treatments-xml/data/B1/11/6A/B1116A44190EAF48E996FD5CFB91FE6C.xml

<document id="C579CCFE8142D3E28416D9950D3D5001" ID="10.11646/zootaxa.4174.1.10" ID-CLB-Dataset="38056" ID-DOI="10.11646/zootaxa.4174.1.10" ID-GBIF-Dataset="5718305a-3088-49ca-8424-8af30a99c614" ID-ISSN="1175-5326" ID-Zenodo-Dep="160032" ID-ZooBank="E2DFA756-B6B2-44F3-BC03-40A08CB89646" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476175700238" checkinUser="plazi" docAuthor="Irigoitia, Manuel M., Cantatore, Delfina M. P., Incorvaia, Inés S. &amp; Timi, Juan T." docDate="2016" docId="B1116A44190EAF48E996FD5CFB91FE6C" docLanguage="en" docName="zootaxa.4174.1.10.pdf" docOrigin="Zootaxa 4174 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kroeyerina sudamericana Irigoitia, Cantatore, Incorvaia &amp; Timi, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="147" masterDocId="4D28123C190CAF42E901FFF6FFE5FF99" masterDocTitle="Parasitic copepods infesting the olfactory sacs of skates from the southwestern Atlantic with the description of a new species of Kroeyerina Wilson, 1932" masterLastPageNumber="152" masterPageNumber="137" pageNumber="139" updateTime="1698429532103" updateUser="ExternalLinkService">
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<mods:title id="C51D2E870E5F366C86A0995146BEB2CE">Parasitic copepods infesting the olfactory sacs of skates from the southwestern Atlantic with the description of a new species of Kroeyerina Wilson, 1932</mods:title>
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<mods:namePart id="3F785887CEDF877B01FCD72C25299F48">Irigoitia, Manuel M.</mods:namePart>
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<mods:namePart id="59B358E569E919370BF63C8265656C00">Cantatore, Delfina M. P.</mods:namePart>
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<mods:namePart id="995261BD6A9D021E545A62511FA0DF80">Incorvaia, Inés S.</mods:namePart>
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<mods:namePart id="A558242568F9B930F8530EF764CCBC15">Timi, Juan T.</mods:namePart>
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<mods:date id="ED568C2F9445654EC282A36936C30337">2016</mods:date>
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<mods:number id="E4658A0D6AAD1772729715229AF4505F">4174</mods:number>
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<treatment id="B1116A44190EAF48E996FD5CFB91FE6C" ID-DOI="http://doi.org/10.5281/zenodo.5662704" ID-GBIF-Taxon="124997228" ID-Zenodo-Dep="5662704" LSID="urn:lsid:plazi:treatment:B1116A44190EAF48E996FD5CFB91FE6C" httpUri="http://treatment.plazi.org/id/B1116A44190EAF48E996FD5CFB91FE6C" lastPageId="10" lastPageNumber="147" pageId="2" pageNumber="139">
<subSubSection id="71A288D9190EAF40E996FD5CFEFAFD7F" pageId="2" pageNumber="139" type="nomenclature">
<paragraph id="3907DB52190EAF40E996FD5CFDD0FD5D" blockId="2.[151,565,682,742]" box="[151,565,682,708]" pageId="2" pageNumber="139">
<heading id="624F6C3E190EAF40E996FD5CFDD0FD5D" bold="true" box="[151,565,682,708]" fontSize="11" level="1" pageId="2" pageNumber="139" reason="1">
<emphasis id="0BCC0740190EAF40E996FD5CFDD0FD5D" bold="true" box="[151,565,682,708]" pageId="2" pageNumber="139">
<taxonomicName id="FEB8A0D1190EAF40E996FD5CFE34FD5D" box="[151,465,682,708]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC0740190EAF40E996FD5CFE34FD5D" bold="true" box="[151,465,682,708]" italics="true" pageId="2" pageNumber="139">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10FFBA3B190EAF40E8D6FD5CFDD0FD5D" box="[471,565,682,708]" pageId="2" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3907DB52190EAF40E996FD3BFEFAFD7F" blockId="2.[151,565,682,742]" box="[151,287,717,742]" pageId="2" pageNumber="139">
(
<figureCitation id="A183C7D7190EAF40E99EFD3BFEF2FD7F" box="[159,279,717,742]" captionStart-0="FIGURES 1 – 11" captionStart-1="FIGURES 12 – 15" captionStart-2="FIGURES 16 – 27" captionStart-3="FIGURES 28 – 31" captionStart-4="FIGURES 32 – 39" captionStart-5="FIGURES 40 – 47" captionStartId-0="3.[151,264,1924,1947]" captionStartId-1="4.[151,264,1571,1594]" captionStartId-2="5.[151,264,1909,1932]" captionStartId-3="6.[151,264,1784,1807]" captionStartId-4="7.[151,264,1862,1885]" captionStartId-5="8.[151,264,1868,1891]" captionTargetBox-0="[157,1436,198,1894]" captionTargetBox-1="[162,1430,198,1544]" captionTargetBox-2="[165,1435,201,1879]" captionTargetBox-3="[157,1419,204,1753]" captionTargetBox-4="[161,1424,202,1831]" captionTargetBox-5="[156,1432,204,1836]" captionTargetId-0="figure@3.[151,1436,193,1903]" captionTargetId-1="figure@4.[151,1435,193,1550]" captionTargetId-2="figure@5.[151,1435,193,1888]" captionTargetId-3="figure@6.[150,1436,193,1764]" captionTargetId-4="figure@7.[150,1436,193,1842]" captionTargetId-5="figure@8.[151,1436,193,1848]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." captionText-1="FIGURES 12 – 15. Kroeyerina sudamericana sp. nov. (adult female). 12, first leg; 13, second leg; 14, third leg; 15, fourth leg. Scale bars: 100 µm." captionText-2="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." captionText-3="FIGURES 28 – 31. Kroeyerina sudamericana sp. nov. (adult male). 28, first leg; 29, second leg; 30, third leg; 31, fourth leg. Scale bars: 100 µm." captionText-4="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." captionText-5="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri-0="https://zenodo.org/record/160034/files/figure.png" httpUri-1="https://zenodo.org/record/160035/files/figure.png" httpUri-2="https://zenodo.org/record/160036/files/figure.png" httpUri-3="https://zenodo.org/record/160037/files/figure.png" httpUri-4="https://zenodo.org/record/256586/files/figure.png" httpUri-5="https://zenodo.org/record/256587/files/figure.png" pageId="2" pageNumber="139">Figs. 1–47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="71A288D9190EAF40E996FCE3FCD1FCEF" pageId="2" pageNumber="139" type="materials_examined">
<paragraph id="3907DB52190EAF40E996FCE3FBD4FCB7" blockId="2.[151,1436,789,2002]" box="[151,1073,789,814]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E996FCE3FEF4FCB7" bold="true" box="[151,273,789,814]" pageId="2" pageNumber="139">Type host.</emphasis>
Spotback skate 
<taxonomicName id="FEB8A0D1190EAF40E8C9FCE3FD26FCB4" box="[456,707,789,813]" class="Elasmobranchii" family="Arhynchobatidae" genus="Atlantoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="castelnaui">
<emphasis id="0BCC0740190EAF40E8C9FCE3FD26FCB4" box="[456,707,789,813]" italics="true" pageId="2" pageNumber="139">Atlantoraja castelnaui</emphasis>
</taxonomicName>
(
<taxonomicName id="FEB8A0D1190EAF40EBD3FCE3FCB0FCB7" box="[722,853,789,814]" class="Elasmobranchii" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="order">Rajiformes</taxonomicName>
: 
<taxonomicName id="FEB8A0D1190EAF40EA5EFCE3FBC3FCB7" box="[863,1062,789,814]" class="Elasmobranchii" family="Arhynchobatidae" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="family">Arhynchobatidae</taxonomicName>
).
</paragraph>
<paragraph id="3907DB52190EAF40E9C6FCCFFCD1FCEF" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FCCFFEBCFCCB" bold="true" box="[199,345,825,850]" pageId="2" pageNumber="139">Other hosts.</emphasis>
Smallnose fanskate 
<taxonomicName id="FEB8A0D1190EAF40EB3FFCCCFCA3FCC8" box="[574,838,825,849]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="bonapartii">
<emphasis id="0BCC0740190EAF40EB3FFCCCFCA3FCC8" box="[574,838,825,849]" italics="true" pageId="2" pageNumber="139">Sympterygia bonapartii</emphasis>
</taxonomicName>
, bignose fanskate 
<taxonomicName id="FEB8A0D1190EAF40ED17FCCCFB03FCC8" box="[1046,1254,826,849]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="acuta">
<emphasis id="0BCC0740190EAF40ED17FCCCFB03FCC8" box="[1046,1254,826,849]" italics="true" pageId="2" pageNumber="139">Sympterygia acuta</emphasis>
</taxonomicName>
and zipper sand skate 
<taxonomicName id="FEB8A0D1190EAF40E9D6FCABFE23FCEC" box="[215,454,861,885]" class="Elasmobranchii" family="Arhynchobatidae" genus="Psammobatis" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="extenta">
<emphasis id="0BCC0740190EAF40E9D6FCABFE23FCEC" box="[215,454,861,885]" italics="true" pageId="2" pageNumber="139">Psammobatis extenta</emphasis>
</taxonomicName>
(
<taxonomicName id="FEB8A0D1190EAF40E8D4FCABFDBDFCEF" box="[469,600,861,886]" class="Elasmobranchii" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="order">Rajiformes</taxonomicName>
: 
<taxonomicName id="FEB8A0D1190EAF40EB63FCABFCCCFCEF" box="[610,809,861,886]" class="Elasmobranchii" family="Arhynchobatidae" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="family">Arhynchobatidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="71A288D9190EAF40E9C6FC77FDDFFC00" box="[199,570,897,922]" pageId="2" pageNumber="139" type="description">
<paragraph id="3907DB52190EAF40E9C6FC77FDDFFC00" blockId="2.[151,1436,789,2002]" box="[199,570,897,922]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FC77FE6EFC03" bold="true" box="[199,395,897,922]" pageId="2" pageNumber="139">Attachment site.</emphasis>
Olfactory sacs.
