23 - 40, DOI: https://doi.org/10.3897/zookeys.410.7488" title="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)." url="https://doi.org/10.3897/zookeys.410.7488" volume="410" year="2014">Jochum et al., 2014</bibRefCitation>
: 27.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Thompson &amp; Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevata">Angustopila elevata</taxonomicName>
- 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.523.6114" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="31 - 64" refId="B39" refString="Pall-Gergely, B, Hunyadi, A, Jochum, A, Asami, T, 2015a. Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra). ZooKeys 523: 31 - 64, DOI: https://doi.org/10.3897/zookeys.523.6114" title="Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra)." url="https://doi.org/10.3897/zookeys.523.6114" volume="523" year="2015 a">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al., 2015a
</bibRefCitation>
: 33, fig. 11 (silhouette for size comparison with other species).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila subelevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subelevata">Angustopila subelevata</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.523.6114" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="31 - 64" refId="B39" refString="Pall-Gergely, B, Hunyadi, A, Jochum, A, Asami, T, 2015a. Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra). ZooKeys 523: 31 - 64, DOI: https://doi.org/10.3897/zookeys.523.6114" title="Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra)." url="https://doi.org/10.3897/zookeys.523.6114" volume="523" year="2015 a">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al., 2015a
</bibRefCitation>
: 39, fig. 4. new synonym.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila subelevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subelevata">Angustopila subelevata</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.17109/AZH.63.3.327.2017" author="Pall-Gergely, B" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="0" pageNumber="1" pagination="143 - 157" refId="B41" refString="Pall-Gergely, B, Jochum, A, Asami, T, 2017. Three new species and a new genus of Hypselostomatidae from Cong Troi Cave, Northern Vietnam (Gastropoda: Pulmonata). Acta Zoologica Academiae Scientiarum Hungaricae 63 (2): 143 - 157, DOI: https://doi.org/10.17109/AZH.63.3.327.2017" title="Three new species and a new genus of Hypselostomatidae from Cong Troi Cave, Northern Vietnam (Gastropoda: Pulmonata)." url="https://doi.org/10.17109/AZH.63.3.327.2017" volume="63" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al., 2017
</bibRefCitation>
: 332, figs 1B, 2A-G, 7E, F.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;Thailand, Chiang Mae Province, Doi Chiang Dao (Mountain), 7 km west of Chiang Dao; 600 m altitude (
<geoCoordinate degrees="19" direction="north" minutes="24.3" orientation="latitude" precision="92" value="19.405">19°24.3'N</geoCoordinate>
, 
<geoCoordinate degrees="98" direction="east" minutes="54.2" orientation="longitude" precision="92" value="98.903336">98°54.2'E</geoCoordinate>
)&quot; (
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
); China, Guangxi (广西), Hechi Shi (河池市), Bama Xian (巴马县), cliffs at the southern edge of Jiaole Cun (交乐村), 590 m, 
<geoCoordinate degrees="24" direction="north" minutes="7.045" orientation="latitude" precision="1" value="24.117416">24°7.045'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="7.847" orientation="longitude" precision="1" value="107.13078">107°7.847'E</geoCoordinate>
&quot; (
<taxonomicName lsidName="A. subelevata" pageId="0" pageNumber="1" rank="species" species="subelevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. subelevata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2013-09-08" collectingDateMax="2013-09-17" collectingDateMin="2013-09-08" collectorName="A. Hunyadi, M. Szekeres" country="China" county="Bose Shi" elevation="590" latitude="25.0995" location="Leye" longLatPrecision="12" longitude="107.96066" municipality="Leye Xian" specimenCount="1" stateProvince="Guangxi">
<collectingCountry name="China">China</collectingCountry>
• 23 shells; 
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
, 
<collectingCounty>Bose Shi</collectingCounty>
, 
<collectingMunicipality>Leye Xian</collectingMunicipality>
, ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
from 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:A317CD5B1B00102D7F4C6A199617A922" country="China" county="Bose Shi" latitude="25.0995" longLatPrecision="12" longitude="107.96066" municipality="Leye Xian" name="Leye" stateProvince="Guangxi">Leye</location>
