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102 lines
14 KiB
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<document id="2C717C660988E294E5DF5592817DBEDD" ID-DOI="10.1016/j.jcz.2023.02.004" ID-ISSN="1873-2674" ID-Zenodo-Dep="8164274" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1689773830888" checkinUser="felipe" docAuthor="Gąsiorek, Piotr" docDate="2023" docId="03EE752FFF91FFD8FFF6FA64FBE0FAE8" docLanguage="en" docName="ZoolAnz.304.49-60.pdf" docOrigin="Zoologischer Anzeiger 304" docSource="http://dx.doi.org/10.1016/j.jcz.2023.02.004" docStyle="DocumentStyle:32483D6D135DE2FF927BA3F1400E0FFD.1:ZoolAnz.2022-.journal_article" docStyleId="32483D6D135DE2FF927BA3F1400E0FFD" docStyleName="ZoolAnz.2022-.journal_article" docStyleVersion="1" docTitle="Echiniscus becki" docType="treatment" docVersion="3" lastPageNumber="55" masterDocId="FFD70D57FF97FFDEFF92FFFDFFC1FFA9" masterDocTitle="Eventually tested: Phylogenetic position of “ Testechiniscus ” meridionalis (Murray, 1906) (Heterotardigrada) revealed" masterLastPageNumber="60" masterPageNumber="49" pageNumber="55" updateTime="1689935945562" updateUser="tatiana">
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<mods:title id="FE40AF4EE940E853AF82AB9A31D7C234">Eventually tested: Phylogenetic position of “ Testechiniscus ” meridionalis (Murray, 1906) (Heterotardigrada) revealed</mods:title>
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<mods:namePart id="7560F37902EEA4708D1C826E9702A4A7">Gąsiorek, Piotr</mods:namePart>
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<mods:title id="75023E520BBC772784336B1350A698D2">Zoologischer Anzeiger</mods:title>
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<mods:part id="28580D7228A67520A90EAAAB842879EE">
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<mods:date id="D3C463B7C76A6B27C2C18313A2CAB131">2023</mods:date>
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<mods:number id="538172DE778B7BFE85F60787A4BE065F">2023-05-31</mods:number>
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<mods:number id="479055498A9ED685012DEA68D8F90A3A">304</mods:number>
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<mods:url id="D271C37030DC89E0B6D41E93B8B35078">http://dx.doi.org/10.1016/j.jcz.2023.02.004</mods:url>
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<mods:identifier id="232A20753A9450BFD2DC2958524B7973" type="DOI">10.1016/j.jcz.2023.02.004</mods:identifier>
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<mods:identifier id="045995EA8B942CB9859F4E4583FD7EA1" type="ISSN">1873-2674</mods:identifier>
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<treatment id="03EE752FFF91FFD8FFF6FA64FBE0FAE8" LSID="urn:lsid:plazi:treatment:03EE752FFF91FFD8FFF6FA64FBE0FAE8" httpUri="http://treatment.plazi.org/id/03EE752FFF91FFD8FFF6FA64FBE0FAE8" lastPageNumber="55" pageId="6" pageNumber="55">
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<subSubSection id="C35D97B2FF91FFD8FFF6FA64FF15FA6E" pageId="6" pageNumber="55" type="nomenclature">
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<paragraph id="8BF8C439FF91FFD8FFF6FA64FF15FA6E" blockId="6.[100,754,1432,1479]" pageId="6" pageNumber="55">
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<heading id="D0B07355FF91FFD8FFF6FA64FF15FA6E" centered="true" fontSize="36" level="2" pageId="6" pageNumber="55" reason="3">
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<emphasis id="B933182BFF91FFD8FFF6FA64FF15FA6E" bold="true" italics="true" pageId="6" pageNumber="55">
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3.3. Revised morphological characters for
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<taxonomicName id="4C47BFBAFF91FFD8FE4CFA65FDAAFA02" ID-CoL="6DW2M" baseAuthorityName="Schuster and Grigarick" baseAuthorityYear="1966" box="[478,619,1432,1451]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="55" phylum="Tardigrada" rank="species" species="becki">Echiniscus becki</taxonomicName>
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and
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<taxonomicName id="4C47BFBAFF91FFD8FD05FA65FF15FA6E" authorityName="Gąsiorek" authorityYear="2023" baseAuthorityName="Ito" baseAuthorityYear="1993" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="55" phylum="Tardigrada" rank="species" species="laterosetosus">Echiniscus laterosetosus</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D97B2FF91FFD8FF16FA11FBE0FAE8" pageId="6" pageNumber="55" type="discussion">
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<paragraph id="8BF8C439FF91FFD8FF16FA11FC1CFBD7" blockId="6.[100,770,1516,1982]" lastBlockId="6.[818,1487,1131,1345]" pageId="6" pageNumber="55">
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An analysis of the type material of
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<taxonomicName id="4C47BFBAFF91FFD8FE56FA10FDC6FA56" baseAuthorityName="Schuster and Grigarick" baseAuthorityYear="1966" box="[452,519,1516,1536]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="55" phylum="Tardigrada" rank="species" species="becki">
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<emphasis id="B933182BFF91FFD8FE56FA10FDC6FA56" bold="true" box="[452,519,1516,1536]" italics="true" pageId="6" pageNumber="55">E. becki</emphasis>
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</taxonomicName>