</paragraph>
</subSubSection>
<subSubSection id="71A288D9190EAF40E9C6FC53FDDCFABF" pageId="2" pageNumber="139" type="materials_examined">
<paragraph id="3907DB52190EAF40E9C6FC53FB9DFC27" blockId="2.[151,1436,789,2002]" box="[199,1144,933,958]" pageId="2" pageNumber="139">
<materialsCitation id="89D0D10F190EAF40E9C6FC53FB9DFC27" ID-GBIF-Occurrence="1316538288" box="[199,1144,933,958]" country="Argentina" location="Waters" pageId="2" pageNumber="139" specimenCount="1" stateProvince="Buenos Aires province" typeStatus="holotype">
<emphasis id="0BCC0740190EAF40E9C6FC53FE80FC27" bold="true" box="[199,357,933,958]" pageId="2" pageNumber="139">
<typeStatus id="E60365F0190EAF40E9C6FC53FEE7FC27" box="[199,258,933,958]" pageId="2" pageNumber="139">Type</typeStatus>
locality.
</emphasis>
<location id="3C678D89190EAF40E86DFC50FE5CFC24" LSID="urn:lsid:plazi:treatment:B1116A44190EAF48E996FD5CFB91FE6C:3C678D89190EAF40E86DFC50FE5CFC24" box="[364,441,934,957]" country="Argentina" name="Waters" pageId="2" pageNumber="139" stateProvince="Buenos Aires province">Waters</location>
off 
<collectingRegion id="FB7C15B0190EAF40E8E6FC50FD0EFC27" box="[487,747,933,958]" country="Argentina" name="Buenos Aires" pageId="2" pageNumber="139">Buenos Aires province</collectingRegion>
, 
<collectingCountry id="41AF9BC2190EAF40EBF7FC53FC80FC27" box="[758,869,933,958]" name="Argentina" pageId="2" pageNumber="139">Argentina</collectingCountry>
(34°–41°S; 53°–62°W).
</materialsCitation>
</paragraph>
<paragraph id="3907DB52190EAF40E9C6FC3FFAF0FBB3" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FC3FFE57FC7B" bold="true" box="[199,434,969,994]" pageId="2" pageNumber="139">Material examined.</emphasis>
17 females and six males from 
<taxonomicName id="FEB8A0D1190EAF40EA23FC3CFC53FC78" box="[802,950,969,993]" class="Elasmobranchii" family="Arhynchobatidae" genus="Atlantoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="castelnaui">
<emphasis id="0BCC0740190EAF40EA23FC3CFCD6FC78" box="[802,819,970,993]" italics="true" pageId="2" pageNumber="139">A</emphasis>
. 
<emphasis id="0BCC0740190EAF40EA42FC3FFC53FC78" box="[835,950,969,993]" italics="true" pageId="2" pageNumber="139">castelnaui</emphasis>
</taxonomicName>
(collected 
<date id="4D06FD92190EAF40ED37FC3FFB09FC7B" box="[1078,1260,969,994]" pageId="2" pageNumber="139" value="2011-11">November 2011</date>
and 
<date id="4D06FD92190EAF40EC27FC3FFF36FB9F" pageId="2" pageNumber="139" value="2013-11">November 2013</date>
), 14 females and two males from 
<taxonomicName id="FEB8A0D1190EAF40EB5DFC18FD14FB9C" box="[604,753,1005,1029]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="bonapartii">
<emphasis id="0BCC0740190EAF40EB5DFC18FD8FFB9C" box="[604,618,1006,1029]" italics="true" pageId="2" pageNumber="139">S</emphasis>
. 
<emphasis id="0BCC0740190EAF40EB7AFC1BFD14FB9C" box="[635,753,1005,1029]" italics="true" pageId="2" pageNumber="139">bonapartii</emphasis>
</taxonomicName>
(collected from 
<date id="4D06FD92190EAF40EAB0FC18FBA8FB9F" box="[945,1101,1006,1030]" pageId="2" pageNumber="139" value="2010" valueMax="2013">2010 to 2013</date>
), 12 females and eight males from 
<taxonomicName id="FEB8A0D1190EAF40E9D5FBE4FECBFBB0" box="[212,302,1042,1065]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="acuta">
<emphasis id="0BCC0740190EAF40E9D5FBE4FECBFBB0" box="[212,302,1042,1065]" italics="true" pageId="2" pageNumber="139">S. acuta</emphasis>
</taxonomicName>
(Collected 
<date id="4D06FD92190EAF40E8B0FBE7FD8DFBB3" box="[433,616,1041,1066]" pageId="2" pageNumber="139" value="2014-11">November 2014</date>
) and one female from 
<emphasis id="0BCC0740190EAF40EA67FBE4FC35FBB0" box="[870,976,1042,1065]" italics="true" pageId="2" pageNumber="139">P. ex t en t a</emphasis>
(collected 
<date id="4D06FD92190EAF40ED4EFBE7FAECFBB3" box="[1103,1289,1041,1066]" pageId="2" pageNumber="139" value="2014-11">November 2014</date>
).
</paragraph>
<paragraph id="3907DB52190EAF40E9C6FBC3FEB1FB47" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FBC3FE91FBD7" bold="true" box="[199,372,1077,1102]" pageId="2" pageNumber="139">Type material.</emphasis>
Holotype MLP-Cr 26.980 (female), host: 
<taxonomicName id="FEB8A0D1190EAF40EA4CFBC0FC3AFBD4" box="[845,991,1077,1101]" class="Elasmobranchii" family="Arhynchobatidae" genus="Atlantoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="castelnaui">
<emphasis id="0BCC0740190EAF40EA4CFBC0FCBBFBD4" box="[845,862,1078,1101]" italics="true" pageId="2" pageNumber="139">A</emphasis>
. 
<emphasis id="0BCC0740190EAF40EA6DFBC3FC3AFBD4" box="[876,991,1077,1101]" italics="true" pageId="2" pageNumber="139">castelnaui</emphasis>
</taxonomicName>
. Allotype MLP-Cr 26.981 (male), host: 
<taxonomicName id="FEB8A0D1190EAF40E996FBACFECDFBE8" box="[151,296,1113,1137]" class="Elasmobranchii" family="Arhynchobatidae" genus="Atlantoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="castelnaui">
<emphasis id="0BCC0740190EAF40E996FBACFF4DFBE8" box="[151,168,1114,1137]" italics="true" pageId="2" pageNumber="139">A</emphasis>
. 
<emphasis id="0BCC0740190EAF40E9B4FBAFFECDFBE8" box="[181,296,1113,1137]" italics="true" pageId="2" pageNumber="139">castelnaui</emphasis>
</taxonomicName>
. 
<materialsCitation id="89D0D10F190EAF40E837FBACFEB1FB47" ID-GBIF-Occurrence="1316538285" collectionCode="CCMLP" collectorName="Voucher &amp; All &amp; Carcinological Collection &amp; Museo de La Plata &amp; La Plata" country="Argentina" location="La Plata" pageId="2" pageNumber="139" specimenCount="1" typeStatus="paratype">
<typeStatus id="E60365F0190EAF40E837FBACFE46FBEB" box="[310,419,1114,1138]" pageId="2" pageNumber="139" type="paratype">Paratypes</typeStatus>
MLP-Cr 26.982 (five females and two males from 
<emphasis id="0BCC0740190EAF40EAE0FBACFB99FBE8" box="[993,1148,1113,1137]" italics="true" pageId="2" pageNumber="139">
<taxonomicName id="FEB8A0D1190EAF40EAE0FBACFB91FBE8" box="[993,1140,1113,1137]" class="Elasmobranchii" family="Arhynchobatidae" genus="Atlantoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="castelnaui">A. castelnaui</taxonomicName>
)
</emphasis>
. 
<collectorName id="944DBE84190EAF40ED8BFBAFFB02FBEB" box="[1162,1255,1113,1138]" pageId="2" pageNumber="139">Voucher</collectorName>
specimens (two females and two males from 
<taxonomicName id="FEB8A0D1190EAF40E8E8FB88FD9BFB0C" box="[489,638,1149,1173]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="bonapartii">
<emphasis id="0BCC0740190EAF40E8E8FB88FD9BFB0C" box="[489,638,1149,1173]" italics="true" pageId="2" pageNumber="139">S. bonapartii</emphasis>
</taxonomicName>
MLP-Cr 26.983; two females and two males from 
<taxonomicName id="FEB8A0D1190EAF40EDD7FB88FAD6FB0C" box="[1238,1331,1150,1173]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="acuta">
<emphasis id="0BCC0740190EAF40EDD7FB88FAD6FB0C" box="[1238,1331,1150,1173]" italics="true" pageId="2" pageNumber="139">S. acuta</emphasis>
</taxonomicName>
MLP-Cr 26.984). 
<collectorName id="944DBE84190EAF40E9FFFB57FEC7FB23" box="[254,290,1185,1210]" pageId="2" pageNumber="139">All</collectorName>
specimens are deposited in the 
<collectorName id="944DBE84190EAF40EB9BFB57FC5BFB23" box="[666,958,1185,1210]" pageId="2" pageNumber="139">Carcinological Collection</collectorName>
of the 
<collectorName id="944DBE84190EAF40ED17FB54FB17FB23" box="[1046,1266,1185,1210]" pageId="2" pageNumber="139">Museo de La Plata</collectorName>
(
<collectionCode id="5FA94397190EAF40EC05FB54FA8CFB23" box="[1284,1385,1186,1210]" pageId="2" pageNumber="139">CCMLP</collectionCode>
), 
<location id="3C678D89190EAF40EC7EFB54FF37FB47" LSID="urn:lsid:plazi:treatment:B1116A44190EAF48E996FD5CFB91FE6C:3C678D89190EAF40EC7EFB54FF37FB47" country="Argentina" name="La Plata" pageId="2" pageNumber="139">
<collectorName id="944DBE84190EAF40EC7EFB54FF37FB47" pageId="2" pageNumber="139">La Plata</collectorName>
</location>
, 
<collectingCountry id="41AF9BC2190EAF40E9DCFB33FEB5FB47" box="[221,336,1221,1246]" name="Argentina" pageId="2" pageNumber="139">Argentina</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="3907DB52190EAF40E9C6FB1FFDDCFABF" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FB1FFEB5FA9B" bold="true" box="[199,336,1257,1282]" pageId="2" pageNumber="139">Prevalence.</emphasis>
<taxonomicName id="FEB8A0D1190EAF40E858FB1FFDB2FA98" box="[345,599,1257,1281]" class="Elasmobranchii" family="Arhynchobatidae" genus="Atlantoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="castelnaui">
<emphasis id="0BCC0740190EAF40E858FB1FFDB2FA98" box="[345,599,1257,1281]" italics="true" pageId="2" pageNumber="139">Atlantoraja castelnaui</emphasis>
</taxonomicName>
= 28.6% (CL = 13.3–50.6), 
<taxonomicName id="FEB8A0D1190EAF40EAA6FB1CFB56FA98" box="[935,1203,1257,1281]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="bonapartii">
<emphasis id="0BCC0740190EAF40EAA6FB1CFB56FA98" box="[935,1203,1257,1281]" italics="true" pageId="2" pageNumber="139">Sympterygia bonapartii</emphasis>
</taxonomicName>
= 7.5% (CL = 3.9– 12.8), 
<taxonomicName id="FEB8A0D1190EAF40E9DEFAF8FEDCFABC" box="[223,313,1294,1317]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="acuta">
<emphasis id="0BCC0740190EAF40E9DEFAF8FEDCFABC" box="[223,313,1294,1317]" italics="true" pageId="2" pageNumber="139">S. acuta</emphasis>
</taxonomicName>
18.2% (CL = 3.3–50).
