242 lines
32 KiB
XML
242 lines
32 KiB
XML
<document ID-CLB-Dataset="55380" ID-DOI="http://dx.doi.org/10.3897/zookeys.1176.104980" ID-GBIF-Dataset="9ab98cc2-dd43-49f3-97bf-5aca4e7b3e61" ID-PMC="PMC10466110" ID-Pensoft-Pub="1313-2970-1176-13" ID-Pensoft-UUID="C178263C62205EE685739607736AAB36" ID-PubMed="37654984" ID-ZooBank="5C628D2388934955B260F4052F4D8193" ModsDocID="1313-2970-1176-13" checkinTime="1692723655473" checkinUser="pensoft" docAuthor="Miller, Kelly B., Benetti, Cesar J. & Michat, Mariano C." docDate="2023" docId="B983A1366A1456A08AFB68A7C53FB4A3" docLanguage="en" docName="ZooKeys 1176: 13-28" docOrigin="ZooKeys 1176" docPubDate="2023-08-22" docSource="http://dx.doi.org/10.3897/zookeys.1176.104980" docTitle="Miradessus rikae Miller, Benetti & Michat 2023, sp. nov." docType="treatment" docUuid="DE04E790-A93F-4ED8-8D63-4B6BA7653AE3" docUuidSource="ZooBank" docVersion="4" id="C178263C62205EE685739607736AAB36" lastPageNumber="13" masterDocId="C178263C62205EE685739607736AAB36" masterDocTitle="Miradessus gen. nov. from South America described for two species previously in Amarodytes Regimbart, 1900 and two new species (Arthropoda, Insecta, Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Bidessini)" masterLastPageNumber="28" masterPageNumber="13" pageNumber="13" updateTime="1732710388969" updateUser="ExternalLinkService">
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<mods:title id="0D24656C3B7774C58D8637E542651986">Miradessus gen. nov. from South America described for two species previously in Amarodytes Regimbart, 1900 and two new species (Arthropoda, Insecta, Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Bidessini)</mods:title>
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<mods:name id="D6600FCDD0ABA1A2C6D08A7ED95DA051" type="personal">
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<mods:roleTerm id="8DB0512126EF0E13430EF92C077EAF0D">Author</mods:roleTerm>
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<mods:namePart id="7886BC102BE8D3936F51472A50DF533B">Miller, Kelly B.</mods:namePart>
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<mods:affiliation id="83BE9F9A1DD562FCC313701F23EA0954">Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
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<mods:nameIdentifier id="9320CBA40F84C67A873A74EB6514A273" type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:name id="B3A58916F214D3AC8FD59392AF97D680" type="personal">
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<mods:roleTerm id="42B4E992B9716FD3DB56B5144AE9BB28">Author</mods:roleTerm>
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<mods:namePart id="DEEDE20FF3F001B9215A362DC504A44A">Benetti, Cesar J.</mods:namePart>
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<mods:nameIdentifier id="3D9AF3C4D6E0F065B73AA3D9CFE2A13E" type="ORCID">https://orcid.org/0000-0001-6795-2194</mods:nameIdentifier>
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<mods:affiliation id="D868345611169D3100EE8496D60749E2">Departamento de Biodiversidad y Gestion Ambiental, Facultad de Ciencias Biologicas y Ambientales, Universidad de Leon, Campus de Vegazana, 24071, Leon, Spain & Coordenacao de Biodiversidade, Programa de Pos-Graduacao em Entomologia (PPGEnt), Instituto Nacional de Pesquisas da Amazonia (INPA), Av. Andre Araujo 2936, CEP 69067 - 375, Manaus, AM, Brazil</mods:affiliation>
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<mods:namePart id="31130AECFD4AD6F8F9836EEB5787E27A">Michat, Mariano C.</mods:namePart>
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<mods:nameIdentifier id="843F2DB147A73AA346413240BC57F85A" type="ORCID">https://orcid.org/0000-0002-1962-7976</mods:nameIdentifier>
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<mods:affiliation id="0421B1B2B83088EEBC24DD2EF1EE1054">Faculty of Exact and Natural Sciences, Department of Biodiversity and Experimental Biology, Laboratory of Entomology, Institute of Biodiversity and Experimental and Applied Biology (IBBEA), CONICET-University of Buenos Aires, Buenos Aires, Argentina</mods:affiliation>
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<mods:typeOfResource id="035A01F0A32C94B016B3C433BF686704">text</mods:typeOfResource>
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<mods:title id="D2F599623AA411EB552BC0E8FECACBFB">ZooKeys</mods:title>
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<mods:date id="C0E978EF5C53F1D0F38D8E0431BFA97B">2023</mods:date>
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<mods:number id="A1FBDAA17BF7CBF5BA20679B91EB1344">2023-08-22</mods:number>
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<mods:number id="E083BD84BF1A5F6DF9CB0D8CFB2102ED">1176</mods:number>
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<mods:url id="3D1854A68DDE7FE5B917F12316F798F4">http://dx.doi.org/10.3897/zookeys.1176.104980</mods:url>
