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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.627.8629" ID-GBIF-Dataset="23f2e74c-7c54-4e0f-97a6-a0361dfabb49" ID-PMC="PMC5118560" ID-Pensoft-Pub="1313-2970-627-1" ID-PubMed="27895525" ID-ZBK="3B7E6EA7C15C415B80A8ED4041525A40" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-627-1" ModsDocOrigin="ZooKeys 627" ModsDocTitle="A survey of the spider family Nesticidae (Arachnida, Araneae) in Asia and Madagascar, with the description of forty-three new species" checkinTime="1477637854587" checkinUser="pensoft" docAuthor="Lin, Yucheng, Ballarin, Francesco &amp; Li, Shuqiang" docDate="2016" docId="6D1E478090614948B6ED50F6DCFC91A7" docLanguage="en" docName="ZooKeys 627: 1-168" docOrigin="ZooKeys 627" docSource="http://dx.doi.org/10.3897/zookeys.627.8629" docTitle="Hamus cornutus Lin, Ballarin &amp; Li, 2016, sp. n." docType="treatment" docUuid="7CF47809-19D3-4348-9876-0D63D8EF95AB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="6" masterDocId="B4077777F170FF89FFAFD838C17AFFF3" masterDocTitle="A survey of the spider family Nesticidae (Arachnida, Araneae) in Asia and Madagascar, with the description of forty-three new species" masterLastPageNumber="168" masterPageNumber="1" pageNumber="6" updateTime="1668163652877" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A survey of the spider family Nesticidae (Arachnida, Araneae) in Asia and Madagascar, with the description of forty-three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Yucheng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ballarin, Francesco</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>627</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>168</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.627.8629</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.627.8629</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-627-1</mods:identifier>
<mods:identifier type="ZBK">3B7E6EA7C15C415B80A8ED4041525A40</mods:identifier>
<mods:identifier type="ZooBank">3B7E6EA7C15C415B80A8ED4041525A40</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127859446" LSID="urn:lsid:zoobank.org:act:7CF47809-19D3-4348-9876-0D63D8EF95AB" httpUri="http://treatment.plazi.org/id/6D1E478090614948B6ED50F6DCFC91A7" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection pageId="5" pageNumber="6" type="multiple">
<paragraph pageId="5" pageNumber="6">Taxon classification Animalia Araneae Nesticidae</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="nomenclature">
<paragraph pageId="5" pageNumber="6">
<taxonomicName LSID="http://zoobank.org/7CF47809-19D3-4348-9876-0D63D8EF95AB" class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus cornutus" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cornutus">Hamus cornutus</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
Figs 1, 2, 81
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="type material">
<paragraph pageId="5" pageNumber="6">Type material.</paragraph>
<paragraph pageId="5" pageNumber="6">
Holotype ♂ and paratypes 3♂3♀ (IZCAS), CHINA: Guangxi Zhuang Autonomous Region: Chongzuo City, Pingxiang, Sanzhishan Cave (
<geoCoordinate direction="north" orientation="latitude" precision="1" value="22.07567">22.07567°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="106.73773">106.73773°E</geoCoordinate>
, 257 m), 7.V.2015, Z. Chen &amp; Y. Li leg. Paratype 1♂ (IZCAS), LAOS: Bolikhamxay Province, Khamkeut District, 17.11 km west of Ban Laksao Town, Tham Mankone (
<geoCoordinate direction="north" orientation="latitude" precision="1" value="18.22156">18.22156°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.81268">104.81268°E</geoCoordinate>
, 495 m), 27.IX.2012, S. Li leg.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="etymology">
<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="6">
The specific name derives from the Latin word &quot;
<taxonomicName lsidName="cornutus" pageId="5" pageNumber="6" rank="species" species="cornutus">cornutus</taxonomicName>
&quot; = horned, and refers to the horn-like distal process of the paracymbium; adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="diagnosis">
<paragraph pageId="5" pageNumber="6">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="6">
Males can be distinguished from those of
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus bowoensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bowoensis">Hamus bowoensis</taxonomicName>
,
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus mangunensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mangunensis">Hamus mangunensis</taxonomicName>
sp. n. and
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus kangdingensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kangdingensis">Hamus kangdingensis</taxonomicName>
