treatments-xml/data/46/69/1A/46691AB6ED1719FC6A28F54C882EF52E.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.349.6164" ID-GBIF-Dataset="cd871e7e-7e75-4f2e-8c8b-7af9cb4803d4" ID-PMC="PMC3837407" ID-Pensoft-Pub="1313-2970-349-81" ID-PubMed="24294080" ID-ZBK="45895E0B79E74794A0155CD74FEDDF22" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-349-81" ModsDocOrigin="ZooKeys 349" ModsDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" checkinTime="1451246687136" checkinUser="pensoft" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2013" docId="46691AB6ED1719FC6A28F54C882EF52E" docLanguage="en" docName="ZooKeys 349: 81-100" docOrigin="ZooKeys 349" docSource="http://dx.doi.org/10.3897/zookeys.349.6164" docTitle="Orthogonius euthyphallus subsp. bolavenensis Tian &amp; Deuve, ssp. n." docType="treatment" docUuid="10A4E8EF-DC6B-48A9-AB91-0C1D3D75FE41" docUuidSource="ZooBank" docVersion="5" lastPageNumber="89" masterDocId="B203D41F1D41C83E8603FFE5D1499035" masterDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" masterLastPageNumber="100" masterPageNumber="81" pageNumber="88" updateTime="1668156835216" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deuve, Thierry</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>349</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>100</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.349.6164</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.349.6164</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-349-81</mods:identifier>
<mods:identifier type="ZBK">45895E0B79E74794A0155CD74FEDDF22</mods:identifier>
<mods:identifier type="ZooBank">45895E0B79E74794A0155CD74FEDDF22</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152049512" LSID="urn:lsid:zoobank.org:act:10A4E8EF-DC6B-48A9-AB91-0C1D3D75FE41" httpUri="http://treatment.plazi.org/id/46691AB6ED1719FC6A28F54C882EF52E" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88">
<subSubSection pageId="7" pageNumber="88" type="nomenclature">
<paragraph pageId="7" pageNumber="88">
<taxonomicName LSID="http://zoobank.org/10A4E8EF-DC6B-48A9-AB91-0C1D3D75FE41" authority="Tian &amp; Deuve" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius euthyphallus" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="subSpecies" species="euthyphallus" subSpecies="bolavenensis">Orthogonius euthyphallus bolavenensis Tian &amp; Deuve</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="88">ssp. n.</taxonomicNameLabel>
Figs 1, 6-7, 16-17
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88" type="description">
<paragraph pageId="7" pageNumber="88">Description.</paragraph>
<paragraph pageId="7" pageNumber="88">Body stout, strongly shiny. Habitus as in Figs 6-7.</paragraph>
<paragraph pageId="7" pageNumber="88">Length: 16.5 mm; width: 6.5 mm.</paragraph>
<paragraph pageId="7" pageNumber="88">Black, antennomeres 4-11, mandibles, palps and tarsomeres dark brown to brown.</paragraph>
<paragraph pageId="7" pageNumber="88">Macrosculpture: Head densely punctured in both sexes, elytra densely punctured in male, (rather sparser in female); pronotum much less punctured in male, without puncture in female; head obscure striate on frons, smooth on vertex; pronotum faintly striate or not; underside surface smooth and glabrous, impunctate.</paragraph>
<paragraph pageId="7" pageNumber="88">Microsculpture: Engraved meshes densely isodiametric on elytra, faint on head and pronotum.</paragraph>
<paragraph pageId="7" pageNumber="88">Head longer than wide, HL/HW=1.17-1.18, eyes large and prominent, frons and vertex convex, frontal impressions large and deep, extending on clypeus; clypeus bisetose, basal portion deeply furrowed; labrum slightly broad at apical margin, sexsetose; palps rather stout, subcylindrical, maxillary palpomere 3 as long as 4 which is glabrous, palpomere 3 with two short setae at apex; labial palpomere 2 longer than 3, bisetose on inner margin, with several additional short setae at subapex and apex, palpomere 3 with a few tiny setae; ligula small, bisetose at apex; mentum without tooth, each of mentum and submentum bisetose, palpiger asetose. Antennae rather stout, not extending over the middle of elytra; pubescent from basal 1/4 of antennomere 4; antennomere 3 almost as long as 4, and 1.55 times longer than 2; antennomere 1 unisetose on subapex.</paragraph>