towards 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:346D3FAFFAE554EA4C4D6ED55C1396E2" country="China" county="Bose Shi" latitude="25.0995" longLatPrecision="12" longitude="107.96066" municipality="Leye Xian" name="Huaping Zhen" stateProvince="Guangxi">Huaping Zhen</location>
, cave above 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:2B3A79F063F689AC6858A9CF9B4AF45E" country="China" county="Bose Shi" latitude="25.0995" longLatPrecision="12" longitude="107.96066" municipality="Leye Xian" name="Niuping" stateProvince="Guangxi">Niuping</location>
junction (locality code: 2013/85); 
<geoCoordinate degrees="24" direction="north" minutes="47.61" orientation="latitude" precision="9" value="24.7935">24°47.61'N</geoCoordinate>
<geoCoordinate degrees="106" direction="east" minutes="31.89" orientation="longitude" precision="9" value="106.5315">106°31.89'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.9" unit="m" value="590.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.9" unit="m" value="590.0">590 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2013-09-08">8 Sep. 2013</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
&amp; 
<collectorName>M. Szekeres</collectorName>
leg.; coll. HA • 3 shells; 
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
, Hechi Shi, Huanjiang Maonanzu Zizhixian, south of 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:D200E219B8E06D2FF181AE56A0FDBDC7" country="China" county="Bose Shi" latitude="25.0995" longLatPrecision="12" longitude="107.96066" municipality="Leye Xian" name="Mulun Guojiaji Ziran Baohuqu" stateProvince="Guangxi">Mulun Guojiaji Ziran Baohuqu</location>
, rocks above Dongning (locality code: 2013/99); 
<geoCoordinate degrees="25" direction="north" minutes="05.97" orientation="latitude" precision="9" value="25.0995">25°05.97'N</geoCoordinate>
<geoCoordinate degrees="107" direction="east" minutes="57.64" orientation="longitude" precision="9" value="107.96066">107°57.64'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.3" unit="m" value="530.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.3" unit="m" value="530.0">530 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2013-09-17">17 September 2013</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
&amp; 
<collectorName>M. Szekeres</collectorName>
leg.; coll. HA
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2018-10-06" collectingDateMax="2019-02-02" collectingDateMin="2018-10-06" collectorName="A. Hunyadi, K. Okubo, J. U. Otani, Hopong, Hopong Spring, J. Grego" country="Myanmar" county="Hoponh" elevation="1170" latitude="20.8175" location="Parpant Cave" longLatPrecision="12" longitude="97.22483" municipality="Namsang" specimenCount="1" stateProvince="Shan State">
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
• 1 figured shell; 
<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.4" unit="km" value="7.4">7.4 km</quantity>
from the centre of 
<collectingCounty>Hoponh</collectingCounty>
towards 
<collectingMunicipality>Namsang</collectingMunicipality>
along road no. 4, ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
north, 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:273FD0CD48A193A391AE383312DDF46B" country="Myanmar" county="Hoponh" latitude="20.8175" longLatPrecision="12" longitude="97.22483" municipality="Namsang" name="Parpant Cave" stateProvince="Shan State">Parpant Cave</location>
(locality code: 2018/36); 
<geoCoordinate degrees="20" direction="north" minutes="50.96" orientation="latitude" precision="9" value="20.849333">20°50.96'N</geoCoordinate>
, 
<geoCoordinate degrees="97" direction="east" minutes="14.26" orientation="longitude" precision="9" value="97.23766">97°14.26'E</geoCoordinate>
; 
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.17" unit="m" value="1170.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.17" unit="m" value="1170.0">1170 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2018-10-06">6 Oct. 2018</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
, 
<collectorName>K. Okubo</collectorName>
&amp; 
<collectorName>J.U. Otani</collectorName>
leg.; HNHM 103484 • 6 shells + 2 j/b shells; same data as for preceding shell; coll. HA • 1 shell; 
<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
, 
<collectorName>Hopong</collectorName>
, 
<collectorName>Hopong Spring</collectorName>
and cave (locality code: JG/2019/2); 
<geoCoordinate degrees="20" direction="north" minutes="49.05" orientation="latitude" precision="9" value="20.8175">20°49.05'N</geoCoordinate>
, 