sp. n. by the absence of the terminal apophysis process I (Ta-I) (Fig. 1A), the sharp, hooked terminal apophysis process II (Ta-II) (Fig. 1A, D) and the horn-like distal process of the paracymbium (Dp) (Fig. 1
<normalizedToken originalValue="AB">A-B</normalizedToken>
), as opposed to a long Ta-I and a needle-like, very short or caniniform Dp in
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus bowoensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bowoensis">Hamus bowoensis</taxonomicName>
(see
<bibRefCitation author="Ballarin, F" journalOrPublisher="Zoological Systematics" pageId="67" pageNumber="68" pagination="179 - 190" title="Three new genera of the family Nesticidae (Arachnida: Araneae) from Tibet and Yunnan, China." volume="40" year="2015">Ballarin and Li 2015</bibRefCitation>
: 180, fig. 1
<normalizedToken originalValue="AD">A-D</normalizedToken>
) and
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus mangunensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mangunensis">Hamus mangunensis</taxonomicName>
sp. n. (Fig. 5
<normalizedToken originalValue="AB">A-B</normalizedToken>
, D) and as opposed to a blunt Ta-I and a pointed Dp in
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus kangdingensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kangdingensis">Hamus kangdingensis</taxonomicName>
sp. n. (Fig. 3D, E, G). Females can be separated from those of the similar species
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus bowoensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bowoensis">Hamus bowoensis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus luzon" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="luzon">Hamus luzon</taxonomicName>
sp. n. by the more pear-shaped spermathecae (Fig. 2
<normalizedToken originalValue="EF">E-F</normalizedToken>
) and the relatively straight copulatory ducts (Cd) (Fig. 2G), as opposed to the almost round spermathecae and the strongly twisted Cd in
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus bowoensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bowoensis">Hamus bowoensis</taxonomicName>
(see
<bibRefCitation author="Ballarin, F" journalOrPublisher="Zoological Systematics" pageId="67" pageNumber="68" pagination="179 - 190" title="Three new genera of the family Nesticidae (Arachnida: Araneae) from Tibet and Yunnan, China." volume="40" year="2015">Ballarin and Li 2015</bibRefCitation>
: 180, fig. 2
<normalizedToken originalValue="AC">A-C</normalizedToken>
with Cd reported as Fd) and
<taxonomicName class="Arachnida" family="Nesticidae" genus="Hamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hamus luzon" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="luzon">Hamus luzon</taxonomicName>
sp. n. (Fig. 4
<normalizedToken originalValue="DF">D-F</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="description">
<paragraph pageId="5" pageNumber="6">Description.</paragraph>
<paragraph pageId="5" pageNumber="6">
Habitus as in Fig. 2
<normalizedToken originalValue="AD">A-D</normalizedToken>
. Carapace yellowish. Mouthparts yellow in the male, light brown in the female. Sternum pale yellow in males, yellow in females. Legs uniformly pale yellow. Opisthosoma yellowish, slightly dark in males.
</paragraph>
<paragraph pageId="5" pageNumber="6">
Male palp (Fig. 1
<normalizedToken originalValue="AD">A-D</normalizedToken>
): paracymbium with a flat, semi-transparent dorsal apophysis and a strongly sclerotized, horn-like distal process (Fig. 1A). Ventral apophysis nearly semi-circular (Fig. 1C). Terminal apophysis almost round and strongly sclerotized (Fig. 1A). Short, sclerotized horned process (Cp-I) at the subapical part of the bulb together with a laminar, elongate process (Cp-II) forming a curved groove (Fig. 1A, D).
</paragraph>
<paragraph pageId="5" pageNumber="6">
Epigyne (Fig. 2
<normalizedToken originalValue="EG">E-G</normalizedToken>
): square, fertilization ducts long and twisted, originating at the ventro-lateral base of the spermathecae (Fig. 2F). (Fig. 2G). Spermathecae wide, pear-shaped, and close to each other.
</paragraph>
<paragraph pageId="5" pageNumber="6">Male (holotype). Total length 1.56. Carapace 0.80 long, 0.73 wide. Opisthosoma 0.77 long, 0.69 wide. Clypeus height 0.15. Sternum 0.47 long, 0.47 wide. Leg measurements: see Appendix A.</paragraph>
<paragraph pageId="5" pageNumber="6">Female (one of the paratypes). Total length 2.02. Carapace 0.90 long, 0.78 wide. Opisthosoma 1.22 long, 0.87 wide. Clypeus height 0.18. Sternum 0.56 long, 0.49 wide. Leg measurements: see Appendix A.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="habitat">
<paragraph pageId="5" pageNumber="6">Habitat.</paragraph>
<paragraph pageId="5" pageNumber="6">Forest leaf litter, cave.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="distribution">
<paragraph pageId="5" pageNumber="6">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="6">China (Guangxi), Laos (Fig. 81).</paragraph>
</subSubSection>
</treatment>
</document>