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<document id="DC509C3739ED0826FDB71E95E13870D1" ID-DOI="10.11646/zootaxa.2618.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5304087" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629699783393" checkinUser="felipe" docAuthor="Oláh, János &amp; Johanson, Kjell Arne" docDate="2010" docId="03E487BBFFCE2B10FF3CFCC63B5F1FBD" docLanguage="en" docName="zt02618p035.pdf" docOrigin="Zootaxa 2618 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Setodes trikuthia Oláh &amp; Johanson 2010, new species" docType="treatment" docVersion="4" lastPageNumber="32" masterDocId="FFDDFFC3FFD32B0FFFABFF9B39571F41" masterDocTitle="Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia" masterLastPageNumber="35" masterPageNumber="1" pageNumber="30" updateTime="1699097725960" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="2441FBC1EE7762D7907C41974A128246" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14CCCB655A221DB29242CC2C40E450FA">
<mods:title id="C7DD8A1162F7D8C29443178A3EC9E908">Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="9D20CD88CE424EFB125E3EC4692AC0CD" type="personal">
<mods:role id="46338DB96A7E85DA181B5EB18A97E255">
<mods:roleTerm id="B05FD433DCBBBC10EE0B080CE5C35613">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D24B9490241A5EDD74AFF2C044DABDA7">Oláh, János</mods:namePart>
<mods:affiliation id="EB8788C8DF42EDD98D403B79F4B871A1">Szent István University, Gödöll, Centre of Environmental Health. Residence postal address: Tarján u. 28, H- 4032 Debrecen, Hungary. E-mail: profolah @ gmail. com</mods:affiliation>
</mods:name>
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<mods:role id="C8180A516993B79BFB9A3017A0CBE8B9">
<mods:roleTerm id="3BAAEE5D8EC8B57849B0BA8492000A9F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="28DC6388EF8DC744157C98BD7C6B056D">Johanson, Kjell Arne</mods:namePart>
<mods:affiliation id="AAB3DC4C19430F0CA5B29D4981B0805E">Swedish Museum of Natural History, Entomology Department, Box 50007, S- 10405 Stockholm, Sweden.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EDEBCA0167A61D777B9F53A74A3EA706">text</mods:typeOfResource>
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<mods:titleInfo id="C4D5376FA6A7E663D21D927520488B70">
<mods:title id="A4C2E22149FA9D3F84BAA5FE72B1037B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AA112E1B71C46C560FEA3B87D2980AA8">
<mods:date id="F2C32978960946EDB5CFA21EFAC0634F">2010</mods:date>
<mods:detail id="D091B976840D1D1AE3C3A5C7B293CD33" type="pubDate">
<mods:number id="F97FEF631467F523C48F3988D07553B8">2010-09-20</mods:number>
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<mods:detail id="E493FBCE0425DB1073950FD0EFCF4693" type="volume">
<mods:number id="8D24A8C757B4B1FFEA239432D11FCF9C">2618</mods:number>
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<mods:detail id="DEA3DA2E27591317FB4E737A2782FCC2" type="issue">
<mods:number id="B9D4B15016514E241601A190CC6B4147">1</mods:number>
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<mods:start id="EE54A0658FD0ACA7F14E5091DE139A4F">1</mods:start>
<mods:end id="B01C6F00AD5045E68D867B5569FA63DA">35</mods:end>
</mods:extent>
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<mods:location id="7370FEBA703E1A35C63C0E88CEA566FC">
<mods:url id="280F81E77E3D8A856FE68CDD2E56AAA6">https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1</mods:url>
</mods:location>