<geoCoordinate degrees="97" direction="east" minutes="13.49" orientation="longitude" precision="9" value="97.22483">97°13.49'E</geoCoordinate>
; 
<collectingDate value="2019-02-02">2 Feb. 2019</collectingDate>
; 
<collectorName>J. Grego</collectorName>
leg.; coll. JG
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2015-02-07" collectingDateMax="2015-02-19" collectingDateMin="2015-02-07" collectorName="A. Hunyadi" country="Thailand" elevation="140" latitude="8.421333" location="Tha Song Yang" longLatPrecision="13" longitude="99.37933" specimenCount="7" specimenCount-adult="7" stateProvince="Tak Province">
<collectingCountry name="Thailand">Thailand</collectingCountry>
• 1 figured shell; 
<collectingRegion country="Thailand" name="Tak">Tak Province</collectingRegion>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
NNW 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:9668CF91CE86B628257390EE8202C61C" country="Thailand" latitude="8.421333" longLatPrecision="13" longitude="99.37933" name="Tha Song Yang" stateProvince="Tak Province">Tha Song Yang</location>
, left side of road no. 105 (locality code: 2015/23); 
<geoCoordinate degrees="17" direction="north" minutes="15.36" orientation="latitude" precision="9" value="17.256">17°15.36'N</geoCoordinate>
, 
<geoCoordinate degrees="98" direction="east" minutes="12.65" orientation="longitude" precision="9" value="98.21083">98°12.65'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">140 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2015-02-15">15 Feb. 2015</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HNHM 100183 • 7 shells; same data as for preceding shell; coll. HA • 1 figured shell; 
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
, north-eastern part of Doi Chiang Dao, Wat Tham Chiang Dao northwest 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
; 
<geoCoordinate degrees="19" direction="north" minutes="24.02" orientation="latitude" precision="9" value="19.400333">19°24.02'N</geoCoordinate>
, 
<geoCoordinate degrees="98" direction="east" minutes="54.68" orientation="longitude" precision="9" value="98.91133">98°54.68'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.35" unit="m" value="835.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.35" unit="m" value="835.0">835 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2015-02-07">7 Feb. 2015</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HNHM 100184; • 11 shells; same data as for preceding shell; coll. HA • 18 shells + 4 j/b shells; 
<collectingRegion country="Thailand" name="Chiang Rai">Chiang Rai Province</collectingRegion>
, 
<locationDeviation location="Mae Sai">
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" unit="km" value="8.0" valueMax="9.0" valueMin="7.0">7 to 9 km</quantity>
SSW of Mae Sai
</locationDeviation>
, around Wat Tham Pla (locality code: 2015/18); 
<geoCoordinate degrees="20" direction="north" minutes="19.72" orientation="latitude" precision="9" value="20.328667">20°19.72'N</geoCoordinate>
, 
<geoCoordinate degrees="99" direction="east" minutes="51.82" orientation="longitude" precision="9" value="99.86367">99°51.82'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2015-02-12">12 Feb. 2015</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 1 figured shell; 
<collectingRegion country="Thailand" name="Nakhon Si Thammarat">Nakhon Si Thammarat Province</collectingRegion>
, Wat Tham Thong Panara, vicinity of the temple (locality code: 2015/31); 
<geoCoordinate degrees="08" direction="north" minutes="25.28" orientation="latitude" precision="9" value="8.421333">08°25.28'N</geoCoordinate>
, 
<geoCoordinate degrees="99" direction="east" minutes="22.76" orientation="longitude" precision="9" value="99.37933">99°22.76'E</geoCoordinate>
; 
<elevation metricMagnitude="1" metricUnit="m" metricValue="4.0" unit="m" value="40.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="4.0" unit="m" value="40.0">40 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2015-02-19">19 Feb. 2015</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HNHM 100180; • 
<specimenCount type="adult">7 adult</specimenCount>
+ 2 j/b shells; same data as for preceding shell; coll. HA
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2019-10-08" collectorName="A. Hunyadi" country="Laos" elevation="785" latitude="20.536833" location="Ban Phou Lek" longLatPrecision="12" longitude="101.135" municipality="Vieng Phou Kha" specimenCount="56" specimenCount-adult="56" stateProvince="Luang Namtha Province">
<collectingCountry name="Laos">Laos</collectingCountry>
• 
<specimenCount type="adult">56 adult</specimenCount>
+ 6 j/b shells; 
