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<document ID-CLB-Dataset="24980" ID-DOI="http://dx.doi.org/10.3897/zookeys.885.38655" ID-GBIF-Dataset="8e14d288-ddb5-485a-bd43-0cd3f46754ff" ID-PMC="PMC6848236" ID-Pensoft-Pub="1313-2970-885-51" ID-Pensoft-UUID="BCEA8CB1014D5D329A2E4B2CF99881FF" ID-PubMed="31736618" ID-ZooBank="BA152050AF7344CA8CED6D30F963CBC9" ModsDocID="1313-2970-885-51" checkinTime="1572912880629" checkinUser="pensoft" docAuthor="Zlatkov, Boyan &amp; Huemer, Peter" docDate="2019" docId="B9BD8C9E92C159D8BC8D950C3C830553" docLanguage="en" docName="ZooKeys 885: 51-87" docOrigin="ZooKeys 885" docSource="http://dx.doi.org/10.3897/zookeys.885.38655" docTitle="Clepsis eatoniana Ragonot 1881, stat. rev." docType="treatment" docVersion="6" id="BCEA8CB1014D5D329A2E4B2CF99881FF" lastPageNumber="51" masterDocId="BCEA8CB1014D5D329A2E4B2CF99881FF" masterDocTitle="Remarkable confusion in some Western Palearctic Clepsis leads to a revised taxonomic concept (Lepidoptera, Tortricidae)" masterLastPageNumber="87" masterPageNumber="51" pageNumber="51" updateTime="1732932338753" updateUser="ExternalLinkService">
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<mods:title id="7A9B2D4A000A7B3A447EBF62A3AE00D8">Remarkable confusion in some Western Palearctic Clepsis leads to a revised taxonomic concept (Lepidoptera, Tortricidae)</mods:title>
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<mods:namePart id="13EBFE8969992C301C0261518A32AD4B">Zlatkov, Boyan</mods:namePart>
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<mods:date id="AC070F3545B04B4D067F9C8BC8CD1D17">2019</mods:date>
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<mods:identifier id="1A2298471432F43A1FAB6418CE9C4381" type="DOI">http://dx.doi.org/10.3897/zookeys.885.38655</mods:identifier>
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<treatment id="B9BD8C9E92C159D8BC8D950C3C830553" ID-GBIF-Taxon="160133794" LSID="urn:lsid:plazi:treatment:B9BD8C9E92C159D8BC8D950C3C830553" httpUri="http://treatment.plazi.org/id/B9BD8C9E92C159D8BC8D950C3C830553" lastPageNumber="51" pageId="0" pageNumber="51" scope_family="Tortricidae" scope_order="Lepidoptera">
<subSubSection id="F96C5F8A01C289E9C2FD7D4D352C027B" pageId="0" pageNumber="51" type="nomenclature">
<paragraph id="9AED383C67B9B37B2BAFADD7E352E95A" pageId="0" pageNumber="51">
<taxonomicName id="205B4854E656FD07DB8CF9ACC4B069C0" ID-CoL="7CJ7N" LSID="B9BD8C9E-92C1-59D8-BC8D-950C3C830553" authority="Ragonot, 1881" authorityName="Ragonot" authorityYear="1881" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis eatoniana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="eatoniana">Clepsis eatoniana (Ragonot, 1881)</taxonomicName>
<taxonomicNameLabel id="02FC4CEB8AE3F5143FDFBAA73E7406EE" pageId="0" pageNumber="51">stat. rev.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A158DFB07A07B7BC0D37AC55914030A7" pageId="0" pageNumber="51" type="reference_group">
<paragraph id="FB0ED626418BCE21AC12F52372BFA830" pageId="0" pageNumber="51">
<taxonomicName id="2D13E6D0EA24F077917D87B0549FB665" class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix eatoniana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="eatoniana">Tortrix eatoniana</taxonomicName>
Ragonot, 1881: 231 (Portugal).
</paragraph>
<paragraph id="6CCF25B3873576A120DE63058ED26F94" pageId="0" pageNumber="51">
<taxonomicName id="15E7D08ACED66C3F1131C82C2DB4E4E4" class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix xylotoma" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="xylotoma">Tortrix xylotoma</taxonomicName>
Meyrick, 1891: 13 (Algeria) syn. nov.
</paragraph>
<paragraph id="3A2335FA0955CB1E786D51112412B169" pageId="0" pageNumber="51">
<taxonomicName id="D87B1FE0AB90EF22EF364CEA5E32D392" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis razowskii" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="razowskii">Clepsis razowskii</taxonomicName>
<normalizedToken id="A2ED773ADBC151DC966888C12B3DAE1B" originalValue="Gastón">Gaston</normalizedToken>
, Vives &amp; Revilla, 2017: 691 (Spain) syn. nov.