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revealed new traits crucial in determining its phylogenetic position. The dorsal plate sculpturing comprises tightly arranged, polygonal endocuticular pillars (
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<figureCitation id="137CD8BCFF91FFD8FD3CF9D9FD35F99E" box="[686,756,1572,1591]" captionStart="Fig" captionStartId="9.[100,130,1015,1032]" captionTargetBox="[107,766,149,986]" captionTargetId="figure-638@9.[106,767,148,987]" captionTargetPageId="9" captionText="Fig. 9. Cephalic body portion of Echiniscus becki (PCM, paratype, female): A – dorsal view, B – ventral view (arrowheads indicate black crystalline eyes; large subcephalic plates visible as dark grey, broad belts bent in a boomerang-like shape). Scale bars = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.8164294" httpUri="https://zenodo.org/record/8164294/files/figure.png" pageId="6" pageNumber="55">Fig. 9A</figureCitation>
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); they lack centrally positioned pores typical for the
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<emphasis id="B933182BFF91FFD8FDF8F9BDFCC3F9FA" bold="true" box="[618,770,1600,1619]" italics="true" pageId="6" pageNumber="55">blumi-canadensis</emphasis>
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complex of species and
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<taxonomicName id="4C47BFBAFF91FFD8FEADF9A1FD50F9C6" authority="(Gasiorek et al., 2018)" baseAuthorityName="Gasiorek" baseAuthorityYear="2018" box="[319,657,1628,1647]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="55" phylum="Tardigrada" rank="genus">
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<emphasis id="B933182BFF91FFD8FEADF9A1FE7AF9C6" bold="true" box="[319,443,1628,1647]" italics="true" pageId="6" pageNumber="55">Testechiniscus</emphasis>
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(
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<bibRefCitation id="EFD6B9C8FF91FFD8FE5BF9A1FD46F9C6" author="Gasiorek, P. & Stec, D. & Zawierucha, K. & Kristensen, R. M. & Michalczyk, L." box="[457,647,1628,1647]" pageId="6" pageNumber="55" pagination="261 - 297" refId="ref10116" refString="Gasiorek, P., Stec, D., Zawierucha, K., Kristensen, R. M., Michalczyk, L., 2018. Revision of Testechiniscus Kristensen, 1987 (Heterotardigrada: Echiniscidae) refutes the polartemperate distribution of the genus. Zootaxa 4472, 261 - 297. https: // doi. org / 10.11646 / zootaxa. 4472.2.3." type="journal article" year="2018">Gąsiorek et al., 2018</bibRefCitation>
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)
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</taxonomicName>
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. In contrary to the original description (
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<bibRefCitation id="EFD6B9C8FF91FFD8FEFCF985FD44F922" author="Schuster, R. O. & Grigarick, A. A." box="[366,645,1656,1675]" pageId="6" pageNumber="55" pagination="127 - 130" refId="ref12551" refString="Schuster, R. O., Grigarick, A. A., 1966. New Tardigrada from Western North America: II, Echiniscus. Proc. Biol. Soc. Wash. 79, 127 - 130." type="journal article" year="1966">Schuster and Grigarick, 1966</bibRefCitation>
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), the species does not have dark red eye spots, which always dissolve in Hoyer’ s medium, but large black crystalline eyes identifiable in all examined
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<typeStatus id="54FC7A9BFF91FFD8FFF6F931FF7EF976" box="[100,191,1740,1759]" pageId="6" pageNumber="55">paratypes</typeStatus>
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(
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<figureCitation id="137CD8BCFF91FFD8FF40F936FED8F976" box="[210,281,1739,1759]" captionStart="Fig" captionStartId="9.[100,130,1015,1032]" captionTargetBox="[107,766,149,986]" captionTargetId="figure-638@9.[106,767,148,987]" captionTargetPageId="9" captionText="Fig. 9. Cephalic body portion of Echiniscus becki (PCM, paratype, female): A – dorsal view, B – ventral view (arrowheads indicate black crystalline eyes; large subcephalic plates visible as dark grey, broad belts bent in a boomerang-like shape). Scale bars = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.8164294" httpUri="https://zenodo.org/record/8164294/files/figure.png" pageId="6" pageNumber="55">Fig. 9B</figureCitation>
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). An erroneous ascribing of the red colour most probably resulted from a strong orange-reddish glow enclosing eyes. In fact, already the description in
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<bibRefCitation id="EFD6B9C8FF91FFD8FE08F8FEFD01F8BF" author="Schuster, R. O. & Grigarick, A. A." box="[410,704,1795,1814]" pageId="6" pageNumber="55" pagination="127 - 130" refId="ref12551" refString="Schuster, R. O., Grigarick, A. A., 1966. New Tardigrada from Western North America: II, Echiniscus. Proc. Biol. Soc. Wash. 79, 127 - 130." type="journal article" year="1966">Schuster and Grigarick (1966)</bibRefCitation>
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raised suspicions as the authors drew well-identifiable eyes in the mounted holotypic female, whereas red carotenoid eyes in echiniscids in the vast majority of cases vanish within an hour from preparing a permanent slide with a tardigrade.
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<taxonomicName id="4C47BFBAFF91FFD8FEDDF88FFE21F82C" baseAuthorityName="Schuster and Grigarick" baseAuthorityYear="1966" box="[335,480,1906,1925]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="55" phylum="Tardigrada" rank="species" species="becki">
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<emphasis id="B933182BFF91FFD8FEDDF88FFE21F82C" bold="true" box="[335,480,1906,1925]" italics="true" pageId="6" pageNumber="55">Echiniscus becki</emphasis>
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</taxonomicName>
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exhibits also a pair of exceptionally shaped subcephalic plates, which extend from the lateral portions of the head and twist in a boomerang-like manner at the level of pharynx (
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<figureCitation id="137CD8BCFF91FFD8FC1FFB96FC0EFBD7" box="[909,975,1131,1150]" captionStart="Fig" captionStartId="9.[100,130,1015,1032]" captionTargetBox="[107,766,149,986]" captionTargetId="figure-638@9.[106,767,148,987]" captionTargetPageId="9" captionText="Fig. 9. Cephalic body portion of Echiniscus becki (PCM, paratype, female): A – dorsal view, B – ventral view (arrowheads indicate black crystalline eyes; large subcephalic plates visible as dark grey, broad belts bent in a boomerang-like shape). Scale bars = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.8164294" httpUri="https://zenodo.org/record/8164294/files/figure.png" pageId="6" pageNumber="55">Fig. 9B</figureCitation>
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).
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</paragraph>
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<paragraph id="8BF8C439FF91FFD8FCC3FB7BFBE0FAE8" blockId="6.[818,1487,1131,1345]" pageId="6" pageNumber="55">
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Representatives of
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<taxonomicName id="4C47BFBAFF91FFD8FB93FB7AFB49FB30" authorityName="Gąsiorek" authorityYear="2023" baseAuthorityName="Ito" baseAuthorityYear="1993" box="[1025,1160,1158,1178]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="55" phylum="Tardigrada" rank="species" species="laterosetosus">
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<emphasis id="B933182BFF91FFD8FB93FB7AFB49FB30" bold="true" box="[1025,1160,1158,1178]" italics="true" pageId="6" pageNumber="55">E. laterosetosus</emphasis>
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</taxonomicName>
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, collected in the high mountains of Honshu, allowed for providing the morphological details relevant to its taxonomy. Eye spots, drawn in the original description without stating their colour (
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<bibRefCitation id="EFD6B9C8FF91FFD8FC3FFB26FBC3FB44" author="Ito, M." box="[941,1026,1242,1262]" pageId="6" pageNumber="55" pagination="1 - 13" refId="ref10737" refString="Ito, M., 1993. Taxonomic study on the class Heterotardigrada (Tardigrada) from the northern slope of Mt. Fuji, Central Japan. Edaphologia 50, 1 - 13." type="journal article" year="1993">Ito, 1993</bibRefCitation>
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), are small and black (
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<figureCitation id="137CD8BCFF91FFD8FB45FB27FAE7FB44" box="[1239,1318,1242,1261]" captionStart="Fig" captionStartId="9.[100,130,1939,1956]" captionTargetBox="[332,1255,1524,1910]" captionTargetId="figure-686@9.[331,1256,1523,1911]" captionTargetPageId="9" captionText="Fig. 10. Habitus of Echiniscus laterosetosus (PCM, female): A – cephalic body portion with arrowheads pointing out black crystalline eyes, B – specimen in toto. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.8164296" httpUri="https://zenodo.org/record/8164296/files/figure.png" pageId="6" pageNumber="55">Fig. 10A</figureCitation>
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). The dorsal plate sculpturing consists of well-developed endocuticular pillars, each joined by
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<emphasis id="B933182BFF91FFD8FCC3FAEFFC40FA8C" bold="true" box="[849,897,1298,1317]" italics="true" pageId="6" pageNumber="55">striae</emphasis>
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with four or five neighbouring pillars, and pores distributed between them (
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<figureCitation id="137CD8BCFF91FFD8FC57FAD3FBD5FAE8" box="[965,1044,1326,1345]" captionStart="Fig" captionStartId="9.[100,130,1939,1956]" captionTargetBox="[332,1255,1524,1910]" captionTargetId="figure-686@9.[331,1256,1523,1911]" captionTargetPageId="9" captionText="Fig. 10. Habitus of Echiniscus laterosetosus (PCM, female): A – cephalic body portion with arrowheads pointing out black crystalline eyes, B – specimen in toto. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.8164296" httpUri="https://zenodo.org/record/8164296/files/figure.png" pageId="6" pageNumber="55">Fig. 10B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |