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<document id="822F0670265B5FE0AABC64AD7E20B094" ID-DOI="10.5281/zenodo.199772" ID-GBIF-Dataset="1550a465-e5fd-42fc-9206-0d3f09622d8c" ID-ISSN="1175-5326" ID-Zenodo-Dep="199772" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460346219133" checkinUser="plazi" docAuthor="Köhler, Gunther, Dehling, Matthias &amp; Köhler, Johannes" docDate="2010" docId="462C878BFFCCFF9AFF22F932571F43B3" docLanguage="en" docName="zt02718p038.pdf" docOrigin="Zootaxa 2718" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anolis osa Köhler, Dehling &amp; Köhler, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="33" masterDocId="BA15FFF3FFC9FF90FFB5FFD4535C4247" masterDocTitle="Cryptic species and hybridization in the Anolis polylepis complex, with the description of a new species from the Osa Peninsula, Costa Rica (Squamata: Polychrotidae)" masterLastPageNumber="38" masterPageNumber="23" pageNumber="28" updateTime="1698262087057" updateUser="plazi">
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<mods:title id="419322CA8F66BD52212A83B823701BB2">Cryptic species and hybridization in the Anolis polylepis complex, with the description of a new species from the Osa Peninsula, Costa Rica (Squamata: Polychrotidae)</mods:title>
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<mods:namePart id="3B289BC71017576052B8EB4CC0DC927B">Köhler, Gunther</mods:namePart>
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<mods:namePart id="37EC8DF31DEA6912A0211E053D637E89">Dehling, Matthias</mods:namePart>
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<treatment id="462C878BFFCCFF9AFF22F932571F43B3" ID-DOI="http://doi.org/10.5281/zenodo.5621820" ID-GBIF-Taxon="119407258" ID-Zenodo-Dep="5621820" LSID="urn:lsid:plazi:treatment:462C878BFFCCFF9AFF22F932571F43B3" httpUri="http://treatment.plazi.org/id/462C878BFFCCFF9AFF22F932571F43B3" lastPageId="10" lastPageNumber="33" pageId="5" pageNumber="28">
<subSubSection id="869F6516FFCCFF95FF22F93252504566" pageId="5" pageNumber="28" type="nomenclature">
<paragraph id="CE3A369DFFCCFF95FF22F93252264547" blockId="5.[151,378,1766,1825]" box="[151,378,1766,1792]" pageId="5" pageNumber="28">
<heading id="957281F1FFCCFF95FF22F93252264547" bold="true" box="[151,378,1766,1792]" fontSize="11" level="1" pageId="5" pageNumber="28" reason="1">
<emphasis id="FCF1EA8FFFCCFF95FF22F93252264547" bold="true" box="[151,378,1766,1792]" pageId="5" pageNumber="28">
<taxonomicName id="09854D1EFFCCFF95FF22F93252494547" box="[151,277,1766,1792]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="28" phylum="Chordata" rank="species" species="osa" status="sp. nov.">
<emphasis id="FCF1EA8FFFCCFF95FF22F93252494547" bold="true" box="[151,277,1766,1792]" italics="true" pageId="5" pageNumber="28">Anolis osa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7C257F4FFCCFF95FEA9F93252264547" box="[284,378,1766,1792]" pageId="5" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CE3A369DFFCCFF95FF22F8DD52504566" blockId="5.[151,378,1766,1825]" box="[151,268,1801,1825]" pageId="5" pageNumber="28">
<figureCitation id="56BE2A18FFCCFF95FF22F8DD52504566" box="[151,268,1801,1825]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="12.[151,255,1896,1920]" captionStartId-1="13.[151,255,1050,1074]" captionStartId-2="13.[151,255,2003,2027]" captionTargetBox-1="[151,1436,178,1028]" captionTargetBox-2="[151,1436,1125,1981]" captionTargetId-1="figure@13.[151,1436,172,1029]" captionTargetId-2="figure@13.[151,1436,1125,1982]" captionTargetPageId-1="13" captionTargetPageId-2="13" captionText-0="FIGURE 9. Head of Anolis osa (SMF 80645). Scale bars = 1.0 mm." captionText-1="FIGURE 10. Adult male of Anolis osa (not collected) at Puerto Escondido in life with extended dewlap." captionText-2="FIGURE 11. Adult female of Anolis osa (not collected) at Puerto Escondido in life." httpUri-1="https://zenodo.org/record/199780/files/figure.png" httpUri-2="https://zenodo.org/record/199781/files/figure.png" pageId="5" pageNumber="28">Figs. 911</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="869F6516FFCCFF95FF22F885514D4520" box="[151,529,1873,1895]" pageId="5" pageNumber="28" type="reference_group">
<paragraph id="CE3A369DFFCCFF95FF22F885514D4520" blockId="5.[151,529,1873,1895]" box="[151,529,1873,1895]" pageId="5" pageNumber="28">
<taxonomicName id="09854D1EFFCCFF95FF22F88551524520" authority="Poe" authorityName="Poe" authorityYear="2004" box="[151,526,1873,1895]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="28" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCCFF95FF22F88552664520" box="[151,314,1873,1895]" italics="true" pageId="5" pageNumber="28">Anolis polylepis</emphasis>
:
<bibRefCitation id="AA144B6CFFCCFF95FEF2F88651524520" author="Poe" box="[327,526,1873,1895]" pageId="5" pageNumber="28" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe (2004, in part.)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="869F6516FFCCFF96FF22F840513C449E" lastPageId="6" lastPageNumber="29" pageId="5" pageNumber="28" type="materials_examined">
<paragraph id="CE3A369DFFCCFF95FF22F840517A45B3" blockId="5.[151,1436,1940,2037]" pageId="5" pageNumber="28">
<emphasis id="FCF1EA8FFFCCFF95FF22F840525745EB" bold="true" box="[151,267,1940,1964]" pageId="5" pageNumber="28">
<typeStatus id="113E883FFFCCFF95FF22F840525A45EB" box="[151,262,1940,1964]" pageId="5" pageNumber="28" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="A894AE58FFCCFF95FEA7F840521645EB" LSID="urn:lsid:biocol.org:col:34838" box="[274,330,1940,1964]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="28">SMF</collectionCode>
80645, an adult male from about
<quantity id="097D9B78FFCCFF95FD71F840504D45EB" box="[708,785,1940,1964]" metricMagnitude="3" metricUnit="m" metricValue="6.3" pageId="5" pageNumber="28" unit="km" value="6.3">6.3 km</quantity>
WSW Rincón de
<taxonomicName id="09854D1EFFCCFF95FC69F840575645EB" box="[988,1034,1940,1964]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="osa">Osa</taxonomicName>
,
<geoCoordinate id="ABB1505AFFCCFF95FBA3F84057F045EB" box="[1046,1196,1940,1964]" direction="north" orientation="latitude" pageId="5" pageNumber="28" precision="1" value="8.676862">8°4036.7N</geoCoordinate>
,
<geoCoordinate id="ABB1505AFFCCFF95FB0DF840560F45EB" box="[1208,1363,1940,1964]" direction="west" orientation="longitude" pageId="5" pageNumber="28" precision="1" value="-83.53553">83°327.9W</geoCoordinate>
, about
<quantity id="097D9B78FFCCFF95FF22F86C53BC4597" box="[151,224,1976,2000]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="28" unit="m" value="150.0">150 m</quantity>
elevation, Puntarenas Province,
<collectingCountry id="B692760DFFCCFF95FDE3F86C51894597" box="[598,725,1976,2000]" name="Costa Rica" pageId="5" pageNumber="28">Costa Rica</collectingCountry>
, collected
<date id="BA3B105DFFCCFF95FCE5F86C57474597" box="[848,1051,1976,2000]" pageId="5" pageNumber="28" value="2008-02-22">22 February 2008</date>
by Gunther Köhler and Johannes Köhler. Field tag number GK 2092.
