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<mods:title id="2432A90C146BEEA888A5973D4D574EF6">Hydroporus sejilashan sp. n., a new diving beetle of the acutangulus - complex from Xizang, China (Qinghai-Tibet Plateau), and notes on other taxa of the genus (Coleoptera, Dytiscidae, Hydroporinae)</mods:title>
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<treatment id="952587BC1C7BFF94FF32FCA66037F9F8" ID-DOI="http://doi.org/10.5281/zenodo.5914733" ID-GBIF-Taxon="119510197" ID-Zenodo-Dep="5914733" LSID="urn:lsid:plazi:treatment:952587BC1C7BFF94FF32FCA66037F9F8" httpUri="http://treatment.plazi.org/id/952587BC1C7BFF94FF32FCA66037F9F8" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
<subSubSection id="559665211C7BFF95FF32FCA66636FC0D" box="[151,585,871,897]" pageId="6" pageNumber="61" type="nomenclature">
<paragraph id="1D3336AA1C7BFF95FF32FCA66636FC0D" blockId="6.[151,585,871,897]" box="[151,585,871,897]" pageId="6" pageNumber="61">
<heading id="467B81C61C7BFF95FF32FCA66636FC0D" bold="true" box="[151,585,871,897]" fontSize="11" level="1" pageId="6" pageNumber="61" reason="1">
<taxonomicName id="DA8C4D291C7BFF95FF32FCA66636FC0D" ID-CoL="3N8YJ" authority="Zaitzev, 1953" authorityName="Zaitzev" authorityYear="1953" box="[151,585,871,897]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7BFF95FF32FCA66636FC0D" bold="true" box="[151,585,871,897]" pageId="6" pageNumber="61">
<emphasis id="2FF8EAB81C7BFF95FF32FCA665E2FC0D" bold="true" box="[151,413,871,897]" italics="true" pageId="6" pageNumber="61">Hydroporus tibetanus</emphasis>
<bibRefCitation id="791D4B5B1C7BFF95FE01FCA66636FC0D" author="Zaitzev" box="[420,585,871,897]" pageId="6" pageNumber="61" refString="Zaitzev, F. A. (1953) Nasekomye zhestkokrylye. Plavuntsovye ivertyachki. Fauna SSSR, 58, 1 - 376. [in Russian]" type="journal article" year="1953">Zaitzev, 1953</bibRefCitation>
</emphasis>
</taxonomicName>
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<paragraph id="1D3336AA1C7BFF95FF32FC6F67F6F9C2" blockId="6.[151,1437,942,2010]" pageId="6" pageNumber="61">
<taxonomicName id="DA8C4D291C7BFF95FF32FC6F65F1FC4A" box="[151,398,942,966]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7BFF95FF32FC6F65F1FC4A" box="[151,398,942,966]" italics="true" pageId="6" pageNumber="61">Hydroporus tibetanus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="791D4B5B1C7BFF95FDC2FC6F6775FC4A" author="Zaitzev" box="[615,778,942,966]" pageId="6" pageNumber="61" refString="Zaitzev, F. A. (1953) Nasekomye zhestkokrylye. Plavuntsovye ivertyachki. Fauna SSSR, 58, 1 - 376. [in Russian]" type="journal article" year="1953">Zaitzev (1953</bibRefCitation>
: 169; see also the English translation
<bibRefCitation id="791D4B5B1C7BFF95FB67FC6F6124FC4A" author="Zaitzev" box="[1218,1371,942,966]" pageId="6" pageNumber="61" refString="Zaitzev, F. A. (1972) Fauna of the U. S. S. R. Coleoptera. Families: Amphizoidae, Hygrobiidae, Haliplidae, Dytiscidae, Gyrinidae. Jerusalem: Israel Program for Scientific Translations, 401 pp." type="book" year="1972">Zaitzev 1972</bibRefCitation>
: 179) from Amdo (
<collectingCountry id="659B763A1C7BFF95FE89FC13650CFC66" box="[300,371,978,1002]" name="China" pageId="6" pageNumber="61">China</collectingCountry>
, Qinghai [Tibet]; ca.
<geoCoordinate id="78B8506D1C7BFF95FDCDFC1366C4FC66" box="[616,699,978,1002]" direction="north" orientation="latitude" pageId="6" pageNumber="61" precision="555" value="32.27">32.27N</geoCoordinate>
<geoCoordinate id="78B8506D1C7BFF95FD61FC136768FC66" box="[708,791,978,1002]" direction="east" orientation="longitude" pageId="6" pageNumber="61" precision="555" value="91.68">91.68E</geoCoordinate>
; altitude ca.