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<mods:classification id="CE972C9F470799A1737396338BAFBBB7">journal article</mods:classification>
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<mods:identifier id="5CF381874A7BFE2692CFBC6D38C4A77C" type="DOI">http://dx.doi.org/10.3897/zookeys.1176.104980</mods:identifier>
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<treatment id="B983A1366A1456A08AFB68A7C53FB4A3" ID-GBIF-Taxon="212450752" LSID="urn:lsid:zoobank.org:act:DE04E790-A93F-4ED8-8D63-4B6BA7653AE3" httpUri="http://treatment.plazi.org/id/B983A1366A1456A08AFB68A7C53FB4A3" lastPageNumber="13" pageId="0" pageNumber="13" scope_class="Insecta" scope_family="Dytiscidae" scope_order="Coleoptera" scope_phylum="Arthropoda" scope_subFamily="Hydroporinae" scope_tribe="Bidessini">
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<subSubSection id="0A6CCCE7F98E66C9F2CDE47D782DEBDF" pageId="0" pageNumber="13" type="nomenclature">
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<paragraph id="FB69C5444336D56E569069F378AC9C0F" pageId="0" pageNumber="13">
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<taxonomicName id="AA8E047DEB8EBEBBCBA68690D6999E26" LSID="https://zoobank.org/DE04E790-A93F-4ED8-8D63-4B6BA7653AE3" authority="Miller, Benetti & Michat" authorityName="Miller, Benetti & Michat" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Miradessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miradessus rikae" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="rikae" status="sp. nov.">Miradessus rikae Miller, Benetti & Michat</taxonomicName>
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<taxonomicNameLabel id="4C933D28E5ECB87DE035F3F8DDC5EE55" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="A307249A7A1C0BCC78B2C7317CDA6A5C" pageId="0" pageNumber="13" type="description">
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<paragraph id="7FCDB640BFC52241BEE6CC694F41B460" pageId="0" pageNumber="13">
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<figureCitation id="E31429A5FB8C8F5BF1139077F1D7CC17" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">Figs 4</figureCitation>
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<figureCitation id="802C306F18DC43451BE641F284862158" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">, 8</figureCitation>
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<figureCitation id="203979306BF3BD13445A845DA7611F86" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">, 15</figureCitation>
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<figureCitation id="A0C1BC902501C99807A3B347F2B390A7" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">, 16</figureCitation>
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<figureCitation id="F11907BD9545EEF9BD9319529ADBEED1" captionStart="Figures 18–29" captionStartId="F3" captionText="Figures 18 - 29. Miradessus species, primary type specimens and labels 18 - 20 M. beni, holotype specimen 18 dorsal habitus 19 ventral habitus 20 specimen labels 21 - 23 M. plaumanni, lectotype specimen 21 dorsal habitus 22 ventral habitus 23 specimen labels 24 - 26 M. pulchellus, holotype specimen 24 dorsal habitus 25 ventral habitus 26 specimen labels 27 - 29 M. rikae, holotype specimen 27 dorsal habitus 28 ventral habitus 29 specimen labels. 21 - 23 courtesy of M. Schwarz (OLML) 24 - 26 courtesy of M. A. Balke (ZSMG)." figureDoi="10.3897/zookeys.1176.104980.figures18-29" httpUri="https://binary.pensoft.net/fig/895236" pageId="0" pageNumber="13">, 27-29</figureCitation>
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<figureCitation id="00A0F80E856EE32D5DAFEFEDF8AEDF19" captionStart="Figure 30" captionStartId="F4" captionText="Figure 30. Miradessus species, distributions." figureDoi="10.3897/zookeys.1176.104980.figure30" httpUri="https://binary.pensoft.net/fig/895237" pageId="0" pageNumber="13">, 30</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="1E4F575D2BA66EC98FFBDB39E7209443" pageId="0" pageNumber="13" type="type locality">
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<paragraph id="CD186360459EBED6633A38222DB220BB" pageId="0" pageNumber="13">Type locality.</paragraph>
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<paragraph id="3EE23BA95320DC06F48197DAEB4B66AF" pageId="0" pageNumber="13">Ecuador, Pastaza, Puyo.</paragraph>
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</subSubSection>
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||
<subSubSection id="5E0F0B1E2E10D88A2E2D3136F971A439" pageId="0" pageNumber="13" type="diagnosis">
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<paragraph id="8BE4830A44D14955A6B4E5A48C1F910B" pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph id="004A27158C79CA9D91E7B4B9E697A1F6" pageId="0" pageNumber="13">
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This species is shorter and more robust (Fig.
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<figureCitation id="05A7BAD2D85BA2CEA9891E85D9F24E45" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">4</figureCitation>
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) than either
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<taxonomicName id="9B9C6B288A9A8235150AFBD34415FE4A" lsidName="M. pulchellus" pageId="0" pageNumber="13" rank="species" species="pulchellus">
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<emphasis id="21DBDAA4A4B41262EC5B023CE65821D7" italics="true" pageId="0" pageNumber="13">M. pulchellus</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4F2E12A015D50194480A1D12A0C2C5E3" lsidName="M. beni" pageId="0" pageNumber="13" rank="species" species="beni">
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<emphasis id="C346C38B2BCCB37AEB7F7C12977E9EEC" italics="true" pageId="0" pageNumber="13">M. beni</emphasis>
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</taxonomicName>
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(Figs
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<figureCitation id="EBAF6ABD4962DCFB79879826CFC9839A" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">1</figureCitation>
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,
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<figureCitation id="41EA15A4685918363DE8B0C4DE62BDBB" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">3</figureCitation>
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) although the dorsal color pattern is similar to them (Fig.
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<figureCitation id="6F99E8835878654D31FBCC774D3B6033" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">4</figureCitation>
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). The male genitalia are diagnostic. The median lobe in
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<taxonomicName id="03D72E3E4B221624BE6B58FDB0185172" lsidName="M. rikae" pageId="0" pageNumber="13" rank="species" species="rikae">
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<emphasis id="03358D4C1A9659106F9FFF1931B6FA74" italics="true" pageId="0" pageNumber="13">M. rikae</emphasis>
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</taxonomicName>
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is trilobed like other
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<taxonomicName id="8A202B443B6FD0ECF492D7EEAB7488BE" authorityName="Miller, Benetti & Michat" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Miradessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miradessus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="3259A763D5F6E30B65953FC5E4371AB7" italics="true" pageId="0" pageNumber="13">Miradessus</emphasis>
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</taxonomicName>
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, but the median portion is uniquely short, broad, and apically broadly subtruncate with the ventral portions elongate, slender and apically narrowly rounded (Fig.
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<figureCitation id="E79D9CEF694286B8DC6CB0F066F46F63" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">15</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="A1CFB119B4444E96C18298AF1AB44B6D" pageId="0" pageNumber="13" type="description">
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<paragraph id="B1D35952C7798BAE7A30B7D67C46FAD0" pageId="0" pageNumber="13">Description.</paragraph>
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<paragraph id="BEBD95F306A4C8A3D9869DEB17F79EFE" pageId="0" pageNumber="13">
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<emphasis id="962B3D7FB81299F2FA5297FEE1E0165B" bold="true" pageId="0" pageNumber="13">
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<emphasis id="1029E5867D6A42D35A98BC59384FCE2E" italics="true" pageId="0" pageNumber="13">Measurements</emphasis>
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.
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</emphasis>
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TL = 2.2 mm, GW = 1.2 mm, PW = 0.9 mm, HW = 0.6 mm, EW = 0.4 mm, TL/GW = 1.8, HW/EW = 1.5, FL/RL = 2.0.
