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<document id="8053FAEA630716D380AC6DA17F53CB96" ID-CLB-Dataset="34707" ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="6D76C5CFDAD5B210C708408C691B407F" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus fossipygus Wenzel 1944" docType="treatment" docVersion="5" lastPageNumber="245" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="242" updateTime="1732934305285" updateUser="ExternalLinkService">
<mods:mods id="9BC38D357566B125A6804635E19E406B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1D56B11B782040035BE2AEB4B89054AC">A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="4596A19A6DD232966042FDA1A53F981B">Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name id="E4098F8E3AD984B110B92DB1792B6748" type="personal">
<mods:role id="AF14D73B57E0F4898FB0D9053B2D53D7">
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</mods:role>
<mods:namePart id="89158636163C5E2DCBD59C31AC8BB71F">Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9C1BD7536A32EECEC0278AE025B6972F">text</mods:typeOfResource>
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<mods:part id="17FF8669D0BFD1DC2BE9C9F0CCEC9059">
<mods:date id="34042D1541B27283188E546512F89A60">2013</mods:date>
<mods:detail id="F51153AC57A4538103A7F52EF34DAE8F" type="volume">
<mods:number id="516512B2C307FB8CB4FCBE85AE352827">271</mods:number>
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<mods:classification id="3D69DDF732DA5494F4853A319E20149D">journal article</mods:classification>
<mods:identifier id="D2D7CC9B99B604E81E49C6E232F0C47C" type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier id="68A0CA9E6682DE820AD48671F6877CE6" type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment id="6D76C5CFDAD5B210C708408C691B407F" ID-GBIF-Taxon="152040943" LSID="urn:lsid:plazi:treatment:6D76C5CFDAD5B210C708408C691B407F" httpUri="http://treatment.plazi.org/id/6D76C5CFDAD5B210C708408C691B407F" lastPageId="244" lastPageNumber="245" pageId="241" pageNumber="242" scope_family="Histeridae" scope_order="Coleoptera" scope_tribe="Exosternini">
<subSubSection id="68C0BCC7751FB15B1C1CC1F69DB993E1" pageId="241" pageNumber="242" type="nomenclature">
<paragraph id="D6684FAA35E777459647A2ED62A2281A" pageId="241" pageNumber="242">
<taxonomicName id="AB2FAFD21EA69B6D83D95587E3275028" ID-CoL="74NL8" LSID="http://species-id.net/wiki/Operclipygus_fossipygus" authority="Wenzel, 1944" authorityName="Wenzel" authorityYear="1944" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus (Wenzel, 1944)</taxonomicName>
Figs 6E64
<normalizedToken id="8BA84414DAD2CBB0C57AC40394A4B20F" originalValue="A–B65A–D">A-B65A-D</normalizedToken>
, GMap 24
</paragraph>
</subSubSection>
<subSubSection id="08FF0827B77813955F03510E947575E6" pageId="241" pageNumber="242" type="reference_group">
<paragraph id="CABD0BEBF8D71BF74BA24141A3B38B02" pageId="241" pageNumber="242">
<taxonomicName id="085241CBDB8F087CDF16BF73994F388B" genus="Phelisteroides" lsidName="Phelisteroides fossipygus" pageId="241" pageNumber="242" rank="species" species="fossipygus">Phelisteroides fossipygus</taxonomicName>
Wenzel, 1944: 144;
<taxonomicName id="75C0ED16D5B61955E6328C7F61791E25" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
:
<bibRefCitation id="C41EA32702FDD2A37FBF479910327B89" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel (1976</bibRefCitation>
: 258).