<quantity id="DA749B4F1C7BFF95FC0CFC13607DFC66" box="[937,1026,978,1002]" metricMagnitude="3" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="61" unit="m" value="4700.0">4700 m</quantity>
; star &quot;a&quot; in
<figureCitation id="85B72A2F1C7BFF95FB2FFC13616EFC66" box="[1162,1297,978,1002]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="10.[151,250,932,955]" captionStartId-1="10.[151,250,1793,1816]" captionTargetBox-0="[264,1322,233,911]" captionTargetBox-1="[265,1321,1099,1772]" captionTargetId-0="figure@10.[264,1322,233,911]" captionTargetId-1="figure@10.[265,1321,1099,1772]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionText-0="FIGURE 13. Distribution of: Hydroporus acutangulus (circles), Hydroporus polaris (squares), Hydroporus tuvaensis (triangle), Hydroporus tibetanus (stars), and Hydroporus sejilashan sp. n. (encircled star); the circle with question mark refers to the type locality of Hydroporus sumakovi (see text). Red squares indicate capitals of countries / provinces or other important and well-known cities." captionText-1="FIGURE 14. Distribution of (enlarged map): Hydroporus tuvaensis (triangle), Hydroporus tibetanus (stars), and Hydroporus sejilashan sp. n. (encircled star). Red squares indicate capitals of countries / provinces or other important and well-known cities." httpUri-0="https://zenodo.org/record/210837/files/figure.png" httpUri-1="https://zenodo.org/record/210838/files/figure.png" pageId="6" pageNumber="61">Figs. 1314</figureCitation>
) as &quot;closely related to&quot;
<taxonomicName id="DA8C4D291C7BFF95FEB6FC3766C9FB82" authority="Fabricius, 1972" authorityName="Fabricius" authorityYear="1972" box="[275,694,1014,1038]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="2FF8EAB81C7BFF95FEB6FC376592FB82" box="[275,493,1014,1038]" italics="true" pageId="6" pageNumber="61">Hydroporus nigrita</emphasis>
(Fabricius, 1972)
</taxonomicName>
and, thus, was treated as member of the
<taxonomicName id="DA8C4D291C7BFF95FB21FC3660AFFB82" box="[1156,1232,1015,1038]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="2FF8EAB81C7BFF95FB21FC3660AFFB82" box="[1156,1232,1015,1038]" italics="true" pageId="6" pageNumber="61">nigrita</emphasis>
</taxonomicName>
-group by
<bibRefCitation id="791D4B5B1C7BFF95FAE3FC376556FBBE" author="Nilsson" pageId="6" pageNumber="61" refString="Nilsson, A. N. (2001) Dytiscidae. World catalogue of insects. Vol. 3, 1 - 395. Stenstrup: Apollo Books." type="journal article" year="2001">Nilsson (2001: 163)</bibRefCitation>
. Zaitzev seemingly overlooked the shape of the metacoxal lines and the setation on the metatrochanters; he wrote &quot;... dorsum and venter more shining; body markedly more weakly shagreened [compared with
<taxonomicName id="DA8C4D291C7BFF95FEEBFBA265C3FBF6" box="[334,444,1123,1146]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="2FF8EAB81C7BFF95FEEBFBA265C3FBF6" box="[334,444,1123,1146]" italics="true" pageId="6" pageNumber="61">H. nigrita</emphasis>
</taxonomicName>
] ...&quot;
<bibRefCitation id="791D4B5B1C7BFF95FE51FBA36697FBF6" author="Shaverdo" box="[500,744,1122,1146]" pageId="6" pageNumber="61" refString="Shaverdo, H. V. (2004) Revision of the nigrita - group of Hydroporus Clairville, 1806 (Insecta: Coleoptera: Dytiscidae). Annalen des Naturhistorischen Museums in Wien, 105 B, 217 - 263." type="journal article" year="2004">Shaverdo (2004: 260)</bibRefCitation>
studied the two
<typeStatus id="C23788081C7BFF95FC05FBA2607DFBF7" box="[928,1026,1123,1147]" pageId="6" pageNumber="61" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="DA8C4D291C7BFF95FB82FBA260CFFBF6" box="[1063,1200,1122,1146]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7BFF95FB82FBA260CFFBF6" box="[1063,1200,1122,1146]" italics="true" pageId="6" pageNumber="61">H. tibetanus</emphasis>
</taxonomicName>
(ZISP), designated a male
<typeStatus id="C23788081C7BFF95FF73FB476542FB12" box="[214,317,1158,1182]" pageId="6" pageNumber="61" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="85B72A2F1C7BFF95FEE8FB4765EDFB12" box="[333,402,1158,1182]" captionStart="FIGURES 1 5" captionStartId="2.[151,264,1824,1847]" captionTargetBox="[284,1307,543,1722]" captionTargetId="figure@2.[264,1323,523,1809]" captionTargetPageId="2" captionText="FIGURES 1 5. Habitus of: (1) Hydroporus acutangulus, (2) Hydroporus polaris, (3) Hydroporus tuvaensis, (4) Hydroporus tibetanus (lectotype), (5) Hydroporus sejilashan sp. n." httpUri="https://zenodo.org/record/210834/files/figure.png" pageId="6" pageNumber="61">Fig. 4</figureCitation>
) and found out that—due to the widely smooth elytra—this species belongs to the
<taxonomicName id="DA8C4D291C7BFF95FAEDFB4761EDFB12" box="[1352,1426,1158,1182]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="2FF8EAB81C7BFF95FAEDFB4761EDFB12" box="[1352,1426,1158,1182]" italics="true" pageId="6" pageNumber="61">planus</emphasis>
</taxonomicName>
- group and, in particular, to the
<taxonomicName id="DA8C4D291C7BFF95FE54FB6B6605FB4E" box="[497,634,1194,1218]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="2FF8EAB81C7BFF95FE54FB6B6605FB4E" box="[497,634,1194,1218]" italics="true" pageId="6" pageNumber="61">acutangulus</emphasis>
</taxonomicName>
-complex because of the parallel metacoxal lines and the setal fringe on each metatrochanter. We have also studied the
<typeStatus id="C23788081C7BFF95FD16FB0E6691FB6B" box="[691,750,1231,1255]" pageId="6" pageNumber="61">types</typeStatus>
of this taxon and can confirm Shaverdo's observations, but want to add that the base of the median lobe as well as the right paramere of the
<typeStatus id="C23788081C7BFF95FBF2FB3360C1FA86" box="[1111,1214,1266,1290]" pageId="6" pageNumber="61" type="lectotype">lectotype</typeStatus>
are both damaged. Additionally, it must be mentioned that the
<typeStatus id="C23788081C7BFF95FDDEFAD7669DFAA2" box="[635,738,1302,1326]" pageId="6" pageNumber="61" type="lectotype">lectotype</typeStatus>
lacks the posterior claw of the left protarsus (the anterior claw is present and clearly not strongly curved near base), the anterior claw of the right protarsus, the last three tarsomeres of the left mesotarsus, the last three tarsomeres of the left metatarsus and the complete right metatarsus. We want to point to the fact that the
<typeStatus id="C23788081C7BFF95FD9DFA4366E0FA16" box="[568,671,1410,1434]" pageId="6" pageNumber="61" type="lectotype">lectotype</typeStatus>
is reticulated more or less as typical
<taxonomicName id="DA8C4D291C7BFF95FBE2FA42608CFA16" box="[1095,1267,1410,1434]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="2FF8EAB81C7BFF95FBE2FA42608CFA16" box="[1095,1267,1410,1434]" italics="true" pageId="6" pageNumber="61">H. acutangulus</emphasis>
</taxonomicName>
: entire surface shiny; pronotum smooth on disc, reticulate laterally; elytra smooth in first third, behind with some indistinct short lines between punctures, before apex with distinct small meshes, but also here not matted. In the female
<typeStatus id="C23788081C7BFF95FF32FA2F6551F98A" box="[151,302,1518,1542]" pageId="6" pageNumber="61" type="paralectotype">paralectotype</typeStatus>
the elytral reticulation with small meshes is already present in the second third and becomes more impressed towards the apex; the smooth areas on the pronotum are even more extended than in the male. The entire upper surface appears by no means less shiny than that of the male.
</paragraph>
<paragraph id="1D3336AA1C7BFF94FF62F99B657FFE90" blockId="6.[151,1437,942,2010]" lastBlockId="7.[151,1437,151,428]" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
After Zaitzev's description (1953), no subsequent specimens of
<taxonomicName id="DA8C4D291C7BFF95FC3DF99A605DF9FE" box="[920,1058,1626,1650]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7BFF95FC3DF99A605DF9FE" box="[920,1058,1626,1650]" italics="true" pageId="6" pageNumber="61">H. tibetanus</emphasis>
</taxonomicName>
have been collected. The species was only listed in some catalogues (
<bibRefCitation id="791D4B5B1C7BFF95FD8CF9BF66E5F91A" author="Hua" box="[553,666,1662,1686]" pageId="6" pageNumber="61" refString="Hua, L. - Z. (2002) List of Chinese insects. Vol. 2. Guangzhou: Zhongshan University Press, 612 pp." type="book" year="2002">Hua 2002</bibRefCitation>
, Nilsson 1995, 2001, 2003, 2011,
<bibRefCitation id="791D4B5B1C7BFF95FBBCF9BF6178F91A" author="Nilsson" box="[1049,1287,1662,1687]" pageId="6" pageNumber="61" refString="Nilsson, A. N. &amp; Fery, H. (2006) World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 76, 55 - 74." type="journal article" year="2006">Nilsson &amp; Fery 2006</bibRefCitation>
) or treated in overviews of members of the
<taxonomicName id="DA8C4D291C7BFF95FE55F9626643F936" box="[496,572,1699,1722]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="2FF8EAB81C7BFF95FE55F9626643F936" box="[496,572,1699,1722]" italics="true" pageId="6" pageNumber="61">nigrita</emphasis>
</taxonomicName>
- or
<taxonomicName id="DA8C4D291C7BFF95FDD5F96366C4F936" box="[624,699,1698,1722]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="2FF8EAB81C7BFF95FDD5F96366C4F936" box="[624,699,1698,1722]" italics="true" pageId="6" pageNumber="61">planus</emphasis>
</taxonomicName>
-groups (
<bibRefCitation id="791D4B5B1C7BFF95FC84F9636781F936" author="Fery" box="[801,1022,1698,1722]" pageId="6" pageNumber="61" refString="Fery, H. &amp; Pesic, V. (2006) Hydroporus macedonicus nov. spec., a new member of the Hydroporus planus - group (Coleoptera, Dytiscidae). Linzer Biologische Beitrage, 38 (1), 595 - 604." type="journal article" year="2006">Fery &amp; Pesic 2006</bibRefCitation>
,
<bibRefCitation id="791D4B5B1C7BFF95FBAEF96360ACF936" author="Gueorguiev" box="[1035,1235,1698,1722]" pageId="6" pageNumber="61" refString="Gueorguiev, V. B. (1966) Un nouveau Hydroporus (Coleoptera, Dytiscidae) de Thrace - Bulgarie meridionale. Fauna na Trakija, 3, 71 - 75. [in Bulgarian, with French Resume]" type="journal article" year="1966">Guéorguiev 1966</bibRefCitation>
,
<bibRefCitation id="791D4B5B1C7BFF95FB45F96361EFF936" author="Shaverdo" box="[1248,1424,1698,1722]" pageId="6" pageNumber="61" refString="Shaverdo, H. V. (2004) Revision of the nigrita - group of Hydroporus Clairville, 1806 (Insecta: Coleoptera: Dytiscidae). Annalen des Naturhistorischen Museums in Wien, 105 B, 217 - 263." type="journal article" year="2004">Shaverdo 2004</bibRefCitation>
). Recently, however, we have studied several specimens which we could determine as this species. The collecting data are as follows:
<date id="6932106A1C7BFF95FE22F92B6593F88E" box="[391,492,1770,1794]" pageId="6" pageNumber="61" value="1999-07-03">3.7.1999</date>
, Sichuan, Ganzi (= Garzê) Tibetan Autonomous Region, Batang County, Shaluli Mountains,
<quantity id="DA749B4F1C7BFF95FE84F8CF6513F8AA" box="[289,364,1806,1830]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="6" pageNumber="61" unit="km" value="57.0">57 km</quantity>
NE Batang, pools, ca.
<quantity id="DA749B4F1C7BFF95FD23F8CF669EF8AA" box="[646,737,1806,1830]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="61" unit="m" value="4500.0">4500 m</quantity>
, A. Pütz leg. (CAP, CHF) (star &quot;b&quot; in
<figureCitation id="85B72A2F1C7BFF95FB11F8CF613EF8AA" box="[1204,1345,1806,1830]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="10.[151,250,932,955]" captionStartId-1="10.[151,250,1793,1816]" captionTargetBox-0="[264,1322,233,911]" captionTargetBox-1="[265,1321,1099,1772]" captionTargetId-0="figure@10.[264,1322,233,911]" captionTargetId-1="figure@10.[265,1321,1099,1772]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionText-0="FIGURE 13. Distribution of: Hydroporus acutangulus (circles), Hydroporus polaris (squares), Hydroporus tuvaensis (triangle), Hydroporus tibetanus (stars), and Hydroporus sejilashan sp. n. (encircled star); the circle with question mark refers to the type locality of Hydroporus sumakovi (see text). Red squares indicate capitals of countries / provinces or other important and well-known cities." captionText-1="FIGURE 14. Distribution of (enlarged map): Hydroporus tuvaensis (triangle), Hydroporus tibetanus (stars), and Hydroporus sejilashan sp. n. (encircled star). Red squares indicate capitals of countries / provinces or other important and well-known cities." httpUri-0="https://zenodo.org/record/210837/files/figure.png" httpUri-1="https://zenodo.org/record/210838/files/figure.png" pageId="6" pageNumber="61">Figs. 1314</figureCitation>
). These specimens are similar to the lecto- and
<typeStatus id="C23788081C7BFF95FDC4F8F36687F8C6" box="[609,760,1842,1866]" pageId="6" pageNumber="61" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="DA8C4D291C7BFF95FC86F8F267D0F8C6" box="[803,943,1842,1866]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7BFF95FC86F8F267D0F8C6" box="[803,943,1842,1866]" italics="true" pageId="6" pageNumber="61">H. tibetanus</emphasis>
</taxonomicName>
in most respects, in particular, their upper surface is shiny in both sexes. They are, however, somewhat darker than the
<typeStatus id="C23788081C7BFF95FC48F8966053F8E3" box="[1005,1068,1879,1903]" pageId="6" pageNumber="61">types</typeStatus>
. Some females are reticulated on the entire elytra and large parts of the pronotum. Other females have the upper surface reticulated as the female
<typeStatus id="C23788081C7BFF95FF32F85F6551F83A" box="[151,302,1950,1974]" pageId="6" pageNumber="61" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="DA8C4D291C7BFF95FEFFF8616598F83A" box="[346,487,1950,1975]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7BFF95FEFFF8616598F83A" box="[346,487,1950,1975]" italics="true" pageId="6" pageNumber="61">H. tibetanus</emphasis>
</taxonomicName>
. The median lobe of the males in ventral view (
<figureCitation id="85B72A2F1C7BFF95FB80F85F6015F83A" box="[1061,1130,1950,1974]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1898,1921]" captionTargetBox="[241,1330,811,1851]" captionTargetId="figure@5.[235,1352,803,1877]" captionTargetPageId="5" captionText="FIGURES 6 10. (6) Median lobe of Hydroporus sejilashan sp. n. in ventral (a) and in lateral view (b); (7) left paramere of Hydroporus sejilashan sp. n.; (8) median lobe of Hydroporus tibetanus in ventral view: (a) lectotype, (b) specimen from Ganzi; same in lateral view: (c) specimen from Ganzi; left paramere of Hydroporus tibetanus; (9) specimen from Ganzi, (10) lectotype." httpUri="https://zenodo.org/record/210835/files/figure.png" pageId="6" pageNumber="61">Fig. 8</figureCitation>
b) is similar to that of the
<typeStatus id="C23788081C7BFF95FF32F8036481F856" box="[151,254,1986,2010]" pageId="6" pageNumber="61" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="DA8C4D291C7BFF95FE80F80265D0F856" box="[293,431,1986,2010]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7BFF95FE80F80265D0F856" box="[293,431,1986,2010]" italics="true" pageId="6" pageNumber="61">H. tibetanus</emphasis>
</taxonomicName>
(
<figureCitation id="85B72A2F1C7BFF95FE64F803667BF856" box="[449,516,1986,2010]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1898,1921]" captionTargetBox="[241,1330,811,1851]" captionTargetId="figure@5.[235,1352,803,1877]" captionTargetPageId="5" captionText="FIGURES 6 10. (6) Median lobe of Hydroporus sejilashan sp. n. in ventral (a) and in lateral view (b); (7) left paramere of Hydroporus sejilashan sp. n.; (8) median lobe of Hydroporus tibetanus in ventral view: (a) lectotype, (b) specimen from Ganzi; same in lateral view: (c) specimen from Ganzi; left paramere of Hydroporus tibetanus; (9) specimen from Ganzi, (10) lectotype." httpUri="https://zenodo.org/record/210835/files/figure.png" pageId="6" pageNumber="61">Fig. 8</figureCitation>
a). We are not absolutely sure about the identity of Pütz' specimens, but since no other material from the region around Amdo has become available, we assume that this population belongs to
<taxonomicName id="DA8C4D291C7AFF94FA24FF586481FF58" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FA24FF586481FF58" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
. Measurements:
<typeStatus id="C23788081C7AFF94FE61FF7C664BFF59" box="[452,564,189,213]" pageId="7" pageNumber="62" type="lectotype">Lectotype</typeStatus>
(male): TL:
<quantity id="DA749B4F1C7AFF94FD68FF7D6758FF58" box="[717,807,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="62" unit="mm" value="3.2">3.2 mm</quantity>
, MW:
<quantity id="DA749B4F1C7AFF94FCDFFF7D67AAFF58" box="[890,981,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="62" unit="mm" value="1.6">1.6 mm</quantity>
, TL/MW: 2.00;
<typeStatus id="C23788081C7AFF94FB30FF7D6153FF58" box="[1173,1324,188,212]" pageId="7" pageNumber="62" type="paralectotype">paralectotype</typeStatus>
(female): TL:
<quantity id="DA749B4F1C7AFF94FF62FF1E655FFF7B" box="[199,288,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="62" unit="mm" value="3.2">3.2 mm</quantity>
, MW:
<quantity id="DA749B4F1C7AFF94FECEFF1E65ADFF7B" box="[363,466,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="7" pageNumber="62" unit="mm" value="1.65">1.65 mm</quantity>
, TL/MW: 1.94; specimens from Ganzi: TL:
<quantity id="DA749B4F1C7AFF94FC64FF1E6026FF7B" box="[961,1113,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="3.125" metricValueMax="3.3" metricValueMin="2.95" pageId="7" pageNumber="62" unit="mm" value="3.125" valueMax="3.3" valueMin="2.95">2.953.3 mm</quantity>
, MW:
<quantity id="DA749B4F1C7AFF94FB01FF1E6152FF7B" box="[1188,1325,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="7" pageNumber="62" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 mm</quantity>
, TL/MW: 1.92.03.
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF62FEE96718FE20" blockId="7.[151,1437,151,428]" pageId="7" pageNumber="62">
We prefer not to present a key to species at this time because any key for this group would be rather primitive and not very useful. Construction of a key for this species group should be postponed until the variability of all group members is known sufficiently. We compare below the new species with the
<typeStatus id="C23788081C7AFF94FB9BFEB16006FE04" box="[1086,1145,368,392]" pageId="7" pageNumber="62">types</typeStatus>
of
<taxonomicName id="DA8C4D291C7AFF94FB3BFEB06157FE04" box="[1182,1320,368,392]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FB3BFEB06157FE04" box="[1182,1320,368,392]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
as well as with specimens from the population of
<taxonomicName id="DA8C4D291C7AFF94FDEAFE5466A8FE20" box="[591,727,404,428]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FDEAFE5466A8FE20" box="[591,727,404,428]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
from Ganzi:
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FE1C6026FE78" blockId="7.[151,1113,476,500]" box="[151,1113,476,500]" pageId="7" pageNumber="62">
-
<taxonomicName id="DA8C4D291C7AFF94FF62FE1D65B9FE78" box="[199,454,476,500]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sejilashan" status="sp. nov.">
<emphasis id="2FF8EAB81C7AFF94FF62FE1D65B9FE78" box="[199,454,476,500]" italics="true" pageId="7" pageNumber="62">Hydroporus sejilashan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="34CB57C31C7AFF94FE69FE1C6677FE78" box="[460,520,477,500]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
is larger and slightly less elongate than
<taxonomicName id="DA8C4D291C7AFF94FC6CFE1C602DFE78" box="[969,1106,476,500]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FC6CFE1C602DFE78" box="[969,1106,476,500]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FDE466ABFDD3" blockId="7.[151,1436,548,607]" pageId="7" pageNumber="62">
- In dorsal view the sides of the pronotum are less curved in
<taxonomicName id="DA8C4D291C7AFF94FCC0FDE46785FDB0" box="[869,1018,548,572]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sejilashan" status="sp. nov.">
<emphasis id="2FF8EAB81C7AFF94FCC0FDE46785FDB0" box="[869,1018,548,572]" italics="true" pageId="7" pageNumber="62">H. sejilashan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="34CB57C31C7AFF94FBA7FDE46041FDB0" box="[1026,1086,549,572]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
In
<taxonomicName id="DA8C4D291C7AFF94FBC0FDE46091FDB0" box="[1125,1262,548,572]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FBC0FDE46091FDB0" box="[1125,1262,548,572]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
the base of the pronotum is not broader than the base of the elytra.
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FD51614FFD40" blockId="7.[151,1436,655,716]" pageId="7" pageNumber="62">
- The new species has the entire surface reticulated and matt.
<taxonomicName id="DA8C4D291C7AFF94FCCBFD51601CFD24" box="[878,1123,656,680]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FCCBFD51601CFD24" box="[878,1123,656,680]" italics="true" pageId="7" pageNumber="62">Hydroporus tibetanus</emphasis>
</taxonomicName>
has a shiny surface (also in reticulated areas) and in the males and most females the reticulation of the elytra is restricted to the apex.
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FD3C6737FCF3" blockId="7.[151,1436,764,895]" pageId="7" pageNumber="62">
-
<taxonomicName id="DA8C4D291C7AFF94FF62FD3D65B6FC98" box="[199,457,764,788]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sejilashan" status="sp. nov.">
<emphasis id="2FF8EAB81C7AFF94FF62FD3D65B6FC98" box="[199,457,764,788]" italics="true" pageId="7" pageNumber="62">Hydroporus sejilashan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="34CB57C31C7AFF94FE77FD3C666EFC98" box="[466,529,765,788]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
has the dorsal and ventral surface almost totally black, only the tarsi and first antennomeres are dark brownish. The
<typeStatus id="C23788081C7AFF94FDEBFCDE66CBFCBB" box="[590,692,799,823]" pageId="7" pageNumber="62" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="DA8C4D291C7AFF94FD78FCE06718FCB4" box="[733,871,800,824]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FD78FCE06718FCB4" box="[733,871,800,824]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
has the tarsi somewhat lighter, the antennomeres are all more or less uniformly brown, and the elytra are dark brown; the female
<typeStatus id="C23788081C7AFF94FB89FC8560BCFCD0" box="[1068,1219,836,860]" pageId="7" pageNumber="62" type="paralectotype">paralectotype</typeStatus>
and the specimens from Ganzi, however, are generally darker than the
<typeStatus id="C23788081C7AFF94FD7FFCA6673BFCF3" box="[730,836,871,895]" pageId="7" pageNumber="62" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FC7165E2FBF0" blockId="7.[151,1436,943,1148]" pageId="7" pageNumber="62">
- The punctation of the surface is distinctly coarser in
<taxonomicName id="DA8C4D291C7AFF94FCB5FC7067E7FC44" box="[784,920,944,968]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FCB5FC7067E7FC44" box="[784,920,944,968]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
, and also denser on head; in the posterolateral depression of the pronotum and before the posterior margin (except centrally) it is very coarse and longitudinally deformed; the punctures of the irregular puncture line behind the anterior pronotal margin are also very coarse and in part longitudinally deformed. In the new species, however, the punctation is generally simple and by far not so coarse in the respective areas. Due to the coarser elytral punctation, the elytral puncture lines appear slightly less distinct in
<taxonomicName id="DA8C4D291C7AFF94FEA8FBA465EAFBF0" box="[269,405,1124,1148]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FEA8FBA465EAFBF0" box="[269,405,1124,1148]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FB6C6635FAF4" blockId="7.[151,1436,1196,1400]" pageId="7" pageNumber="62">
- The first three protarsomeres are more dilated in
<taxonomicName id="DA8C4D291C7AFF94FCA5FB6C67E9FB48" box="[768,918,1196,1220]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sejilashan" status="sp. nov.">
<emphasis id="2FF8EAB81C7AFF94FCA5FB6C67E9FB48" box="[768,918,1196,1220]" italics="true" pageId="7" pageNumber="62">H. sejilashan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="34CB57C31C7AFF94FC05FB6C679FFB48" box="[928,992,1197,1220]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
; especially the second protarsomere is almost two times as broad as long and the third one more or less of round shape; the differences between male and female protarsomeres are obvious. In the
<typeStatus id="C23788081C7AFF94FDC8FB3566ACFA80" box="[621,723,1268,1292]" pageId="7" pageNumber="62" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="DA8C4D291C7AFF94FD5FFB3467FBFA80" box="[762,900,1268,1292]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FD5FFB3467FBFA80" box="[762,900,1268,1292]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
the second protarsomere is only one and a half times as broad as long and the third one is more elongate; differences between the protarsomeres of males and females are rather slight. The male mesotarsomeres of the new species are also broader than those of the females and broader than those of
<taxonomicName id="DA8C4D291C7AFF94FE1FFAA0663CFAF4" box="[442,579,1376,1400]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FE1FFAA0663CFAF4" box="[442,579,1376,1400]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FA6967C8FA33" blockId="7.[151,951,1447,1472]" box="[151,951,1447,1472]" pageId="7" pageNumber="62">- Antennomeres five to eleven are slightly longer in the new species.</paragraph>
<paragraph id="1D3336AA1C7AFF94FF32FA316037F9F8" blockId="7.[151,1436,1519,1652]" pageId="7" pageNumber="62">
- The median lobe of
<taxonomicName id="DA8C4D291C7AFF94FE03FA306645F984" box="[422,570,1520,1544]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sejilashan" status="sp. nov.">
<emphasis id="2FF8EAB81C7AFF94FE03FA306645F984" box="[422,570,1520,1544]" italics="true" pageId="7" pageNumber="62">H. sejilashan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="34CB57C31C7AFF94FDE5FA316604F98B" box="[576,635,1520,1543]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
is in ventral view slightly narrower in the apical third and more pointed (compare
<figureCitation id="85B72A2F1C7AFF94FEADF9D5652AF9A0" box="[264,341,1556,1580]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1898,1921]" captionTargetBox="[241,1330,811,1851]" captionTargetId="figure@5.[235,1352,803,1877]" captionTargetPageId="5" captionText="FIGURES 6 10. (6) Median lobe of Hydroporus sejilashan sp. n. in ventral (a) and in lateral view (b); (7) left paramere of Hydroporus sejilashan sp. n.; (8) median lobe of Hydroporus tibetanus in ventral view: (a) lectotype, (b) specimen from Ganzi; same in lateral view: (c) specimen from Ganzi; left paramere of Hydroporus tibetanus; (9) specimen from Ganzi, (10) lectotype." httpUri="https://zenodo.org/record/210835/files/figure.png" pageId="7" pageNumber="62">Figs. 6</figureCitation>
a with 8ab). In lateral view (
<figureCitation id="85B72A2F1C7AFF94FD01F9D56698F9A0" box="[676,743,1556,1580]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1898,1921]" captionTargetBox="[241,1330,811,1851]" captionTargetId="figure@5.[235,1352,803,1877]" captionTargetPageId="5" captionText="FIGURES 6 10. (6) Median lobe of Hydroporus sejilashan sp. n. in ventral (a) and in lateral view (b); (7) left paramere of Hydroporus sejilashan sp. n.; (8) median lobe of Hydroporus tibetanus in ventral view: (a) lectotype, (b) specimen from Ganzi; same in lateral view: (c) specimen from Ganzi; left paramere of Hydroporus tibetanus; (9) specimen from Ganzi, (10) lectotype." httpUri="https://zenodo.org/record/210835/files/figure.png" pageId="7" pageNumber="62">Fig. 6</figureCitation>
b) a comparison is impossible because the median lobe of the
<typeStatus id="C23788081C7AFF94FF32F9F66481F9C3" box="[151,254,1591,1615]" pageId="7" pageNumber="62" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="DA8C4D291C7AFF94FE86F9F865D3F9DC" box="[291,428,1592,1616]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="2FF8EAB81C7AFF94FE86F9F865D3F9DC" box="[291,428,1592,1616]" italics="true" pageId="7" pageNumber="62">H. tibetanus</emphasis>
</taxonomicName>
is partly destroyed. A comparison with that of a male from Ganzi (
<figureCitation id="85B72A2F1C7AFF94FB38F9F660A0F9C3" box="[1181,1247,1591,1615]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1898,1921]" captionTargetBox="[241,1330,811,1851]" captionTargetId="figure@5.[235,1352,803,1877]" captionTargetPageId="5" captionText="FIGURES 6 10. (6) Median lobe of Hydroporus sejilashan sp. n. in ventral (a) and in lateral view (b); (7) left paramere of Hydroporus sejilashan sp. n.; (8) median lobe of Hydroporus tibetanus in ventral view: (a) lectotype, (b) specimen from Ganzi; same in lateral view: (c) specimen from Ganzi; left paramere of Hydroporus tibetanus; (9) specimen from Ganzi, (10) lectotype." httpUri="https://zenodo.org/record/210835/files/figure.png" pageId="7" pageNumber="62">Fig. 8</figureCitation>
c) shows that the lobe of the new species is less bent near the base and slightly broader in apical third.
</paragraph>
</subSubSection>
</treatment>
</document>