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</paragraph>
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<paragraph id="CB603A5496490FD83BAC7CDB9A860946" pageId="0" pageNumber="13">
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<emphasis id="66D66E9048D2C70EBE59A0D4DF0E0BA6" bold="true" pageId="0" pageNumber="13">
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<emphasis id="84C2B5247C7A1872FCB1243C29029AB8" italics="true" pageId="0" pageNumber="13">Habitus</emphasis>
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.
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</emphasis>
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Body shape elongate-oval, lateral outline discontinuous between pronotum and elytron, posterior apex narrowly rounded (Fig.
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<figureCitation id="1037E9050BA4ACFB356E7F9E9806E8DC" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">4</figureCitation>
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).
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</paragraph>
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<paragraph id="F59D9FB390D44D8EF700491FB2ED5258" pageId="0" pageNumber="13">
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<emphasis id="BC4BE4977CA79E991AABF45CD92648ED" bold="true" italics="true" pageId="0" pageNumber="13">Coloration</emphasis>
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(Fig.
|
||
<figureCitation id="C50EA20D7C5035C9BE1D262DB8A807B4" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">4</figureCitation>
|
||
). Head yellow-orange. Pronotum yellow-orange with a narrow, rounded lobe of black broadly along each side of posterior margin. Elytron maculate, yellow-orange and black, margins of maculae strongly demarcated; with large, transverse yellow-orange regions anteriorly, medially and apically, not extending medially to suture, with narrow band of black along entire length of suture, margins of maculae irregular and variously lobed. Ventral surfaces mostly orange; legs, epipleuron, and lateral areas of prothorax and head lighter orange-yellow.
|
||
</paragraph>
|
||
<paragraph id="980E04799B13545AD8197E7C9F294A1D" pageId="0" pageNumber="13">
|
||
<emphasis id="88A2347037B457D448F65CD7C0BECA0E" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="448F82CD5D0EFDA36FBB549BB9C697C8" italics="true" pageId="0" pageNumber="13">Sculpture and structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head shiny and smooth, nearly impunctate, with small micropunctures sparsely distributed; eyes large (HW/EW = 1.5); antennae slender, unmodified. Pronotum with lateral margins moderately curved; with narrow bead along entire margin; surface shiny, moderately and evenly punctate; lateral pronotal plica distinctly impressed, sublinear, extending about 1/3 distance across pronotum. Elytron with lateral margin strongly curved anteriorly in dorsal aspect, evenly curved to apex; surface shiny, moderately and evenly punctate. Prosternum medially moderately broad, medially not protruberant, mediolaterally somewhat granular; prosternal process moderately broad medially with slight tubercle, apical blade large, laterally with prominent rounded ridges along entire length, medially longitudinally sulcate, lateral margins slightly convergent to narrowly rounded apex. Metaventrite and metaventral wings smooth and shiny, covered with shallow punctures (Fig.
|
||
<figureCitation id="C706870D31C4B3E0BB6673FF49C3AC54" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">8</figureCitation>
|
||
). Metacoxa with medial portion moderately broad, metacoxal lines distinct, evenly divergent anteriorly to posterior margin of metaventrite; lateral portion large, medially and anteriorly covered with small, shallow punctures; metatrochanter about 1/3 length of metafemur (Fig.
|
||
<figureCitation id="2622FC5F3E0DE8CFEE3EC5405F0E40DE" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">8</figureCitation>
|
||
). Abdominal ventrites nearly impunctate except II and III laterally with shallow, indistinct punctures; VI with apex pointed.
|
||
</paragraph>
|
||
<paragraph id="348725474BC60953454BD98003F1C40B" pageId="0" pageNumber="13">
|
||
<emphasis id="7DB4C74F200ED02AB08BEA44CF9C03D9" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="B1BEE14326191B397AD203D0999AA9C5" italics="true" pageId="0" pageNumber="13">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Median lobe in ventral aspect conspicuously trilobed, with medial portion short and broad, apically expanded and subtruncate, apicolateral angles with short setae, ventral portions elongate, slender, apically narrowly rounded (Fig.
|
||
<figureCitation id="AB7837F1C2EE373CFA6EF7E392C55655" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">15</figureCitation>
|
||
); lateral lobe in lateral aspect robust, basal segment somewhat broad and robust, apical segment broad basally, with deep emargination along dorsal margin, apex rounded with series of setae (Fig.