<collectingRegion country="Laos" name="Louangnamtha">Luang Namtha Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.06" unit="km" value="20.6">20.6 km</quantity>
southeast from centre of 
<collectingMunicipality>Vieng Phou Kha</collectingMunicipality>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="km" value="1.5">1.5 km</quantity>
south of 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:71919A857C4218FFFB96B23D7AEDAEA4" country="Laos" latitude="20.536833" longLatPrecision="12" longitude="101.135" municipality="Vieng Phou Kha" name="Ban Phou Lek" stateProvince="Luang Namtha Province">Ban Phou Lek</location>
, right side of road (locality code: 2019/121); 
<geoCoordinate degrees="20" direction="north" minutes="32.21" orientation="latitude" precision="9" value="20.536833">20°32.21'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="08.10" orientation="longitude" precision="9" value="101.135">101°08.10'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.85" unit="m" value="785.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.85" unit="m" value="785.0">785 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-08">8 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
• 
<materialsCitation collectingDate="2019-10-07" collectingDateMax="2019-10-08" collectingDateMin="2019-10-07" collectorName="A. Hunyadi" country="Laos" county="Vieng Phou Kha" elevation="780" latitude="20.8895" location="Na Mor" longLatPrecision="12" longitude="101.816" municipality="Ban Phou Lek" specimenCount="70" specimenCount-adult="70" stateProvince="Luang Namtha Province">
<specimenCount type="adult">70 adult</specimenCount>
+ 22 j/b shells; 
<collectingRegion country="Laos" name="Louangnamtha">Luang Namtha Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.98" unit="km" value="19.8">19.8 km</quantity>
southeast from centre of 
<collectingCounty>Vieng Phou Kha</collectingCounty>
, south from 
<collectingMunicipality>Ban Phou Lek</collectingMunicipality>
, southern edge of quarry (locality code: 2019/122); 
<geoCoordinate degrees="20" direction="north" minutes="32.54" orientation="latitude" precision="9" value="20.542334">20°32.54'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="07.96" orientation="longitude" precision="9" value="101.13267">101°07.96'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">780 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-08">8 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 21 shells; 
<collectingRegion country="Laos" name="Oudomxai">Udomxai Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
south of centre of 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:97C8445887E0512E5FEB415B466BDCBA" country="Laos" county="Vieng Phou Kha" latitude="20.8895" longLatPrecision="12" longitude="101.816" municipality="Ban Phou Lek" name="Na Mor" stateProvince="Luang Namtha Province">Na Mor</location>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.8" unit="km" value="3.8">3.8 km</quantity>
east-southeast of 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:F969481E9A6EA2BBB807432C2CAEB622" country="Laos" county="Vieng Phou Kha" latitude="20.8895" longLatPrecision="12" longitude="101.816" municipality="Ban Phou Lek" name="Na Xay" stateProvince="Luang Namtha Province">Na Xay</location>
, rock wall facing north (locality code: 2019/119); 
<geoCoordinate degrees="20" direction="north" minutes="53.37" orientation="latitude" precision="9" value="20.8895">20°53.37'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="48.96" orientation="longitude" precision="9" value="101.816">101°48.96'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.6" unit="m" value="660.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.6" unit="m" value="660.0">660 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-07">7 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
• 
<materialsCitation collectingDate="2019-10-08" collectorName="A. Hunyadi" country="Laos" elevation="100" latitude="20.742167" location="Phou Lan" longLatPrecision="12" longitude="101.185" municipality="Vieng Phou Kha" specimenCount="141" specimenCount-adult="141" stateProvince="Luang Namtha Province">
<specimenCount type="adult">141 adult</specimenCount>
shells + 21 j/b shells; 
<collectingRegion country="Laos" name="Louangnamtha">Luang Namtha Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="4.38" unit="km" value="43.8">43.8 km</quantity>
from centre of 
<collectingRegion country="Laos" name="Louangnamtha">Luang Namtha</collectingRegion>
towards 