</paragraph>
</subSubSection>
<subSubSection id="18A39A945E359FA7AABF88452BA34118" pageId="241" pageNumber="242" type="type locality">
<paragraph id="9F675F309DC3D3F932EAA3A383F2AD04" pageId="241" pageNumber="242">Type locality.</paragraph>
<paragraph id="8EDAC7D95806B7E0F05A9A0CDD7EC20D" pageId="241" pageNumber="242">
BRAZIL:
<normalizedToken id="C8A90955DB56AAAA36ED3562C164128A" originalValue="Pará">Para</normalizedToken>
:
<normalizedToken id="C8771C6CF2EA8A93952BC5DD0E5FB5A4" originalValue="Santarém">Santarem</normalizedToken>
[
<geoCoordinate id="1F361F1C085788EFFCF020D7CBF8644D" direction="south" orientation="latitude" precision="925" value="-2.4333334">2°26'S</geoCoordinate>
,
<geoCoordinate id="F743FBDFBEB41793000E2127E262168F" direction="west" orientation="longitude" precision="925" value="-54.7">54°42'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="C6D10FF43043B65855E6458491C02CA1" pageId="241" pageNumber="242" type="type material">
<paragraph id="5CCC5689B5F976CA9A1F6695DD1D351B" pageId="241" pageNumber="242">Type material.</paragraph>
<paragraph id="E23517EB965AE8C2E1A18039F0B77D13" pageId="241" pageNumber="242">
Holotype: &quot;Santarem Brazil Acc. No. 2966&quot; / &quot;Type
<taxonomicName id="1B91F7BE5E3F266ADBC708B573EC2043" genus="Phelisteroides" lsidName="Phelisteroides" pageId="241" pageNumber="242" rank="genus">Phelisteroides</taxonomicName>
fossipygus Wenzel&quot; (CMNH), examined, 2012.
</paragraph>
</subSubSection>
<subSubSection id="433AE618D3F7C927ADACDA23DB70F87F" lastPageId="243" lastPageNumber="244" pageId="241" pageNumber="242" type="other material">
<paragraph id="6DBD0E1F1F5F2F784441E65F60B87378" pageId="241" pageNumber="242">Other material.</paragraph>
<paragraph id="06EDEE8061EB35E3A19BFB45F3E3AA1B" lastPageId="243" lastPageNumber="244" pageId="241" pageNumber="242">
BRAZIL:
<normalizedToken id="002E518B0575C87416AE7069B2393600" originalValue="Amapá">Amapa</normalizedToken>
: 1: Serra do Navio,
<geoCoordinate id="74FE08B4BCE7A854193965599954AA53" direction="north" orientation="latitude" precision="925" value="0.98333335">0°59'N</geoCoordinate>
,
<geoCoordinate id="7693FE19B204F7512417A8EF942891B6" direction="west" orientation="longitude" precision="925" value="-52.0">52°00'W</geoCoordinate>
, 1-14.v. 1991, FIT (CHND); Mato Grosso: 1: Tangara da Serra,
<geoCoordinate id="9EDBC433F06265EC398354D4176C96CB" direction="south" orientation="latitude" precision="9" value="-14.3175">14°19.05'S</geoCoordinate>
,
<geoCoordinate id="25ECA12394DBCB50C3CEB8F3B581E20F" direction="west" orientation="longitude" precision="92" value="-57.731667">57°43.9'W</geoCoordinate>
, 640m, 22-29.i.2009, FIT, R.S. Silva (CEMT);
<normalizedToken id="AA1FA0F2B1910B1E188A4D6F3BF6B26E" originalValue="Pará">Para</normalizedToken>
: 2:
<normalizedToken id="14A0E34C067071238D26E441354EB606" originalValue="Tucuruí">Tucurui</normalizedToken>
,
<geoCoordinate id="2E5FDD2A4A06C41C7A43222AA1AECE77" direction="south" orientation="latitude" precision="925" value="-3.75">3°45'S</geoCoordinate>
,
<geoCoordinate id="344AF99F8C676E1278D7898B2EA51811" direction="west" orientation="longitude" precision="925" value="-49.666668">49°40'W</geoCoordinate>
, iv.1986,
<pageBreakToken id="A967DF84666F963B56CBF00F36642D81" pageId="242" pageNumber="243" start="start">FIT</pageBreakToken>