</paragraph>
<paragraph id="CC65B63E219A4729E3A8A4D39783FA8A" pageId="0" pageNumber="51">
non
<taxonomicName id="E3B638DA1130AEA13CE770233CE28D6D" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">Clepsis consimilana</taxonomicName>
(
<normalizedToken id="861A93DFD483C4680AC2E4F1B433A2D3" originalValue="Hübner">Huebner</normalizedToken>
, 1817)
</paragraph>
</subSubSection>
<subSubSection id="AEFB8A131B63850D1383B7C3D0771DBD" pageId="0" pageNumber="51" type="materials_examined">
<paragraph id="7C53C97BEADD8AE36738B5CE2C782FB4" pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph id="DBF53EAE46A55F55B09BF8F9CF4D0801" pageId="0" pageNumber="51">
Lectotype ♂ by designation of
<bibRefCitation id="9533561649BE5238753B76C844A219E5" author="Razowski, J" journalOrPublisher="Acta zoologica cracoviensia" pageId="0" pageNumber="51" pagination="101 - 198" refId="B34" refString="Razowski, J, 1979. Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I. Acta zoologica cracoviensia 23: 101 - 198" title="Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I." volume="23" year="1979">Razowski (1979)</bibRefCitation>
, pinned, with 7 labels: &quot;Eatoniana / Rag. n. sp. / Portugal&quot; [handwritten; the same label on backside:] &quot;port Olivaes / Port. [ugal] / 24.4.80&quot; &quot;
<emphasis id="7ECA28E6DD82D16D6D91E56F7C6A26EA" italics="true" pageId="0" pageNumber="51">Tortrix / eatoniana</emphasis>
Rag. / Ent. month. Mag., / 1881, 17, p. 231&quot; [handwritten] &quot;Mus. Paris / coll. Ragonot / 15997&quot; [printed and handwritten]
<normalizedToken id="0F9F37ECACC67E891959EB92A2E620B0" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
[red printed]
<normalizedToken id="A4EA661E2CDF1F565F30288B9D4E1D4F" originalValue="“communiqué">&quot;communique</normalizedToken>
/
<normalizedToken id="99F70B65ED5E2DB9AF9C020293A27357" originalValue="á">a</normalizedToken>
M. J. Kennel&quot; [handwritten] &quot;Genitalia ♂ / P. Viette /
<normalizedToken id="0D696BE880CE325000EB39D6E30BD35F" originalValue="prép">prep</normalizedToken>
. No. 3699&quot; [printed] &quot;1901 / coll. E. L. Ragonot /
<normalizedToken id="DAB6925F7CA7436EBF74BCEB2DA76110" originalValue="Muséum">Museum</normalizedToken>
Paris&quot; [printed]; male genitalia on a slide with two labels: &quot;♂ /
<emphasis id="688ED08CDE09BB97AB89350DB9B44A5F" italics="true" pageId="0" pageNumber="51">Tortrix</emphasis>
/
<emphasis id="76263FB29211B512F19997A2C453E5FA" italics="true" pageId="0" pageNumber="51">eatoniana</emphasis>
/ Rag. / Type&quot;
<normalizedToken id="08D8A1AA7841BF2AE619462CDF38E304" originalValue="“Prép">&quot;Prep</normalizedToken>
. P. Viette / # 3699 /
<normalizedToken id="8BB459B70EC1F5053CAE120CA90F1D83" originalValue="Muséum">Museum</normalizedToken>
/ Paris / Type&quot; [both handwritten, with red border].
</paragraph>
<paragraph id="4FDE07F651A3264EBC71A56F6AC22E56" pageId="0" pageNumber="51">PORTUGAL • 1 ♂; Lisbon, Olivais; 24 Apr. 1880; Ragonot leg.; GS 3699; MNHN.</paragraph>
<paragraph id="7AA88A8D9CF993DCF19376A6EBE77356" pageId="0" pageNumber="51">
Lectotype ♂ of
<taxonomicName id="F134B59130806D2E2BE38FA066A9C789" class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix xylotoma" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="xylotoma">
<emphasis id="5CD8D165D372F1C95D74FE927286059C" italics="true" pageId="0" pageNumber="51">Tortrix xylotoma</emphasis>
</taxonomicName>
by designation of
<bibRefCitation id="A989B4B929246EBD7DA6000DF62E8C42" author="Razowski, J" journalOrPublisher="Acta zoologica cracoviensia" pageId="0" pageNumber="51" pagination="101 - 198" refId="B34" refString="Razowski, J, 1979. Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I. Acta zoologica cracoviensia 23: 101 - 198" title="Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I." volume="23" year="1979">Razowski (1979)</bibRefCitation>
, pinned, with 7 labels: &quot;Bougie / Algeria / 25/4/90&quot; [handwritten] &quot;Tortrix / xylotoma / Meyr[ick] / Type ♂&quot; [handwritten and printed] &quot;Lecto- / type&quot; [circular with violet border]
<normalizedToken id="33B0BA1C307F5033BA35D0712791249A" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
[circular with red border] &quot;♂ genitalia on / slide 6.IV.1949 / J.F.G.C. 9364&quot; [printed and handwritten] &quot;Meyrick coll. / B. M. 1938
<normalizedToken id="F0A9D17654E353D0398D5E3BABAF1DF9" originalValue="290”">-290&quot;</normalizedToken>
&quot;NHMUK010219594 [QR code]&quot;.
</paragraph>
<paragraph id="40E39FA91A1EACB44B28D9023346A15C" pageId="0" pageNumber="51">
ALGERIA • 1 ♂;
<normalizedToken id="975CC0ADF5C67910FE637F0C8BC45066" originalValue="Béjaïa">Bejaia</normalizedToken>
; 25 Apr. 1890; Meyrick leg.; BMNH NHMUK010219594.