<paragraph pageId="7" pageNumber="88">Pronotum rather broad though transverse, PW/PL=1.49-1.52, disc moderately convex, fore and basal margins well beaded, sides evenly expanded, widest at about middle; lateral expanded margin wide and hardly reflexed, better defined in female than in male; both transversal impressions indistinct, basal foveae shallow, median line clear.</paragraph>
<paragraph pageId="7" pageNumber="88">Elytra elongate ovate, EL/EW=1.57-1.60, widest at about middle, where sides slightly expanded, base completely bordered, apex broadly sinuate, inner angle nearly rectangular, faintly denticulate; striae deep, intervals convex, subequal in width in middle portion, interval 3 with basal and middle setiferous pores, the subapical one absent.</paragraph>
<paragraph pageId="8" pageNumber="89">
<pageBreakToken pageId="8" pageNumber="89" start="start">Legs</pageBreakToken>
slender, fore tibia distinctly expanded at apex, apex obliquely truncate, outer margin smooth, not subserrate; middle and hind coxae smooth and glabrous in median portion; middle tibia slender and slightly expanded at apex, strongly curved in male, while normal in female; hind femur moderately expanded, with two setae posteriorly; hind tibia slightly expanded at apex, apical spurs long and sharp; fore tarsi distinctly wider than middle and hind ones, middle wider than the hind; hind tarsomeres 1 and 3 much longer than 2 and 4 respectively, length ratio of tarsomeres 1-4 as 2.57, 1.86, 1.28 and 1.0; tarsomere 4 asymmetrically emarginat, with longer lobe as deep as 1/3 of the joint; all tarsal claws strongly pectinate.
</paragraph>
<paragraph pageId="8" pageNumber="89">Prosternal process well bordered at apex, abdominal ventrite VII of male narrowly and deeply emarginate at apical margin.</paragraph>
<paragraph pageId="8" pageNumber="89">Male genitalia (Figs 16-17): Slender and quite straight, expanded medially, gently sinuate on ventral margin, apex stout and blunt in profile; in dorsal view, broad at subapical portion, the apical lamella slightly wider than long, broad at tip.</paragraph>
<caption pageId="8" pageNumber="89">
<paragraph pageId="8" pageNumber="89">
Figures 16-19. Male genitalia of the lancangjiang species group. 16
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius euthyphallus subsp. bolavenensis" order="Coleoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="subSpecies" species="euthyphallus" subSpecies="bolavenensis">Orthogonius euthyphallus bolavenensis</taxonomicName>
ssp. n., lateral view 17 idem, dorsal view of apical portion 18
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius carinatus" order="Coleoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="carinatus">Orthogonius carinatus</taxonomicName>
sp. n., lateral view 19 idem, dorsal view of apical portion.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="89" type="remarks">
<paragraph pageId="8" pageNumber="89">Remarks.</paragraph>
<paragraph pageId="8" pageNumber="89">Similar to the nominate subspecies, but darker and broader (in particular pronotum and elytra), and the apical lamella is more slender in dorsal view.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="89" type="material examined">
<paragraph pageId="8" pageNumber="89">Material examined.</paragraph>
<paragraph pageId="8" pageNumber="89">Holotype: male, &quot;Coll. I. R. Sc.N.B/ purchased from Mr. B. Makovsky, I.G. 31.969&quot; and &quot;Laos South, Bolaven Plateau, Ban Itou, 23.-27. V. 2007, B. Makovsky Lat.&quot;, in IRSNB.</paragraph>
<paragraph pageId="8" pageNumber="89">Paratype. 1 female, idem, in IRSNB.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="89" type="etymology">
<paragraph pageId="8" pageNumber="89">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="89">Refers to its type locality.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="89" type="distribution">
<paragraph pageId="8" pageNumber="89">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="89">Southern Laos. Known only from the type locality (Fig. 1).</paragraph>
</subSubSection>
</treatment>
</document>