(UFPR), 4: vi.1985, FIT (CHND), 1: 16-29.vii.1985, FIT, human dung (CHND); 1: IPEAN, Utinga,
<normalizedToken id="9F5385F7809828386175ADD1AEE5AB2C" originalValue="Belém">Belem</normalizedToken>
,
<geoCoordinate id="6AE8D7BD948DBCE3E912D9F13D843237" direction="south" orientation="latitude" precision="925" value="-1.45">1°27'S</geoCoordinate>
,
<geoCoordinate id="1DFC6C484C2AD50DF12308EDA23FD252" direction="west" orientation="longitude" precision="925" value="-48.433334">48°26'W</geoCoordinate>
, v.1985, FIT (CHND), 2: viii.1985, FIT (CHND); 1:
<normalizedToken id="71C80B6AC1BFCCDA7646FF20F517346C" originalValue="Melgaço">Melgaco</normalizedToken>
, Rio
<normalizedToken id="1436F21718BF4CA8719EB904A12485BE" originalValue="Marinaú">Marinau</normalizedToken>
,
<geoCoordinate id="6BE6DCFBF31919CCDE378005C286FC40" direction="south" orientation="latitude" precision="925" value="-1.85">1°51'S</geoCoordinate>
,
<geoCoordinate id="ED6924D6595FDB23BEF49345F41B1D3A" direction="west" orientation="longitude" precision="925" value="-51.333332">51°20'W</geoCoordinate>
, 31.
<normalizedToken id="0A1E4517E73467C86D732B650853E9DC" originalValue="x–">x-</normalizedToken>
13.xi.1993, FIT (CHND). COLOMBIA:
<normalizedToken id="3D497C5A4A8946F627033E198454B739" originalValue="Vaupés">Vaupes</normalizedToken>
: 1: Est. Biol.
<normalizedToken id="A59B9DE5CAC609310EE395E193DF1420" originalValue="Caparú">Caparu</normalizedToken>
, Rio Apoporis,
<geoCoordinate id="202A9C38D31CD0F1E33402295F9E025C" direction="south" orientation="latitude" precision="5555" value="-1.1">1.1°S</geoCoordinate>
,
<geoCoordinate id="A8DC01D24B6CD021F5EBBFD26CCCB7BE" direction="west" orientation="longitude" precision="5555" value="-69.5">69.5°W</geoCoordinate>
, 27.ix-1.xii.1995, FIT, Black-water terrace forest on sandy soils, B.D. Gill (BDGC); 2: Parque Nac. Mosiro-Itajura (
<normalizedToken id="CE9A55D58E476185B94536BBAAD03960" originalValue="Caparú">Caparu</normalizedToken>
), Centro Ambiental,
<geoCoordinate id="C22AE1681B4666EEA64C7F9CFD349D1D" direction="south" orientation="latitude" precision="925" value="-1.0666667">1°04'S</geoCoordinate>
,
<geoCoordinate id="6D4C757189454D341512C2FCFA5B3D1E" direction="west" orientation="longitude" precision="925" value="-69.51667">69°31'W</geoCoordinate>
, 60m, 20-30.i.2003, FIT, D. Arias &amp; M. Sharkey (IAVH). ECUADOR: Orellana: 3:
<normalizedToken id="8C09EFE2D15F500D8358A3139CABCAF1" originalValue="Yasuní">Yasuni</normalizedToken>
Res. Stn., mid.Rio Tiputini,
<geoCoordinate id="3953607A176DD5DF7B1065B7FC5AB41D" direction="south" orientation="latitude" precision="92" value="-0.675">0°40.5'S</geoCoordinate>
,
<geoCoordinate id="B62ADB6F03DACE64BDE4E5700DECB39A" direction="west" orientation="longitude" precision="925" value="-76.4">76°24'W</geoCoordinate>
, 20-29.vi.1999, FIT, C.E. Carlton &amp;
<pageBreakToken id="4A8AE9BD9BCF0A368CAB64444F9BFDF8" pageId="243" pageNumber="244" start="start">V</pageBreakToken>
. Moseley (LSAM), 2: 17-23.vi.1999 (LSAM), 1: 22-28.vi.1999 (LSAM), 3: 5-12.vii.1999 (LSAM, FMNH), 1: 7-13.vii.1999 (LSAM), 3: 23-30.vi.1999 (LSAM), 5: 28.vi-5.vii.1999 (LSAM), 3: 4-17.vii.1999 (LSAM), 2: 17-26.vii.1999 (LSAM); 1: 5-10.ix.1999, FIT, primary forest, E.G. Riley (TAMU); 3: Parque Nac.
<normalizedToken id="73B46B535E970CB6D64492357371397A" originalValue="Yasuní">Yasuni</normalizedToken>
, Via Maxus at Puente
<normalizedToken id="89774CA541164B952B8A7EF5C995186D" originalValue="Piraña">Pirana</normalizedToken>
,
<geoCoordinate id="E84B35ADA1737958E7E8E184D2F26F32" direction="south" orientation="latitude" precision="92" value="-0.65833336">0°39.5'S</geoCoordinate>
,
<geoCoordinate id="5227C3E96D7F9112B856903D966B0030" direction="west" orientation="longitude" precision="925" value="-76.433334">76°26'W</geoCoordinate>
, 14-20.vii.2008, FIT, A.K. Tishechkin (AKTC), 1: 20-24.vii.2008, FIT, A.K. Tishechkin (AKTC); 1: Tiputini Biodiversity Station, 30.vii.2008, Day FIT, A.K. Tishechkin, DNA Extract MSC-1896 (AKTC); 3: Tiputini Biodiversity Station,
<geoCoordinate id="68409AF107397F63C09D265904DC0CAF" direction="south" orientation="latitude" precision="5" value="-0.6376">0.6376°S</geoCoordinate>
,
<geoCoordinate id="84FEB3386E57376AC3E86DE79834BF54" direction="west" orientation="longitude" precision="5" value="-76.1499">76.1499°W</geoCoordinate>
, 4-9.vi.2011, FIT, M.S. Caterino &amp; A.K. Tishechkin, DNA Extracts MSC-2289, MSC-2175, MSC-2291 (SBMNH, MSCC, USFQ); 2: Tiputini Biodiversity Station,
<geoCoordinate id="B12F523D2D805E724DE20973FA1D5C99" direction="south" orientation="latitude" precision="15" value="-0.6333333">0°38'0&quot;S</geoCoordinate>
,
<geoCoordinate id="D642D6C555BCB0E7B5A42089E1C202AA" direction="west" orientation="longitude" precision="15" value="-76.15">76°9'0&quot;W</geoCoordinate>
, 220m, 5-25.ix.2000, D.J. Inward &amp; K.A. Jackson (BMNH). FRENCH GUIANA: 1:
<normalizedToken id="6FE3FAB2764A42DF0C234DAFC887C00E" originalValue="Régina">Regina</normalizedToken>
,
<normalizedToken id="1C12F5AF9DBFC9777B6568A59D023EC1" originalValue="Réserve">Reserve</normalizedToken>
des Nouragues,
<geoCoordinate id="2F3144E76FD7D06201DF728D320B60E0" direction="north" orientation="latitude" precision="9" value="4.037833">4°2.27'N</geoCoordinate>
,
<geoCoordinate id="0D252EECC8AFE7C463330CE75D5073C1" direction="west" orientation="longitude" precision="9" value="-52.6725">52°40.35'W</geoCoordinate>
, 28.i.2010, FIT, SEAG (CHND); 1:
<normalizedToken id="288376ED2640FA3F13AAB431752C0AC9" originalValue="Régina">Regina</normalizedToken>
,
<normalizedToken id="A15D414071281F0F9269FB407724D272" originalValue="Réserve">Reserve</normalizedToken>
des Nouragues,
<geoCoordinate id="44C4D805BD86D26D39BE217F5376929F" direction="north" orientation="latitude" precision="9" value="4.037833">4°2.27'N</geoCoordinate>
,
<geoCoordinate id="D940ADD1ED39B37CDADFBA917F8B5BCA" direction="west" orientation="longitude" precision="9" value="-52.6725">52°40.35'W</geoCoordinate>
, 19.ii.2010, FIT, SEAG (CHND), 2: 3.xi.2009, FIT, SEAG (CHND); 1: Roura, 27.4km SSE,
<geoCoordinate id="BA1175BE73752A2C8C2D48D399D14C50" direction="north" orientation="latitude" precision="15" value="4.738889">4°44'20&quot;N</geoCoordinate>
,
<geoCoordinate id="A863CB3892DC904168A048F3533B1057" direction="west" orientation="longitude" precision="15" value="-52.22361">52°13'25&quot;W</geoCoordinate>
, 280m, 10.vi.1997, FIT, J. Ashe, R. Brooks (SEMC); 1:
<normalizedToken id="430285D95D8FD08C19CCFF1D0DD271EE" originalValue="Belvèdére">Belvedere</normalizedToken>
de
<normalizedToken id="8ECC6D6151F789C0699E2052E0679951" originalValue="Saül">Sauel</normalizedToken>
,
<geoCoordinate id="993ECBAC731BEF8924F07D611F60D909" direction="north" orientation="latitude" precision="15" value="3.0227778">3°1'22&quot;N</geoCoordinate>
,
<geoCoordinate id="295A0D64D7FB9B76E8277977A1E74512" direction="west" orientation="longitude" precision="15" value="-53.209446">53°12'34&quot;W</geoCoordinate>
, 30.xi.2010, Window trap, SEAG (MNHN). PERU: Loreto: 1: Campamento San Jacinto,
<geoCoordinate id="F7B03E31F6619BC0AD687AEB845CA8B6" direction="south" orientation="latitude" precision="9" value="-2.3125">2°18.75'S</geoCoordinate>
,
<geoCoordinate id="FBB706EDCC43552BE3E8CE9CB986C276" direction="west" orientation="longitude" precision="9" value="-75.86283">75°51.77'W</geoCoordinate>
, 175-215m, 12.vii.1993, FIT, R. Leschen (SEMC); 1: 1.5km N Teniente Lopez,
<geoCoordinate id="2C5A10D2A0921D53421D3DB373232950" direction="south" orientation="latitude" precision="9" value="-2.5943334">2°35.66'S</geoCoordinate>
,
<geoCoordinate id="0C9F499E9016A696012040104F26018C" direction="west" orientation="longitude" precision="9" value="-76.11533">76°06.92'W</geoCoordinate>
, 210-240m, 22.vii.1993, FIT, R. Leschen (SEMC); Madre de Dios: 1: Manu National Park, Zona res., Rio Manu, Cocha Juarez, trail nr. Manu Lodge, 18-24.ix.1991, FIT, A. Hartman (FMNH); 4: Tambopata, Reserva Cuzco Amazonico, 15km NE Pto. Maldonado,
<geoCoordinate id="F305C3934C41710AF546F117815DC51D" direction="south" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
,
<geoCoordinate id="2092CE4970F4AD8C38E403C9FE952CA0" direction="west" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
, 200m, 22.vi.1989, FIT, J. Ashe &amp; R. Leschen (SEMC), 1: 28.vi.1989, FIT, D. Silva, R.A. Leschen (SEMC), 1: 24.vi.1989, FIT, J. Ashe &amp; R. Leschen (SEMC). SURINAME: Saramacca: 1: West Suriname Road, 108km WSW Zanderij Airport,
<geoCoordinate id="FC9A255016363FE65E56332EEE2985E3" direction="north" orientation="latitude" precision="15" value="5.2269444">5°13'37&quot;N</geoCoordinate>
,
<geoCoordinate id="0A3651A882E6CCFD5E29EC20E70D0504" direction="west" orientation="longitude" precision="15" value="-55.881664">55°52'54&quot;W</geoCoordinate>
, 30m, 10-14.vi.1999, FIT, Z. Falin, B. DeDijn (SEMC); Sipaliwini: 5: CI-RAP Surv. Camp 2: Sipaliwini River,
<geoCoordinate id="6C6F9B1425560222B6872D5CBD47DD27" direction="north" orientation="latitude" precision="1" value="2.17535">2°10.521'N</geoCoordinate>
,
<geoCoordinate id="0E20263236E8826B70CEDAEDE4DE2DBE" direction="west" orientation="longitude" precision="1" value="-56.7874">56°47.244'W</geoCoordinate>
, 210m, 27.viii-1.ix.2010, FIT, T. Larsen &amp; A.E.Z. Short (SEMC); 1: CI-RAP Surv. Camp 3: Wehepai SE Kwamala,
<geoCoordinate id="C28B1C78AF8F2728025F01EFAF401165" direction="north" orientation="latitude" precision="1" value="2.3629334">2°21.776'N</geoCoordinate>
,
<geoCoordinate id="F39FFAF66E5380673DA3C99EE8165B4E" direction="west" orientation="longitude" precision="1" value="-56.697685">56°41.861'W</geoCoordinate>
, 237m, 3-7.ix.2010, FIT, T. Larsen &amp; A.E.Z. Short (SEMC).
</paragraph>
</subSubSection>
<subSubSection id="C4C0C2D0901EEEF615C99A3C399A0DFC" lastPageId="244" lastPageNumber="245" pageId="243" pageNumber="244" type="description">
<paragraph id="4867E84BFEF99F64DD71B479CE5AE053" pageId="243" pageNumber="244">Diagnostic description.</paragraph>
<paragraph id="0FC1D58820D5A36BCE36DB301E6A3359" lastPageId="244" lastPageNumber="245" pageId="243" pageNumber="244">
Length: 1.87-2.25 mm, width: 1.68-2.00 mm; body rufopiceous, ovoid; frons depressed at middle; frontal stria rounded, weakly divergent at sides, generally interrupted over antennal bases, strongly rounded anterad at middle; supraorbital stria weak, fragmented, detached from sides of frontal stria; epistoma convex; labrum about twice as wide as long, asymmetrically emarginate apically; left mandible untoothed, right mandible with small acute basal tooth; pronotum broadly depressed at base, but lacking distinct prescutellar impression, with fine, sparse ground punctation, with few or no larger lateral punctures; anterior pronotal margin distinctly projecting at middle; marginal pronotal stria broadly interrupted behind head; submarginal pronotal stria continuous, complete along lateral and anterior margins; median pronotal gland openings simple, about two-thirds pronotal length from anterior margin; elytra with two complete epipleural striae, outer subhumeral stria complete or interrupted at middle, inner subhumeral stria present in apical half, often shortened at apex, dorsal striae 1-3 complete, 4th and 5th striae similar in length, present in
<pageBreakToken id="156FF5D1FD91503B03FCFCFC9BE95C3D" pageId="244" pageNumber="245" start="start">about</pageBreakToken>
apical half, sutural stria present in apical three-fourths; prosternal keel weakly projecting at base, carinal striae generally complete, free or united anteriorly; anterior mesoventral margin shallowly emarginate, marginal stria complete; mesometaventral stria weakly arched forward at middle one-third, sinuate near mesocoxa, continued by lateral metaventral stria which curves laterad toward outer corner of metacoxa; 1st abdominal ventrite with two abbreviated lateral striae; postmetacoxal fovea not evident; propygidium with large punctures confined to middle and along basal margin, with fine, dense ground punctation elsewhere; pygidium with fine, dense ground punctation, lacking coarser punctures except along extreme basal margin; marginal pygidial sulcus deep, ending in very large, transversely elongate basal foveae. Male genitalia (Figs 65
<normalizedToken id="67A2C005EC27FB3990795E856F79D9F1" originalValue="A–D">A-D</normalizedToken>
, G): accessory sclerites absent; T8 more or less parallel-sided, slightly convergent apically, with narrow apical emargination, basal emargination evenly arcuate, basal membrane attachment line distad basal emargination by about one-third its depth, ventrolateral apodemes most strongly developed at their bases, narrowed apically; S8 parallel-sided, with apical guides narrow, evenly developed over most of length, with halves fused; T9 parallel sided in basal two-thirds, convergent to broad, subtruncate apices; T10 with halves finely divided; S9 rather short and broad, sclerotized along midline and along distal edges, with base widened, truncate to weakly emarginate, apical emargination narrow, apical flanges separate; tegmen widest near middle, narrowed to base and apex, strongly curved dorsoventrally, with medioventral process composed of inwardly projecting lateral flanges that are not fused along midline, together forming strong ventral projection; median lobe about three-fourths tegmen length, with proximal apodemes strongly differentiated into thick and thinner proximal portions; basal piece about one-third tegmen length.
</paragraph>
</subSubSection>
<subSubSection id="D30A36F8D09C213654DBEC8EA4705630" pageId="244" pageNumber="245" type="remarks">
<paragraph id="B99A25CA6EA8312C22462571D11DDE7E" pageId="244" pageNumber="245">Remarks.</paragraph>
<paragraph id="F8C75A61B148AC9F1B7A8720970D1845" pageId="244" pageNumber="245">
This is a highly distinctive species, easily recognized by the deep, basal pygidial foveae (Fig. 64B), the dense ground punctation on the sides of the propygidium, and the projecting anterior pronotal margin (Fig. 64A). There is some variation in genitalic structure, exhibited in an individual from Amazonian Peru, especially, in which the spiculum gastrale (S9) is sclerotized along the midline rather than along the margins. However, the unusual form of the aedeagus and its medioventral process is identical with typical
<taxonomicName id="A9B5D1424D8AAA27A61543733D48C4BD" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
.
</paragraph>
<caption id="B29B0E52BA7915DEE1E57D5B6D2B5D11" pageId="244" pageNumber="245">
<paragraph id="F809269E0CA29CEA7CE754F7E23A9AD4" pageId="244" pageNumber="245">
Figure 64.
<taxonomicName id="B0E70DFDEB7FC616CC845DF8FE6B3BFF" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
group. A Dorsal habitus of
<taxonomicName id="F221C57705E1A47DE0F5BD6047EA0F8D" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
B Pygidia of
<taxonomicName id="C7124795AA46D14F87E46B8D154897A5" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
C Ventral habitus of
<taxonomicName id="6B964F71682899A67340579EED26CE59" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus foveipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="foveipygus">Operclipygus foveipygus</taxonomicName>
D Pygidia of
<taxonomicName id="7CD957A7DF3F8D6FF93F69295E4AD9FD" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus foveipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="foveipygus">Operclipygus foveipygus</taxonomicName>
E Ventral habitus of
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F Pygidia of
<taxonomicName id="0C94B9724E4B2E37FC06BBED342755A4" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus gibbulus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="gibbulus">Operclipygus gibbulus</taxonomicName>
G Pronotum of
<taxonomicName id="44FACDB24784E2815FBB62AB328D2496" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fungicolus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fungicolus">Operclipygus fungicolus</taxonomicName>
H Pygidia of
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.
</paragraph>
</caption>
<caption id="01778D2B79CA8AD71E31A7FB2E6B2FBE" pageId="244" pageNumber="245">
<paragraph id="110DCDD4F3CF0CD1BEE63C682C4C98BE" pageId="244" pageNumber="245">
Figure 65. Male genitalia of
<taxonomicName id="4704A2DCC54DB5FD2283FF54CD8D62DA" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
group. A T8 of
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B S8 of
<taxonomicName id="9052B66E81D8A3AC1D3107E5F1D6698D" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
C T9 &amp; T10 of
<taxonomicName id="2D707D1D1452C2253DCFA74A209C2BD9" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
D S9 of
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E S8 of
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F S9 of
<taxonomicName id="75B33A7F45BE350E90BE603D781C1D06" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus foveipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="foveipygus">Operclipygus foveipygus</taxonomicName>
G Aedeagus, dorsal and lateral views, of
<taxonomicName id="FEFA896A22D0C74B33FC068129A2FA4E" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
H Aedeagus, dorsal and lateral views, of
<taxonomicName id="22427F841B76432C2FD8A9899492D90E" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus foveipygus" order="Coleoptera" pageId="244" pageNumber="245" phylum="Arthropoda" rank="species" species="foveipygus">Operclipygus foveipygus</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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