</paragraph>
<caption id="9AFA6615FFCFFF96FF22FCF851864103" box="[151,730,812,836]" httpUri="https://zenodo.org/record/199774/files/figure.png" pageId="6" pageNumber="29" targetBox="[151,1427,201,773]" targetPageId="6">
<paragraph id="CE3A369DFFCFFF96FF22FCF851864103" blockId="6.[151,730,812,836]" box="[151,730,812,836]" pageId="6" pageNumber="29">
<emphasis id="FCF1EA8FFFCFFF96FF22FCF852454103" bold="true" box="[151,281,812,836]" pageId="6" pageNumber="29">FIGURE 2.</emphasis>
Designation of dorsal color pattern types.
</paragraph>
</caption>
<caption id="9AFA6615FFCFFF96FF22FA7250B747FA" box="[151,1003,1446,1470]" httpUri="https://zenodo.org/record/199775/files/figure.png" pageId="6" pageNumber="29" targetBox="[287,1293,896,1421]" targetPageId="6">
<paragraph id="CE3A369DFFCFFF96FF22FA7250B747FA" blockId="6.[151,1003,1446,1470]" box="[151,1003,1446,1470]" pageId="6" pageNumber="29">
<emphasis id="FCF1EA8FFFCFFF96FF22FA72524547F9" bold="true" box="[151,281,1446,1470]" pageId="6" pageNumber="29">FIGURE 3.</emphasis>
Hemipenis of
<taxonomicName id="09854D1EFFCFFF96FE02FA73513E47F9" box="[439,610,1447,1470]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCFFF96FE02FA73513E47F9" box="[439,610,1447,1470]" italics="true" pageId="6" pageNumber="29">Anolis polylepis</emphasis>
</taxonomicName>
(MHCH 1024). Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="CE3A369DFFCFFF96FF7CFA3E53A944F2" blockId="6.[151,1436,1513,2006]" pageId="6" pageNumber="29">
<emphasis id="FCF1EA8FFFCFFF96FF7CFA3E52144445" bold="true" box="[201,328,1514,1538]" pageId="6" pageNumber="29">
<typeStatus id="113E883FFFCFFF96FF7CFA3E521F4445" box="[201,323,1514,1538]" pageId="6" pageNumber="29" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="A894AE58FFCFFF96FEE6FA3D52D74446" LSID="urn:lsid:biocol.org:col:34838" box="[339,395,1513,1537]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="29">SMF</collectionCode>
8922732, same collecting data as
<typeStatus id="113E883FFFCFFF96FC83FA3D50C04446" box="[822,924,1513,1537]" pageId="6" pageNumber="29" type="holotype">holotype</typeStatus>
.
<collectionCode id="A894AE58FFCFFF96FC1FFA3D50BE4446" LSID="urn:lsid:biocol.org:col:34838" box="[938,994,1513,1537]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="29">SMF</collectionCode>
8921521, UCR 2073335, 2 km W Rincón de
<taxonomicName id="09854D1EFFCFFF96FEA1F9D9521E4462" box="[276,322,1549,1573]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="36" phylum="Chordata" rank="species" species="osa">Osa</taxonomicName>
, about
<quantity id="097D9B78FFCFFF96FE2DF9D9528F4462" box="[408,467,1549,1573]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="29" unit="m" value="10.0">10 m</quantity>
elevation, Puntarenas Province,
<collectingCountry id="B692760DFFCFFF96FCFEF9D950964462" box="[843,970,1549,1573]" name="Costa Rica" pageId="6" pageNumber="29">Costa Rica</collectingCountry>
, collected
<date id="BA3B105DFFCFFF96FBF3F9D9564E4462" box="[1094,1298,1549,1573]" pageId="6" pageNumber="29" value="2008-02-22">22 February 2008</date>
by Gunther Köhler and Johannes Köhler.
<collectionCode id="A894AE58FFCFFF96FE5BF9E5517A440E" LSID="urn:lsid:biocol.org:col:34838" box="[494,550,1585,1609]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="29">SMF</collectionCode>
8926061, 8942729, Parque Nacional Corcovado, Estación Sirena,
<geoCoordinate id="ABB1505AFFCFFF96FAF1F9E556C4440E" box="[1348,1432,1585,1609]" direction="north" orientation="latitude" pageId="6" pageNumber="29" precision="555" value="8.48">8.48°N</geoCoordinate>
,
<geoCoordinate id="ABB1505AFFCFFF96FF22F98153A0442A" box="[151,252,1621,1645]" direction="west" orientation="longitude" pageId="6" pageNumber="29" precision="555" value="-83.58">83.58°W</geoCoordinate>
, Puntarenas Province,
<collectingCountry id="B692760DFFCFFF96FDB7F98151DD442A" box="[514,641,1621,1645]" name="Costa Rica" pageId="6" pageNumber="29">Costa Rica</collectingCountry>
, collected
<date id="BA3B105DFFCFFF96FD4BF9815082442A" box="[766,990,1621,1646]" pageId="6" pageNumber="29" value="2005-08-10" valueMax="2005-08-11" valueMin="2005-08-10">1011 August 2005</date>
by Matthias Dehling.
<collectionCode id="A894AE58FFCFFF96FB51F9815640442A" LSID="urn:lsid:biocol.org:col:34838" box="[1252,1308,1621,1645]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="29">SMF</collectionCode>
8922729,
<collectionCode id="A894AE58FFCFFF96FF22F9AD539344D6" LSID="urn:lsid:biocol.org:col:34838" box="[151,207,1657,1681]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="29">SMF</collectionCode>
8926061, 8942729, UCR 2073335 are adult males with everted hemipenes.
<collectionCode id="A894AE58FFCFFF96FBCFF9AD57EE44D6" LSID="urn:lsid:biocol.org:col:34838" box="[1146,1202,1657,1681]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="29">SMF</collectionCode>
8923032 are adult females.
</paragraph>
<paragraph id="CE3A369DFFCFFF96FF7CF915513C449E" blockId="6.[151,1436,1513,2006]" box="[201,608,1729,1753]" pageId="6" pageNumber="29">
<emphasis id="FCF1EA8FFFCFFF96FF7CF91552C0449E" box="[201,412,1729,1753]" italics="true" pageId="6" pageNumber="29">Referred Specimen</emphasis>
: See Appendix I.
</paragraph>
</subSubSection>
<subSubSection id="869F6516FFCFFF97FF7CF93150FC4388" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29" type="diagnosis">
<paragraph id="CE3A369DFFCFFF97FF7CF93150FC4388" blockId="6.[151,1436,1513,2006]" lastBlockId="7.[151,1436,151,464]" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29">
<emphasis id="FCF1EA8FFFCFFF96FF7CF931521F44B9" bold="true" box="[201,323,1765,1790]" pageId="6" pageNumber="29">Diagnosis.</emphasis>
A medium-sized species (SVL in largest specimen
<quantity id="097D9B78FFCFFF96FC27F93150B844BA" box="[914,996,1765,1789]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="6" pageNumber="29" unit="mm" value="57.0">57 mm</quantity>
) of the genus
<taxonomicName id="09854D1EFFCFFF96FB3DF931578C44BA" box="[1160,1232,1765,1789]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="genus">
<emphasis id="FCF1EA8FFFCFFF96FB3DF931578C44BA" box="[1160,1232,1765,1789]" italics="true" pageId="6" pageNumber="29">Anolis</emphasis>
</taxonomicName>
(sensu
<bibRefCitation id="AA144B6CFFCFFF96FA9DF93256C944BA" author="Poe" box="[1320,1429,1765,1789]" pageId="6" pageNumber="29" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe 2004</bibRefCitation>
) that differs from all other Lower Central American beta anoles (sensu
<bibRefCitation id="AA144B6CFFCFFF96FC11F8DD570C4566" author="Etheridge" box="[932,1104,1801,1825]" pageId="6" pageNumber="29" refString="Etheridge, R. (1967) Lizard caudal vertebrae. Copeia, 1967, 699 - 721." type="journal article" year="1967">Etheridge 1967</bibRefCitation>
) except
<emphasis id="FCF1EA8FFFCFFF96FB1BF8DD52364502" italics="true" pageId="6" pageNumber="29">
<taxonomicName id="09854D1EFFCFFF96FB1BF8DD56CB4566" box="[1198,1431,1801,1825]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="apletophallus">Anolis apletophallus</taxonomicName>
,
<taxonomicName id="09854D1EFFCFFF96FF22F8F9523B4502" box="[151,359,1837,1861]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="cryptolimifrons">A. cryptolimifrons</taxonomicName>
,
</emphasis>
and
<taxonomicName id="09854D1EFFCFFF96FE11F8F951754502" box="[420,553,1837,1861]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="limifrons">
<emphasis id="FCF1EA8FFFCFFF96FE11F8F951754502" box="[420,553,1837,1861]" italics="true" pageId="6" pageNumber="29">A. limifrons</emphasis>
</taxonomicName>
in that it is long-legged (longest toe of adpressed hind leg reaches to at least center of eye), has smooth ventral scales, and a slender habitus, often delicate.
<taxonomicName id="09854D1EFFCFFF96FBA1F88557D7452E" box="[1044,1163,1873,1897]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCFFF96FBA1F88557D7452E" box="[1044,1163,1873,1897]" italics="true" pageId="6" pageNumber="29">Anolis osa</emphasis>
</taxonomicName>
differs from the species in the cluster listed above, except
<taxonomicName id="09854D1EFFCFFF96FDADF8A151C245CA" box="[536,670,1909,1933]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCFFF96FDADF8A151C245CA" box="[536,670,1909,1933]" italics="true" pageId="6" pageNumber="29">A. polylepis</emphasis>
</taxonomicName>
, by having two elongate, overlapping superciliaries (one such scale in
<emphasis id="FCF1EA8FFFCFFF96FF01F84D511045F6" box="[180,588,1945,1969]" italics="true" pageId="6" pageNumber="29">
<taxonomicName id="09854D1EFFCFFF96FF01F84D523345F6" box="[180,367,1945,1969]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="apletophallus">A. apletophallus</taxonomicName>
,
<taxonomicName id="09854D1EFFCFFF96FECFF84D511545F6" box="[378,585,1945,1969]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="cryptolimifrons">A. cryptolimifrons</taxonomicName>
,
</emphasis>
and
<taxonomicName id="09854D1EFFCFFF96FD36F84D505B45F6" box="[643,775,1945,1969]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="limifrons">
<emphasis id="FCF1EA8FFFCFFF96FD36F84D505B45F6" box="[643,775,1945,1969]" italics="true" pageId="6" pageNumber="29">A. limifrons</emphasis>
</taxonomicName>
), usually two anterior prenasal scales (versus one elongate prenasal scale) and dewlap coloration (usually orange with a darker orange basal portion in
<taxonomicName id="09854D1EFFCFFF96FB76F86956504592" box="[1219,1292,1981,2005]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="29" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCFFF96FB76F86956504592" box="[1219,1292,1981,2005]" italics="true" pageId="6" pageNumber="29">A. osa</emphasis>
</taxonomicName>
versus dirty white with a basal orange-yellow blotch in
<taxonomicName id="09854D1EFFCEFF97FDCBFF4C501642F7" box="[638,842,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="cryptolimifrons">
<emphasis id="FCF1EA8FFFCEFF97FDCBFF4C501642F7" box="[638,842,152,176]" italics="true" pageId="7" pageNumber="30">A. cryptolimifrons</emphasis>
</taxonomicName>
and
<emphasis id="FCF1EA8FFFCEFF97FC34FF4C575042F7" box="[897,1036,152,176]" italics="true" pageId="7" pageNumber="30">
<taxonomicName id="09854D1EFFCEFF97FC34FF4C575B42F7" box="[897,1031,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="limifrons">A. limifrons</taxonomicName>
,
</emphasis>
almost uniformly orange-yellow in
<taxonomicName id="09854D1EFFCEFF97FF22FF68520C4293" box="[151,336,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="apletophallus">
<emphasis id="FCF1EA8FFFCEFF97FF22FF68520C4293" box="[151,336,188,212]" italics="true" pageId="7" pageNumber="30">A. apletophallus</emphasis>
</taxonomicName>
).
<taxonomicName id="09854D1EFFCEFF97FEDFFF6852BD4293" box="[362,481,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCEFF97FEDFFF6852BD4293" box="[362,481,188,212]" italics="true" pageId="7" pageNumber="30">Anolis osa</emphasis>
</taxonomicName>
differs from
<taxonomicName id="09854D1EFFCEFF97FDCEFF68505D4293" box="[635,769,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FDCEFF68505D4293" box="[635,769,188,212]" italics="true" pageId="7" pageNumber="30">A. polylepis</emphasis>
</taxonomicName>
by having a unilobed hemipenis (bilobed in
<taxonomicName id="09854D1EFFCEFF97FAB0FF6856D74293" box="[1285,1419,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FAB0FF6856D74293" box="[1285,1419,188,212]" italics="true" pageId="7" pageNumber="30">A. polylepis</emphasis>
</taxonomicName>
). Other anole species from Lower Central
<collectingCountry id="B692760DFFCEFF97FDDEFF0B519742B0" box="[619,715,223,247]" name="United States of America" pageId="7" pageNumber="30">America</collectingCountry>
that are somewhat similar in appearance to both
<taxonomicName id="09854D1EFFCEFF97FAB7FF34561542BF" box="[1282,1353,224,248]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCEFF97FAB7FF34561542BF" box="[1282,1353,224,248]" italics="true" pageId="7" pageNumber="30">A. osa</emphasis>
</taxonomicName>
and
<taxonomicName id="09854D1EFFCEFF97FA31FF3453A0435B" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FA31FF3453A0435B" italics="true" pageId="7" pageNumber="30">A. polylepis</emphasis>
</taxonomicName>
are
<taxonomicName id="09854D1EFFCEFF97FE87FED052B0435B" box="[306,492,260,284]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="tropidogaster">
<emphasis id="FCF1EA8FFFCEFF97FE87FED052B0435B" box="[306,492,260,284]" italics="true" pageId="7" pageNumber="30">A. tropidogaster</emphasis>
</taxonomicName>
and
<taxonomicName id="09854D1EFFCEFF97FD92FED051FC435B" box="[551,672,260,284]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="FCF1EA8FFFCEFF97FD92FED051FC435B" box="[551,672,260,284]" italics="true" pageId="7" pageNumber="30">A. cupreus</emphasis>
</taxonomicName>
, the latter of which occurs syntopically at some localities with
<taxonomicName id="09854D1EFFCEFF97FA31FED053A04307" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FA31FED053A04307" italics="true" pageId="7" pageNumber="30">A. polylepis</emphasis>
</taxonomicName>
. However, both
<taxonomicName id="09854D1EFFCEFF97FE00FEFC51374307" box="[437,619,296,320]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="tropidogaster">
<emphasis id="FCF1EA8FFFCEFF97FE00FEFC51374307" box="[437,619,296,320]" italics="true" pageId="7" pageNumber="30">A. tropidogaster</emphasis>
</taxonomicName>
and
<taxonomicName id="09854D1EFFCEFF97FD14FEFC504A4307" box="[673,790,296,320]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="FCF1EA8FFFCEFF97FD14FEFC504A4307" box="[673,790,296,320]" italics="true" pageId="7" pageNumber="30">A. cupreus</emphasis>
</taxonomicName>
have strongly keeled ventral scales (smooth in
<taxonomicName id="09854D1EFFCEFF97FA92FEFC56314307" box="[1319,1389,296,320]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCEFF97FA92FEFC56314307" box="[1319,1389,296,320]" italics="true" pageId="7" pageNumber="30">A. osa</emphasis>
</taxonomicName>
and
<taxonomicName id="09854D1EFFCEFF97FF22FE9852424323" box="[151,286,332,356]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FF22FE9852424323" box="[151,286,332,356]" italics="true" pageId="7" pageNumber="30">A. polylepis</emphasis>
</taxonomicName>
). Also, the majority of males in
<taxonomicName id="09854D1EFFCEFF97FD15FE9850064323" box="[672,858,332,356]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="tropidogaster">
<emphasis id="FCF1EA8FFFCEFF97FD15FE9850064323" box="[672,858,332,356]" italics="true" pageId="7" pageNumber="30">A. tropidogaster</emphasis>
</taxonomicName>
have a pair of greatly enlarged postcloacal scales (these scales usually not enlarged in
<taxonomicName id="09854D1EFFCEFF97FDFEFEA451C943CF" box="[587,661,368,392]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCEFF97FDFEFEA451C943CF" box="[587,661,368,392]" italics="true" pageId="7" pageNumber="30">A. osa</emphasis>
</taxonomicName>
and
<taxonomicName id="09854D1EFFCEFF97FD61FEA4500143CF" box="[724,861,368,392]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FD61FEA4500143CF" box="[724,861,368,392]" italics="true" pageId="7" pageNumber="30">A. polylepis</emphasis>
</taxonomicName>
). In the field, most individuals of
<taxonomicName id="09854D1EFFCEFF97FB4FFEA4561943CF" box="[1274,1349,368,392]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCEFF97FB4FFEA4561943CF" box="[1274,1349,368,392]" italics="true" pageId="7" pageNumber="30">A. osa</emphasis>
</taxonomicName>
and
<taxonomicName id="09854D1EFFCEFF97FA31FEA453A043EB" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FA31FEA453A043EB" italics="true" pageId="7" pageNumber="30">A. polylepis</emphasis>
</taxonomicName>
have a pale longitudinal lateral stripe (absent in Costa Rican populations of
<taxonomicName id="09854D1EFFCEFF97FBE8FE40578843EB" box="[1117,1236,404,428]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="FCF1EA8FFFCEFF97FBE8FE40578843EB" box="[1117,1236,404,428]" italics="true" pageId="7" pageNumber="30">A. cupreus</emphasis>
</taxonomicName>
). For variation in selected morphometric and scalation characters of
<taxonomicName id="09854D1EFFCEFF97FD7BFE6C50484397" box="[718,788,440,464]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCEFF97FD7BFE6C50484397" box="[718,788,440,464]" italics="true" pageId="7" pageNumber="30">A. osa</emphasis>
</taxonomicName>
see
<tableCitation id="83070326FFCEFF97FCF3FE6350C74388" box="[838,923,439,463]" captionStart="TABLE 3" captionStartId="9.[151,239,151,175]" captionText="TABLE 3. Selected measurements, proportions and scale characters of Anolis polylepis and A. osa. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." pageId="7" pageNumber="30">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="9AFA6615FFCEFF97FF22FBAD50FF46D7" box="[151,931,1145,1169]" httpUri="https://zenodo.org/record/199776/files/figure.png" pageId="7" pageNumber="30" targetBox="[297,1295,529,1116]" targetPageId="7">
<paragraph id="CE3A369DFFCEFF97FF22FBAD50FF46D7" blockId="7.[151,931,1145,1169]" box="[151,931,1145,1169]" pageId="7" pageNumber="30">
<emphasis id="FCF1EA8FFFCEFF97FF22FBAD524546D6" bold="true" box="[151,281,1145,1169]" pageId="7" pageNumber="30">FIGURE 4.</emphasis>
Hemipenis of
<taxonomicName id="09854D1EFFCEFF97FE02FBAD517B46D7" box="[439,551,1145,1168]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFCEFF97FE02FBAD517B46D7" box="[439,551,1145,1168]" italics="true" pageId="7" pageNumber="30">Anolis osa</emphasis>
</taxonomicName>
(SMF 89260). Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="9AFA6615FFCEFF97FF22F85956DD45E3" box="[151,1409,1933,1957]" httpUri="https://zenodo.org/record/199777/files/figure.png" pageId="7" pageNumber="30" targetBox="[293,1296,1225,1902]" targetPageId="7">
<paragraph id="CE3A369DFFCEFF97FF22F85956DD45E3" blockId="7.[151,1409,1933,1957]" box="[151,1409,1933,1957]" pageId="7" pageNumber="30">
<emphasis id="FCF1EA8FFFCEFF97FF22F859524545E2" bold="true" box="[151,281,1933,1957]" pageId="7" pageNumber="30">FIGURE 5.</emphasis>
Hemipenis of a suspected hybrid (SMF 89181) between
<taxonomicName id="09854D1EFFCEFF97FCCFF85A577945E2" box="[890,1061,1934,1957]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFCEFF97FCCFF85A577945E2" box="[890,1061,1934,1957]" italics="true" pageId="7" pageNumber="30">Anolis polylepis</emphasis>
</taxonomicName>
and
<emphasis id="FCF1EA8FFFCEFF97FBECF85A57FF45E2" box="[1113,1187,1934,1957]" italics="true" pageId="7" pageNumber="30">
<taxonomicName id="09854D1EFFCEFF97FBECF85A57C345E2" box="[1113,1183,1934,1957]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="30" phylum="Chordata" rank="species" species="osa">A. osa</taxonomicName>
.
</emphasis>
Scale bar = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="869F6516FFC1FF98FF7CFF4357794537" pageId="8" pageNumber="31" type="description">
<paragraph id="CE3A369DFFC1FF98FF7CFF43525046AF" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="31">
<emphasis id="FCF1EA8FFFC1FF98FF7CFF43514742F7" bold="true" box="[201,539,151,176]" pageId="8" pageNumber="31">
Description of the
<typeStatus id="113E883FFFC1FF98FE18FF4C514A42F7" box="[429,534,152,176]" pageId="8" pageNumber="31" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male as indicated by everted hemipenes and presence of large dewlap; SVL
<quantity id="097D9B78FFC1FF98FF67FF6852654293" box="[210,313,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" pageId="8" pageNumber="31" unit="mm" value="50.5">50.5 mm</quantity>
; tail length
<quantity id="097D9B78FFC1FF98FE0FFF6851454293" box="[442,537,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="1.06" pageId="8" pageNumber="31" unit="mm" value="106.0">106 mm</quantity>
, tail complete; tail compressed in cross section, tail height 2.0 mm, tail width
<quantity id="097D9B78FFC1FF98FACCFF68539A42B0" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="31" unit="mm" value="1.4">1.4 mm</quantity>
; axilla to groin distance
<quantity id="097D9B78FFC1FF98FE56FF0B511142B0" box="[483,589,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="8" pageNumber="31" unit="mm" value="20.8">20.8 mm</quantity>
; head length
<quantity id="097D9B78FFC1FF98FD5DFF0B500C42B0" box="[744,848,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="8" pageNumber="31" unit="mm" value="13.3">13.3 mm</quantity>
, head length/SVL ratio 0.26; snout length
<quantity id="097D9B78FFC1FF98FA89FF0B56C442B0" box="[1340,1432,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="8" pageNumber="31" unit="mm" value="6.6">6.6 mm</quantity>
; head width
<quantity id="097D9B78FFC1FF98FEA8FED05224435B" box="[285,376,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="31" unit="mm" value="8.2">8.2 mm</quantity>
; longest toe of adpressed hind limb reaching to a point between eye and snout; shank length
<quantity id="097D9B78FFC1FF98FF22FEF3525C4378" box="[151,256,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="8" pageNumber="31" unit="mm" value="14.6">14.6 mm</quantity>
, shank length/head length ratio 1.1; longest finger of extended forelimb reaching tip of snout; longest finger of adpressed forelimb reaches to anterior insertion of hind limbs. Scales on snout keeled (
<figureCitation id="56BE2A18FFC1FF98FAAEFE9856284323" box="[1307,1396,332,356]" captionStart="FIGURE 10" captionStartId="13.[151,255,1050,1074]" captionTargetBox="[151,1436,178,1028]" captionTargetId="figure@13.[151,1436,172,1029]" captionTargetPageId="13" captionText="FIGURE 10. Adult male of Anolis osa (not collected) at Puerto Escondido in life with extended dewlap." httpUri="https://zenodo.org/record/199780/files/figure.png" pageId="8" pageNumber="31">Fig. 10</figureCitation>
); 9 postrostrals; 10 scales between nasals; scales in distinct prefrontal depression slightly wrinkled; supraorbital semicircles well developed, separated by a minimum of 2 scales; supraorbital disc composed of 6 distinctly enlarged keeled scales; two elongated, strongly overlapping superciliaries, the anterior one much larger than the posterior scale; about 34 rows of small keeled scales extending between enlarged supraorbitals and superciliaries; interparietal scale well developed, 1.8 x 1.0 mm (length x width), surrounded by scales of moderate size; 2 scales present between interparietal and supraorbital semicircles; canthal ridge distinct, composed of 3 large (posterior two largest) and 6 small anterior canthal scales; 11 scales present between second canthals; 12 scales present between posterior canthals; 70 (right)63 (left) mostly keeled loreal scales in a maximum of 8 (right)9 (left) horizontal rows, with the scales of lower rows mostly keeled, and those of upper rows mostly tuberculated; subocular scales keeled, narrowly in contact with supralabials; 8 supralabials to level below center of eye; ear opening 0.7 x
<quantity id="097D9B78FFC1FF98FE8BFD0352CB40A8" box="[318,407,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="31" unit="mm" value="1.4">1.4 mm</quantity>
(length x height); mental distinctly wider than long, almost completely divided medially, bordered posteriorly by 8 postmentals (outer larger than inner); 7 infralabials to level below center of eye; sublabials undifferentiated; slightly keeled granular scales present on chin and throat; dewlap extending well onto chest, anterior insertion at level of posterior canthals, posterior insertion about 6.0 mm beyond level of axilla; dorsum of body with keeled granular scales, 2 medial rows of slightly enlarged scales, largest dorsal scales about 0.28 x
<quantity id="097D9B78FFC1FF98FF53FC58521741E3" box="[230,331,908,932]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="31" unit="mm" value="0.22">0.22 mm</quantity>