<mods:classification id="C514CD8D4E35088C30520DF86850C62D">journal article</mods:classification>
<mods:identifier id="2E96A472A2087A0323C830891547941B" type="DOI">10.11646/zootaxa.2618.1.1</mods:identifier>
<mods:identifier id="A217908420CEC726BAB01644ECD66EC9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4C00E02785E1FABB0CAD9557B3EF305B" type="Zenodo-Dep">5304087</mods:identifier>
</mods:mods>
<treatment id="03E487BBFFCE2B10FF3CFCC63B5F1FBD" ID-DOI="http://doi.org/10.5281/zenodo.5311124" ID-Zenodo-Dep="5311124" LSID="urn:lsid:plazi:treatment:03E487BBFFCE2B10FF3CFCC63B5F1FBD" httpUri="http://treatment.plazi.org/id/03E487BBFFCE2B10FF3CFCC63B5F1FBD" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="C3576526FFCE2B12FF3CFCC63B561C36" box="[151,513,861,887]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BF236ADFFCE2B12FF3CFCC63B561C36" blockId="29.[151,513,861,922]" box="[151,513,861,887]" pageId="29" pageNumber="30">
<heading id="D0BA81C1FFCE2B12FF3CFCC63B561C36" bold="true" box="[151,513,861,887]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B939EABFFFCE2B12FF3CFCC63B561C36" bold="true" box="[151,513,861,887]" pageId="29" pageNumber="30">
<taxonomicName id="4C4D4D2EFFCE2B12FF3CFCC638341C36" authority="Oláh &amp; Johanson, 2010" authorityName="Oláh &amp; Johanson" authorityYear="2010" box="[151,355,861,887]" class="Insecta" family="Leptoceridae" genus="Setodes" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trikuthia" status="sp. nov.">
<emphasis id="B939EABFFFCE2B12FF3CFCC638341C36" bold="true" box="[151,355,861,887]" italics="true" pageId="29" pageNumber="30">Setodes trikuthia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20A57C4FFCE2B12FED9FCC63B561C36" box="[370,513,861,887]" pageId="29" pageNumber="30" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3576526FFCE2B11FF3CFC1B3A061EF6" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="8BF236ADFFCE2B12FF3CFC1B384D1CDB" blockId="29.[151,513,861,922]" box="[151,282,896,922]" pageId="29" pageNumber="30">
<figureCitation id="13762A28FFCE2B12FF3CFC1B384D1CDB" box="[151,282,896,922]" captionStart="FIGURES 60–65" captionStartId="30.[151,275,1971,1995]" captionTargetBox="[197,1384,639,1920]" captionTargetId="figure-193@30.[151,1436,605,1951]" captionTargetPageId="30" captionText="FIGURES 60–65. Setodes trikuthia, new species, holotype. 60—head, dorsal; 61—head, frontal; 62—pro- and mesonotum, dorsal; 63—genitalia, left lateral; 64—right gonocoxite, ventral; 65—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5304135" httpUri="https://zenodo.org/record/5304135/files/figure.png" pageId="29" pageNumber="30">Figs 60–65</figureCitation>
</paragraph>
<paragraph id="8BF236ADFFCE2B12FF3CFC5138951BDE" blockId="29.[151,1436,970,2004]" pageId="29" pageNumber="30">
This species is similar to
<taxonomicName id="4C4D4D2EFFCE2B12FE6BFC513AFC1CA5" authority="Malicky &amp; Chantaramongkol" authorityName="Malicky &amp; Chantaramongkol" box="[448,939,970,996]" class="Insecta" family="Leptoceridae" genus="Setodes" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nausikaa">
<emphasis id="B939EABFFFCE2B12FE6BFC513B1F1CA2" box="[448,584,970,995]" italics="true" pageId="29" pageNumber="30">S. nausikaa</emphasis>
Malicky &amp; Chantaramongkol
</taxonomicName>
, known from several localities in northern
<collectingRegion id="4989F84FFFCE2B12FF3CFC6B39AB1B4B" box="[151,252,1008,1034]" country="Indonesia" name="Sumatera Utara" pageId="29" pageNumber="30">Sumatra</collectingRegion>
.
<taxonomicName id="4C4D4D2EFFCE2B12FEA3FC6B38991B48" authority="Oláh &amp; Johanson, 2010" authorityName="Oláh &amp; Johanson" authorityYear="2010" box="[264,462,1008,1033]" class="Insecta" family="Leptoceridae" genus="Setodes" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trikuthia" status="sp. nov.">
<emphasis id="B939EABFFFCE2B12FEA3FC6B38991B48" box="[264,462,1008,1033]" italics="true" pageId="29" pageNumber="30">Setodes trikuthia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20A57C4FFCE2B12FE75FC6B3B381B4B" box="[478,623,1008,1034]" pageId="29" pageNumber="30" rank="species">new species</taxonomicNameLabel>
, is easily separated from
<taxonomicName id="4C4D4D2EFFCE2B12FC30FC6B3D741B48" box="[923,1059,1008,1033]" class="Insecta" family="Leptoceridae" genus="Setodes" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nausikaa">
<emphasis id="B939EABFFFCE2B12FC30FC6B3D741B48" box="[923,1059,1008,1033]" italics="true" pageId="29" pageNumber="30">S. nausikaa</emphasis>
</taxonomicName>
by the following male genitalic characters: segment IX in lateral view is vertically elongate and nearly rectangular, not pyriform; shorter modified cerci are fused with the juncture of tergum IX, segment X, and the paraproct; and the apical pattern of gonopods is characterized by presence of 3 rounded, sinuous, excisions visible among the apical lobes, not by only a single excision.
</paragraph>
<paragraph id="8BF236ADFFCE2B12FF6EFB313BBC19C5" blockId="29.[151,1436,970,2004]" pageId="29" pageNumber="30">Male. Body pale, eyes black. Scapes shorter than head, without elongated setal tufts; long setae present on mesal surfaces of scapes. Maxillary palp formula V-IV-I-II-III. Anterior surface of head (or cephalic face) characterized by narrow frontal area and large genal areas; anterior tentorial pits almost indiscernible; wide frontogenal areas almost fully occupied by frontogenal compact setose warts. Clypeogenal compact setose warts small, rounded, present on clypeogenal cranial area above articulation of each maxillary palp segment I. Labrum elongate axially, posterior part covered with diffuse setal wart. Haustellum narrow at base; broad at mid-length; with mesodistal hump; posterior half with parallel channels. Head dorsum with pair of large, ovoid compact occipital setose warts; pair of rounded, vertexal ocellar compact setose warts; and single, large, nearly diamond-shaped vertexal medioantennal compact setose wart dominating on head dorsum between and behind scapes. Postgenal compact setose warts small, ventrally attached to ocular groove as narrow warts, invisible in dorsal view. Coronal, lateral vertexal, occipital and postoccipital grooves forming dark brown lines on head dorsum. Cranial areas and warts medium brown. Swollen proepisternal setal wart absent. Katepisternum of mesopleura acute anterodorsally.</paragraph>
<paragraph id="8BF236ADFFCE2B12FF6EF90B398B19B5" blockId="29.[151,1436,970,2004]" pageId="29" pageNumber="30">Spur formula 0,2,2; middle leg anteroapical spurs each 3 times longer than posterior spurs; hind leg anteroapical spurs each half as long as posteroapical spurs; middle leg apical spurs shorter than hind leg apical spurs.</paragraph>
<paragraph id="8BF236ADFFCE2B12FF6EF89B38B9185B" blockId="29.[151,1436,970,2004]" box="[197,494,1792,1818]" pageId="29" pageNumber="30">Forewing length 6.2 mm.</paragraph>