|
||
<figureCitation id="DA4BB6647238758F25F48D7892E68038" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF476815FC73EAF632DF7FA25E71BAA0" pageId="0" pageNumber="13">
|
||
<emphasis id="DB1D84990F04698CB5D662A2E4142D88" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="A1209DA33F16234E16C173C5F604DA00" italics="true" pageId="0" pageNumber="13">Sexual dimorphism</emphasis>
|
||
.
|
||
</emphasis>
|
||
Only males are known.
|
||
</paragraph>
|
||
<paragraph id="C16C8186084CE5D88D17ADBBCF42BBC4" pageId="0" pageNumber="13">
|
||
<emphasis id="E2D7CD83A8908B5642F4385EC942F8A8" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="B3547C7BD2A30FBAAFF2702B02B1159C" italics="true" pageId="0" pageNumber="13">Variation</emphasis>
|
||
.
|
||
</emphasis>
|
||
The two specimens exhibit slight variation in the shape and extent of maculation on the dorsal surface, but they are otherwise similar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8BDCF7EB112EB4BFB6D14EE75C3C9866" pageId="0" pageNumber="13" type="etymology">
|
||
<paragraph id="C31152415C73C23B9357F41C1C999442" pageId="0" pageNumber="13">Etymology.</paragraph>
|
||
<paragraph id="F02B5AEBD4E8BBD83013697DC13F5F06" pageId="0" pageNumber="13">
|
||
This species is named
|
||
<emphasis id="4DE22DB8CD5358360DD68458FD78138B" italics="true" pageId="0" pageNumber="13">rikae</emphasis>
|
||
after Ms Rikelle Timpe, close friend of the first author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E363B85E4F3820FC8113F2A2ADCA9563" pageId="0" pageNumber="13" type="distribution">
|
||
<paragraph id="178E72510CA7666BAB79173E91E58135" pageId="0" pageNumber="13">Distribution.</paragraph>
|
||
<paragraph id="D8FC2C2942C01C754277569CF381D42F" pageId="0" pageNumber="13">
|
||
<taxonomicName id="C2A44FC8A5A2F1D4813D9B65662DB371" authorityName="Miller, Benetti & Michat" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Miradessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miradessus rikae" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="rikae">
|
||
<emphasis id="93926D52687F4A70E581D9C4FCED1C72" italics="true" pageId="0" pageNumber="13">Miradessus rikae</emphasis>
|
||
</taxonomicName>
|
||
is known from two sites in Ecuador (Fig.
|
||
<figureCitation id="9D810C6382728B3E8765BD301D00A60D" captionStart="Figure 30" captionStartId="F4" captionText="Figure 30. Miradessus species, distributions." figureDoi="10.3897/zookeys.1176.104980.figure30" httpUri="https://binary.pensoft.net/fig/895237" pageId="0" pageNumber="13">30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E1059B432757C91D33AC3D6D1334C7F3" pageId="0" pageNumber="13" type="habitat">
|
||
<paragraph id="2654FB5C33A6DFC7C3D8B149E0CB6C68" pageId="0" pageNumber="13">Habitat.</paragraph>
|
||
<paragraph id="BB5A68E51A4094F346F70EA37FDFDF1E" pageId="0" pageNumber="13">The two known specimens were collected at blacklights, so nothing is known of the specific habitat. The two collection localities are in forested regions of lowland Ecuador.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B91E58EE272CBA48808C01FFA10130C0" pageId="0" pageNumber="13" type="materials_examined">
|
||
<paragraph id="54A8E6588ED24161E9BF4E7F55339C7D" pageId="0" pageNumber="13">Material examined.</paragraph>
|
||
<paragraph id="24BB03D5B2E400DC0AEC035774439732" pageId="0" pageNumber="13">
|
||
<emphasis id="41FAF739D362F752A92AB6C3C8F0F9A2" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="952D13BE1A31CC6F55B13BB308929EA0" italics="true" pageId="0" pageNumber="13">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="AA856BF35E34A1F4AA1019E8C9709CD9" collectingDate="1975-09-26" collectingDateMax="1976-02-07" collectingDateMin="1975-09-26" collectorName="Both" country="ECUADOR" location="Figs" specimenCount="1" stateProvince="NAPO" typeStatus="Holotype">