(length x width); about 54 medial dorsal scales in one head length; about 85 medial dorsal scales between axilla and groin; lateral scales keeled, granular and homogeneous, average size
<quantity id="097D9B78FFC1FF98FB15FC7B565B418F" box="[1184,1287,943,968]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="31" unit="mm" value="0.15">0.15 mm</quantity>
in diameter; ventrals at midbody smooth, bulging, subimbricate, about 0.23 x
<quantity id="097D9B78FFC1FF98FCC5FC00508F41AB" box="[880,979,980,1004]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="31" unit="mm" value="0.23">0.23 mm</quantity>
(length x width); about 54 ventral scales in one head length; about 94 ventral scales between axilla and groin; about 176 scales around midbody; caudal scales strongly keeled; caudal middorsal scales slightly enlarged, without whorls of enlarged scales; precloacal scales pointed and granular; postcloacal scales not enlarged; no tube-like axillary pocket present; scales on dorsal surface of forelimb slightly keeled, imbricate, about 0.25 x
<quantity id="097D9B78FFC1FF98FC8DFBB050C1463B" box="[824,925,1124,1148]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="31" unit="mm" value="0.27">0.27 mm</quantity>
(length x width); digital pads dilated, dilated pad about 2 times width of non-dilated scales on distal phalanx; distal phalanx narrower than and raised from dilated pad; 25 (right)25 (left) lamellae under phalanges IIIV of fourth toe; 9 scales under distal phalanx of fourth toe.
</paragraph>
<paragraph id="CE3A369DFFC1FF98FF7CFB2057134713" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="31">The completely everted hemipenis is a small unilobed organ; sulcus spermaticus bordered by well developed sulcal lips and opens at base of apex into a concave field without ornamentation; a small asulcate processus present; asulcal side of apex covered with fine calyces, truncus with transverse folds.</paragraph>
<paragraph id="CE3A369DFFC1FF98FF7CFAB457224787" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="31">The extended dewlap in life has 67 horizontal gorgetal-sternal rows with 2642 scales per row, rows somewhat irregular, some of them with pairs or triplets of scales, apical portion of dewlap between marginal pairs and last gorgetal-sternal row free of scales; modal number of anterior marginal pairs 45.</paragraph>
<paragraph id="CE3A369DFFC1FF98FF7CFA18560144FB" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="31">Coloration in life was recorded as follows: Dorsal ground color Dark Drab (119B) with a Pale Horn Color (92) broken longitudinal lateral stripe, edged diffusely by Mars Brown (223A) pigment; a Sepie (119) shoulder spot, edged posteriorly by a Yellow Ocher (123C) blotch; dorsum of head Dark Drab (119B) with a Hair Brown (119A) interorbital bar; dorsal surfaces of limbs Army Brown (219B) with indistinct Tawny Olive (223D) markings; tail Brick Red (132A) with indistinct Tawny Olive (223D) banding; tip of tail Tawny Olive (223D); ventral surfaces Glaucus (79); dewlap Orange Yellow (18), basal portion suffused with Raw Umber (23), gorgetals dirty white; iris Mars Brown (219A). Coloration after one year preservation in 70% ethanol agrees well with this description.</paragraph>
<paragraph id="CE3A369DFFC1FF98FF7CF91357794537" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="31">
<emphasis id="FCF1EA8FFFC1FF98FF7CF913526244A7" bold="true" box="[201,318,1735,1760]" pageId="8" pageNumber="31">Variation.</emphasis>
The
<typeStatus id="113E883FFFC1FF98FECAF91D52B644A6" box="[383,490,1737,1761]" pageId="8" pageNumber="31" type="paratype">paratypes</typeStatus>
agree well with the
<typeStatus id="113E883FFFC1FF98FD69F91C506244A7" box="[732,830,1736,1760]" pageId="8" pageNumber="31" type="holotype">holotype</typeStatus>
in general appearance, morphometrics and scalation (see
<tableCitation id="83070326FFC1FF98FF79F93852424543" box="[204,286,1772,1796]" captionStart="TABLE 1" captionStartId="3.[151,239,543,567]" captionTargetBox="[151,1298,594,2025]" captionTargetPageId="3" captionText="TABLE 1. PCA variable loadings for the first and second principal components. For abbreviations see text." httpUri="http://table.plazi.org/id/9AFA6615FFCAFF93FF22FDCB564E4070" pageId="8" pageNumber="31" tableUuid="9AFA6615FFCAFF93FF22FDCB564E4070">Table 1</tableCitation>
). We documented variation in relative hind leg length in this species. Of
<specimenCount id="D883FD14FFC1FF98FBEBF93857AB4543" box="[1118,1271,1772,1796]" pageId="8" pageNumber="31" type="generic">66 specimens</specimenCount>
examined, the longest toe of the adpressed hind leg reached to mid-eye in 5 individuals (7.6%), to anterior margin of eye in 33 individuals (50.0%), and to a point between eye and snout in 28 individuals (42.4%). For variation in nasal region see
<tableCitation id="83070326FFC1FF98FF77F88C52494537" box="[194,277,1880,1904]" captionStart="TABLE 2" captionStartId="5.[151,239,1085,1109]" captionTargetBox="[151,1271,1180,1672]" captionTargetPageId="5" captionText="TABLE 2. Frequency distribution of nasal region and dorsal pattern types in Anolis osa and A. polylepis." httpUri="http://table.plazi.org/id/9AFA6615FFCCFF95FF22FBE957AB4612" pageId="8" pageNumber="31" tableUuid="9AFA6615FFCCFF95FF22FBE957AB4612">Table 2</tableCitation>
and
<figureCitation id="56BE2A18FFC1FF98FEFEF88C52F64537" box="[331,426,1880,1904]" captionStart="FIGURE 1" captionStartId="2.[151,255,1687,1711]" captionTargetBox="[292,1293,208,1658]" captionTargetId="figure@2.[287,1300,194,1666]" captionTargetPageId="2" captionText="FIGURE 1. Designation of nasal scalation types. (a) Type A (UCR 20713); (b) Type B (SMF 89430); (c) Type C (SMF 89210); (d) Type D (UCR 14257); (e) Type E (UCR 20732). Scale bar = 1.0 mm. See Text for details." httpUri="https://zenodo.org/record/199773/files/figure.png" pageId="8" pageNumber="31">Figure 1</figureCitation>
. For variation in dorsal pattern see
<tableCitation id="83070326FFC1FF98FC82F88C50D64537" box="[823,906,1880,1904]" captionStart="TABLE 2" captionStartId="5.[151,239,1085,1109]" captionTargetBox="[151,1271,1180,1672]" captionTargetPageId="5" captionText="TABLE 2. Frequency distribution of nasal region and dorsal pattern types in Anolis osa and A. polylepis." httpUri="http://table.plazi.org/id/9AFA6615FFCCFF95FF22FBE957AB4612" pageId="8" pageNumber="31" tableUuid="9AFA6615FFCCFF95FF22FBE957AB4612">Table 2</tableCitation>
and
<figureCitation id="56BE2A18FFC1FF98FC75F88C577D4537" box="[960,1057,1880,1904]" captionStart="FIGURE 2" captionStartId="6.[151,255,812,836]" captionTargetBox="[151,1427,201,773]" captionTargetId="figure@6.[151,1436,194,791]" captionTargetPageId="6" captionText="FIGURE 2. Designation of dorsal color pattern types." httpUri="https://zenodo.org/record/199774/files/figure.png" pageId="8" pageNumber="31">Figure 2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="869F6516FFC1FF9AFF7CF8A8571F43B3" lastPageId="10" lastPageNumber="33" pageId="8" pageNumber="31" type="etymology">
<paragraph id="CE3A369DFFC1FF98FF7CF8A8508445FF" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="31">
<emphasis id="FCF1EA8FFFC1FF98FF7CF8A8520C45D3" bold="true" box="[201,336,1916,1940]" pageId="8" pageNumber="31">Etymology.</emphasis>
The name
<taxonomicName id="09854D1EFFC1FF98FE7BF8A952A945D3" box="[462,501,1917,1940]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="31" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFC1FF98FE7BF8A952A945D3" box="[462,501,1917,1940]" italics="true" pageId="8" pageNumber="31">osa</emphasis>
</taxonomicName>
is used as a noun in apposition and in reference to the Peninsula de
<taxonomicName id="09854D1EFFC1FF98FB4FF8A8567A45D3" box="[1274,1318,1916,1940]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="36" phylum="Chordata" rank="species" species="osa">Osa</taxonomicName>
where the
<typeStatus id="113E883FFFC1FF98FF22F875539B45FE" box="[151,199,1953,1977]" pageId="8" pageNumber="31">type</typeStatus>
series of the species was collected and where it is probably restricted.
