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<document id="C2E144BAE977548239DAAE782D746D1A" ID-CLB-Dataset="46726" ID-DOI="10.11646/zootaxa.3753.4.4" ID-GBIF-Dataset="59df87e1-3da4-43ef-8c4c-b8a7cdd580ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="249874" ID-ZooBank="394ADA8F-E7A8-4751-941F-FE8E6A3921A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365347958" checkinUser="plazi" docAuthor="Ribeiro, Guilherme C. &amp; Lukashevich, Elena D." docDate="2014" docId="C431860C5838DD3DB4DAFF76B400FC9B" docLanguage="en" docName="zt03753p363.pdf" docOrigin="Zootaxa 3753 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptotarsus contractus Ribeiro &amp; Lukashevich, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="360" masterDocId="3808FE745835DD30B44DFFE1B570FFD9" masterDocTitle="New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)" masterLastPageNumber="363" masterPageNumber="347" pageNumber="360" updateTime="1698295164094" updateUser="ExternalLinkService">
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<mods:title id="6CDF6E55EB81A52D6F27FCA27D0EA3F5">New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)</mods:title>
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<mods:namePart id="CEA48EC8C20619945A4E3A0EA294CBFB">Ribeiro, Guilherme C.</mods:namePart>
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<mods:namePart id="73FCD79D7E1FEEB1458072E6BE0B0892">Lukashevich, Elena D.</mods:namePart>
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<mods:title id="9B193C806D106C3DA9AA672A033C459C">Zootaxa</mods:title>
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<mods:date id="CA61BBE048DF467A601BCF6B1AFC1F18">2014</mods:date>
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<mods:number id="EE78B10FB5ADAE5B7D2467BBA9054211">3753</mods:number>
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<treatment id="C431860C5838DD3DB4DAFF76B400FC9B" ID-DOI="http://doi.org/10.5281/zenodo.6143105" ID-GBIF-Taxon="119410703" ID-Zenodo-Dep="6143105" LSID="urn:lsid:plazi:treatment:C431860C5838DD3DB4DAFF76B400FC9B" httpUri="http://treatment.plazi.org/id/C431860C5838DD3DB4DAFF76B400FC9B" lastPageNumber="360" pageId="13" pageNumber="360">
<subSubSection id="048264915838DD3DB4DAFF76B469FF0A" pageId="13" pageNumber="360" type="nomenclature">
<paragraph id="4C27371A5838DD3DB4DAFF76B77FFF6B" blockId="13.[151,527,151,211]" box="[151,527,151,178]" pageId="13" pageNumber="360">
<heading id="176F80765838DD3DB4DAFF76B77FFF6B" bold="true" box="[151,527,151,178]" fontSize="11" level="1" pageId="13" pageNumber="360" reason="1">
<emphasis id="7EECEB085838DD3DB4DAFF76B77FFF6B" bold="true" box="[151,527,151,178]" pageId="13" pageNumber="360">
<taxonomicName id="8B984C995838DD3DB4DAFF76B4DAFF68" box="[151,426,151,177]" class="Insecta" family="Tipulidae" genus="Leptotarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="360" phylum="Arthropoda" rank="species" species="contractus" status="sp. nov.">
<emphasis id="7EECEB085838DD3DB4DAFF76B4DAFF68" bold="true" box="[151,426,151,177]" italics="true" pageId="13" pageNumber="360">Leptotarsus contractus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65DF56735838DD3DB5FFFF79B77FFF6B" box="[434,527,152,178]" pageId="13" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4C27371A5838DD3DB4DAFF5BB469FF0A" blockId="13.[151,527,151,211]" box="[151,281,186,211]" pageId="13" pageNumber="360">
(