<paragraph id="8BF236ADFFCE2B11FF6DF8BE3A061EF6" blockId="29.[151,1436,970,2004]" lastBlockId="30.[151,1436,152,551]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">Male genitalia. Segment IX short, quadrangular in lateral view, tergal region constricted; lateroventrally setose; setal row present behind borders of segment IX and paraproctal plates. Segment X fused with tergum IX; long, trapezoid, slightly excised apicomesally in dorsal view; weakly sclerotized, covered with few minute setae; bordered laterally by well-developed paraproctal plates. Paraproctal complex forming phallic guide; fused to tergum IX and laterally to venter of segment X into pair of broad plates. Paraproctal plates broad basally, protruding apicad; each with apex rounded and with pair of dark ventrolateral spines. Cerci setose, reduced to setose foliform surfaces in lateral view. Gonopods short, each with 4 apical lobes with elevated alveolar tubercles of various sizes; 3 sinuous excisions present among apical lobes; gonopods bilobed in ventral view. Phallic apparatus with short phallobase; endotheca membranous, amorphous; with paired, slender arciform parameres, right paramere short, not reaching lateral flange of phallicata. Phallicata arching similarly to parameres; slender, with pair of subapical dorsal flanges; with phallotrema in trough between flanges and inconspicuous phallotremal sclerites. Phallicata excised mesally at apex in dorsal view, producing 2 dorsad-turning, hook-shaped, terminal corners.</paragraph>
</subSubSection>
<subSubSection id="C3576526FFCD2B11FF6EFE593AF01D66" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BF236ADFFCD2B11FF6EFE593BF31D43" blockId="30.[151,1436,152,551]" pageId="30" pageNumber="31">
<materialsCitation id="3B253CF0FFCD2B11FF6EFE593BF31D43" collectingDate="2008-07-10" collectorName="J. Olah" country="Indonesia" elevation="1000" location="Tapan Road" municipality="Kerinci National Park" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="B939EABFFFCD2B11FF6EFE59382D1E9D" bold="true" box="[197,378,450,476]" pageId="30" pageNumber="31">
<typeStatus id="54F6880FFFCD2B11FF6EFE5938601E9D" box="[197,311,450,476]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
male
</emphasis>
:
<emphasis id="B939EABFFFCD2B11FE20FE593B661E9D" bold="true" box="[395,561,450,476]" pageId="30" pageNumber="31">
<collectingCountry id="F35A763DFFCD2B11FE20FE593B661E9D" box="[395,561,450,476]" name="Indonesia" pageId="30" pageNumber="31">INDONESIA</collectingCountry>
</emphasis>
(
<collectingRegion id="4989F84FFFCD2B11FDEBFE593BFE1E9D" box="[576,681,450,476]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="31">Sumatra</collectingRegion>
):
<collectingMunicipality id="6B96ACD7FFCD2B11FD16FE593A951E9D" box="[701,962,450,476]" pageId="30" pageNumber="31">Kerinci National Park</collectingMunicipality>
,
<location id="8E926076FFCD2B11FC65FE593D091E9D" LSID="urn:lsid:plazi:treatment:03E487BBFFCE2B10FF3CFCC63B5F1FBD:8E926076FFCD2B11FC65FE593D091E9D" box="[974,1118,450,476]" country="Indonesia" municipality="Kerinci National Park" name="Tapan Road" pageId="30" pageNumber="31" stateProvince="Sumatera Utara">Tapan Road</location>
,
<quantity id="4CB59B48FFCD2B11FBC1FE593D921E9D" box="[1130,1221,450,476]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="m" value="1000.0">
<elevation id="0060D19EFFCD2B11FBC1FE593D921E9D" box="[1130,1221,450,476]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFF3106DFFCD2B11FB79FE593C0E1E9D" box="[1234,1369,450,476]" pageId="30" pageNumber="31" value="2008-07-10">
<collectingDate id="EFB7E985FFCD2B11FB79FE593C0E1E9D" box="[1234,1369,450,476]" pageId="30" pageNumber="31" value="2008-07-10">10.vii.2008</collectingDate>
</date>
,
<collectingMethod id="520C4EBAFFCD2B11FACDFE5939921D43" pageId="30" pageNumber="31">light trap</collectingMethod>
[
<collectorName id="26B8537BFFCD2B11FF7FFE73387C1D43" box="[212,299,488,514]" pageId="30" pageNumber="31">J. Oláh</collectorName>
, Jr.]—
<specimenCount id="9D4BFD24FFCD2B11FEDDFE7338921D43" box="[374,453,488,514]" pageId="30" pageNumber="31" type="male">1 male</specimenCount>
(OPC, in alcohol).