</paragraph>
<caption id="9AFA6615FFC0FF99FF22FF4350934288" pageId="9" pageNumber="32">
<paragraph id="CE3A369DFFC0FF99FF22FF4350934288" blockId="9.[151,1436,151,833]" pageId="9" pageNumber="32">
<emphasis id="FCF1EA8FFFC0FF99FF22FF43525442E8" bold="true" box="[151,264,151,175]" pageId="9" pageNumber="32">TABLE 3.</emphasis>
Selected measurements, proportions and scale characters of
<taxonomicName id="09854D1EFFC0FF99FC38FF4C576442E8" box="[909,1080,152,175]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="32" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="FCF1EA8FFFC0FF99FC38FF4C576442E8" box="[909,1080,152,175]" italics="true" pageId="9" pageNumber="32">Anolis polylepis</emphasis>
</taxonomicName>
and
<emphasis id="FCF1EA8FFFC0FF99FBDEFF4C57E842E8" box="[1131,1204,152,175]" italics="true" pageId="9" pageNumber="32">
<taxonomicName id="09854D1EFFC0FF99FBDEFF4C57EC42E8" box="[1131,1200,152,175]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="32" phylum="Chordata" rank="species" species="osa">A. osa</taxonomicName>
.
</emphasis>
Range is followed by mean value and standard deviation in parentheses. For abbreviations see text.
</paragraph>
</caption>
<paragraph id="CE3A369DFFC0FF99FD40FF3E57254346" blockId="9.[151,1436,151,833]" box="[757,1145,234,257]" pageId="9" pageNumber="32">
<emphasis id="FCF1EA8FFFC0FF99FD40FF3E57254346" box="[757,1145,234,257]" italics="true" pageId="9" pageNumber="32">
<taxonomicName id="09854D1EFFC0FF99FD40FF3E502F4346" box="[757,883,234,257]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="32" phylum="Chordata" rank="species" species="polylepis">A. polylepis</taxonomicName>
<taxonomicName id="09854D1EFFC0FF99FB82FF3E57254346" box="[1079,1145,234,257]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="32" phylum="Chordata" rank="species" species="osa">A. osa</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="CE3A369DFFC0FF99FD40FEDE57304366" blockId="9.[151,1436,151,833]" box="[757,1132,266,290]" pageId="9" pageNumber="32">
<date id="BA3B105DFFC0FF99FD40FEDE57174365" box="[757,1099,266,290]" pageId="9" pageNumber="32" value="1947-03-03">3 47 3</date>
22
</paragraph>
<paragraph id="CE3A369DFFC0FF99FD40FEFE57304306" blockId="9.[151,1436,151,833]" box="[757,1132,298,321]" pageId="9" pageNumber="32">Ƥ 24 Ƥ 19</paragraph>
<paragraph id="CE3A369DFFC0FF9AFF2AFE81517D43C0" blockId="9.[151,1436,151,833]" lastBlockId="10.[151,1436,151,501]" lastPageId="10" lastPageNumber="33" pageId="9" pageNumber="32">
SVL 3 42.259.4 (51.3 ± 3.76) 49.955.4 (50.2 ± 2.28) Ƥ 39.653.5 (47.3 ± 3.52) 41.950.7 (46.4 ± 2.60) Tail length 3 80.1117.0 (102.1 ± 10.28) 84.9107.2 (97.3 ± 6.88) Ƥ 70.0101.5 (86.3 ± 8.29) 73.193.0 (82.0 ± 4.59) HL 3 10.814.4 (12.8 ± 0.88) 11.313.9 (12.4 ± 0.62) Ƥ 10.113.7 (11.9 ± 0.78) 10.412.8 (11.5 ± 0.60) HW 3 6.78.5 (7.7 ± 0.48) 7.08.2 (7.5 ± 0.35) Ƥ 6.38.5 (7.3 ± 0.56) 6.77.9 (7.1 ± 0.36) Shank length 3 11.616.0 (13.9 ± 1.12) 12.215.0 (13.6 ± 0.66) Ƥ 10.514.0 (12.4 ± 0.98) 11.013.2 (12.3 ± 0.65) Axillagroin distance 3 16.326.1 (21.2 ± 2.16) 18.524.4 (21.3 ± 1.32) Ƥ 15.524.1 (19.87 ± 2.14) 17.829.4 (20.7 ± 2.60)
<emphasis id="FCF1EA8FFFC3FF9AFF7CFF4C528E42F7" bold="true" box="[201,466,151,176]" pageId="10" pageNumber="33">Natural history notes.</emphasis>
<taxonomicName id="09854D1EFFC3FF9AFE6FFF4C510D42F7" box="[474,593,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="33" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFC3FF9AFE6FFF4C510D42F7" box="[474,593,152,176]" italics="true" pageId="10" pageNumber="33">Anolis osa</emphasis>
</taxonomicName>
is an extremely common anole that reaches high population densities. At most sites on the Peninsula de
<taxonomicName id="09854D1EFFC3FF9AFE4EFF68517B4293" box="[507,551,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="36" phylum="Chordata" rank="species" species="osa">Osa</taxonomicName>
it was the most abundant species of anole. During daytime, most individuals were observed perching head down on the lower parts (
<quantity id="097D9B78FFC3FF9AFCBFFF0B50D542B0" box="[778,905,223,247]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="10" pageNumber="33" unit="cm" value="75.0" valueMax="120.0" valueMin="30.0">30120 cm</quantity>
above the ground) of trunks of small trees and other plants such as heliconias (
<taxonomicName id="09854D1EFFC3FF9AFE4AFED051FD435B" box="[511,673,260,284]" class="Liliopsida" family="Heliconiaceae" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="33" phylum="Tracheophyta" rank="family">Heliconiaceae</taxonomicName>
) and banana plants (
<taxonomicName id="09854D1EFFC3FF9AFC3FFED0575D435B" box="[906,1025,260,284]" class="Liliopsida" family="Musaceae" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="33" phylum="Tracheophyta" rank="family">Musaceae</taxonomicName>
). Males of
<taxonomicName id="09854D1EFFC3FF9AFBCAFED0579A435B" box="[1151,1222,260,284]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="33" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFC3FF9AFBCAFED0579A435B" box="[1151,1222,260,284]" italics="true" pageId="10" pageNumber="33">A. osa</emphasis>
</taxonomicName>
are very territorial and will show aggressive display behavior whenever another adult male is placed in reaching distance (less than about
<quantity id="097D9B78FFC3FF9AFF69FE9852694323" box="[220,309,332,356]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="33" unit="cm" value="150.0">150 cm</quantity>
). The species was studied in detail by
<bibRefCitation id="AA144B6CFFC3FF9AFD53FE9850CB4323" author="Andrews" box="[742,919,332,356]" pageId="10" pageNumber="33" refString="Andrews, R. M. (1971) Structural habitat and time budget of a tropical Anolis lizard. Ecology, 52, 262 - 279." type="journal article" year="1971">Andrews (1971</bibRefCitation>
,
<bibRefCitation id="AA144B6CFFC3FF9AFC16FE9850824323" author="Andrews" box="[931,990,332,356]" pageId="10" pageNumber="33" refString="Andrews, R. M. (1983) Norops polylepis (lagartija, anole, anolis lizard.). Pp. 409 - 410. In Janzen, D. H. (Ed.), Costa Rican Natural History. University of Chicago Press, Chicago, Illinois, USA." type="book chapter" year="1983">1983</bibRefCitation>
), and
<bibRefCitation id="AA144B6CFFC3FF9AFB96FE98579A4323" author="Savage" box="[1059,1222,332,356]" pageId="10" pageNumber="33" refString="Savage, J. M. (2002) The amphibians and Reptiles of Costa Rica. A Herpetofauna between two Continents, between two Seas. University of Chicago Press, Chicago, Illinois, USA." type="book" year="2002">Savage (2002)</bibRefCitation>
summarized much of the information provided by her.