<collectingMunicipality>Vieng Phou Kha</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:1AA04318AED690B27C0CEB29993900BB" country="Laos" latitude="20.742167" longLatPrecision="12" longitude="101.185" municipality="Vieng Phou Kha" name="Phou Lan" stateProvince="Luang Namtha Province">Phou Lan</location>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
from left side of road (locality code: 2019/123); 
<geoCoordinate degrees="20" direction="north" minutes="44.53" orientation="latitude" precision="9" value="20.742167">20°44.53'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="11.10" orientation="longitude" precision="9" value="101.185">101°11.10'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">770 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-08">8 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
• 
<materialsCitation collectingDate="2019-10-05" collectorName="A. Hunyadi" country="Laos" county="Nong Khiaw" elevation="345" latitude="20.553667" location="Tham Pha Toke" longLatPrecision="12" longitude="102.62867" municipality="Pak Xeng" specimenCount="12" specimenCount-adult="12" stateProvince="Luang Prabang Province">
<specimenCount type="adult">12 adult</specimenCount>
shells; 
<collectingRegion country="Laos" name="Louangphabang">Luang Prabang Province</collectingRegion>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.1" unit="km" value="3.1">3.1 km</quantity>
from centre of 
<collectingCounty>Nong Khiaw</collectingCounty>
towards 
<collectingMunicipality>Pak Xeng</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:148183C830390828F792188E0D1AB50A" country="Laos" county="Nong Khiaw" latitude="20.553667" longLatPrecision="12" longitude="102.62867" municipality="Pak Xeng" name="Tham Pha Toke" stateProvince="Luang Prabang Province">Tham Pha Toke</location>
, around the cave (locality code: 2019/114); 
<geoCoordinate degrees="20" direction="north" minutes="33.22" orientation="latitude" precision="9" value="20.553667">20°33.22'N</geoCoordinate>
, 
<geoCoordinate degrees="102" direction="east" minutes="37.72" orientation="longitude" precision="9" value="102.62867">102°37.72'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.45" unit="m" value="345.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.45" unit="m" value="345.0">345 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-05">5 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
• 
<materialsCitation collectingDate="2010-03" collectingDateMax="2019-10-14" collectingDateMin="2010-03" collectorName="A. Hunyadi, A. Reischuetz" country="Laos" county="Tham Kong Lor" elevation="745" latitude="18.9255" location="Vang Vieng" longLatPrecision="13" longitude="102.41583" municipality="Recreational Area" specimenCount="3" specimenCount-adult="3" stateProvince="Luang Namtha Province">
<specimenCount type="adult">3 adult</specimenCount>
shells; 
<collectingRegion country="Laos" name="Louangnamtha">Luang Namtha Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="4.8" unit="km" value="48.0">48 km</quantity>
southwest from centre of 
<collectingRegion country="Laos" name="Louangnamtha">Luang Namtha</collectingRegion>
towards Vieng Phou Kha, Nam Eng, Tham Kao Rao (locality code: 2019/120); 
<geoCoordinate degrees="20" direction="north" minutes="43.45" orientation="latitude" precision="9" value="20.724167">20°43.45'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="09.26" orientation="longitude" precision="9" value="101.154335">101°09.26'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.45" unit="m" value="745.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.45" unit="m" value="745.0">745 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-07">7 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 1 shell; 
<collectingRegion country="Laos" name="Khammouan">Khammouane Province</collectingRegion>
, 
<collectingCounty>Tham Kong Lor</collectingCounty>
(cave), 
<quantity metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
direct 
<locationDeviation location="Thakhek">N of Thakhek</locationDeviation>
, at the base of limestone rocks above 
<collectingMunicipality>Recreational Area</collectingMunicipality>
(locality code: La.2); 
<geoCoordinate degrees="17" direction="north" minutes="57.39" orientation="latitude" precision="9" value="17.9565">17°57.39'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="45.65" orientation="longitude" precision="9" value="104.76083">104°45.65'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2010-03">Mar. 2010</collectingDate>
; 
<collectorName>
A. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
</collectorName>
leg.; NHMW-MO-112005 • 2 complete + 1 broken shells; same data as for preceding; coll. RE • 1 shell; 
