treatments-xml/data/C4/31/86/C431860C583CDD3AB4DAFAB1B166FF21.xml
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<document id="CB5A97AA186A514DF7B6F5C142BF5B81" ID-CLB-Dataset="46726" ID-DOI="10.11646/zootaxa.3753.4.4" ID-GBIF-Dataset="59df87e1-3da4-43ef-8c4c-b8a7cdd580ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="249874" ID-ZooBank="394ADA8F-E7A8-4751-941F-FE8E6A3921A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365347958" checkinUser="plazi" docAuthor="Ribeiro, Guilherme C. &amp; Lukashevich, Elena D." docDate="2014" docId="C431860C583CDD3AB4DAFAB1B166FF21" docLanguage="en" docName="zt03753p363.pdf" docOrigin="Zootaxa 3753 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptotarsus martinsnetoi Ribeiro &amp; Lukashevich, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="357" masterDocId="3808FE745835DD30B44DFFE1B570FFD9" masterDocTitle="New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)" masterLastPageNumber="363" masterPageNumber="347" pageNumber="356" updateTime="1698295164094" updateUser="ExternalLinkService">
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<mods:title id="60D29161B4369BC5670406419932CCC4">New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)</mods:title>
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<mods:namePart id="47AA2FC848B48716D236C3E367DFE319">Ribeiro, Guilherme C.</mods:namePart>
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<mods:namePart id="E8A3BEAB58C025E7D493FE6AA7425B06">Lukashevich, Elena D.</mods:namePart>
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<mods:title id="5F54BE5ABB8C7428A4A6409C18EEE45B">Zootaxa</mods:title>
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<mods:date id="9DA1F5FCC1B4858EEFC838E4E8A62601">2014</mods:date>
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<mods:number id="7EDD86095FE42852986C53AE7ADB8929">3753</mods:number>
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<treatment id="C431860C583CDD3AB4DAFAB1B166FF21" ID-DOI="http://doi.org/10.5281/zenodo.6143099" ID-GBIF-Taxon="119410702" ID-Zenodo-Dep="6143099" LSID="urn:lsid:plazi:treatment:C431860C583CDD3AB4DAFAB1B166FF21" httpUri="http://treatment.plazi.org/id/C431860C583CDD3AB4DAFAB1B166FF21" lastPageId="10" lastPageNumber="357" pageId="9" pageNumber="356">
<subSubSection id="04826491583CDD39B4DAFAB1B432FA54" pageId="9" pageNumber="356" type="nomenclature">
<paragraph id="4C27371A583CDD39B4DAFAB1B759FAB2" blockId="9.[151,553,1360,1421]" box="[151,553,1360,1387]" pageId="9" pageNumber="356">
<heading id="176F8076583CDD39B4DAFAB1B759FAB2" bold="true" box="[151,553,1360,1387]" fontSize="11" level="1" pageId="9" pageNumber="356" reason="1">
<emphasis id="7EECEB08583CDD39B4DAFAB1B759FAB2" bold="true" box="[151,553,1360,1387]" pageId="9" pageNumber="356">
<taxonomicName id="8B984C99583CDD39B4DAFAB1B4B4FAB3" box="[151,452,1360,1386]" class="Insecta" family="Tipulidae" genus="Leptotarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="martinsnetoi" status="sp. nov.">
<emphasis id="7EECEB08583CDD39B4DAFAB1B4B4FAB3" bold="true" box="[151,452,1360,1386]" italics="true" pageId="9" pageNumber="356">Leptotarsus martinsnetoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65DF5673583CDD39B581FAB0B759FAB2" box="[460,553,1361,1387]" pageId="9" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4C27371A583CDD39B4DAFA95B432FA54" blockId="9.[151,553,1360,1421]" box="[151,322,1396,1421]" pageId="9" pageNumber="356">
(
<figureCitation id="D4A32B9F583CDD39B4D2FA95B597FA55" box="[159,231,1396,1421]" captionStart="FIGURES 1 4" captionStartId="3.[151,264,1955,1977]" captionTargetBox="[156,1422,198,1926]" captionTargetId="figure@3.[153,1433,195,1933]" captionTargetPageId="3" captionText="FIGURES 1 4. Wing venation of Leptotarsus spp. from Santana Formation, Early Cretaceous, Brazil. 1 — L. grimaldii sp. nov., holotype AMNH SA 46358, right wing. 2 — L. grimaldii sp. nov., paratype AMNH SA 46366, left wing. 3 — L. cretaceus sp. nov., holotype AMNH SA 46354, right wing. 4 — L. martinsnetoi sp. nov., holotype MZUSP 1580, left wing." httpUri="https://zenodo.org/record/249875/files/figure.png" pageId="9" pageNumber="356">Figs 4</figureCitation>