</paragraph>
<paragraph id="CE3A369DFFC0FF99FF2AFC8157AC47AB" pageId="9" pageNumber="32">
<table id="BC85C43DFFC0006FFF22FC8056C047AB" box="[151,1436,852,1516]" gridcols="4" gridrows="6" pageId="9" pageNumber="32">
<tr id="70B534DFFFC0006FFF22FC8056C0412B" box="[151,1436,852,876]" gridrow="0" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22FC805118412B" box="[151,580,852,876]" gridcol="0" gridrow="0" pageId="9" pageNumber="32">Tail length / SVL</th>
<th id="33645DA3FFC0006FFDCAFC8051CF412B" box="[639,659,852,876]" gridcol="1" gridrow="0" pageId="9" pageNumber="32">3</th>
<th id="33645DA3FFC0006FFD40FC8050AE412B" box="[757,1010,852,876]" gridcol="2" gridrow="0" pageId="9" pageNumber="32">1.802.16 (2.00 ± 0.09)</th>
<th id="33645DA3FFC0006FFB82FC8056C0412B" box="[1079,1436,852,876]" gridcol="3" gridrow="0" pageId="9" pageNumber="32">1.742.06 (1.94 ± 0.10)</th>
</tr>
<tr id="70B534DFFFC0006FFF22FC5456C0462B" box="[151,1436,896,1132]" gridrow="1" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22FC545118462B" box="[151,580,896,1132]" gridcol="0" gridrow="1" pageId="9" pageNumber="32">Tail diameter vertical / horizontal HL / SVL HL / HW</th>
<td id="33645DA3FFC0006FFDCAFC5451CF462B" box="[639,659,896,1132]" gridcol="1" gridrow="1" pageId="9" pageNumber="32">Ƥ 3 Ƥ 3 Ƥ 3</td>
<td id="33645DA3FFC0006FFD40FC5450AE462B" box="[757,1010,896,1132]" gridcol="2" gridrow="1" pageId="9" pageNumber="32">1.691.91 (1.82 ± 0.06) 1.201.85 (1.46 ± 0.14) 1.231.55 (1.36 ± 0.10) 0.240.27 (0.25 ± 0.01) 0.240.27 (0.25 ± 0.01) 1.551.80 (1.66 ± 0.06)</td>
<td id="33645DA3FFC0006FFB82FC5456C0462B" box="[1079,1436,896,1132]" gridcol="3" gridrow="1" pageId="9" pageNumber="32">1.681.87 (1.78 ± 0.06 1.261.69 (1.47 ± 0.16) 1.201.46(1.33 ± 0.08) 0.230.26 (0.25 ± 0.01) 0.24 0.26 (0.25 ± 0.01) 1.531.76 (1.65 ± 0.06)</td>
</tr>
<tr id="70B534DFFFC0006FFF22FB5456C04706" box="[151,1436,1152,1345]" gridrow="2" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22FB5451184706" box="[151,580,1152,1345]" gridcol="0" gridrow="2" pageId="9" pageNumber="32">Shank length / SVL Axillagroin distance / SVL</th>
<td id="33645DA3FFC0006FFDCAFB5451CF4706" box="[639,659,1152,1345]" gridcol="1" gridrow="2" pageId="9" pageNumber="32">Ƥ 3 Ƥ 3 Ƥ</td>
<td id="33645DA3FFC0006FFD40FB5450AE4706" box="[757,1010,1152,1345]" gridcol="2" gridrow="2" pageId="9" pageNumber="32">1.521.77 (1.63 ± 0.06) 0.250.30 (0.27 ± 0.01) 0.230.28 (0.26 ± 0.01) 0.370.46 (0.41 ± 0.02) 0.390.46 (0.42 ± 0.02)</td>
<td id="33645DA3FFC0006FFB82FB5456C04706" box="[1079,1436,1152,1345]" gridcol="3" gridrow="2" pageId="9" pageNumber="32">1.521.68 (1.61 ± 0.04) 0.250.30 (0.27 ± 0.01) 0.250.28 (0.26 ± 0.01) 0.380.46 (0.42 ± 0.02) 0.400.60 (0.45 ± 0.04)</td>
</tr>
<tr id="70B534DFFFC0006FFF22FA8156C0472B" box="[151,1436,1365,1388]" gridrow="3" pageId="9" pageNumber="32" rowspan-1="1">
<th id="33645DA3FFC0006FFF22FA815118472B" box="[151,580,1365,1388]" gridcol="0" gridrow="3" pageId="9" pageNumber="32">Subdigital lamellae of 4th toe</th>
<td id="33645DA3FFC0006FFD40FA8150AE472B" box="[757,1010,1365,1388]" gridcol="2" gridrow="3" pageId="9" pageNumber="32">2129 (24.66 ± 1.52)</td>
<td id="33645DA3FFC0006FFB82FA8156C0472B" box="[1079,1436,1365,1388]" gridcol="3" gridrow="3" pageId="9" pageNumber="32">2228 (25.34 ± 1.71)</td>
</tr>
<tr id="70B534DFFFC0006FFF22FA5456C04786" box="[151,1436,1408,1473]" gridrow="4" pageId="9" pageNumber="32" rowspan-1="1">
<th id="33645DA3FFC0006FFF22FA5451184786" box="[151,580,1408,1473]" gridcol="0" gridrow="4" pageId="9" pageNumber="32">Number of scales between SS Number of scales between IP and SS</th>
<td id="33645DA3FFC0006FFD40FA5450AE4786" box="[757,1010,1408,1473]" gridcol="2" gridrow="4" pageId="9" pageNumber="32">15 (2.69 ± 0.83) 27 (3.80 ± 1.03)</td>
<td id="33645DA3FFC0006FFB82FA5456C04786" box="[1079,1436,1408,1473]" gridcol="3" gridrow="4" pageId="9" pageNumber="32">14 (2.85 ± 0.73) 15 (3.98 ± 0.94)</td>
</tr>
<tr id="70B534DFFFC0006FFF22FA0156C047AB" box="[151,1436,1493,1516]" gridrow="5" pageId="9" pageNumber="32" rowspan-1="1">
<th id="33645DA3FFC0006FFF22FA01511847AB" box="[151,580,1493,1516]" gridcol="0" gridrow="5" pageId="9" pageNumber="32">Number of scales between SO and SPL</th>
<td id="33645DA3FFC0006FFD40FA0150AE47AB" box="[757,1010,1493,1516]" gridcol="2" gridrow="5" pageId="9" pageNumber="32">01 (0.75 ± 0.44)</td>
<td id="33645DA3FFC0006FFB82FA0156C047AB" box="[1079,1436,1493,1516]" gridcol="3" gridrow="5" pageId="9" pageNumber="32">01 (0.41 ± 0.50)</td>
</tr>
</table>
</paragraph>
<paragraph id="CE3A369DFFC0FF99FF2AF9D456444586" pageId="9" pageNumber="32">
<table id="BC85C43DFFC0006FFF22F9D456C04586" box="[151,1436,1536,1985]" gridcols="3" gridrows="7" pageId="9" pageNumber="32">
<tr id="70B534DFFFC0006FFF22F9D456C04406" box="[151,1436,1536,1601]" gridrow="0" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22F9D451994406" box="[151,709,1536,1601]" gridcol="0" gridrow="0" pageId="9" pageNumber="32">Number of SPL to level below center of eye Number of INL to level below center of eye</th>
<th id="33645DA3FFC0006FFD40F9D450BF4406" box="[757,995,1536,1601]" gridcol="1" gridrow="0" pageId="9" pageNumber="32">69 (7.17 ± 0.65) 59 (7.34 ± 0.81)</th>
<th id="33645DA3FFC0006FFB82F9D456C04406" box="[1079,1436,1536,1601]" gridcol="2" gridrow="0" pageId="9" pageNumber="32">79 (7.56 ± 0.59) 610 (7.66 ± 0.85)</th>