</paragraph>
<paragraph id="DF7D47E700CEDCCAED94F6F578B7E852" pageId="0" pageNumber="51">Other material: SPAIN • 8 ♂♂, 1 ♀; Valencia, El Saler, Albufera; 39.3278°N, -0.3078°W; alt. 5 m; 18 May 2004; P. Huemer leg.; GS 1/18.10.2017, 2/18.10.2017, 3/18.10.2017; TLMF • 5 ♂♂, same collection data; 8 Sep. 2005; P. Huemer leg.; TLMF • 1 ♂, 1 ♀; Valencia, 5 km NE Albufera, Sierra de Crevillente; alt. 450 m; 26 May 2004; P. Huemer leg.; TLMF • 1 ♀; Valencia, Santa Pola, Playa del Pinet; 38.1585°N, -0.6256°W; alt. 5 m; 5Sep. 2005; P. Huemer leg.; GS 4/18.10.2017; TLMF.</paragraph>
</subSubSection>
<subSubSection id="E5415DD4DE96E677B1E3972CE3377635" pageId="0" pageNumber="51" type="diagnosis">
<paragraph id="A991D65B0D7B8FCD21AAE495BE5F3B56" pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph id="EC629DF6839325024192A3DDB7C73FF1" pageId="0" pageNumber="51">
The wing pattern in males resembles those of
<taxonomicName id="DD5B47EE189A1FE4B0CBB4D35D5A5E10" lsidName="C. trivia" pageId="0" pageNumber="51" rank="species" species="trivia">
<emphasis id="443FD254A3CEA0163ECE3D5C12790AFF" italics="true" pageId="0" pageNumber="51">C. trivia</emphasis>
</taxonomicName>
,
<taxonomicName id="9C2399E36439C6110102B40968641F70" lsidName="C. acclivana" pageId="0" pageNumber="51" rank="species" species="acclivana">
<emphasis id="F03CC21BE20F8E7E5544529872A764D0" italics="true" pageId="0" pageNumber="51">C. acclivana</emphasis>
</taxonomicName>
, and
<taxonomicName id="03CDF09431D907464BCBFE54606728D3" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="6ABCB58D50F1E630577B4C60861613AF" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
.
<taxonomicName id="302B85842D9F1958BB070E75FECAD470" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis eatoniana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="eatoniana">
<emphasis id="70F128C12F881C4BB2188FA5BCE7753A" italics="true" pageId="0" pageNumber="51">Clepsis eatoniana</emphasis>
</taxonomicName>
differs from
<taxonomicName id="6E1CF66231337543B5C4E69566348374" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="79273DE12A25E7335C32937C44243CBC" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
, by the more yellowish instead of fulvous forewing ground colour. The most characteristic feature in the external morphology of
<taxonomicName id="D3FF99913E81E7C14DD1E36BC768DE37" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="B6F2727006D17DEFACA56106B8660E27" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
is the absence of a forewing costal fold, in contrast to
<taxonomicName id="FB26DFB3037CF6E7DE506C6B1F2AB978" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="2068C194BAD58C814062505ED1771E0E" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
. The male genitalia are similar to those in
<taxonomicName id="5A46423266DB1CBE5B4700EA8D1F2099" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="F44DB36369375D4171C4827F86EF569E" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
but the valva is more slender, elongate distally, and the modified large setae are more numerous and more slender. The phallus is adorned with spines, with the keel and terminal process overpassing its tip; in
<taxonomicName id="C0EC3ECD9DE356DDD233224DC3B204D6" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="4147D92DCA41B735E0DC98302DDD8199" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
the phallus lack lateral spines and a keel, and the phallic process is shorter. The caulis in
<taxonomicName id="73EDE01F4B238B049B9A9B3110F31181" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="EE1036C17BFFF16E8CCA25EF0F65F751" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
is larger, the vesica is bent at nearly a right angle to the phallus, its diverticulum has different location, and the cornuti are more slender and smoother. Females of
<taxonomicName id="6AD6532A263840B4BD9AA397807A253B" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="C2A718BDBBC2BB62FE0D64D8F860529E" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
are not distinguishable externally from the females of
<taxonomicName id="2E6409E0AC8F2FC9FF8062089649F02E" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="F282D7D508A41A4001D1A0914205A9DE" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
but differ from
<taxonomicName id="D7EFEAAF5B2724192E5849C722B03749" lsidName="C. striolana" pageId="0" pageNumber="51" rank="species" species="striolana">
<emphasis id="A7C4091C2F051B547645E235D383CCEA" italics="true" pageId="0" pageNumber="51">C. striolana</emphasis>
</taxonomicName>
by the presence of two brown dots on the forewing. The female genitalia of
<taxonomicName id="B1E00D0AF153B605127CD5F5AF2B3C76" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="0F699B513D15BB58A7EE583C9CA80813" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
are similar to those in both the
<taxonomicName id="F92482F33C7C581D962A21BE0446CBD8" lsidName="C. neglectana" pageId="0" pageNumber="51" rank="species" species="neglectana">
<emphasis id="667C6127E04E82D3BFF5FB640982EC1A" italics="true" pageId="0" pageNumber="51">C. neglectana</emphasis>
</taxonomicName>
and
<taxonomicName id="3F9AAF84ECE239B09D15DD8348806E03" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="88468890ABFA5772C37BF0A0EF019B2B" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
species groups, but the protrusion on the right side of the colliculum is much larger and elongated, and the sterigma has larger lateral pockets.