</paragraph>
</subSubSection>
<subSubSection id="71A288D9190EAF40E9C6FAC7FE34FAF7" pageId="2" pageNumber="139" type="description">
<paragraph id="3907DB52190EAF40E9C6FAC7FE34FAF7" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FAC7FE3AFAD3" bold="true" box="[199,479,1329,1354]" pageId="2" pageNumber="139">Mean intensity (range).</emphasis>
<taxonomicName id="FEB8A0D1190EAF40E8E7FAC7FD04FAD0" box="[486,737,1329,1353]" class="Elasmobranchii" family="Arhynchobatidae" genus="Atlantoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="castelnaui">
<emphasis id="0BCC0740190EAF40E8E7FAC7FD04FAD0" box="[486,737,1329,1353]" italics="true" pageId="2" pageNumber="139">Atlantoraja castelnaui</emphasis>
</taxonomicName>
= 4.8 ± 5.6 (1–15), 
<taxonomicName id="FEB8A0D1190EAF40EAC5FAC4FB2BFAD0" box="[964,1230,1329,1353]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="bonapartii">
<emphasis id="0BCC0740190EAF40EAC5FAC4FB2BFAD0" box="[964,1230,1329,1353]" italics="true" pageId="2" pageNumber="139">Sympterygia bonapartii</emphasis>
</taxonomicName>
= 1.7 ± 1.2 (1–5), 
<taxonomicName id="FEB8A0D1190EAF40E996FAA0FF14FAF4" box="[151,241,1366,1389]" class="Elasmobranchii" family="Arhynchobatidae" genus="Sympterygia" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="acuta">
<emphasis id="0BCC0740190EAF40E996FAA0FF14FAF4" box="[151,241,1366,1389]" italics="true" pageId="2" pageNumber="139">S. acuta</emphasis>
</taxonomicName>
= 10 ± 11.3 (2–18).
</paragraph>
</subSubSection>
<subSubSection id="71A288D9190EAF40E9C6FA8FFD36FA2F" pageId="2" pageNumber="139" type="etymology">
<paragraph id="3907DB52190EAF40E9C6FA8FFD36FA2F" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FA8FFEABFA0B" bold="true" box="[199,334,1401,1426]" pageId="2" pageNumber="139">Etymology.</emphasis>
The specific name refers to the subcontinent where the parasites were found, alluding to the first record of the genus 
<emphasis id="0BCC0740190EAF40E874FA68FE11FA2C" box="[373,500,1438,1461]" italics="true" pageId="2" pageNumber="139">Kroeyerina</emphasis>
for South America.
</paragraph>
</subSubSection>
<subSubSection id="71A288D9190EAF4BE9C6FA37FCD4FACB" lastPageId="9" lastPageNumber="146" pageId="2" pageNumber="139" type="description">
<paragraph id="3907DB52190EAF40E9C6FA37FCFFF8FC" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
<emphasis id="0BCC0740190EAF40E9C6FA37FE34FA43" bold="true" box="[199,465,1473,1498]" pageId="2" pageNumber="139">Description of female.</emphasis>
(
<figureCitation id="A183C7D7190EAF40E8E3FA37FDBBFA43" box="[482,606,1473,1498]" captionStart-0="FIGURES 1 – 11" captionStart-1="FIGURES 12 – 15" captionStartId-0="3.[151,264,1924,1947]" captionStartId-1="4.[151,264,1571,1594]" captionTargetBox-0="[157,1436,198,1894]" captionTargetBox-1="[162,1430,198,1544]" captionTargetId-0="figure@3.[151,1436,193,1903]" captionTargetId-1="figure@4.[151,1435,193,1550]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." captionText-1="FIGURES 12 – 15. Kroeyerina sudamericana sp. nov. (adult female). 12, first leg; 13, second leg; 14, third leg; 15, fourth leg. Scale bars: 100 µm." httpUri-0="https://zenodo.org/record/160034/files/figure.png" httpUri-1="https://zenodo.org/record/160035/files/figure.png" pageId="2" pageNumber="139">Figs. 1–15</figureCitation>
, 
<figureCitation id="A183C7D7190EAF40EB6DFA34FD50FA43" box="[620,693,1474,1498]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="2" pageNumber="139">32–39</figureCitation>
; measurements based on 
<specimenCount id="2FBE10DB190EAF40EADBFA37FB9CFA43" box="[986,1145,1473,1498]" pageId="2" pageNumber="139" type="generic">15 specimens</specimenCount>
). Overall length in dorsal view 3797 ± 568 µm (2500–4,420 µm). Cephalothorax 583 ± 50 µm (490–650 µm) long and 612 ± 85 µm (420– 700 µm) wide. Cephalothoracic sutures (
<figureCitation id="A183C7D7190EAF40EB5FF9FFFD7BF9BB" box="[606,670,1545,1570]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
) arising anterolaterally, uniting posteromedially. Three free thoracic somites with overlapping terga, decreasing in width posteriorly. Genital complex cylindrical (
<figureCitation id="A183C7D7190EAF40EDC4F9DBFAD1F9DF" box="[1221,1332,1581,1606]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="2" pageNumber="139">Figs. 1, 2</figureCitation>
), 1915 ± 391 µm (1020–2420 µm) long, 531 ± 120 µm (380–750 µm) wide, comprising approximately 50% of total body length, with lateral rows of spinules along entire length. Fifth leg (
<figureCitation id="A183C7D7190EAF40EA79F983FC22F917" box="[888,967,1653,1678]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="2" pageNumber="139">Figs. 4</figureCitation>
, 
<figureCitation id="A183C7D7190EAF40EAD2F983FC15F917" box="[979,1008,1653,1678]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="2" pageNumber="139">35</figureCitation>
) located near posterior third of genital complex. Abdomen (
<figureCitation id="A183C7D7190EAF40E883F96FFE0EF92B" box="[386,491,1689,1714]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="2" pageNumber="139">Figs. 1, 2</figureCitation>
) 477 ± 61 µm (320–540 µm) long, indistinctly 3-segmented, first somite with pair of ventral setules, third somite with membranes on ventral distal margin, all somites with patches of spinules on ventral surface. Caudal ramus (
<figureCitation id="A183C7D7190EAF40E8F6F917FDA3F963" box="[503,582,1761,1786]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="2" pageNumber="139">Figs. 3</figureCitation>
, 
<figureCitation id="A183C7D7190EAF40EB52F914FD7EF963" box="[595,667,1762,1786]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="2" pageNumber="139">33, 34</figureCitation>
) about three times as long as wide, 195 ± 55 µm (90–250 µm) long, fringed with setules on medial margin and bearing membranes on distal margin of ventral side and six setae as follows: three distal setae, lateral one naked, short and curved, others long and pinnate; two ventral pinnate distolateral setae and one dorsal pinnate distomedial seta.
</paragraph>
<paragraph id="3907DB52190EAF40E9C6F887FC68F84B" blockId="2.[151,1436,789,2002]" pageId="2" pageNumber="139">
Antennule (
<figureCitation id="A183C7D7190EAF40E84AF887FE68F813" box="[331,397,1905,1930]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="2" pageNumber="139">Fig. 5</figureCitation>
) 7-segmented, armature (base to apex) as follows: 11, 5, 2, 3, 1, 1, 12 + 1 aesthetasc. Ventral surface between bases of antennules bearing two small rostral lobes (
<figureCitation id="A183C7D7190EAF40EAA7F863FC12F837" box="[934,1015,1941,1966]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="2" pageNumber="139">Figs. 6</figureCitation>
, 
<figureCitation id="A183C7D7190EAF40ED02F863FBC5F837" box="[1027,1056,1941,1966]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="2" pageNumber="139">36</figureCitation>
) with lateral setule; an additional pair of setules located on dorsal shield near bases of the antennules.
</paragraph>
<caption id="6DC78BDA190FAF41E996F872FCCCF841" httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="3" pageNumber="140" targetBox="[157,1436,198,1894]" targetPageId="3">
<paragraph id="3907DB52190FAF41E996F872FCCCF841" blockId="3.[151,1436,1924,2008]" pageId="3" pageNumber="140">
<emphasis id="0BCC0740190FAF41E996F872FEA2F803" bold="true" box="[151,327,1924,1947]" pageId="3" pageNumber="140">FIGURES 1–11.</emphasis>
<taxonomicName id="FEB8A0D1190FAF41E84EF870FDBCF803" box="[335,601,1924,1947]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="140" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC0740190FAF41E84EF870FDBCF803" box="[335,601,1924,1947]" italics="true" pageId="3" pageNumber="140">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<emphasis id="0BCC0740190FAF41EB60F873FD51F803" bold="true" box="[609,692,1925,1946]" pageId="3" pageNumber="140">
<taxonomicNameLabel id="10FFBA3B190FAF41EB60F873FD51F803" box="[609,692,1925,1946]" pageId="3" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1–2 = 200 µm; 3–4, 7–11 = 50 µm; 5 = 100 µm; 6 = 20 µm.
</paragraph>
</caption>
<caption id="6DC78BDA1908AF46E996F9D5FE85F9C1" httpUri="https://zenodo.org/record/160035/files/figure.png" pageId="4" pageNumber="141" targetBox="[162,1430,198,1544]" targetPageId="4">
<paragraph id="3907DB521908AF46E996F9D5FE85F9C1" blockId="4.[151,1436,1571,1624]" pageId="4" pageNumber="141">
<emphasis id="0BCC07401908AF46E996F9D5FEABF9A1" bold="true" box="[151,334,1571,1594]" pageId="4" pageNumber="141">FIGURES 12–15</emphasis>
. 