|
||
<emphasis id="9DAD0213702E6B88073CF4A4E0335D22" italics="true" pageId="0" pageNumber="13">
|
||
<typeStatus id="D3B2A86CEAF273A1A3FE8D96238986C4">Holotype</typeStatus>
|
||
</emphasis>
|
||
male (
|
||
<location id="62B38D2B0D03093C96853780C58A1257" LSID="urn:lsid:plazi:treatment:B983A1366A1456A08AFB68A7C53FB4A3:62B38D2B0D03093C96853780C58A1257" country="ECUADOR" name="Figs" stateProvince="NAPO">Figs</location>
|
||
<figureCitation id="783C2656D2B8D66587D3FB67ED4B7D98" captionStart="Figures 18–29" captionStartId="F3" captionText="Figures 18 - 29. Miradessus species, primary type specimens and labels 18 - 20 M. beni, holotype specimen 18 dorsal habitus 19 ventral habitus 20 specimen labels 21 - 23 M. plaumanni, lectotype specimen 21 dorsal habitus 22 ventral habitus 23 specimen labels 24 - 26 M. pulchellus, holotype specimen 24 dorsal habitus 25 ventral habitus 26 specimen labels 27 - 29 M. rikae, holotype specimen 27 dorsal habitus 28 ventral habitus 29 specimen labels. 21 - 23 courtesy of M. Schwarz (OLML) 24 - 26 courtesy of M. A. Balke (ZSMG)." figureDoi="10.3897/zookeys.1176.104980.figures18-29" httpUri="https://binary.pensoft.net/fig/895236" pageId="0" pageNumber="13">27-29</figureCitation>
|
||
) in USNM labeled, "
|
||
<collectingCountry id="4C8F7064068E175799DAAFCC1BE37370" name="Ecuador">ECUADOR</collectingCountry>
|
||
,Past. Puyo.
|
||
<collectingDate id="0B633AA6B81D2BE729927794B6EA00AC" value="1976-02-01" valueMax="1976-02-07" valueMin="1976-02-01">1-7 Feb. 1976</collectingDate>
|
||
at blacklight Spangler,et al./ ECUADOR-PEACE CORPS- SMITHSONIAN INSTITUTION AQUATIC INSECT SURVEY/
|
||
<typeStatus id="16BC9F12D6B51513B8D395D86F98ADC4">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="17C75B3DB15CE33A776C3C5F8597D77E" authorityName="Miller, Benetti & Michat" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Miradessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miradessus rikae" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="rikae">
|
||
<emphasis id="F4C5DD41F11BBA1DAB97D2CA4C90A8FD" italics="true" pageId="0" pageNumber="13">Miradessus rikae</emphasis>
|
||
</taxonomicName>
|
||
Miller, Benetti & Michat, 2023 [red label with double black line border]." One
|
||
<typeStatus id="2DED5D81307EBA12B0AB1476D52C3A25">paratype</typeStatus>
|
||
male in USNM labeled "
|
||
<collectingCountry id="B2995EB35773A7FAAB257DE305C66E71" name="Ecuador">ECUADOR</collectingCountry>
|
||
,
|
||
<collectingRegion id="9D101C5B601530E935324F32DF296116" country="Ecuador" name="Napo">NAPO</collectingRegion>
|
||
, Lago Agrio(5 Km N)
|
||
<collectingDate id="FAED4D0E4612CFC1DFAE2842F3264ECC" value="1975-09-26">26 Sept.1975</collectingDate>
|
||
at blacklight Andrea Langley/
|
||
<typeStatus id="5C4AB3DC9EDC5FD44567AE961DC203AE">PARATYPE</typeStatus>
|
||
<taxonomicName id="5D5F0B5380B1213FEF4D5829532A39F3" authorityName="Miller, Benetti & Michat" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Miradessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miradessus rikae" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="rikae">
|
||
<emphasis id="52A08C928E0D59D74666F4AFB3C70136" italics="true" pageId="0" pageNumber="13">Miradessus rikae</emphasis>
|
||
</taxonomicName>
|
||
Miller, Benetti & Michat, 2023 [blue label with black line border]."
|
||
<collectorName id="1D7C6E790F39149FC79B3211A3D29A3C">Both</collectorName>
|
||
the
|
||
<typeStatus id="C8BF5F00B018004DFE5138E75FE2172B">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="AB00E9760EA5A3E8BD532F0885A986E1">paratype</typeStatus>
|
||
have the male genitalia dissected and placed in microvials mounted on the pins.
|
||
</materialsCitation>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |