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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. &amp; Dowling, Ashley P. G." docDate="2017" docId="6D98F30878C979E4CBA327D09CC84276" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola tysoni Fisher &amp; Dowling, sp. n." docType="treatment" docUuid="3F9D2DE0-5A5E-4055-8EE6-99F6070AA127" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, J. Ray</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Danielle M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skvarla, Michael J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nelson, Whitney A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>701</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>496</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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<treatment ID-GBIF-Taxon="134839691" LSID="urn:lsid:zoobank.org:act:3F9D2DE0-5A5E-4055-8EE6-99F6070AA127" httpUri="http://treatment.plazi.org/id/6D98F30878C979E4CBA327D09CC84276" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/3F9D2DE0-5A5E-4055-8EE6-99F6070AA127" authority="Fisher &amp; Dowling" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola tysoni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tysoni">Torrenticola tysoni Fisher &amp; Dowling</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
HOLOTYPE (♀): from USA, Alabama, Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="34.941944">34°56'31&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-87.82806">87°49'41&quot;W</geoCoordinate>
), 24 Sep 2009, by IM Smith, IMS090121, DNA 2871.
</paragraph>
<paragraph pageId="0" pageNumber="1">
PARATYPES (6 ♀; 5 ♂): Alabama, USA: 1 ♂ (ALLOTYPE) from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="34.941944">34°56'31&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-87.82806">87°49'41&quot;W</geoCoordinate>
), 24 Sep 2009, by IM Smith, IMS090121, DNA 2870
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="34.941944">34°56'31&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-87.82806">87°49'41&quot;W</geoCoordinate>
), 24 Sep 2009, by IM Smith, IMS090121
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ and 2 ♂ from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="34.942223">34°56'32&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-87.82861">87°49'43&quot;W</geoCoordinate>
), 27 Sep 2010, by IM Smith, IMS100163
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="34.941944">34°56'31&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-87.82806">87°49'41&quot;W</geoCoordinate>
), 27 Sep 2010, by IM Smith, IMS100162
<normalizedToken originalValue="•">*</normalizedToken>
Tennessee, USA: 3 ♀ and 1 ♂ from Wayne County, Glenrock Branch Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="35.26389">35°15'50&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-87.626114">87°37'34&quot;W</geoCoordinate>
), 24 Sep 2009, by IM Smith, IMS090124.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type deposition">
<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
<paragraph pageId="0" pageNumber="1">Holotype (♀), allotype (♂), and most paratypes (4 ♀; 2 ♂) deposited in the CNC; other paratypes (2 ♀; 2 ♂) deposited in ACUA.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola tysoni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tysoni">Torrenticola tysoni</taxonomicName>
are similar to other members of the
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Rusetria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rusetria" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Rusetria</taxonomicName>
&quot;Eastern 2-Plates&quot; group (
<taxonomicName lsidName="T. biscutella" pageId="0" pageNumber="1" rank="species" species="biscutella">T. biscutella</taxonomicName>
,
<taxonomicName lsidName="T. caerulea" pageId="0" pageNumber="1" rank="species" species="caerulea">T. caerulea</taxonomicName>
,
<taxonomicName lsidName="T. delicatexa" pageId="0" pageNumber="1" rank="species" species="delicatexa">T. delicatexa</taxonomicName>
,
<taxonomicName lsidName="T. indistincta" pageId="0" pageNumber="1" rank="species" species="indistincta">T. indistincta</taxonomicName>
,
<taxonomicName lsidName="T. malarkeyorum" pageId="0" pageNumber="1" rank="species" species="malarkeyorum">T. malarkeyorum</taxonomicName>
,
<taxonomicName lsidName="T. pendula" pageId="0" pageNumber="1" rank="species" species="pendula">T. pendula</taxonomicName>
,
<taxonomicName lsidName="T. sellersorum" pageId="0" pageNumber="1" rank="species" species="sellersorum">T. sellersorum</taxonomicName>
,
<taxonomicName lsidName="T. ululata" pageId="0" pageNumber="1" rank="species" species="ululata">T. ululata</taxonomicName>
,
<taxonomicName lsidName="T. whitneyae" pageId="0" pageNumber="1" rank="species" species="whitneyae">T. whitneyae</taxonomicName>
,
<taxonomicName lsidName="T. microbiscutella" pageId="0" pageNumber="1" rank="species" species="microbiscutella">T. microbiscutella</taxonomicName>
, and
<taxonomicName lsidName="T. feminellai" pageId="0" pageNumber="1" rank="species" species="feminellai">T. feminellai</taxonomicName>
) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except
<taxonomicName lsidName="T. ululata" pageId="0" pageNumber="1" rank="species" species="ululata">T. ululata</taxonomicName>
and
<taxonomicName lsidName="T. indistincta" pageId="0" pageNumber="1" rank="species" species="indistincta">T. indistincta</taxonomicName>
), and being distributed in the east. It is one of only four Eastern 2-Plates that have dark, bold, bluish-purple coloration (also
<taxonomicName lsidName="T. biscutella" pageId="0" pageNumber="1" rank="species" species="biscutella">T. biscutella</taxonomicName>
,
<taxonomicName lsidName="T. sellersorum" pageId="0" pageNumber="1" rank="species" species="sellersorum">T. sellersorum</taxonomicName>
, and
<taxonomicName lsidName="T. pendula" pageId="0" pageNumber="1" rank="species" species="pendula">T. pendula</taxonomicName>
).
<taxonomicName lsidName="T. tysoni" pageId="0" pageNumber="1" rank="species" species="tysoni">T. tysoni</taxonomicName>
can be further differentiated from other Eastern 2-Plates by having a more elongate rostrum (length/width = ♀ = 3.06-3.31 in
<taxonomicName lsidName="T. tysoni" pageId="0" pageNumber="1" rank="species" species="tysoni">T. tysoni</taxonomicName>
, 2.33-3.0 in others; ♂ = 3.14-3.50 in
<taxonomicName lsidName="T. tysoni" pageId="0" pageNumber="1" rank="species" species="tysoni">T. tysoni</taxonomicName>
, 2.50-3.05 in others), except
<taxonomicName lsidName="T. feminellai" pageId="0" pageNumber="1" rank="species" species="feminellai">T. feminellai</taxonomicName>
(3.05-3.38) and female
<taxonomicName lsidName="T. pendula" pageId="0" pageNumber="1" rank="species" species="pendula">T. pendula</taxonomicName>
(3.0-3.06).
<taxonomicName lsidName="T. tysoni" pageId="0" pageNumber="1" rank="species" species="tysoni">T. tysoni</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. feminellai" pageId="0" pageNumber="1" rank="species" species="feminellai">T. feminellai</taxonomicName>
and
<taxonomicName lsidName="T. pendula" pageId="0" pageNumber="1" rank="species" species="pendula">T. pendula</taxonomicName>
by dorsal coloration and pattern.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female (Figure 259) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (610-670 (670) long; 450-475 (475) wide) ovoid with bold bluish-purple coloration separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (125-137.5 (135) long; 42.5-52.5 (52.5) wide). Anterio-lateral platelets (162.5-175 (172.5) long; 55-67.5 (67.5) wide) fused to dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 320-330 (330)). Dorsal plate proportions: dorsum length/width 1.35-1.41 (1.41); dorsal width/distance between Dgl-4 1.38-1.44 (1.44); anterio-medial platelet length/width 2.50-3.18 (2.57); anterio-lateral platelet length/width 2.56-3.18 (2.56); anterio-lateral/anterio-medial length 1.18-1.40 (1.28).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (310-327.5 (327.5) long (ventral); 230-245 (245) long (dorsal); 130-135 (135) tall) mostly colorless. Rostrum (125-132.5 (130) long; 40-42.5 (42.5) wide). Chelicerae (315-380 (350) long) with curved fangs (57.5-60 (60) long). Subcapitular proportions: ventral length/height 2.30-2.46 (2.43); rostrum length/width 3.06-3.31 (3.06). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5-47.5 (45) long); femur (112.5-120 (120) long); genu (62.5-70 (70) long); tibia (80-90 (87.5) long; 22.5-25 (25) wide); tarsus (17.5-22.5 (20) long). Palpomere proportions: femur/genu 1.61-1.80 (1.71); tibia/femur 0.70-0.80 (0.73); tibia length/width 3.50-3.67 (3.50).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter - (715-790 (790) long; 505-560 (540) wide) with bold bluish-purple coloration. Gnathosomal bay (162.5-172.5 (172.5) long; 72.5-100 (85) wide). Cxgl-4 subapical. Medial suture (13.75-22.5 (22.5) long). Genital plates (175-185 (180) long; 152.5-155 (155) wide). Additional measurements: Cx-1 (280-315 (310) long (total); 115-145 (140) long (medial)); Cx-3 (310-350 (335) wide); anterior venter (150-172.5 (172.5) long). Ventral proportions: gnathosomal bay length/width 1.65-2.24 (2.03); anterior venter/genital field length 0.81-0.99 (0.96); anterior venter length/genital field width 0.97-1.11 (1.11); anterior venter/medial suture 7.67-11.27 (7.67).</paragraph>
<paragraph pageId="0" pageNumber="1">Male (Figure 260) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (430-485 (460) long; 310-340 (320) wide) ovoid with bold bluish-purple coloration separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (97.5-127.5 (127.5) long; 35-40 (40) wide). Anterio-lateral platelets (127.5-137.5 (127.5) long; 42.5-50 (47.5) wide) fused to dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 235-245 (240)). Dorsal plate proportions: dorsum length/width 1.39-1.48 (1.44); dorsal width/distance between Dgl-4 1.32-1.39 (1.33); anterio-medial platelet length/width 2.60-3.19 (3.19); anterio-lateral platelet length/width 2.65-3.00 (2.68); anterio-lateral/anterio-medial length 1.00-1.31 (1.00).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (250-265 (250) long (ventral); 192.5-200 (195) long (dorsal); 77.5-95 (90) tall) mostly colorless. Rostrum (100-110 (100) long; 30-35 (30) wide). Chelicerae (237.5-260 (250) long) with curved fangs (45-47.5 (45) long). Subcapitular proportions: ventral length/height 2.76-3.23 (2.78); rostrum length/width 3.14-3.50 (3.33). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (33.75-48.75 (37.5) long); femur (87.5-97.5 (92.5) long); genu (52.5-62.5 (57.5) long); tibia (67.5-75 (75) long; 20-21.25 (21.25) wide); tarsus (15-17.5 (15) long). Palpomere proportions: femur/genu 1.56-1.68 (1.61); tibia/femur 0.72-0.81 (0.81); tibia length/width 3.18-3.63 (3.53).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter- (540-585 (560) long; 350-410 (360) wide) with bold bluish-purple coloration. Gnathosomal bay (110-122.5 (112.5) long; 62.5-67.5 (62.5) wide). Cxgl-4 subapical. Medial suture (85-115 (85) long). Genital plates (110-117.5 (110) long; 87.5-92.5 (92.5) wide). Additional measurements: Cx-1 (240-250 (240) long (total); 115-127.5 (122.5) long (medial)); Cx-3 (250-290 (265) wide); anterior venter (227.5-250 (232.5) long). Ventral proportions: gnathosomal bay length/width 1.69-1.85 (1.80); anterior venter/genital field length 1.98-2.13 (2.11); anterior venter length/genital field width 2.51-2.86 (2.51); anterior venter/medial suture 2.17-2.74 (2.74).</paragraph>
<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Specific epithet (
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea tysoni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tysoni">tysoni</taxonomicName>
) named in honor of Neil Degrasse Tyson for his efforts in popularizing cosmology and science in general with Cosmos: A Spacetime Odyssey (2014), which was a worthy update to Carl
<normalizedToken originalValue="Sagans">Sagan's</normalizedToken>
Cosmos: A Personal Voyage (1980).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Known only from several localities along Natchez Trace Parkway (Figure 258).</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 258.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola tysoni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tysoni">Torrenticola tysoni</taxonomicName>
sp. n. distribution.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 259.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola tysoni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tysoni">Torrenticola tysoni</taxonomicName>
sp. n. female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 260.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola tysoni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tysoni">Torrenticola tysoni</taxonomicName>
sp. n. male: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola tysoni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tysoni">Torrenticola tysoni</taxonomicName>
groups with other members of the
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Rusetria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rusetria" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Rusetria</taxonomicName>
Complex with high support. Specimens of this species from Tennessee and Alabama are less than 1% different in COI sequence from each other and the specimen from Arkansas is less than 2% different from those. This species is greater than 10% different from sister species.
</paragraph>
<paragraph pageId="0" pageNumber="1">
In all analyses,
<taxonomicName lsidName="T. tysoni" pageId="0" pageNumber="1" rank="species" species="tysoni">T. tysoni</taxonomicName>
groups with two other species (
<taxonomicName lsidName="T. sellersorum" pageId="0" pageNumber="1" rank="species" species="sellersorum">T. sellersorum</taxonomicName>
and
<taxonomicName lsidName="T. pendula" pageId="0" pageNumber="1" rank="species" species="pendula">T. pendula</taxonomicName>
) as a paraphyletic grade at the base of the Eastern 2-Plate Identification Group. Based upon overall similarity, lateral platelets fused with the dorsal shield, and distribution, we were able to place this species within the Eastern 2-Plate Identification Group.
</paragraph>
<paragraph pageId="0" pageNumber="1">This species hypothesis is supported by low COI divergence within the species (0-2%) and high divergence between species (3-15%), and by the morphological characters outlined in the diagnosis.</paragraph>
</subSubSection>
</treatment>
</document>