<taxonomicName id="FEB8A0D11908AF46E85AF9D2FD86F9A0" box="[347,611,1571,1593]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="141" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC07401908AF46E85AF9D2FD86F9A0" box="[347,611,1571,1593]" italics="true" pageId="4" pageNumber="141">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<emphasis id="0BCC07401908AF46EB6BF9D2FD5EF9A0" bold="true" box="[618,699,1572,1593]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="10FFBA3B1908AF46EB6BF9D2FD5EF9A0" box="[618,699,1572,1593]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(adult female). 12, first leg; 13, second leg; 14, third leg; 15, fourth leg. Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="3907DB521908AF46E9C6F973FBCDF95B" blockId="4.[151,1437,1668,1982]" pageId="4" pageNumber="141">
Antenna (
<figureCitation id="A183C7D71908AF46E837F972FE9CF904" box="[310,377,1668,1693]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="4" pageNumber="141">Fig. 7</figureCitation>
) robust, chelate, 4-segmented; third segment expanded distally into narrow arm with terminal receptacle to receive tip of fourth segment; fourth segment forming thickset claw.
</paragraph>
<paragraph id="3907DB521908AF46E9C6F938FDF1F8CB" blockId="4.[151,1437,1668,1982]" pageId="4" pageNumber="141">
Postantennary process papilliform, lateral to bases of mouthparts. Mouth tube (
<figureCitation id="A183C7D71908AF46ED65F93AFB5FF97C" box="[1124,1210,1740,1765]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="4" pageNumber="141">Fig. 37</figureCitation>
) siphonostomatoid; labrum with two setules, slightly anterior to curved transverse fringe of spinules; two lateral patches of spinules separated by scattered smaller spinules and patch of setules; apical border fringed; labium with two lateral rows of setules and fringing apical border.
</paragraph>
<paragraph id="3907DB521908AF46E9C6F8ABFC2FF8EF" blockId="4.[151,1437,1668,1982]" box="[199,970,1885,1910]" pageId="4" pageNumber="141">
Mandible (
<figureCitation id="A183C7D71908AF46E840F8ABFE61F8EC" box="[321,388,1885,1910]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="4" pageNumber="141">Fig. 8</figureCitation>
) siphonostomatoid, with eight equal teeth on blade.
</paragraph>
<paragraph id="3907DB521908AF46E9C6F877FDB3F824" blockId="4.[151,1437,1668,1982]" pageId="4" pageNumber="141">
Maxillule (
<figureCitation id="A183C7D71908AF46E842F877FE77F800" box="[323,402,1921,1946]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="4" pageNumber="141">Figs. 9</figureCitation>
, 
<figureCitation id="A183C7D71908AF46E89CF877FE5FF800" box="[413,442,1921,1945]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="4" pageNumber="141">37</figureCitation>
) biramous, composed of broad base and two lobes; outer lobe with two short setae, inner lobe ending in two longer pinnate setae.
</paragraph>
<caption id="6DC78BDA1909AF47E996F883FF5AF871" httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="5" pageNumber="142" targetBox="[165,1435,201,1879]" targetPageId="5">
<paragraph id="3907DB521909AF47E996F883FF5AF871" blockId="5.[151,1436,1909,2024]" pageId="5" pageNumber="142">
<emphasis id="0BCC07401909AF47E996F883FEA8F812" bold="true" box="[151,333,1909,1932]" pageId="5" pageNumber="142">FIGURES 16–27</emphasis>
. 
<taxonomicName id="FEB8A0D11909AF47E859F881FD85F815" box="[344,608,1910,1932]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="142" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC07401909AF47E859F881FD85F815" box="[344,608,1910,1932]" italics="true" pageId="5" pageNumber="142">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<emphasis id="0BCC07401909AF47EB67F880FD53F812" bold="true" box="[614,694,1910,1931]" pageId="5" pageNumber="142">
<taxonomicNameLabel id="10FFBA3B1909AF47EB67F880FD53F812" box="[614,694,1910,1931]" pageId="5" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16–17 = 200 µm; 18, 21–27 = 50 µm; 19–20 = 25 µm; 22 = 20 µm.
</paragraph>
</caption>
<caption id="6DC78BDA190AAF44E996F90EFE85F8B4" httpUri="https://zenodo.org/record/160037/files/figure.png" pageId="6" pageNumber="143" targetBox="[157,1419,204,1753]" targetPageId="6">
<paragraph id="3907DB52190AAF44E996F90EFE85F8B4" blockId="6.[151,1436,1784,1837]" pageId="6" pageNumber="143">
<emphasis id="0BCC0740190AAF44E996F90EFEAAF897" bold="true" box="[151,335,1784,1807]" pageId="6" pageNumber="143">FIGURES 28–31</emphasis>
. 
<taxonomicName id="FEB8A0D1190AAF44E85DF90CFD82F896" box="[348,615,1785,1807]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC0740190AAF44E85DF90CFD82F896" box="[348,615,1785,1807]" italics="true" pageId="6" pageNumber="143">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<emphasis id="0BCC0740190AAF44EB6FF90FFD25F897" bold="true" box="[622,704,1785,1806]" pageId="6" pageNumber="143">
<taxonomicNameLabel id="10FFBA3B190AAF44EB6FF90FFD25F897" box="[622,704,1785,1806]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(adult male). 28, first leg; 29, second leg; 30, third leg; 31, fourth leg. Scale bars: 100 µm.
</paragraph>
</caption>
<caption id="6DC78BDA190BAF45E996F8B0FB63F803" httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="7" pageNumber="144" targetBox="[161,1424,202,1831]" targetPageId="7">
<paragraph id="3907DB52190BAF45E996F8B0FB63F803" blockId="7.[151,1436,1862,1946]" pageId="7" pageNumber="144">
<emphasis id="0BCC0740190BAF45E996F8B0FEB6F8C5" bold="true" box="[151,339,1862,1885]" pageId="7" pageNumber="144">FIGURES 32–39.</emphasis>
<taxonomicName id="FEB8A0D1190BAF45E858F8BEFD84F8C4" box="[345,609,1863,1885]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC0740190BAF45E858F8BEFD84F8C4" box="[345,609,1863,1885]" italics="true" pageId="7" pageNumber="144">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<emphasis id="0BCC0740190BAF45EB67F8B1FD53F8C5" bold="true" box="[614,694,1863,1884]" pageId="7" pageNumber="144">
<taxonomicNameLabel id="10FFBA3B190BAF45EB67F8B1FD53F8C5" box="[614,694,1863,1884]" pageId="7" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34–36 = 10 µm; 38 = 20 µm.
</paragraph>
</caption>
<caption id="6DC78BDA1904AF4AE996F8BAFA63F838" httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="8" pageNumber="145" targetBox="[156,1432,204,1836]" targetPageId="8">
<paragraph id="3907DB521904AF4AE996F8BAFA63F838" blockId="8.[151,1436,1868,1953]" pageId="8" pageNumber="145">
<emphasis id="0BCC07401904AF4AE996F8BAFEB0F8FB" bold="true" box="[151,341,1868,1891]" pageId="8" pageNumber="145">FIGURES 40–47.</emphasis>
<taxonomicName id="FEB8A0D11904AF4AE85AF8B8FD81F8FA" box="[347,612,1869,1891]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="145" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC07401904AF4AE85AF8B8FD81F8FA" box="[347,612,1869,1891]" italics="true" pageId="8" pageNumber="145">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<emphasis id="0BCC07401904AF4AEB6DF8B8FD58F8FA" bold="true" box="[620,701,1870,1891]" pageId="8" pageNumber="145">
<taxonomicNameLabel id="10FFBA3B1904AF4AEB6DF8B8FD58F8FA" box="[620,701,1870,1891]" pageId="8" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42–44 = 10 µm; 45–47 = 20 µm.
</paragraph>
</caption>
<paragraph id="3907DB521905AF4BE9C6FF61FAE6FF61" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="146">
Maxilla (
<figureCitation id="A183C7D71905AF4BE82EFF61FE6BFF29" box="[303,398,151,176]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="9" pageNumber="146">Figs. 10</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BE89DFF6EFE5CFF29" box="[412,441,152,176]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="9" pageNumber="146">38</figureCitation>
) brachiform, lacertus robust, distal half of brachium with depression on medial border, distomedial patch of spinules and two distodorsal rows of setules (not easily visible on ventral view), distal process bearing recurved spine. Calamus scoop-like, bearing two medial and two dorsal serrated membranes.
</paragraph>
<paragraph id="3907DB521905AF4BE9C6FEF2FADCFE84" blockId="9.[151,1436,151,501]" box="[199,1337,260,285]" pageId="9" pageNumber="146">
Maxilliped (
<figureCitation id="A183C7D71905AF4BE850FEF2FE48FE85" box="[337,429,260,285]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[157,1436,198,1894]" captionTargetId="figure@3.[151,1436,193,1903]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Kroeyerina sudamericana sp. nov. (adult female). 1, general habitus, dorsal; 2, general habitus, ventral; 3, caudal ramus; 4, fifth leg; 5, antennule; 6, rostral area; 7, antenna; 8, mandible; 9, maxillule; 10, maxilla; 11, maxilliped. Scale bars: 1 – 2 = 200 µm; 3 – 4, 7 – 11 = 50 µm; 5 = 100 µm; 6 = 20 µm." httpUri="https://zenodo.org/record/160034/files/figure.png" pageId="9" pageNumber="146">Figs. 11</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BE8B8FEF2FE33FE85" box="[441,470,260,284]" captionStart="FIGURES 32 – 39" captionStartId="7.[151,264,1862,1885]" captionTargetBox="[161,1424,202,1831]" captionTargetId="figure@7.[150,1436,193,1842]" captionTargetPageId="7" captionText="FIGURES 32 – 39. Kroeyerina sudamericana sp. nov. SEM micrographs (adult female). 32, general habitus, ventral; 33, caudal rami; 34, detail of distal armature of caudal ramus; 35, fifth leg; 36, rostral processes; 37, mouth tube and maxillules; 38, maxilla; 39, maxilliped. Scale bars: 32 = 500 µm; 33, 37, 39 = 50 µm; 34 – 36 = 10 µm; 38 = 20 µm." httpUri="https://zenodo.org/record/256586/files/figure.png" pageId="9" pageNumber="146">39</figureCitation>
) subchelate, four-segmented, corpus unarmed and subchela with small spine.