,
<figureCitation id="D4A32B9F583CDD39B4BEFA95B44AFA54" box="[243,314,1396,1421]" captionStart-0="FIGURES 16 17" captionStart-1="FIGURE 18" captionStartId-0="8.[151,264,1959,1981]" captionStartId-1="9.[151,250,1238,1260]" captionTargetBox-0="[198,1372,193,1920]" captionTargetBox-1="[153,1433,193,1217]" captionTargetId-0="figure@8.[198,1372,193,1921]" captionTargetId-1="figure@9.[153,1433,193,1217]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 16 17. Leptotarsus martinsnetoi, sp. nov., from Santana Formation, Early Cretaceous, Brazil, holotype (part) MZUSP 1580. 16 — Total view. 17 — Detail of right wing. Abbreviations: comp eye: compound eye; scp: scape." captionText-1="FIGURE 18. Leptotarsus martinsnetoi, sp. nov., from Santana Formation, Early Cretaceous, Brazil, paratype MZUSP 1581, total view." httpUri-0="https://zenodo.org/record/249880/files/figure.png" httpUri-1="https://zenodo.org/record/249881/files/figure.png" pageId="9" pageNumber="356">1618</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="04826491583CDD39B4DAFA5DB6D3F999" pageId="9" pageNumber="356" type="materials_examined">
<paragraph id="4C27371A583CDD39B4DAFA5DB6D3F999" blockId="9.[151,1436,1468,2011]" pageId="9" pageNumber="356">
<emphasis id="7EECEB08583CDD39B4DAFA5DB435FA0C" bold="true" box="[151,325,1468,1493]" pageId="9" pageNumber="356">
<typeStatus id="932389B8583CDD39B4DAFA5DB5A2FA0C" box="[151,210,1468,1493]" pageId="9" pageNumber="356">Type</typeStatus>
material.
</emphasis>
<typeStatus id="932389B8583CDD39B501FA5DB4C3FA0C" box="[332,435,1468,1493]" pageId="9" pageNumber="356" type="holotype">
<emphasis id="7EECEB08583CDD39B501FA5DB4C3FA0C" box="[332,435,1468,1493]" italics="true" pageId="9" pageNumber="356">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="2A89AFDF583CDD39B58CFA5DB750FA0C" box="[449,544,1468,1493]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="356">MZUSP</collectionCode>
1580 (part and counterpart of entire female specimen, preserved in dorsal view; head partially preserved.
<typeStatus id="932389B8583CDD39B5FAFA00B76EFA21" box="[439,542,1505,1528]" pageId="9" pageNumber="356" type="paratype">
<emphasis id="7EECEB08583CDD39B5FAFA00B76EFA21" box="[439,542,1505,1528]" italics="true" pageId="9" pageNumber="356">Paratype</emphasis>
</typeStatus>
:
<collectionCode id="2A89AFDF583CDD39B663FA3EB7FFFA21" box="[558,655,1503,1528]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="356">MZUSP</collectionCode>
1581 (entire female specimen, preserved in dorsal view; head poorly preserved). NE
<collectingCountry id="348F778A583CDD39B50AF9E5B4FEF9C4" box="[327,398,1540,1565]" name="Brazil" pageId="9" pageNumber="356">Brazil</collectingCountry>
, Santana Fm., Aptian/Albian, Early Cretaceous. Both specimens are housed in the Museu de Zoologia da Universidade de São Paulo, São Paulo,
<collectingCountry id="348F778A583CDD39B6ADF9C6B654F999" box="[736,804,1575,1600]" name="Brazil" pageId="9" pageNumber="356">Brazil</collectingCountry>
(
<collectionCode id="2A89AFDF583CDD39B77EF9C9B6E7F999" box="[819,919,1576,1600]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="356">MZUSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="04826491583CDD39B48AF9ADB758F951" pageId="9" pageNumber="356" type="etymology">
<paragraph id="4C27371A583CDD39B48AF9ADB758F951" blockId="9.[151,1436,1468,2011]" pageId="9" pageNumber="356">
<emphasis id="7EECEB08583CDD39B48AF9ADB43EF9BC" bold="true" box="[199,334,1612,1637]" pageId="9" pageNumber="356">Etymology.</emphasis>
The species is named after Dr. Rafael Gioia Martins-Neto (
<date id="382611DA583CDD39B7BDF9ADB105F9BC" box="[1008,1141,1612,1637]" pageId="9" pageNumber="356" value="1954" valueMax="2010">19542010</date>
), Brazilian paleontologist, collector of the MZUSP specimens.