<figureCitation id="D4A32B9F5838DD3DB4D2FF5BB597FF0A" box="[159,231,186,211]" captionStart="FIGURES 5 9" captionStartId="4.[151,264,1935,1957]" captionTargetBox="[290,1291,196,1902]" captionTargetId="figure@4.[286,1297,193,1905]" captionTargetPageId="4" captionText="FIGURES 5 9. Wing venation of Leptotarsus spp. from Las Hoyas, Early Cretaceous, Spain. 5 — L. buscalioniae sp. nov., holotype LH 35088, right wing. 6 — L. buscalioniae sp. nov., holotype LH 35088, left wing. 7 — L. ibericus sp. nov., holotype LH 28206. 8 — L. ibericus sp. nov., paratype LH 29995. 9 — L. contractus sp. nov., holotype LH 15419. Scale bar 1 mm." httpUri="https://zenodo.org/record/249876/files/figure.png" pageId="13" pageNumber="360">Figs 9</figureCitation>
,
<figureCitation id="D4A32B9F5838DD3DB4BEFF5BB460FF0A" box="[243,272,186,211]" captionStart="FIGURE 26" captionStartId="13.[151,250,1551,1573]" captionTargetBox="[153,1433,889,1529]" captionTargetId="figure@13.[153,1433,889,1529]" captionTargetPageId="13" captionText="FIGURE 26. Leptotarsus contractus sp. nov., from Las Hoyas, Early Cretaceous, Spain, holotype LH 15419, total view." httpUri="https://zenodo.org/record/249884/files/figure.png" pageId="13" pageNumber="360">26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="048264915838DD3DB4DAFEE3B4E6FEBA" pageId="13" pageNumber="360" type="materials_examined">
<paragraph id="4C27371A5838DD3DB4DAFEE3B4E6FEBA" blockId="13.[151,1436,258,834]" pageId="13" pageNumber="360">
<emphasis id="7EECEB085838DD3DB4DAFEE3B435FEC2" bold="true" box="[151,325,258,283]" pageId="13" pageNumber="360">
<typeStatus id="932389B85838DD3DB4DAFEE3B5A2FEC2" box="[151,210,258,283]" pageId="13" pageNumber="360">Type</typeStatus>
material.
</emphasis>
<emphasis id="7EECEB085838DD3DB501FEE3B4CCFEC2" box="[332,444,258,283]" italics="true" pageId="13" pageNumber="360">
<typeStatus id="932389B85838DD3DB501FEE3B4C6FEC2" box="[332,438,258,283]" pageId="13" pageNumber="360" type="holotype">Holotype</typeStatus>
:
</emphasis>
LH 15419± (part and counterpart of male without head). East-central
<collectingCountry id="348F778A5838DD3DB082FEE3B062FEC2" box="[1231,1298,258,283]" name="Spain" pageId="13" pageNumber="360">Spain</collectingCountry>
, Las Hoyas; La Huérguina Limestone Formation, Late Barremian, Early Cretaceous. Housed in Museo de Ciencias de Castillia—La Mancha.
</paragraph>
</subSubSection>
<subSubSection id="048264915838DD3DB48AFE8FB069FE5E" box="[199,1305,366,391]" pageId="13" pageNumber="360" type="etymology">
<paragraph id="4C27371A5838DD3DB48AFE8FB069FE5E" blockId="13.[151,1436,258,834]" box="[199,1305,366,391]" pageId="13" pageNumber="360">
<emphasis id="7EECEB085838DD3DB48AFE8FB43EFE5E" bold="true" box="[199,334,366,391]" pageId="13" pageNumber="360">Etymology.</emphasis>
From the Latin
<taxonomicName id="8B984C995838DD3DB64EFE8EB709FE5F" box="[515,633,367,390]" class="Insecta" family="Tipulidae" genus="Leptotarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="360" phylum="Arthropoda" rank="species" species="contractus">
<emphasis id="7EECEB085838DD3DB64EFE8EB709FE5F" box="[515,633,367,390]" italics="true" pageId="13" pageNumber="360">contractus</emphasis>
</taxonomicName>
(tightened) in reference to unusually narrow shape of wing.
</paragraph>
</subSubSection>
<subSubSection id="048264915838DD3DB48AFE73B761FE16" pageId="13" pageNumber="360" type="diagnosis">
<paragraph id="4C27371A5838DD3DB48AFE73B761FE16" blockId="13.[151,1436,258,834]" pageId="13" pageNumber="360">
<emphasis id="7EECEB085838DD3DB48AFE73B431FE72" bold="true" box="[199,321,402,427]" pageId="13" pageNumber="360">Diagnosis.</emphasis>
This species differ from all Cretaceous species by longer Rs (Rs origin proximal to connection of
<emphasis id="7EECEB085838DD3DB4DAFE56B5BFFE17" box="[151,207,439,462]" italics="true" pageId="13" pageNumber="360">m-cu</emphasis>
and Cu) and narrower wing.