<collectingRegion country="Laos" name="Viangchan">Vientiane Province</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:EE99E839DBCB8387EF4BB32626F66E52" country="Laos" county="Tham Kong Lor" latitude="18.9255" longLatPrecision="13" longitude="102.41583" municipality="Recreational Area" name="Vang Vieng" stateProvince="Luang Namtha Province">Vang Vieng</location>
, rocks at 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:10B9565B9A4340D9644F5EBB863A30E9" country="Laos" county="Tham Kong Lor" latitude="18.9255" longLatPrecision="13" longitude="102.41583" municipality="Recreational Area" name="Vang Vieng Resort" stateProvince="Luang Namtha Province">Vang Vieng Resort</location>
, by 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:AB16BB1FACB23A235D51D3A1E16534D9" country="Laos" county="Tham Kong Lor" latitude="18.9255" longLatPrecision="13" longitude="102.41583" municipality="Recreational Area" name="river Nam Song" stateProvince="Luang Namtha Province">river Nam Song</location>
, limestone rocks at 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:216AE6DC6953B88E6CFD4DBFBCA13F58" country="Laos" county="Tham Kong Lor" latitude="18.9255" longLatPrecision="13" longitude="102.41583" municipality="Recreational Area" name="Tham Chang cave" stateProvince="Luang Namtha Province">Tham Chang cave</location>
(locality code: La.12); 
<geoCoordinate degrees="18" direction="north" minutes="54.66" orientation="latitude" precision="9" value="18.911">18°54.66'N</geoCoordinate>
, 
<geoCoordinate degrees="102" direction="east" minutes="26.42" orientation="longitude" precision="9" value="102.44033">102°26.42'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">280 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2010-03">Mar. 2010</collectingDate>
; 
<collectorName>
A. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
</collectorName>
leg.; coll. RE • 1 shell; 
<collectingRegion country="Laos" name="Viangchan">Vientiane Province</collectingRegion>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.5" unit="km" value="4.5">4.5 km</quantity>
west from centre of Vang Vieng, Phone Ngeun, Tham Khan Kham (locality code: 2019/133); 
<geoCoordinate degrees="18" direction="north" minutes="55.53" orientation="latitude" precision="9" value="18.9255">18°55.53'N</geoCoordinate>
, 
<geoCoordinate degrees="102" direction="east" minutes="24.95" orientation="longitude" precision="9" value="102.41583">102°24.95'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">280 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-14">14 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2020-02-08" collectingDateMax="2020-02-11" collectingDateMin="2020-02-08" collectorName="Co M, A. Hunyadi, Chau, Hang" country="Vietnam" elevation="900" latitude="20.849333" location="Chau District" longLatPrecision="12" longitude="104.639" municipality="Thu" specimenCount="18" specimenCount-adult="18" stateProvince="Sơn La Province">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
• 
<specimenCount type="adult">18 adult</specimenCount>
shells; 
<collectingRegion country="Vietnam" name="Son La">
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province
</collectingRegion>
, 
<collectingMunicipality>Thu</collectingMunicipality>
ận 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:9A6595CC5BFDE0D274B7B0C334331827" country="Vietnam" latitude="20.849333" longLatPrecision="12" longitude="104.639" municipality="Thu" name="Chau District" stateProvince="Sơn La Province">
<normalizedToken originalValue="Châu">Chau</normalizedToken>
District
</location>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
</quantity>
southeast from centre of 
<collectorName>Co M</collectorName>
ạ, rock wall above unpaved road (locality code: 2020/18); 
<geoCoordinate degrees="21" direction="north" minutes="21.19" orientation="latitude" precision="9" value="21.353167">21°21.19'N</geoCoordinate>
, 
<geoCoordinate degrees="103" direction="east" minutes="31.68" orientation="longitude" precision="9" value="103.528">103°31.68'E</geoCoordinate>
; 
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-08">8 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 1 shell; 
<collectingRegion country="Vietnam" name="Son La">
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province
</collectingRegion>
, Mộc 
<normalizedToken originalValue="Châu">
<collectorName>Chau</collectorName>
</normalizedToken>
, 
<collectorName>Hang</collectorName>
<normalizedToken originalValue="Dơi">Dơi</normalizedToken>
, around the entrance of the cave (locality code: 2020/26); 