</paragraph>
</subSubSection>
<subSubSection id="7D847F05C8B6025FA3ED27FC70971AD6" pageId="0" pageNumber="51" type="description">
<paragraph id="EF8AAA7BF3607E700CF42102382E3F36" pageId="0" pageNumber="51">Description.</paragraph>
<paragraph id="4F9D220FC170F9FA08B55832B075AA2E" pageId="0" pageNumber="51">
Adult. Sexual dimorphism prominent. Male (
<figureCitation id="E288A70B168201B8F39FFE677BF46CAD" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Adults of Clepsis consimilana species group A-H C. consimilana: A illustration from the original description of Tortrix consimilana B neotype of T. consimilana C male, Bulgaria D lectotype of T. unifasciana E Holotype of Siclobola placida F male, Poland G female, Italy H female, Poland I-L C. eatoniana: I Lectotype of T. eatoniana J male, Spain K Lectotype of Tortrix xylotoma (courtesy by D. Lees and Trustees of the Natural History Museum, London) L female, Spain. Scale bar: 5 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure11" httpUri="https://binary.pensoft.net/fig/352389" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure11">
Fig. 11
<normalizedToken id="AD757060B1DB5724F253F71884586793" originalValue="IK">I-K</normalizedToken>
</figureCitation>
). Head. Vertex fulvous, frons and labial palps brown. Antennae with brown scales on scapus and fulvous scales on pedicellus and flagellum, and with numerous sensilla trichodea as long as width of flagellomeres. Thorax. Fulvous dorsally and creamy ventrally, legs brownish. Tegula fulvous with brown costal margin. Forewing with length 6.8-7.4 mm (mean 7.1,
<emphasis id="8C96C2D38CF1EBA0E050B88A5EDC2B0B" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 3), elongated, costa convex at basal half, slightly sinuate apically. Costal fold lacking (
<figureCitation id="2B6AD7CABA59BE3176A67E217B19C180" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Costal folds of males of Clepsis spp. A C. neglectana, Germany B C. striolana, Austria C C. trivia, Crete D C. acclivana, lectotype of Cacoecia acclivana E C. consimilana, neotype of Tortrix consimilana F C. eatoniana, Spain. Scale bar: 1 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure3" httpUri="https://binary.pensoft.net/fig/352381" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure3">Fig. 3F</figureCitation>
). Upperside background dark yellow with fulvous reticulate pattern more prominent in the paler subterminal area, cilia concolourous or paler. Basal blotch atrophied, with remnants of darker scales at costa and dorsum. Median fascia brown with lead refractive tint, almost interrupted at middle of median cell by yellowish scales, often ceasing at vein CuA. Subapical blotch brown, triangular, more or less well defined. Underside pale grey-brown with creamy longitudinal blotch in distal half of costal area. Specimens with paler and darker forewings and incomplete median fascia observed. Hindwing upperside monochrome pale grey with paler cilia, underside whitish with scattered pale grey-brown scales. Abdomen. Grey. Male genitalia (
<figureCitation id="2D36413CDF776142E3C15B8634916E4C" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genitalia of the Clepsis consimilana species group (some without phallus) A-E C. consimilana: A Tortrix consimilana neotype B C. consimilana, Italy C C. consimilana, Albania D Siclobola placida holotype E T. unifasciana lectotype F T. eatoniana lectotype. Scale bar: 500 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure12" httpUri="https://binary.pensoft.net/fig/352390" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure12">Fig. 12F</figureCitation>
,
<figureCitation id="4F03D969AEE490ED2DE6E8C7AD5FF313" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Male genitalia of the Clepsis consimilana species group: C. eatoniana A Tortrix xylotoma lectotype (courtesy by D. Lees and Trustees of the Natural History Museum, London) B C. eatoniana, Spain. Scale bar: 500 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure15" httpUri="https://binary.pensoft.net/fig/352393" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure15">15</figureCitation>
). Uncus variably shaped, more or less trapezoidal, slightly widened distally, rounded, with slightly convex distal edge which may look incised if excessive pressure is applied on coverslip. Gnathos plough-shaped. Socius small, membranous. Valvae pointed dorsolaterally when mounted on slide. Costal sclerite wide, protruded medially into large triangular labis with elongated spinulate tip (
<figureCitation id="3109CFBE84EBD1CEF1C3192A883824CF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Labides of Clepsis spp. A, B C. neglectana: A Southern Germany B Finland C C. striolana, Austria, North Tyrol D C. trivia, Crete E C. acclivana, paralectotype of Cacoecia acclivana F, G C. consimilana: F Germany, neotype of Tortrix consimilana G Italy H C. eatoniana, Spain. Scale bar: 100 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure5" httpUri="https://binary.pensoft.net/fig/352383" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure5">Fig. 5H</figureCitation>
). Basal and apical part of sacculus of nearly equal length, both forming angle of 150-160°, saccular process large, flat, triangular. Distal part of valva membranous, comparatively small, narrow, with parallel costal and saccular margins, apically triangular, without distinct brachiola, with longitudinal fold on the median surface bearing row of 8-12 large modified setae. Apical part may look as brachiola due to deformation during preparation. Phallus robust, coecum medially concave, basal part curved ventrad at ca. 140°, distal part smoothly curved dorsad, with wide keel on left side grading into large sharp tipped lateral process as long as 0.23
<normalizedToken id="39B6E730EE3533FC7FD91CF37174F413" originalValue="×">x</normalizedToken>
distance between anterior opening and tip of phallus, slightly curved laterally and dorsally, overpassing phallic tip. Several more or less prominent spines pointed caudad located at basal part of keel. Caulis large, widely separated from coecum. Vesica cylindrical, curved at 80-90° to phallus, with small expansion basally and fingerlike dorsal diverticulum (
<figureCitation id="F8986008A7F9B1CEDB304D47758E6560" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phalli with vesicae everted of the Clepsis consimilana species group A-B C. consimilana: A neotype of Tortrix consimilana B specimen from Italy C C. eatoniana, Spain. The phallus of each species is shown in three aspects from top to bottom: sclerotised phallic tube (without vesica) in dorsal view, tip of vesica in dorsal view and whole phallus in left view. Scale bar: 250 μm." figureDoi="10.3897/zookeys.885.38655.figure13" httpUri="https://binary.pensoft.net/fig/352391" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure13">Fig. 13C</figureCitation>