</paragraph>
</subSubSection>
<subSubSection id="04826491583CDD39B48AF975B487F909" pageId="9" pageNumber="356" type="diagnosis">
<paragraph id="4C27371A583CDD39B48AF975B487F909" blockId="9.[151,1436,1468,2011]" pageId="9" pageNumber="356">
<emphasis id="7EECEB08583CDD39B48AF975B431F974" bold="true" box="[199,321,1684,1709]" pageId="9" pageNumber="356">Diagnosis.</emphasis>
This species differs from the other Cretaceous species by its dark costal field, wide small discal cell and veins M3 and M4 divergent.
</paragraph>
</subSubSection>
<subSubSection id="04826491583CDD3AB48AF93FB166FF21" lastPageId="10" lastPageNumber="357" pageId="9" pageNumber="356" type="description">
<paragraph id="4C27371A583CDD39B48AF93FB7E7F802" blockId="9.[151,1436,1468,2011]" pageId="9" pageNumber="356">
<emphasis id="7EECEB08583CDD39B48AF93FB429F92E" bold="true" box="[199,345,1758,1783]" pageId="9" pageNumber="356">Description.</emphasis>
<emphasis id="7EECEB08583CDD39B513F93FB4D3F92E" box="[350,419,1758,1783]" italics="true" pageId="9" pageNumber="356">Head.</emphasis>
Scape stout and long, about 2.4x longer than wide.
<emphasis id="7EECEB08583CDD39B7ACF93FB142F92E" box="[993,1074,1758,1783]" italics="true" pageId="9" pageNumber="356">Thorax</emphasis>
stout, ca. 2.1x longer than wide.
<emphasis id="7EECEB08583CDD39B4DAF8E5B5BFF8C2" box="[151,207,1796,1819]" italics="true" pageId="9" pageNumber="356">Wing</emphasis>
transparent except for dark pterostigma and costal field. Sc reaching wing margin shortly beyond the level of first bifurcation of Rs and near the level of the tip of M4; sc-r present, positioned near mid-point between origin of Rs and tip of R1; Rs curved, slightly longer than R2+3; R1 reaching wing margin at the level of bifurcation of R2+3; r- r oblique, linking R1 to R2; R2 oblique; medial vein four-branched; discal cell ca. 1.63x longer than wide; M3 and M4 divergent toward wing margin, similar in shape; m-cu linking Cu to the base of M4.
<emphasis id="7EECEB08583CDD39B00CF87AB1DBF86D" box="[1089,1195,1947,1972]" italics="true" pageId="9" pageNumber="356">Abdomen</emphasis>
stout, ca. 2.0x longer than thorax; ovipositor with short, stout cerci.
</paragraph>
<paragraph id="4C27371A583FDD3AB48AFF76B14EFF69" blockId="10.[151,1436,151,248]" box="[199,1086,151,176]" pageId="10" pageNumber="357">
<emphasis id="7EECEB08583FDD3AB48AFF76B406FF69" bold="true" box="[199,374,151,176]" pageId="10" pageNumber="357">Measurements</emphasis>
. Body length, 10.711.9. Wing length/width, 8.910.5/2.42.8.
</paragraph>
<paragraph id="4C27371A583FDD3AB48AFF5DB166FF21" blockId="10.[151,1436,151,248]" pageId="10" pageNumber="357">
<emphasis id="7EECEB08583FDD3AB48AFF5DB476FF0C" bold="true" box="[199,262,188,213]" pageId="10" pageNumber="357">Note.</emphasis>
The
<typeStatus id="932389B8583FDD3AB509FF5DB4D6FF0C" box="[324,422,188,213]" pageId="10" pageNumber="357" type="holotype">holotype</typeStatus>
is a bit smaller than the
<typeStatus id="932389B8583FDD3AB681FF5CB65FFF0C" box="[716,815,189,213]" pageId="10" pageNumber="357" type="paratype">paratype</typeStatus>
, but since the two specimens have basically the same pattern of wing venation, they are treated here as belonging to the same species.
</paragraph>
</subSubSection>
</treatment>
</document>