</paragraph>
</subSubSection>
<subSubSection id="048264915838DD3DB48AFE3BB400FC9B" pageId="13" pageNumber="360" type="description">
<paragraph id="4C27371A5838DD3DB48AFE3BB046FD76" blockId="13.[151,1436,258,834]" pageId="13" pageNumber="360">
<emphasis id="7EECEB085838DD3DB48AFE3BB429FE2A" bold="true" box="[199,345,474,499]" pageId="13" pageNumber="360">Description.</emphasis>
<emphasis id="7EECEB085838DD3DB52FFE3BB4C3FE2A" box="[354,435,474,499]" italics="true" pageId="13" pageNumber="360">Thorax</emphasis>
stout, as long as wide, transverse suture distinct, praescutal longitudinal stripes clearly delimited.
<emphasis id="7EECEB085838DD3DB543FE1EB437FDCF" box="[270,327,511,534]" italics="true" pageId="13" pageNumber="360">Wing</emphasis>
transparent except for dark pterostigma. Sc reaching wing margin at the level of first bifurcation of Rs and beyond level of the tip of Cu; sc-r present, positioned near the tip of Sc; Rs 1.8 times longer than R2+3; R1 reaching wing margin distal to level of R2+3 bifurcation; r-r oblique, linking R1 to R2, R2 oblique; medial vein fourbranched; discal cell narrowed; distal sections of veins M3 and M4 subparallel, subequal; long m-cu linking Cu to the base of M4.
<emphasis id="7EECEB085838DD3DB508FD77B4DFFD76" box="[325,431,662,687]" italics="true" pageId="13" pageNumber="360">Abdomen</emphasis>
stout, ca. 1.5x longer than thorax; gonocoxite conical, probably as long as wide.
</paragraph>
<paragraph id="4C27371A5838DD3DB48AFD5CB6C3FD0F" blockId="13.[151,1436,258,834]" box="[199,947,701,726]" pageId="13" pageNumber="360">
<emphasis id="7EECEB085838DD3DB48AFD5CB406FD0F" bold="true" box="[199,374,701,726]" pageId="13" pageNumber="360">Measurements</emphasis>
. Body length, 7.1. Wing length/width, 7.17.4/1.6.
</paragraph>
<paragraph id="4C27371A5838DD3DB48AFD00B400FC9B" blockId="13.[151,1436,258,834]" pageId="13" pageNumber="360">
<emphasis id="7EECEB085838DD3DB48AFD00B58FFD23" bold="true" box="[199,255,737,762]" pageId="13" pageNumber="360">Note</emphasis>
. One of the wings is probably distorted judging by the shape of second wing with unclear venation. The virtually straight vein m-cu (
<figureCitation id="D4A32B9F5838DD3DB5A5FCE4B740FCC7" box="[488,560,773,798]" captionStart="FIGURES 5 9" captionStartId="4.[151,264,1935,1957]" captionTargetBox="[290,1291,196,1902]" captionTargetId="figure@4.[286,1297,193,1905]" captionTargetPageId="4" captionText="FIGURES 5 9. Wing venation of Leptotarsus spp. from Las Hoyas, Early Cretaceous, Spain. 5 — L. buscalioniae sp. nov., holotype LH 35088, right wing. 6 — L. buscalioniae sp. nov., holotype LH 35088, left wing. 7 — L. ibericus sp. nov., holotype LH 28206. 8 — L. ibericus sp. nov., paratype LH 29995. 9 — L. contractus sp. nov., holotype LH 15419. Scale bar 1 mm." httpUri="https://zenodo.org/record/249876/files/figure.png" pageId="13" pageNumber="360">Fig. 9</figureCitation>
) is also suggestive of such distortion. In
<taxonomicName id="8B984C995838DD3DB056FCE4B1F5FCC7" box="[1051,1157,773,798]" class="Insecta" family="Tipulidae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="360" phylum="Arthropoda" rank="family">Tipulidae</taxonomicName>
in general, m-cu is not straight but curved.
</paragraph>
</subSubSection>
</treatment>
</document>