). Three deciduous, slender, straight cornuti attached ventroapically (
<figureCitation id="612B4DF0C73CC19ACF2532BE2D6EDBFB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Specialised male setae in Clepsis spp. A-E cornuti: A C. neglectana, Finland B C. striolana, Austria C C. trivia, Crete D C. consimilana, neotype of Tortrix consimilana E C. eatoniana, Spain F modified large seta from the medial surface of valva of C. consimilana, Germany. Scale bar: 100 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure7" httpUri="https://binary.pensoft.net/fig/352385" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure7">Fig. 7E</figureCitation>
). Gonopore located in straight line with cornuti sockets and diverticulum. Female darker than male (
<figureCitation id="F39285EBC47AE7079B72B40348AF4A51" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Adults of Clepsis consimilana species group A-H C. consimilana: A illustration from the original description of Tortrix consimilana B neotype of T. consimilana C male, Bulgaria D lectotype of T. unifasciana E Holotype of Siclobola placida F male, Poland G female, Italy H female, Poland I-L C. eatoniana: I Lectotype of T. eatoniana J male, Spain K Lectotype of Tortrix xylotoma (courtesy by D. Lees and Trustees of the Natural History Museum, London) L female, Spain. Scale bar: 5 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure11" httpUri="https://binary.pensoft.net/fig/352389" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure11">Fig. 11L</figureCitation>
). Head. Vertex rusty, frons and labial palps brown. Whole antennae with brown scales, sensilla trichodea sparse, shorter than width of flagellomeres. Thorax. Rusty dorsally and creamy ventrally, legs brownish. Tegula rusty with brown costal margin. Forewing length 6.7 mm (n = 1), with shape as in males, upperside uniformly rusty with ill-defined reticulate pattern more distinct in distal half and two brown dots at dorsum, underside mainly creamy with scattered pale brown scales, denser in middle of wing. Hindwing upperside grey brown, underside creamy with darker dorsal part. Abdomen grey.
</paragraph>
<caption id="B1438B09E9DF31BCF9E87F43801F423F" doi="10.3897/zookeys.885.38655.figure15" httpUri="https://binary.pensoft.net/fig/352393" pageId="0" pageNumber="51" start="Figure 15" startId="F15">
<paragraph id="2E12E2843CCC0A91EC055723AC614D8A" pageId="0" pageNumber="51">
<emphasis id="0F9CE8222A5642A4DC90EA4A84F704FE" bold="true" pageId="0" pageNumber="51">Figure 15.</emphasis>
Male genitalia of the
<taxonomicName id="213160B6D4FEA6B51753E7A9003F5857" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis id="2C8EEC9DD7B9962181DAB7C10D6A1996" italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
</taxonomicName>
species group:
<taxonomicName id="96F08E73FDD637CB9032B92DB64F35A9" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="6CE62F101866590DFEAB662EFA54B39B" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
<emphasis id="4D897A820E8E865D64B1133214741C97" bold="true" pageId="0" pageNumber="51">A</emphasis>
<taxonomicName id="69FD06252EC773E965B0A19CF7BD8141" class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix xylotoma" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="xylotoma">
<emphasis id="9FD1E879BD81795EE084DF95176510EF" italics="true" pageId="0" pageNumber="51">Tortrix xylotoma</emphasis>
</taxonomicName>
lectotype (courtesy by D. Lees and Trustees of the Natural History Museum, London)
<emphasis id="80235515F80E82EECCB13E473E45C2D6" bold="true" pageId="0" pageNumber="51">B</emphasis>
<taxonomicName id="117B492B47CA5A55BCB0AA6CC0B54EF3" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="FEFDA14E4174874BF4645B614FA67A5D" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
, Spain. Scale bar: 500
<normalizedToken id="82F0901459E46980CD7FA71BF37563C9" originalValue="μm">μm</normalizedToken>
, all to scale.
</paragraph>
</caption>
<paragraph id="E5799A63D3A11AC1912F49C4B360DDC0" pageId="0" pageNumber="51">
Female genitalia (
<figureCitation id="AA292E3716157556690D550624A96BFF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Female genitalia of the Clepsis consimilana species group A C. consimilana, Italy B C. eatoniana, Spain. The sterigma and colliculum are shown enlarged to the right of the whole genitalia. Scale bars: 1 mm (whole genitalia), 250 μm (sterigma and colliculum)." figureDoi="10.3897/zookeys.885.38655.figure14" httpUri="https://binary.pensoft.net/fig/352392" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure14">Fig. 14B</figureCitation>
). Papillae anales not modified. Apophyses anteriores 1.1
<normalizedToken id="6269F43F38F913DF2B2F7BC1C1187A10" originalValue="×">x</normalizedToken>
longer than apophyses posteriores. Sterigma relatively wide, widened cephalad, with large lateral sclerotised pockets, large excavation on dorsal wall and wide v-shaped median part of lamella antevaginalis. Colliculum large, with length 0.25
<normalizedToken id="DA02AA7D9EC79783D8C4067360EB9C8F" originalValue="×">x</normalizedToken>
length of ductus bursae, asymmetrical, funnel-shaped, with well-developed sclerotisation and two evaginations: very large one at right and small one at left, both consisting of colourless thick cuticle. Ductus bursae long and narrow, emerging at left between out-pocketings, with cestum extending along cranial 0.7
<normalizedToken id="69B31AE1E2EAA66AD415748B61F832D9" originalValue="×">x</normalizedToken>
of its length. Ductus seminalis inserted dorsally at caudal end of ductus bursae. Corpus bursae ovoid, with large falcate signum with capitulum and flat signum consisting of sclerotised papillae located near end of cestum (
<figureCitation id="75D83CC7791D92FAA3B7A10BA69981CA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Signa of bursa copulatrix in Clepsis spp. A C. neglectana B C. striolana C C. trivia D C. consimilana E C. eatoniana F A detail of plate shaped signum of C. consimilana consisting of sclerotised papillae. 1 falcate signum, 2 plate shaped signum, 3 cestum. Scale bars: 250 μm (A-E, all to scale), 25 μm (F)." figureDoi="10.3897/zookeys.885.38655.figure9" httpUri="https://binary.pensoft.net/fig/352387" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure9">Fig. 9E</figureCitation>
).