</paragraph>
<paragraph id="3907DB521905AF4BE9C6FEDEFACDFE6C" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="146">
Legs 1–4 (
<figureCitation id="A183C7D71905AF4BE842FED1FE2BFED9" box="[323,462,295,320]" captionStart="FIGURES 12 – 15" captionStartId="4.[151,264,1571,1594]" captionTargetBox="[162,1430,198,1544]" captionTargetId="figure@4.[151,1435,193,1550]" captionTargetPageId="4" captionText="FIGURES 12 – 15. Kroeyerina sudamericana sp. nov. (adult female). 12, first leg; 13, second leg; 14, third leg; 15, fourth leg. Scale bars: 100 µm." httpUri="https://zenodo.org/record/160035/files/figure.png" pageId="9" pageNumber="146">Figs. 12–15</figureCitation>
), coxa 1 fringed with two distal membranes on ventral side, coxae 2–3 fringed with distolateral membrane. Basis 1 with lateral pinnate setae, two medial setae, larger pinnate and smaller naked and two distal membranes. Basis 2 with lateral pinnate setae and two distal membranes. Basis 3 with lateral pinnate setae and medial membrane. Basis 4 with lateral naked seta and medial membrane. All exopodal segments of legs 1–4 with lateral membrane. All endopodal segments of legs 1–4 with lateral membrane and dorsolateral row of setules. Armature formula of legs 1–4 as follows (spines in Roman numerals; setae in Arabic numerals):
</paragraph>
<paragraph id="3907DB521905AF4BE8B0FDD6FCD6FCC0" blockId="9.[433,804,544,566]" lastBlockId="9.[151,1437,797,2010]" pageId="9" pageNumber="146">exopod endopod Larger spines on third exopodal segment with lateral membranes in all legs, those of legs 2–3 also hemipinnate. Leg 5 vestigial, each ramus with pair of setae.</paragraph>
<paragraph id="3907DB521905AF4BE996FDA9FC8AFD49" pageId="9" pageNumber="146">
<table id="4BB829F2190550BDE996FDA8FC8AFD49" box="[151,879,606,720]" gridcols="3" gridrows="4" pageId="9" pageNumber="146">
<tr id="8788D910190550BDE996FDA8FC8AFDED" box="[151,879,606,628]" gridrow="0" pageId="9" pageNumber="146">
<th id="C459B06C190550BDE996FDA8FF34FDED" box="[151,209,606,628]" gridcol="0" gridrow="0" pageId="9" pageNumber="146">Leg 1</th>
<th id="C459B06C190550BDE8B0FDA8FDA7FDED" box="[433,578,606,628]" gridcol="1" gridrow="0" pageId="9" pageNumber="146">I–1; I–1; II–4</th>
<th id="C459B06C190550BDEBCCFDA8FC8AFDED" box="[717,879,606,628]" gridcol="2" gridrow="0" pageId="9" pageNumber="146">0–1; 0–1; 6</th>
</tr>
<tr id="8788D910190550BDE996FD8BFC8AFD0A" box="[151,879,637,659]" gridrow="1" pageId="9" pageNumber="146">
<th id="C459B06C190550BDE996FD8BFF34FD0A" box="[151,209,637,659]" gridcol="0" gridrow="1" pageId="9" pageNumber="146">Leg 2</th>
<td id="C459B06C190550BDE8B0FD8BFDA7FD0A" box="[433,578,637,659]" gridcol="1" gridrow="1" pageId="9" pageNumber="146">I–1; I–1; III–4</td>
<td id="C459B06C190550BDEBCCFD8BFC8AFD0A" box="[717,879,637,659]" gridcol="2" gridrow="1" pageId="9" pageNumber="146">0–1; 0–1; 6</td>
</tr>
<tr id="8788D910190550BDE996FD6AFC8AFD2B" box="[151,879,668,690]" gridrow="2" pageId="9" pageNumber="146">
<th id="C459B06C190550BDE996FD6AFF34FD2B" box="[151,209,668,690]" gridcol="0" gridrow="2" pageId="9" pageNumber="146">Leg 3</th>
<td id="C459B06C190550BDE8B0FD6AFDA7FD2B" box="[433,578,668,690]" gridcol="1" gridrow="2" pageId="9" pageNumber="146">I–1; I–1; III–4</td>
<td id="C459B06C190550BDEBCCFD6AFC8AFD2B" box="[717,879,668,690]" gridcol="2" gridrow="2" pageId="9" pageNumber="146">0–1; 0–1; 1–I–3</td>
</tr>
<tr id="8788D910190550BDE996FD4CFC8AFD49" box="[151,879,698,720]" gridrow="3" pageId="9" pageNumber="146">
<th id="C459B06C190550BDE996FD4CFF34FD49" box="[151,209,698,720]" gridcol="0" gridrow="3" pageId="9" pageNumber="146">Leg 4</th>
<td id="C459B06C190550BDE8B0FD4CFDA7FD49" box="[433,578,698,720]" gridcol="1" gridrow="3" pageId="9" pageNumber="146">I–1; 1–I; III–4</td>
<td id="C459B06C190550BDEBCCFD4CFC8AFD49" box="[717,879,698,720]" gridcol="2" gridrow="3" pageId="9" pageNumber="146">0–1; 0–1; 1–I–2</td>
</tr>
</table>
</paragraph>
<paragraph id="3907DB521905AF4BE9C6FC93FAB4FB7F" blockId="9.[151,1437,797,2010]" pageId="9" pageNumber="146">
<emphasis id="0BCC07401905AF4BE9C6FC93FE59FCE7" bold="true" box="[199,444,869,894]" pageId="9" pageNumber="146">Description of male.</emphasis>
(
<figureCitation id="A183C7D71905AF4BE8CCFC93FDB2FCE7" box="[461,599,869,894]" captionStart-0="FIGURES 16 – 27" captionStart-1="FIGURES 28 – 31" captionStartId-0="5.[151,264,1909,1932]" captionStartId-1="6.[151,264,1784,1807]" captionTargetBox-0="[165,1435,201,1879]" captionTargetBox-1="[157,1419,204,1753]" captionTargetId-0="figure@5.[151,1435,193,1888]" captionTargetId-1="figure@6.[150,1436,193,1764]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." captionText-1="FIGURES 28 – 31. Kroeyerina sudamericana sp. nov. (adult male). 28, first leg; 29, second leg; 30, third leg; 31, fourth leg. Scale bars: 100 µm." httpUri-0="https://zenodo.org/record/160036/files/figure.png" httpUri-1="https://zenodo.org/record/160037/files/figure.png" pageId="9" pageNumber="146">Figs. 16–31</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BEB64FC90FD4AFCE7" box="[613,687,870,894]" captionStart="FIGURES 40 – 47" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[156,1432,204,1836]" captionTargetId="figure@8.[151,1436,193,1848]" captionTargetPageId="8" captionText="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="9" pageNumber="146">40–47</figureCitation>
; measurements based on six specimens). Overall length in dorsal view 2467 ± 178 µm (2160–2660 µm). Cephalothorax 447 ± 28 µm (400–480 µm) long and 447 ± 43 µm (380–500 µm) wide. Cephalothoracic sutures (
<figureCitation id="A183C7D71905AF4BEB39FC5BFD69FC5F" box="[568,652,941,966]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Fig. 16</figureCitation>
) arising anterolaterally, uniting posteromedially. Three free thoracic somites with overlapping terga, decreasing in width posteriorly. Genital complex (
<figureCitation id="A183C7D71905AF4BED3AFC27FB26FC73" box="[1083,1219,977,1002]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Figs. 16, 17</figureCitation>
) cylindrical, 670 ± 113 µm (500–800 µm) long, 258 ± 16 µm (240–280 µm) wide, comprising approximately 27% of total body length, with lateral rows of spinules (
<figureCitation id="A183C7D71905AF4BE8E9FBEFFDDEFBAB" box="[488,571,1049,1074]" captionStart="FIGURES 40 – 47" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[156,1432,204,1836]" captionTargetId="figure@8.[151,1436,193,1848]" captionTargetPageId="8" captionText="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="9" pageNumber="146">Fig. 43</figureCitation>
) along entire length. Fifth leg (
<figureCitation id="A183C7D71905AF4BEA9EFBEFFC14FBAB" box="[927,1009,1049,1074]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Fig. 19</figureCitation>
) located laterally at middle of genital complex, with four spiniform setae. Sixth leg (
<figureCitation id="A183C7D71905AF4BEBB2FBCBFCE2FBCF" box="[691,775,1085,1110]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Fig. 20</figureCitation>
) vestigial, consisting of protuberance with two spiniform setae, located laterally near posterior margin of genital complex. Abdomen (
<figureCitation id="A183C7D71905AF4BEAFBFB97FB61FBE3" box="[1018,1156,1121,1146]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Figs. 16, 17</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BED93FB94FB55FBE3" box="[1170,1200,1122,1146]" captionStart="FIGURES 40 – 47" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[156,1432,204,1836]" captionTargetId="figure@8.[151,1436,193,1848]" captionTargetPageId="8" captionText="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="9" pageNumber="146">40</figureCitation>
) 588 ± 73 µm (460– 660 µm) long, 3-segmented, first two somites with slight constriction at anterior third, third somite with membranes near junction with caudal rami with patches of spinules on ventral surface. Caudal ramus (
<figureCitation id="A183C7D71905AF4BEC11FB5FFA88FB5B" box="[1296,1389,1193,1218]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Figs. 18</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BEC78FB5CFF50FB7F" captionStart="FIGURES 40 – 47" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[156,1432,204,1836]" captionTargetId="figure@8.[151,1436,193,1848]" captionTargetPageId="8" captionText="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="9" pageNumber="146">41, 42</figureCitation>
) as in female, but four times as long as wide, 217 ± 48 µm (140–260 µm) long and with all setae pinnate.
</paragraph>
<paragraph id="3907DB521905AF4BE9C6FB07FCD4FACB" blockId="9.[151,1437,797,2010]" pageId="9" pageNumber="146">
Antennule (
<figureCitation id="A183C7D71905AF4BE84FFB07FE41FA93" box="[334,420,1265,1290]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Fig. 21</figureCitation>
) 7-segmented, armature (base to apex) as follows: 11, 5, 2, 5, 2, 1, 12 + 1 aesthetasc. Rostral lobes (
<figureCitation id="A183C7D71905AF4BE843FAE3FE40FAB7" box="[322,421,1301,1326]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Figs. 22</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BE8B5FAE0FE36FAB7" box="[436,467,1302,1326]" captionStart="FIGURES 40 – 47" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[156,1432,204,1836]" captionTargetId="figure@8.[151,1436,193,1848]" captionTargetPageId="8" captionText="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="9" pageNumber="146">44</figureCitation>
), rest of cephalic appendages (
<figureCitation id="A183C7D71905AF4BEA43FAE3FC2AFAB7" box="[834,975,1301,1326]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Figs. 23–26</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BEADEFAE3FBC9FAB7" box="[991,1068,1301,1326]" captionStart="FIGURES 40 – 47" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[156,1432,204,1836]" captionTargetId="figure@8.[151,1436,193,1848]" captionTargetPageId="8" captionText="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="9" pageNumber="146">45, 46</figureCitation>
), maxilliped (
<figureCitation id="A183C7D71905AF4BEDD1FAE3FAD7FAB7" box="[1232,1330,1301,1326]" captionStart="FIGURES 16 – 27" captionStartId="5.[151,264,1909,1932]" captionTargetBox="[165,1435,201,1879]" captionTargetId="figure@5.[151,1435,193,1888]" captionTargetPageId="5" captionText="FIGURES 16 – 27. Kroeyerina sudamericana sp. nov. (adult male). 16, general habitus, dorsal; 17, general habitus, ventral; 18, posterior portion of abdomen and caudal rami; 19, fifth leg; 20, sixth leg; 21, antennule; 22, rostral area; 23, antenna; 24, mandible; 25, maxillule; 26, maxilla; 27, maxilliped. Scale bars: 16 – 17 = 200 µm; 18, 21 – 27 = 50 µm; 19 – 20 = 25 µm; 22 = 20 µm." httpUri="https://zenodo.org/record/160036/files/figure.png" pageId="9" pageNumber="146">Figs. 27</figureCitation>
, 
<figureCitation id="A183C7D71905AF4BEC42FAE0FA85FAB7" box="[1347,1376,1302,1326]" captionStart="FIGURES 40 – 47" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[156,1432,204,1836]" captionTargetId="figure@8.[151,1436,193,1848]" captionTargetPageId="8" captionText="FIGURES 40 – 47. Kroeyerina sudamericana sp. nov. SEM micrographs (adult male). 40, general habitus, ventral; 41, caudal rami; 42, detail of distal armature of caudal rami; 43, spinulation in lateral fields of genital complex; 44, rostral processes; 45, antenna; 46, mouth tube and maxilla; 47, maxilliped. Scale bars: 40 = 200 µm; 41 = 50 µm; 42 – 44 = 10 µm; 45 – 47 = 20 µm." httpUri="https://zenodo.org/record/256587/files/figure.png" pageId="9" pageNumber="146">47</figureCitation>
) and general morphology of legs 1–4 (
<figureCitation id="A183C7D71905AF4BEB0DFACFFD77FACB" box="[524,658,1337,1362]" captionStart="FIGURES 28 – 31" captionStartId="6.[151,264,1784,1807]" captionTargetBox="[157,1419,204,1753]" captionTargetId="figure@6.[150,1436,193,1764]" captionTargetPageId="6" captionText="FIGURES 28 – 31. Kroeyerina sudamericana sp. nov. (adult male). 28, first leg; 29, second leg; 30, third leg; 31, fourth leg. Scale bars: 100 µm." httpUri="https://zenodo.org/record/160037/files/figure.png" pageId="9" pageNumber="146">Figs. 28–31</figureCitation>
) as in female.
</paragraph>
</subSubSection>
<subSubSection id="71A288D91905AF48E9C6FAABFB91FE6C" lastPageId="10" lastPageNumber="147" pageId="9" pageNumber="146" type="discussion">
<paragraph id="3907DB521905AF4BE9C6FAABFCF8F9EB" blockId="9.[151,1437,797,2010]" pageId="9" pageNumber="146">
<materialsCitation id="89D0D10F1905AF4BE9C6FAABFCF8F9EB" ID-GBIF-Occurrence="1316538287" country="United States of America" location="Mississippi" pageId="9" pageNumber="146" specimenCount="1" stateProvince="Mississippi">
<emphasis id="0BCC07401905AF4BE9C6FAABFEDEFAEF" bold="true" box="[199,315,1373,1398]" pageId="9" pageNumber="146">Remarks.</emphasis>
In his phylogenetic analysis of 
<emphasis id="0BCC07401905AF4BEBD9FAA8FCB3FAEC" box="[728,854,1374,1397]" italics="true" pageId="9" pageNumber="146">Kroeyerina</emphasis>
, 
<bibRefCitation id="5D29A6A31905AF4BEA6FFAA8FBECFAEF" author="Deets" box="[878,1033,1374,1398]" pageId="9" pageNumber="146" refString="Deets, G. B. (1987) Phylogenetic analysis and revision of Kroeyerina Wilson, 1932 (Siphonostomatoida: Kroyeriidae), copepods parasitic on chondrichthyans, with descriptions of four new species and the erection of a new genus, Prokroyeria. Canadian Journal of Zoology, 65, 2121 - 2148. http: // dx. doi. org / 10.1139 / z 87 - 327" type="journal article" year="1987">Deets (1987)</bibRefCitation>
identified two clades presenting differences in the morphology of the rostral lobes (
<bibRefCitation id="5D29A6A31905AF4BEBDAFA74FC78FA03" author="Benz" box="[731,925,1409,1434]" pageId="9" pageNumber="146" refString="Benz, G. W., Smith, B. E. &amp; Bullard, S. A. (2001) Kroeyerina deetsorum n. sp. (Copepoda: Kroyeriidae) from the olfactory sacs of Atlantic sharpnose sharks (Rhizoprionodon terraenovae) captured in the Gulf of Mexico and northwestern Atlantic and first report of copepodids representing Kroyeriidae. Journal of Parasitology, 87, 1279 - 1290. http: // dx. doi. org / 10.1645 / 0022 - 3395 (2001) 087 [1279: kdnsck] 2.0. co; 2" type="journal article" year="2001">
Benz 
<emphasis id="0BCC07401905AF4BEA1EFA74FCB7FA00" box="[799,850,1409,1433]" italics="true" pageId="9" pageNumber="146">et al</emphasis>
. 2001
</bibRefCitation>
). One clade, with upturned rostral processes, infects sharks and the other composed of only three species, with reduced lobes, is parasitic on rays. More recently 
<bibRefCitation id="5D29A6A31905AF4BE996FA3CFE80FA7B" box="[151,357,1481,1506]" pageId="9" pageNumber="146" refString="Benz, G. W., Smith, B. E. &amp; Bullard, S. A. (2001) Kroeyerina deetsorum n. sp. (Copepoda: Kroyeriidae) from the olfactory sacs of Atlantic sharpnose sharks (Rhizoprionodon terraenovae) captured in the Gulf of Mexico and northwestern Atlantic and first report of copepodids representing Kroyeriidae. Journal of Parasitology, 87, 1279 - 1290. http: // dx. doi. org / 10.1645 / 0022 - 3395 (2001) 087 [1279: kdnsck] 2.0. co; 2" type="journal article">
Benz 
<emphasis id="0BCC07401905AF4BE9D8FA3CFEEEFA78" box="[217,267,1481,1505]" italics="true" pageId="9" pageNumber="146">et al</emphasis>
. (2001)
</bibRefCitation>
described a new species, 
<taxonomicName id="FEB8A0D11905AF4BEB8CFA3CFB07FA7B" authority="Benz, Smith &amp; Bullard, 2001" authorityName="Benz, Smith &amp; Bullard" authorityYear="2001" box="[653,1250,1481,1506]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="deetsorum">
<emphasis id="0BCC07401905AF4BEB8CFA3CFC6CFA78" box="[653,905,1481,1505]" italics="true" pageId="9" pageNumber="146">Kroeyerina deetsorum</emphasis>
<bibRefCitation id="5D29A6A31905AF4BEA90FA3CFB07FA7B" author="Benz" box="[913,1250,1481,1506]" pageId="9" pageNumber="146" refString="Benz, G. W., Smith, B. E. &amp; Bullard, S. A. (2001) Kroeyerina deetsorum n. sp. (Copepoda: Kroyeriidae) from the olfactory sacs of Atlantic sharpnose sharks (Rhizoprionodon terraenovae) captured in the Gulf of Mexico and northwestern Atlantic and first report of copepodids representing Kroyeriidae. Journal of Parasitology, 87, 1279 - 1290. http: // dx. doi. org / 10.1645 / 0022 - 3395 (2001) 087 [1279: kdnsck] 2.0. co; 2" type="journal article" year="2001">Benz, Smith &amp; Bullard, 2001</bibRefCitation>
</taxonomicName>
, parasitic on the shark 
<taxonomicName id="FEB8A0D11905AF4BE9DDFA1BFCE7F99F" authority="Richardson, 1837" authorityName="Richardson" authorityYear="1837" box="[220,770,1517,1542]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="terraenovae">
<emphasis id="0BCC07401905AF4BE9DDFA1BFDC5F99C" box="[220,544,1517,1541]" italics="true" pageId="9" pageNumber="146">Rhizoprionodon terraenovae</emphasis>
(
<bibRefCitation id="5D29A6A31905AF4BEB31FA1BFD1FF99F" author="Richardson" box="[560,762,1517,1542]" pageId="9" pageNumber="146" refString="Richardson, J. (1837) The Fish. In: Fauna Boreali-Americana; or the Zoology of the northern Parts of British America: containing Descriptions of the Objects of natural History collected on the late northern Land Expeditions, under the Command of Sir John Franklin, R. N. J. Bentley, London, Part 3, xv + 327 pp., plates 74 - 97." type="book chapter" year="1837">Richardson, 1837</bibRefCitation>
)
</taxonomicName>
from the 
<collectingRegion id="FB7C15B01905AF4BEA75FA1BFC11F99F" box="[884,1012,1517,1542]" country="United States of America" name="Mississippi" pageId="9" pageNumber="146">Mississippi</collectingRegion>
Delta (
<collectingRegion id="FB7C15B01905AF4BED49FA18FB68F99F" box="[1096,1165,1518,1542]" country="United States of America" name="Texas" pageId="9" pageNumber="146">Texas</collectingRegion>
, 
<collectingCountry id="41AF9BC21905AF4BED9BFA18FB32F99F" box="[1178,1239,1518,1542]" name="United States of America" pageId="9" pageNumber="146">USA</collectingCountry>
). This parasite of sharks, however, does not possess rostral lobes or upturned projections. This characteristic was considered as a possible secondary loss due to the diminutive size (body length 
<quantity id="FE4076B71905AF4BEA67F9C0FC58F9D7" box="[870,957,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="146" unit="mm" value="1.2">1.2 mm</quantity>
) of 
<taxonomicName id="FEB8A0D11905AF4BEAEAF9C0FAB3F9D7" authority="Benz et al. 2001" authorityName="Benz et al." authorityYear="2001" box="[1003,1366,1589,1614]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="deetsorum">
<emphasis id="0BCC07401905AF4BEAEAF9C0FC1BF9D4" box="[1003,1022,1590,1613]" italics="true" pageId="9" pageNumber="146">K</emphasis>
. 
<emphasis id="0BCC07401905AF4BED0DF9C3FB64F9D4" box="[1036,1153,1589,1613]" italics="true" pageId="9" pageNumber="146">deetsorum</emphasis>
(
<bibRefCitation id="5D29A6A31905AF4BED91F9C0FAABF9D7" author="Benz" box="[1168,1358,1589,1614]" pageId="9" pageNumber="146" refString="Benz, G. W., Smith, B. E. &amp; Bullard, S. A. (2001) Kroeyerina deetsorum n. sp. (Copepoda: Kroyeriidae) from the olfactory sacs of Atlantic sharpnose sharks (Rhizoprionodon terraenovae) captured in the Gulf of Mexico and northwestern Atlantic and first report of copepodids representing Kroyeriidae. Journal of Parasitology, 87, 1279 - 1290. http: // dx. doi. org / 10.1645 / 0022 - 3395 (2001) 087 [1279: kdnsck] 2.0. co; 2" type="journal article" year="2001">
Benz 
<emphasis id="0BCC07401905AF4BEDD3F9C0FAE1F9D4" box="[1234,1284,1589,1613]" italics="true" pageId="9" pageNumber="146">et al</emphasis>
. 2001
</bibRefCitation>
)
</taxonomicName>
; these both features clearly differentiate it from the new species.
</materialsCitation>
</paragraph>
<paragraph id="3907DB521905AF4BE9CEF988FDD3F8F7" blockId="9.[151,1437,797,2010]" pageId="9" pageNumber="146">
In the possession of reduced rostral lobes, the new species is similar to congeners forming the second clade proposed by 
<bibRefCitation id="5D29A6A31905AF4BE82FF954FE20F923" author="Deets" box="[302,453,1698,1722]" pageId="9" pageNumber="146" refString="Deets, G. B. (1987) Phylogenetic analysis and revision of Kroeyerina Wilson, 1932 (Siphonostomatoida: Kroyeriidae), copepods parasitic on chondrichthyans, with descriptions of four new species and the erection of a new genus, Prokroyeria. Canadian Journal of Zoology, 65, 2121 - 2148. http: // dx. doi. org / 10.1139 / z 87 - 327" type="journal article" year="1987">Deets (1987)</bibRefCitation>
, namely 
<taxonomicName id="FEB8A0D11905AF4BEB30F954FC4EF923" authority="Wilson, 1932" authorityName="Wilson" authorityYear="1932" box="[561,939,1697,1722]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="0BCC07401905AF4BEB30F954FCE0F920" box="[561,773,1698,1721]" italics="true" pageId="9" pageNumber="146">Kroeyerina nasuta</emphasis>
<bibRefCitation id="5D29A6A31905AF4BEA0EF957FC4EF923" author="Wilson" box="[783,939,1697,1722]" pageId="9" pageNumber="146" refString="Wilson, C. B. (1932) The copepods of the Woods Hole region, Massachusetts. Bulletin of the United States National Museum, 158, 1 - 635, plates 1 - 41." type="journal article" year="1932">Wilson, 1932</bibRefCitation>
</taxonomicName>
, a parasite of 
<taxonomicName id="FEB8A0D11905AF4BED55F957FF2AF947" authority="Garman, 1913" authorityName="Garman" authorityYear="1913" class="Elasmobranchii" family="Dasyatidae" genus="Dasybatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="marinus">
<emphasis id="0BCC07401905AF4BED55F957FAD5F920" box="[1108,1328,1697,1721]" italics="true" pageId="9" pageNumber="146">Dasybatus marinus</emphasis>
<bibRefCitation id="5D29A6A31905AF4BEC3AF954FF2AF947" author="Garman" pageId="9" pageNumber="146" refString="Garman, S. (1913) The Plagiostomia (sharks, skates, and rays). Memoirs of the Museum of comparative Zoology, 36, i - xiii + 1 - 515. [also Atlas: 77 plates]" type="book chapter" year="1913">Garman, 1913</bibRefCitation>
</taxonomicName>
(= 
<taxonomicName id="FEB8A0D11905AF4BE9F7F930FD74F947" authority="Mitchill, 1815" authorityName="Mitchill" authorityYear="1815" box="[246,657,1733,1758]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="centroura">
<emphasis id="0BCC07401905AF4BE9F7F930FE28F944" box="[246,461,1734,1757]" italics="true" pageId="9" pageNumber="146">Dasyatis centroura</emphasis>
<bibRefCitation id="5D29A6A31905AF4BE8D5F933FD74F947" author="Mitchill" box="[468,657,1733,1758]" pageId="9" pageNumber="146" refString="Mitchill, S. L. (1815) The fishes of New-York, described and arranged. Transactions of the literary and philosophical Society of New-York, 1 (5), 355 - 492, plates 1 - 6. [for 1814]" type="journal article" year="1815">(Mitchill, 1815))</bibRefCitation>
</taxonomicName>
from the western North Atlantic (Woods Hole, 
<collectingCountry id="41AF9BC21905AF4BEDAAF930FB00F947" box="[1195,1253,1734,1758]" name="United States of America" pageId="9" pageNumber="146">USA</collectingCountry>
) (
<bibRefCitation id="5D29A6A31905AF4BEDFAF933FA75F947" author="Wilson" box="[1275,1424,1733,1758]" pageId="9" pageNumber="146" refString="Wilson, C. B. (1932) The copepods of the Woods Hole region, Massachusetts. Bulletin of the United States National Museum, 158, 1 - 635, plates 1 - 41." type="journal article" year="1932">Wilson 1932</bibRefCitation>
), 
<taxonomicName id="FEB8A0D11905AF4BE996F91CFDC1F89B" authority="Deets, 1987" authorityName="Deets" authorityYear="1987" box="[151,548,1769,1794]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="deborahae">
<emphasis id="0BCC07401905AF4BE996F91CFE73F898" box="[151,406,1769,1793]" italics="true" pageId="9" pageNumber="146">Kroeyerina deborahae</emphasis>
<bibRefCitation id="5D29A6A31905AF4BE89CF91CFDC1F89B" author="Deets" box="[413,548,1770,1794]" pageId="9" pageNumber="146" refString="Deets, G. B. (1987) Phylogenetic analysis and revision of Kroeyerina Wilson, 1932 (Siphonostomatoida: Kroyeriidae), copepods parasitic on chondrichthyans, with descriptions of four new species and the erection of a new genus, Prokroyeria. Canadian Journal of Zoology, 65, 2121 - 2148. http: // dx. doi. org / 10.1139 / z 87 - 327" type="journal article" year="1987">Deets, 1987</bibRefCitation>
</taxonomicName>
parasitic on 
<taxonomicName id="FEB8A0D11905AF4BEBB6F91FFBD8F89B" authority="Ayres, 1854" authorityName="Ayres" authorityYear="1854" box="[695,1085,1769,1794]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="productus">
<emphasis id="0BCC07401905AF4BEBB6F91FFC48F898" box="[695,941,1769,1793]" italics="true" pageId="9" pageNumber="146">Rhinobatos productus</emphasis>
<bibRefCitation id="5D29A6A31905AF4BEAB4F91CFBD8F89B" author="Ayres" box="[949,1085,1769,1794]" pageId="9" pageNumber="146" refString="Ayres, W. O. (1854) [Description of new fishes from California.] (Minutes of Academy meetings were printed in &quot; The Pacific &quot; (a newspaper) shortly after each meeting. New species date to publication in The Pacific. Dates of publication are given in each species account). The Pacific, 3 - 4 (through 6). [newspaper, also as Proceedings of the California Academy of natural Sciences, 1, 3 - 22. (reprinted in 2 nd ed. of Academy's Proceedings, 1873)]." type="journal article" year="1854">Ayres, 1854</bibRefCitation>
</taxonomicName>
from the eastern North Pacific (Palos Verdes, 
<collectingRegion id="FB7C15B01905AF4BE83FF8FBFE54F8BF" box="[318,433,1805,1830]" country="United States of America" name="California" pageId="9" pageNumber="146">California</collectingRegion>
) and 
<taxonomicName id="FEB8A0D11905AF4BE8F1F8F8FC82F8BF" authority="Deets, 1987" authorityName="Deets" authorityYear="1987" box="[496,871,1805,1830]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="mobulae">
<emphasis id="0BCC07401905AF4BE8F1F8F8FD32F8BC" box="[496,727,1805,1829]" italics="true" pageId="9" pageNumber="146">Kroeyerina mobulae</emphasis>
<bibRefCitation id="5D29A6A31905AF4BEBDCF8F8FC82F8BF" author="Deets" box="[733,871,1806,1830]" pageId="9" pageNumber="146" refString="Deets, G. B. (1987) Phylogenetic analysis and revision of Kroeyerina Wilson, 1932 (Siphonostomatoida: Kroyeriidae), copepods parasitic on chondrichthyans, with descriptions of four new species and the erection of a new genus, Prokroyeria. Canadian Journal of Zoology, 65, 2121 - 2148. http: // dx. doi. org / 10.1139 / z 87 - 327" type="journal article" year="1987">Deets, 1987</bibRefCitation>
</taxonomicName>
, a parasite of 
<taxonomicName id="FEB8A0D11905AF4BED02F8FBFF2AF8D3" authority="Beebe &amp; Tee-Van, 1938" authorityName="Beebe &amp; Tee-Van" authorityYear="1938" class="Elasmobranchii" family="Myliobatidae" genus="Mobula" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="lucasana">
<emphasis id="0BCC07401905AF4BED02F8FBFB23F8BC" box="[1027,1222,1805,1829]" italics="true" pageId="9" pageNumber="146">Mobula lucasana</emphasis>
<bibRefCitation id="5D29A6A31905AF4BEDCFF8FBFF2AF8D3" author="Beebe" pageId="9" pageNumber="146" refString="Beebe, W. &amp; Tee-Van, J. (1938) Eastern Pacific expeditions of the New York Zoological Society, XV. Seven new marine fishes from Lower California. Zoologica, Scientific Contributions of the New York Zoological Society, 23 (3), 299 - 312, plates 1 - 3." type="journal article" year="1938">Beebe &amp; Tee-Van, 1938</bibRefCitation>
</taxonomicName>
(= 
<taxonomicName id="FEB8A0D11905AF4BE9FAF8C7FD94F8D3" authority="Lloyd, 1908" authorityName="Lloyd" authorityYear="1908" box="[251,625,1841,1866]" class="Elasmobranchii" family="Myliobatidae" genus="Mobula" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="thurstoni">
<emphasis id="0BCC07401905AF4BE9FAF8C7FE24F8D0" box="[251,449,1841,1865]" italics="true" pageId="9" pageNumber="146">Mobula thurstoni</emphasis>
<bibRefCitation id="5D29A6A31905AF4BE8CBF8C7FD94F8D3" author="Lloyd" box="[458,625,1841,1866]" pageId="9" pageNumber="146" refString="Lloyd, R. E. (1908) On two new species of eagle-rays (Myliobatidae), with notes on the skull of the genus Ceratoptera. Records of the Indian Museum (Calcutta), 2, 175 - 180, plates 4 - 5, 10." type="journal article" year="1908">(Lloyd, 1908))</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="FEB8A0D11905AF4BEBACF8C7FB69F8D3" authority="Muller &amp; Henle, 1841" authorityName="Muller &amp; Henle" authorityYear="1841" box="[685,1164,1841,1866]" class="Elasmobranchii" family="Myliobatidae" genus="Mobula" kingdom="Animalia" order="Myliobatiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="japanica">
<emphasis id="0BCC07401905AF4BEBACF8C7FC95F8D0" box="[685,880,1841,1865]" italics="true" pageId="9" pageNumber="146">Mobula japanica</emphasis>
(
<bibRefCitation id="5D29A6A31905AF4BEA80F8C7FB61F8D3" author="Muller" box="[897,1156,1841,1866]" pageId="9" pageNumber="146" refString="Muller, J. &amp; Henle, F. G. J. (1841) Systematische Beschreibung der Plagiostomen. Veit und Comp., Berlin, xxii + 200 pp., 60 plates. [pp. 1 - 28 published in 1838, reset pp. 27 - 28, 29 - 102 in 1839, pp. i - xxii + 103 - 200 in 1841]" type="book chapter" year="1841">Müller &amp; Henle, 1841</bibRefCitation>
)
</taxonomicName>
from the eastern North Pacific (Sea of Cortez) (
<bibRefCitation id="5D29A6A31905AF4BE8A4F8A0FDCFF8F7" author="Deets" box="[421,554,1878,1902]" pageId="9" pageNumber="146" refString="Deets, G. B. (1987) Phylogenetic analysis and revision of Kroeyerina Wilson, 1932 (Siphonostomatoida: Kroyeriidae), copepods parasitic on chondrichthyans, with descriptions of four new species and the erection of a new genus, Prokroyeria. Canadian Journal of Zoology, 65, 2121 - 2148. http: // dx. doi. org / 10.1139 / z 87 - 327" type="journal article" year="1987">Deets 1987</bibRefCitation>
).
</paragraph>
<paragraph id="3907DB521905AF48E9C6F88FFDB5FEFC" blockId="9.[151,1437,797,2010]" lastBlockId="10.[151,1436,151,501]" lastPageId="10" lastPageNumber="147" pageId="9" pageNumber="146">
Both males and females of the new species are readily distinguished from 
<taxonomicName id="FEB8A0D11905AF4BED35F88CFB5CF808" box="[1076,1209,1913,1937]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="mobulae">
<emphasis id="0BCC07401905AF4BED35F88CFBA2F808" box="[1076,1095,1914,1937]" italics="true" pageId="9" pageNumber="146">K</emphasis>
. 
<emphasis id="0BCC07401905AF4BED59F88FFB5CF808" box="[1112,1209,1913,1937]" italics="true" pageId="9" pageNumber="146">mobulae</emphasis>
</taxonomicName>
, which possesses a maxilliped with a more robust subchela and has extremely reduced setae on the caudal rami. 
<taxonomicName id="FEB8A0D11905AF4BED9EF868FA79F82C" box="[1183,1436,1949,1973]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="deborahae">
<emphasis id="0BCC07401905AF4BED9EF868FA79F82C" box="[1183,1436,1949,1973]" italics="true" pageId="9" pageNumber="146">Kroeyerina deborahae</emphasis>
</taxonomicName>
differs from the new species in having a caudal ramus with all setae naked; an almost straight subchela on the maxilliped, a smaller number of elements on the antennule and a proportionally shorter genital complex (45% of total body length). 
<taxonomicName id="FEB8A0D11906AF48E86DFF4BFDD8FF4D" box="[364,573,189,212]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="0BCC07401906AF48E86DFF4BFDD8FF4D" box="[364,573,189,212]" italics="true" pageId="10" pageNumber="147">Kroeyerina nasuta</emphasis>
</taxonomicName>
has a caudal ramus similar to the new species, but can be distinguished from it by possessing a different armature of the antennule and a proportionally larger genital complex (57% of total body length). Additionally, in both 
<taxonomicName id="FEB8A0D11906AF48EB35FEF3FD35FE85" box="[564,720,260,284]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="deborahae">
<emphasis id="0BCC07401906AF48EB35FEF3FDA2FE85" box="[564,583,261,284]" italics="true" pageId="10" pageNumber="147">K</emphasis>
. 
<emphasis id="0BCC07401906AF48EB59FEF2FD35FE85" box="[600,720,260,284]" italics="true" pageId="10" pageNumber="147">deborahae</emphasis>
</taxonomicName>
and 
<taxonomicName id="FEB8A0D11906AF48EA0CFEF3FC98FE85" box="[781,893,261,284]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="0BCC07401906AF48EA0CFEF3FCC5FE85" box="[781,800,261,284]" italics="true" pageId="10" pageNumber="147">K</emphasis>
. 
<emphasis id="0BCC07401906AF48EA33FEF3FC98FE85" box="[818,893,261,284]" italics="true" pageId="10" pageNumber="147">nasuta</emphasis>
</taxonomicName>
, the chela of the antenna which, when closed, leaves no gap between the corpus and the claw. The claw bears stout spines (
<bibRefCitation id="5D29A6A31906AF48ED39FEDEFB27FED9" author="Deets" box="[1080,1218,296,320]" pageId="10" pageNumber="147" refString="Deets, G. B. (1987) Phylogenetic analysis and revision of Kroeyerina Wilson, 1932 (Siphonostomatoida: Kroyeriidae), copepods parasitic on chondrichthyans, with descriptions of four new species and the erection of a new genus, Prokroyeria. Canadian Journal of Zoology, 65, 2121 - 2148. http: // dx. doi. org / 10.1139 / z 87 - 327" type="journal article" year="1987">Deets 1987</bibRefCitation>
). The males of 
<taxonomicName id="FEB8A0D11906AF48EC83FEDFFEEAFEFD" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="deborahae">
<emphasis id="0BCC07401906AF48EC83FEDFFA70FED9" box="[1410,1429,297,320]" italics="true" pageId="10" pageNumber="147">K</emphasis>
. 
<emphasis id="0BCC07401906AF48E996FEBAFEEAFEFD" box="[151,271,332,356]" italics="true" pageId="10" pageNumber="147">deborahae</emphasis>
</taxonomicName>
and 
<taxonomicName id="FEB8A0D11906AF48E844FEBBFE54FEFD" box="[325,433,333,356]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="0BCC07401906AF48E844FEBBFEBDFEFD" box="[325,344,333,356]" italics="true" pageId="10" pageNumber="147">K</emphasis>
. 
<emphasis id="0BCC07401906AF48E867FEBBFE54FEFD" box="[358,433,333,356]" italics="true" pageId="10" pageNumber="147">nasuta</emphasis>
</taxonomicName>
are unknown.
</paragraph>
<paragraph id="3907DB521906AF48E9C6FE99FB91FE6C" blockId="10.[151,1436,151,501]" pageId="10" pageNumber="147">
Based on these differences a new species, 
<taxonomicName id="FEB8A0D11906AF48EBACFE86FC31FE11" box="[685,980,368,392]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="sudamericana" status="sp. nov.">
<emphasis id="0BCC07401906AF48EBACFE86FC31FE11" box="[685,980,368,392]" italics="true" pageId="10" pageNumber="147">Kroeyerina sudamericana</emphasis>
</taxonomicName>
<emphasis id="0BCC07401906AF48EADDFE99FBD2FE11" bold="true" box="[988,1079,367,392]" pageId="10" pageNumber="147">
<taxonomicNameLabel id="10FFBA3B1906AF48EADDFE99FBD2FE11" box="[988,1079,367,392]" pageId="10" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is proposed. This new species represents the first record of 
<emphasis id="0BCC07401906AF48EB05FE63FD66FE35" box="[516,643,405,428]" italics="true" pageId="10" pageNumber="147">Kroeyerina</emphasis>
parasitizing members of the 
<taxonomicName id="FEB8A0D11906AF48EAF2FE62FAB3FE34" authority="(Rajiformes)" baseAuthorityName="Rajiformes" box="[1011,1366,404,429]" class="Elasmobranchii" family="Arhynchobatidae" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="family">Arhynchobatidae (Rajiformes)</taxonomicName>
. It is noteworthy that the hosts of 
<taxonomicName id="FEB8A0D11906AF48E8D9FE4FFDA1FE49" box="[472,580,440,464]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="0BCC07401906AF48E8D9FE4FFE0EFE49" box="[472,491,441,464]" italics="true" pageId="10" pageNumber="147">K</emphasis>
. 
<emphasis id="0BCC07401906AF48E8F8FE4FFDA1FE49" box="[505,580,441,464]" italics="true" pageId="10" pageNumber="147">nasuta</emphasis>
</taxonomicName>
and 
<taxonomicName id="FEB8A0D11906AF48EB7AFE4FFD19FE49" box="[635,764,440,464]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="mobulae">
<emphasis id="0BCC07401906AF48EB7AFE4FFD6BFE49" box="[635,654,441,464]" italics="true" pageId="10" pageNumber="147">K</emphasis>
. 
<emphasis id="0BCC07401906AF48EB9DFE4EFD19FE49" box="[668,764,440,464]" italics="true" pageId="10" pageNumber="147">mobulae</emphasis>
</taxonomicName>
belong to the order 
<taxonomicName id="FEB8A0D11906AF48EAE1FE41FB7AFE49" box="[992,1183,439,464]" class="Elasmobranchii" kingdom="Animalia" order="Myliobatiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="order">Myliobatiformes</taxonomicName>
, and although the host of 
<taxonomicName id="FEB8A0D11906AF48E9B4FE2BFEABFE6D" box="[181,334,476,500]" class="Elasmobranchii" family="Arhynchobatidae" genus="Kroeyerina" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="deborahae">
<emphasis id="0BCC07401906AF48E9B4FE2BFF2DFE6D" box="[181,200,477,500]" italics="true" pageId="10" pageNumber="147">K</emphasis>
. 
<emphasis id="0BCC07401906AF48E9D7FE2AFEABFE6D" box="[214,334,476,500]" italics="true" pageId="10" pageNumber="147">deborahae</emphasis>
</taxonomicName>
belongs to the 
<taxonomicName id="FEB8A0D11906AF48E8F8FE2AFD9CFE6C" box="[505,633,476,501]" class="Elasmobranchii" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="147" phylum="Chordata" rank="order">Rajiformes</taxonomicName>
, it is a guitarfish of the family Rhynobatidae.
</paragraph>
</subSubSection>
</treatment>
</document>