</tr>
<tr id="70B534DFFFC0006FFF22F98156C0442B" box="[151,1436,1621,1644]" gridrow="1" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22F9815199442B" box="[151,709,1621,1644]" gridcol="0" gridrow="1" pageId="9" pageNumber="32">Total number of loreals</th>
<td id="33645DA3FFC0006FFD40F98150BF442B" box="[757,995,1621,1644]" gridcol="1" gridrow="1" pageId="9" pageNumber="32">3792 (57.69 ± 11.38)</td>
<td id="33645DA3FFC0006FFB82F98156C0442B" box="[1079,1436,1621,1644]" gridcol="2" gridrow="1" pageId="9" pageNumber="32">4183 (56.78 ± 8.91)</td>
</tr>
<tr id="70B534DFFFC0006FFF22F95456C04486" box="[151,1436,1664,1729]" gridrow="2" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22F95451994486" box="[151,709,1664,1729]" gridcol="0" gridrow="2" pageId="9" pageNumber="32">Number of horizontal loreal scale rows Number of postrostrals</th>
<td id="33645DA3FFC0006FFD40F95450BF4486" box="[757,995,1664,1729]" gridcol="1" gridrow="2" pageId="9" pageNumber="32">610 (8.06 ± 1.05) 711 (7.90 ± 0.87)</td>
<td id="33645DA3FFC0006FFB82F95456C04486" box="[1079,1436,1664,1729]" gridcol="2" gridrow="2" pageId="9" pageNumber="32">610 (8.02 ± 0.88) 711 (8.12 ± 0.93)</td>
</tr>
<tr id="70B534DFFFC0006FFF22F90156C044AB" box="[151,1436,1749,1772]" gridrow="3" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22F901519944AB" box="[151,709,1749,1772]" gridcol="0" gridrow="3" pageId="9" pageNumber="32">Number of postmentals</th>
<td id="33645DA3FFC0006FFD40F90150BF44AB" box="[757,995,1749,1772]" gridcol="1" gridrow="3" pageId="9" pageNumber="32">69 (6.82 ± 0.96)</td>
<td id="33645DA3FFC0006FFB82F90156C044AB" box="[1079,1436,1749,1772]" gridcol="2" gridrow="3" pageId="9" pageNumber="32">610 (7.00 ± 1.02)</td>
</tr>
<tr id="70B534DFFFC0006FFF22F8D456C04506" box="[151,1436,1792,1857]" gridrow="4" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22F8D451994506" box="[151,709,1792,1857]" gridcol="0" gridrow="4" pageId="9" pageNumber="32">Number of scales between nasals Number of scales between 2nd canthals</th>
<td id="33645DA3FFC0006FFD40F8D450BF4506" box="[757,995,1792,1857]" gridcol="1" gridrow="4" pageId="9" pageNumber="32">711 (8.52 ± 1.19) 716 (10.49 ± 1.66)</td>
<td id="33645DA3FFC0006FFB82F8D456C04506" box="[1079,1436,1792,1857]" gridcol="2" gridrow="4" pageId="9" pageNumber="32">711 (8.59 ± 1.22) 913 (10.63 ± 1.18)</td>
</tr>
<tr id="70B534DFFFC0006FFF22F88156C0452B" box="[151,1436,1877,1900]" gridrow="5" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22F8815199452B" box="[151,709,1877,1900]" gridcol="0" gridrow="5" pageId="9" pageNumber="32">Number of scales between posterior canthals</th>
<td id="33645DA3FFC0006FFD40F88150BF452B" box="[757,995,1877,1900]" gridcol="1" gridrow="5" pageId="9" pageNumber="32">818 (12.20 ± 1.92)</td>
<td id="33645DA3FFC0006FFB82F88156C0452B" box="[1079,1436,1877,1900]" gridcol="2" gridrow="5" pageId="9" pageNumber="32">916 (12.41 ± 1.43)</td>
</tr>
<tr id="70B534DFFFC0006FFF22F85456C04586" box="[151,1436,1920,1985]" gridrow="6" pageId="9" pageNumber="32">
<th id="33645DA3FFC0006FFF22F85451994586" box="[151,709,1920,1985]" gridcol="0" gridrow="6" pageId="9" pageNumber="32">Number of medial dorsal scales in one head length Number of medial ventral scales in one head length</th>
<td id="33645DA3FFC0006FFD40F85450BF4586" box="[757,995,1920,1985]" gridcol="1" gridrow="6" pageId="9" pageNumber="32">3576 (53.10 ± 7.48) 3662 (47.23 ± 5.97)</td>
<td id="33645DA3FFC0006FFB82F85456C04586" box="[1079,1436,1920,1985]" gridcol="2" gridrow="6" pageId="9" pageNumber="32">4468 (56.29 ± 6.67) 3858 (46.83 ± 4.71)</td>
</tr>
</table>
</paragraph>
<paragraph id="CE3A369DFFC3FF9AFF7CFE47571F43B3" blockId="10.[151,1436,151,501]" pageId="10" pageNumber="33">
<emphasis id="FCF1EA8FFFC3FF9AFF7CFE4752AD43EB" bold="true" box="[201,497,403,428]" pageId="10" pageNumber="33">Geographic distribution.</emphasis>
<collectingCountry id="B692760DFFC3FF9AFE4DFE40514B43EB" box="[504,535,404,428]" name="American Samoa" pageId="10" pageNumber="33">As</collectingCountry>
currently known,
<taxonomicName id="09854D1EFFC3FF9AFD50FE40500743EB" box="[741,859,404,428]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="33" phylum="Chordata" rank="species" species="osa">
<emphasis id="FCF1EA8FFFC3FF9AFD50FE40500743EB" box="[741,859,404,428]" italics="true" pageId="10" pageNumber="33">Anolis osa</emphasis>
</taxonomicName>
is restricted to the Peninsula de
<taxonomicName id="09854D1EFFC3FF9AFB7DFE4057AA43EB" box="[1224,1270,404,428]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="36" phylum="Chordata" rank="species" species="osa">Osa</taxonomicName>
,
<collectingCountry id="B692760DFFC3FF9AFAB7FE4056DC43EB" box="[1282,1408,404,428]" name="Costa Rica" pageId="10" pageNumber="33">Costa Rica</collectingCountry>
. It occurs both in protected areas (Corcovado National Park) and in disturbed areas such as at the
<typeStatus id="113E883FFFC3FF9AFB63FE6C565A4397" box="[1238,1286,440,464]" pageId="10" pageNumber="33">type</typeStatus>
locality. The documented vertical range of the species is from sea level to about
<quantity id="097D9B78FFC3FF9AFC32FE08509243B3" box="[903,974,476,500]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="33" unit="m" value="550.0">550 m</quantity>
elevation.
</paragraph>
</subSubSection>
</treatment>
</document>