</paragraph>
<paragraph id="E799C5CB796B62F4D1DB2629ECF5D02F" pageId="0" pageNumber="51">Preimaginal stages unknown.</paragraph>
</subSubSection>
<subSubSection id="035CC32063ACF85DCAAE0933936DA6E4" pageId="0" pageNumber="51" type="molecular data">
<paragraph id="65CD5F05B7EEAB0DCDF7E4B7DF8CAE03" pageId="0" pageNumber="51">Molecular data</paragraph>
<paragraph id="C1B251E0BDDE0BFA37F681BE8A0B8F55" pageId="0" pageNumber="51">
(
<figureCitation id="4B218ED69511DA2F65F00179E6EE70F8" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Neighbour-joining tree of Clepsis neglectana and C. consimilana species groups (Kimura 2 parameter, built with MEGA 6; cf. Tamura et al. 2013). Note: the scale bar only applies to internal branches between species. The width of the triangles represents the sample size, and the depth the relative genetic variation within the cluster (2 x scale bar). Source: DNA Barcode data from BOLD (Barcode of Life Database, cf. Ratnasingham and Hebert 2007)." figureDoi="10.3897/zookeys.885.38655.figure16" httpUri="https://binary.pensoft.net/fig/352394" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure16">Fig. 16</figureCitation>
). BIN: BOLD:AAJ1025. The intraspecific average of the barcode region is 0.09%, the maximum distance 0.15% (p-dist) (
<emphasis id="50665045C91F4D920E394E2283F8402D" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 2). The minimum distance to the nearest neighbour
<taxonomicName id="B6FA0C4AF48E9BFF7465021EAB35B843" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis id="D3F764183C1A7A60A051513C3FE1140F" italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
</taxonomicName>
is 2.25%.
</paragraph>
<caption id="DFA8B487D8CDB8CF863826286B1CA7AE" doi="10.3897/zookeys.885.38655.figure16" httpUri="https://binary.pensoft.net/fig/352394" pageId="0" pageNumber="51" start="Figure 16" startId="F16">
<paragraph id="EE0BF19D65451B2F2E230E40DE5099C1" pageId="0" pageNumber="51">
<emphasis id="C1D034E26EEFFAAFF379A3CED2187A28" bold="true" pageId="0" pageNumber="51">Figure 16.</emphasis>
Neighbour-joining tree of
<taxonomicName id="29BDFFAC864D175A134BE2990F4D01E9" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis neglectana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="neglectana">
<emphasis id="67F0162A3DDF9213C8E5764E73C8F1D3" italics="true" pageId="0" pageNumber="51">Clepsis neglectana</emphasis>
</taxonomicName>
and
<taxonomicName id="9136542CE59CFD60CC272BF19CBCA6D1" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="023AEF6A2DFAC99A297CE5F5EC9B22B3" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
species groups (Kimura 2 parameter, built with MEGA6; cf.
<bibRefCitation id="ED279424E39E6F60A330E7447BD16A34" DOI="https://doi.org/10.1093/molbev/mst197" author="Tamura, K" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="51" pagination="2725 - 2729" refId="B41" refString="Tamura, K, Stecher, G, Peterson, D, Filipski, A, Kumar, S, 2013. MEGA6: Molecular Evolutionary Genetics Analysis Version 6.0. Molecular Biology and Evolution 30: 2725 - 2729" title="MEGA 6: Molecular Evolutionary Genetics Analysis Version 6.0." url="https://doi.org/10.1093/molbev/mst197" volume="30" year="2013">Tamura et al. 2013</bibRefCitation>
). Note: the scale bar only applies to internal branches between species. The width of the triangles represents the sample size, and the depth the relative genetic variation within the cluster (2
<normalizedToken id="173ADBEA1D62E36DE8708655B2A33C53" originalValue="×">x</normalizedToken>
scale bar). Source: DNA Barcode data from BOLD (Barcode of Life Database, cf.
<bibRefCitation id="90C8039C0F2A17396C61622EF61284AB" author="Ratnasingham, S" journalOrPublisher="The Entomologist's monthly magazine" pageId="0" pageNumber="51" publicationUrl="http://www.barcodinglife.org" refId="B32" refString="Ratnasingham, S, Hebert, PDN, 2007. .http://www.barcodinglife.org" url="http://www.barcodinglife.org" year="2007">Ratnasingham and Hebert 2007</bibRefCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1BBC88D9641EC72922D0E9AB7353F4C4" pageId="0" pageNumber="51" type="distribution">
<paragraph id="FCE2CDDDB2AA3089379AC3A25C52C7BF" pageId="0" pageNumber="51">Distribution</paragraph>
<paragraph id="866CE252F39BD0C8668C2C2D3F19B6CF" pageId="0" pageNumber="51">
(
<figureCitation id="C02357334238916B022B22988577A3B7" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution map of examined material of Clepsis neglectana and C. consimilana species groups. Map created with SimpleMappr (http: // www. simplemappr. net)." figureDoi="10.3897/zookeys.885.38655.figure17" httpUri="https://binary.pensoft.net/fig/352395" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure17">Fig. 17</figureCitation>
). Europe. Portugal: Lisbon, Olivais; Ponte de Mucela (Ponte de Morcellos) (
<bibRefCitation id="B861F6268DE6DB88AE44743BAC653EF5" author="Ragonot, EL" journalOrPublisher="The Entomologist's monthly magazine" pageId="0" pageNumber="51" pagination="229 - 233" refId="B31" refString="Ragonot, EL, 1881. Notes on the entomology of Portugal. IV. Lepidoptera (Continued). Micro-Lepidoptera (except Tineina) collected by the Rev. A. E. Eaton in 1880, with descriptions of new species. The Entomologist's monthly magazine 17: 229 - 233" title="Notes on the entomology of Portugal. IV. Lepidoptera (Continued). Micro-Lepidoptera (except Tineina) collected by the Rev. A. E. Eaton in 1880, with descriptions of new species." volume="17" year="1881">Ragonot 1881</bibRefCitation>
); Lusitania; Spain: Cadiz, Granada, Malaga, Sevilla, Zaragoza; possibly France (
<bibRefCitation id="48EC2139C2E040FAFD4F6B1490C91491" author="Gaston, J" journalOrPublisher="Shilap Revista de lepidopterologia" pageId="0" pageNumber="51" pagination="689 - 698" refId="B15" refString="Gaston, J, Vives Moreno, A, Revilla, Tx, 2017. Descripcion de tres especies nuevas de la Familia Tortricidae en la Peninsula Iberica (Lepidoptera: Tortricidae). Shilap Revista de lepidopterologia 45: 689 - 698" title="Descripcion de tres especies nuevas de la Familia Tortricidae en la Peninsula Iberica (Lepidoptera: Tortricidae)." volume="45" year="2017">
<normalizedToken id="59E223DD46F7DDAD1876A11FE9B33F13" originalValue="Gastón">Gaston</normalizedToken>
et al. 2017
</bibRefCitation>
), Valencia. Africa: Algeria.
</paragraph>
<caption id="8D620E0CD7BDB3B52617779EB5E3278B" doi="10.3897/zookeys.885.38655.figure17" httpUri="https://binary.pensoft.net/fig/352395" pageId="0" pageNumber="51" start="Figure 17" startId="F17">
<paragraph id="C3428D31D779D133374258A2B019A647" pageId="0" pageNumber="51">
<emphasis id="C6A022E3C75C65F726ABC1AB094BA983" bold="true" pageId="0" pageNumber="51">Figure 17.</emphasis>
Distribution map of examined material of
<taxonomicName id="64DDDDCAE43C469B35BA01BD054DD1E8" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis neglectana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="neglectana">
<emphasis id="972706611C35168CE3B125CADFD8E429" italics="true" pageId="0" pageNumber="51">Clepsis neglectana</emphasis>
</taxonomicName>
and
<taxonomicName id="4DDA8080CA28AB29D9A889EBABE8BFB9" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="BCB4C3D6B6D5BB8194267613A96ABF2D" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
species groups. Map created with SimpleMappr (http://www.simplemappr.net).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1ADEB71E14E16C4A7875BAB574F1DD34" pageId="0" pageNumber="51" type="ecology">
<paragraph id="78B1DF921D7EA3D79D9F5A135AB8F8BD" pageId="0" pageNumber="51">Ecology.</paragraph>
<paragraph id="AD3B5DD5DF9BFADF997C1C169C79B46D" pageId="0" pageNumber="51">
The moths were collected in macchie habitat (
<bibRefCitation id="862391C5B29EF6B61B29009097EE5A34" author="Gaston, J" journalOrPublisher="Shilap Revista de lepidopterologia" pageId="0" pageNumber="51" pagination="689 - 698" refId="B15" refString="Gaston, J, Vives Moreno, A, Revilla, Tx, 2017. Descripcion de tres especies nuevas de la Familia Tortricidae en la Peninsula Iberica (Lepidoptera: Tortricidae). Shilap Revista de lepidopterologia 45: 689 - 698" title="Descripcion de tres especies nuevas de la Familia Tortricidae en la Peninsula Iberica (Lepidoptera: Tortricidae)." volume="45" year="2017">
<normalizedToken id="A25341A6D32F3CD59EA0AC23C5872ABA" originalValue="Gastón">Gaston</normalizedToken>
et al. 2017
</bibRefCitation>
) from April to the first half of September. The larval host plant is unknown.
</paragraph>
</subSubSection>
<subSubSection id="4247BD1025F741DD1F1491DB4B0FDAC4" pageId="0" pageNumber="51" type="remarks">
<paragraph id="0F2B8AAE6844DC00DB9CF7C43E07C42A" pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph id="D20CBE6D32318A9E13A62D0DE624185B" pageId="0" pageNumber="51">
The species was described by
<bibRefCitation id="254A0C5B9A1B583750FA49E1F4D69527" author="Ragonot, EL" journalOrPublisher="The Entomologist's monthly magazine" pageId="0" pageNumber="51" pagination="229 - 233" refId="B31" refString="Ragonot, EL, 1881. Notes on the entomology of Portugal. IV. Lepidoptera (Continued). Micro-Lepidoptera (except Tineina) collected by the Rev. A. E. Eaton in 1880, with descriptions of new species. The Entomologist's monthly magazine 17: 229 - 233" title="Notes on the entomology of Portugal. IV. Lepidoptera (Continued). Micro-Lepidoptera (except Tineina) collected by the Rev. A. E. Eaton in 1880, with descriptions of new species." volume="17" year="1881">Ragonot (1881)</bibRefCitation>
after two males from Portugal. He emphasised its resemblance with
<taxonomicName id="22D02B764500B6025A9F785DBC7E07A3" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="171F871E13AB1114307FF8FE8830F44A" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
.
<bibRefCitation id="F6535D9C7714FF02758AA9CFA8314612" author="Razowski, J" journalOrPublisher="Acta zoologica cracoviensia" pageId="0" pageNumber="51" pagination="101 - 198" refId="B34" refString="Razowski, J, 1979. Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I. Acta zoologica cracoviensia 23: 101 - 198" title="Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I." volume="23" year="1979">Razowski (1979)</bibRefCitation>
considered it conspecific with
<taxonomicName id="4F0E4BDA9CE97B6CC977DE87156C584E" lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis id="865A870A028250D50D3AD1365504CC71" italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
, probably relying on the superficial similarity of the modified setae of the valvae of these two taxa. The female remained unstudied until
<bibRefCitation id="48260C93509F62A5812584CD15A4E9C2" author="Gaston, J" journalOrPublisher="Shilap Revista de lepidopterologia" pageId="0" pageNumber="51" pagination="689 - 698" refId="B15" refString="Gaston, J, Vives Moreno, A, Revilla, Tx, 2017. Descripcion de tres especies nuevas de la Familia Tortricidae en la Peninsula Iberica (Lepidoptera: Tortricidae). Shilap Revista de lepidopterologia 45: 689 - 698" title="Descripcion de tres especies nuevas de la Familia Tortricidae en la Peninsula Iberica (Lepidoptera: Tortricidae)." volume="45" year="2017">
<normalizedToken id="CDE25CBDCD9E91AE192426888557026F" originalValue="Gastón">Gaston</normalizedToken>
et al. (2017)
</bibRefCitation>
described it again under the name
<taxonomicName id="A4B660826A7EF85A958028E73319BD31" lsidName="C. razowskii" pageId="0" pageNumber="51" rank="species" species="razowskii">
<emphasis id="F6C4B2CB0314FF66BE286A69084E8F20" italics="true" pageId="0" pageNumber="51">C. razowskii</emphasis>
</taxonomicName>
.
<taxonomicName id="59D95FB60F594D68648F5BCAA76B1B06" class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix xylotoma" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="xylotoma">
<emphasis id="D802013F19854C1C385D435343EE5AE4" italics="true" pageId="0" pageNumber="51">Tortrix xylotoma</emphasis>
</taxonomicName>
was sunk into
<taxonomicName id="61EC77E633587274E0C902342B67387F" lsidName="C. neglectana" pageId="0" pageNumber="51" rank="species" species="neglectana">
<emphasis id="BCA7A662DBD813E9EC4A091BEBBDF67A" italics="true" pageId="0" pageNumber="51">C. neglectana</emphasis>
</taxonomicName>
by
<bibRefCitation id="43111F72E14EFCD6654BB4CB6851C268" author="Razowski, J" journalOrPublisher="Acta zoologica cracoviensia" pageId="0" pageNumber="51" pagination="101 - 198" refId="B34" refString="Razowski, J, 1979. Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I. Acta zoologica cracoviensia 23: 101 - 198" title="Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I." volume="23" year="1979">Razowski (1979)</bibRefCitation>
. This synonym is not justified as the genitalia of the male lectotype (illustrated also by
<bibRefCitation id="B7BE688927545A8B03F3B5FAE6323F14" author="Clarke, JFG" journalOrPublisher="Trustees of the British Museum (N H), London" pageId="0" pageNumber="51" refId="B7" refString="Clarke, JFG, 1958. Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick 3. Trustees of the British Museum (N H), London" title="Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick 3." year="1958">Clarke 1958</bibRefCitation>
: pl. 109) rather resemble
<taxonomicName id="E1BD26F8EE60ECF6A21DC179890F29FC" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="C694795B2FBCF1D72F9B0E9096D6D7EF" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
regarding the shape of labis, presence of modified setae on the valva, and morphology of the phallus. The preparation of the genitalia is poor, therefore the shape of uncus and valvae looks unusual. The wing pattern fits with
<taxonomicName id="3302A0B14C30467B00334B884642850E" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="377A77F3F97460180EC2A84567632FD3" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
. After comparison of material from Spain, the lectotype of
<taxonomicName id="242B664587B56AF8B1CE7A6A8D0A99D4" lsidName="T. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="786028994ACEDEAE2D94BC5A01381AB3" italics="true" pageId="0" pageNumber="51">T. eatoniana</emphasis>
</taxonomicName>
, the lectotype of
<taxonomicName id="9D8A6834D133FFDDCA9F21348A18A2FA" lsidName="T. xylotoma" pageId="0" pageNumber="51" rank="species" species="xylotoma">
<emphasis id="5F28F531F711C930331D8F25367A0B0D" italics="true" pageId="0" pageNumber="51">T. xylotoma</emphasis>
</taxonomicName>
, the original description of
<taxonomicName id="3AAA895664090A9D5A071D6B34502EDF" lsidName="C. razowskii" pageId="0" pageNumber="51" rank="species" species="razowskii">
<emphasis id="36355C191795539E9C9E9E6D666AF405" italics="true" pageId="0" pageNumber="51">C. razowskii</emphasis>
</taxonomicName>
, and numerous photographs of specimens identified as
<taxonomicName id="EDB9A269046EC84C93AFC935A4AA96CC" lsidName="C. razowskii" pageId="0" pageNumber="51" rank="species" species="razowskii">
<emphasis id="6B9C747ADB42B0B6EDE3E05683113B69" italics="true" pageId="0" pageNumber="51">C. razowskii</emphasis>
</taxonomicName>
by the authors of the latter taxon, we concluded that all these specimens are conspecific and the present valid name of the species should be
<taxonomicName id="43FD129586345A32A91BCA7E07139B31" lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis id="C1545F8F3F146D507294BF4A8FDC30A0" italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>