<geoCoordinate degrees="20" direction="north" minutes="50.96" orientation="latitude" precision="9" value="20.849333">20°50.96'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="38.34" orientation="longitude" precision="9" value="104.639">104°38.34'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">865 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-11">11 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
• 
<materialsCitation collectingDate="2020-02-07" collectingDateMax="2020-02-17" collectingDateMin="2020-02-07" collectorName="Di Chi Keo Leng, A. Hunyadi, Qu, Thu, Chau, Chi, Khoang" country="Vietnam" county="Province" elevation="375" latitude="21.557333" location="Di Chi Keo Leng" longLatPrecision="12" longitude="103.68183" municipality="Binh Gia" specimenCount="4" specimenCount-adult="4" stateProvince="Sơn La Province">
<specimenCount type="adult">4 adult</specimenCount>
shells; Lạng 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
<collectingCounty>Province</collectingCounty>
, 
<collectingMunicipality>
<normalizedToken originalValue="Bình">Binh</normalizedToken>
Gia
</collectingMunicipality>
, eastern edge of the village, 
<collectorName>
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:D6E7C9B22B79734D488CCCD0BA0C4226" country="Vietnam" county="Province" latitude="21.557333" longLatPrecision="12" longitude="103.68183" municipality="Binh Gia" name="Di Chi Keo Leng" stateProvince="Sơn La Province">
Di 
<normalizedToken originalValue="Chí">Chi</normalizedToken>
<normalizedToken originalValue="Kéo">Keo</normalizedToken>
<normalizedToken originalValue="Lèng">Leng</normalizedToken>
</location>
</collectorName>
, rock wall (locality code: 2020/48); 
<geoCoordinate degrees="21" direction="north" minutes="56.30" orientation="latitude" precision="9" value="21.938334">21°56.30'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="23.82" orientation="longitude" precision="9" value="106.397">106°23.82'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">375 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-17">17 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 1 shell; 
<collectingRegion country="Vietnam" name="Son La">
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province
</collectingRegion>
, 
<collectorName>Qu</collectorName>
ỳnh 
<location LSID="urn:lsid:plazi:treatment:C4FCA297B86056A5A534B1F02D40B928:DBD1B5840E961D458FD6D017ABA73D3C" country="Vietnam" county="Province" latitude="21.557333" longLatPrecision="12" longitude="103.68183" municipality="Binh Gia" name="Nhai district" stateProvince="Sơn La Province">Nhai district</location>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
north from cross to 
<collectorName>Thu</collectorName>
ận 
<normalizedToken originalValue="Châu">
<collectorName>Chau</collectorName>
</normalizedToken>
, 
<collectorName>Chi</collectorName>
ềng 
<collectorName>Khoang</collectorName>
, cave above the village (locality code: 2020/9); 
<geoCoordinate degrees="21" direction="north" minutes="33.44" orientation="latitude" precision="9" value="21.557333">21°33.44'N</geoCoordinate>
, 
<geoCoordinate degrees="103" direction="east" minutes="40.91" orientation="longitude" precision="9" value="103.68183">103°40.91'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.15" unit="m" value="315.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.15" unit="m" value="315.0">315 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-07">7 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A small to medium-sized 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with relatively irregularly coiled, slightly concave-conical shell, and a toothless, rather subquadrate aperture. Individuals in most populations possess widely spaced radial ribs the on the ventral side and on the body whorl in apertural view.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.8-0.97, D = 0.74-0.88, H/D*100 = 95.4-123 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 58), RUD = 24.1-31.6 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 20).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
See also under 
<taxonomicName lsidName="A. apiostoma" pageId="0" pageNumber="1" rank="species" species="apiostoma">
<emphasis italics="true" pageId="0" pageNumber="1">A. apiostoma</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName lsidName="A. halongensis" pageId="0" pageNumber="1" rank="species" species="halongensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. halongensis</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName lsidName="A. oostoma" pageId="0" pageNumber="1" rank="species" species="oostoma">
<emphasis italics="true" pageId="0" pageNumber="1">A. oostoma</emphasis>
</taxonomicName>
sp. nov., and 
<taxonomicName lsidName="A. werneri" pageId="0" pageNumber="1" rank="species" species="werneri">
<emphasis italics="true" pageId="0" pageNumber="1">A. werneri</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
is the most widely distributed 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species, inhabiting 
<normalizedToken originalValue="Myanmar’s">Myanmar's</normalizedToken>
Shan State, northern and southern Thailand, northern Laos, northern Vietnam, and southern China (Guangxi) (see Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Distribution of Angustopila elevata (Thompson &amp; Upatham, 1997)." figureDoi="10.3897/zookeys.1147.93824.figure14" httpUri="https://binary.pensoft.net/fig/808823" pageId="0" pageNumber="1">14</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure12" httpUri="https://binary.pensoft.net/fig/808821" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
<taxonomicName baseAuthorityName="Thompson &amp; Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila elevata</emphasis>
</taxonomicName>
(Thompson &amp; Upatham, 1997) (specimen from the type locality, HNHM 100184). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) sides of the shell; granular sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure13" httpUri="https://binary.pensoft.net/fig/808822" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
<taxonomicName baseAuthorityName="Thompson &amp; Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila elevata</emphasis>
</taxonomicName>
(Thompson &amp; Upatham, 1997) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Perpant Cave (Myanmar) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Chiang Dao 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
Tha Song Yang 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
Panara 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
2019/123 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
2019/119 
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
2020/18 
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
2020/26 
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
Cong Troi Cave 
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
2020/52 
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
2020/48 
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
<taxonomicName lsidName="A. subelevata" pageId="0" pageNumber="1" rank="species" species="subelevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. subelevata</emphasis>
</taxonomicName>
holotype 
<emphasis bold="true" pageId="0" pageNumber="1">M</emphasis>
La.2.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure14" httpUri="https://binary.pensoft.net/fig/808823" pageId="0" pageNumber="1" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 14.</emphasis>
Distribution of 
<taxonomicName baseAuthorityName="Thompson &amp; Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila elevata</emphasis>
</taxonomicName>
(Thompson &amp; Upatham, 1997).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
When 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila subelevata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subelevata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila subelevata</emphasis>
</taxonomicName>
was described, we relied on the original description of 
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
only. Although it was evident that the two species were very similar, we considered the differences (
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
had a more slender shell, a deeper umbilicus, and was lacking spiral striae on its base) and the nearly 1000-km distance sufficient to describe 
<taxonomicName lsidName="A. subelevata" pageId="0" pageNumber="1" rank="species" species="subelevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. subelevata</emphasis>
</taxonomicName>
as new. The newly collected shells from the type locality of 
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
showed that it had weak spiral striae on the base. Furthermore, the geographical populations examined between the type localities of 
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="A. subelevata" pageId="0" pageNumber="1" rank="species" species="subelevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. subelevata</emphasis>
</taxonomicName>
indicated that only minor conchological differences exist. Although the area spans nearly 2000 kilometres in a straight line from southern Thailand to Guangxi Province in China, the populations listed here show little variation in terms of conchological characters. Thus, we now treat them as a single species.
</paragraph>
</subSubSection>
</treatment>
</document>