</materialsCitation>
</paragraph>
<paragraph id="8BF236ADFFCD2B11FF6EFD963AF01D66" blockId="30.[151,1436,152,551]" box="[197,935,525,551]" pageId="30" pageNumber="31">
<materialsCitation id="3B253CF0FFCD2B11FF6EFD963AF41D66" box="[197,931,525,551]" collectingDate="2008-07-10" collectorName="J. Olah" country="Indonesia" elevation="1000" location="Tapan Road" municipality="Kerinci National Park" pageId="30" pageNumber="31" specimenCount="2" specimenCount-male="2" stateProvince="Sumatera Utara" typeStatus="paratype">
<emphasis id="B939EABFFFCD2B11FF6EFD9638151D66" bold="true" box="[197,322,525,551]" pageId="30" pageNumber="31">
<typeStatus id="54F6880FFFCD2B11FF6EFD9638151D66" box="[197,322,525,551]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
</emphasis>
: same data as holotype—
<specimenCount id="9D4BFD24FFCD2B11FDC5FD963B9E1D66" box="[622,713,525,551]" pageId="30" pageNumber="31" type="male">2 males</specimenCount>
(OPC, in alcohol)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF326625FFCD2B11FF3CF8283C1718AA" ID-DOI="http://doi.org/10.5281/zenodo.5304135" ID-Zenodo-Dep="5304135" httpUri="https://zenodo.org/record/5304135/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,275,1971,1995]" targetBox="[197,1384,639,1920]" targetPageId="30">
<paragraph id="8BF236ADFFCD2B11FF3CF8283C1718AA" blockId="30.[151,1436,1971,2027]" pageId="30" pageNumber="31">
<emphasis id="B939EABFFFCD2B11FF3CF828383E188A" bold="true" box="[151,361,1971,1995]" pageId="30" pageNumber="31">FIGURES 60–65.</emphasis>
<taxonomicName id="4C4D4D2EFFCD2B11FED9F82F3B63188A" authority="Oláh &amp; Johanson, 2010" authorityName="Oláh &amp; Johanson" authorityYear="2010" box="[370,564,1972,1995]" class="Insecta" family="Leptoceridae" genus="Setodes" kingdom="Animalia" order="Trichoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="trikuthia" status="sp. nov.">
<emphasis id="B939EABFFFCD2B11FED9F82F3B63188A" box="[370,564,1972,1995]" italics="true" pageId="30" pageNumber="31">Setodes trikuthia</emphasis>
</taxonomicName>
,
<emphasis id="B939EABFFFCD2B11FDEEF8283B86188A" bold="true" box="[581,721,1971,1995]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A20A57C4FFCD2B11FDEEF8283B86188A" box="[581,721,1971,1995]" pageId="30" pageNumber="31" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype. 60—head, dorsal; 61—head, frontal; 62—pro- and mesonotum, dorsal; 63—genitalia, left lateral; 64—right gonocoxite, ventral; 65—phallic apparatus, left lateral.
</paragraph>
</caption>
<subSubSection id="C3576526FFCC2B10FF6EFF0C3B321FF3" box="[197,613,151,178]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BF236ADFFCC2B10FF6EFF0C3B321FF3" blockId="31.[151,1436,151,252]" box="[197,613,151,178]" pageId="31" pageNumber="32">
<emphasis id="B939EABFFFCC2B10FF6EFF0C38301FF0" bold="true" box="[197,359,151,177]" pageId="31" pageNumber="32">Distribution:</emphasis>
<collectingCountry id="F35A763DFFCC2B10FEC4FF0338B61FF3" box="[367,481,152,178]" name="Indonesia" pageId="31" pageNumber="32">Indonesia</collectingCountry>
(
<collectingRegion id="4989F84FFFCC2B10FE5AFF033B0E1FF3" box="[497,601,152,178]" country="Indonesia" name="Sumatera Utara" pageId="31" pageNumber="32">Sumatra</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3576526FFCC2B10FF6EFF263B5F1FBD" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BF236ADFFCC2B10FF6EFF263B5F1FBD" blockId="31.[151,1436,151,252]" pageId="31" pageNumber="32">
<emphasis id="B939EABFFFCC2B10FF6EFF2638021F96" bold="true" box="[197,341,189,215]" pageId="31" pageNumber="32">Etymology:</emphasis>
<emphasis id="B939EABFFFCC2B10FEF6FF26389F1F97" box="[349,456,189,214]" italics="true" pageId="31" pageNumber="32">Trikuthia</emphasis>
, from
<emphasis id="B939EABFFFCC2B10FDB2FF263B621F97" box="[537,565,189,214]" italics="true" pageId="31" pageNumber="32">tri</emphasis>
, “three,”
<emphasis id="B939EABFFFCC2B10FD0CFF263BA71F97" box="[679,752,189,214]" italics="true" pageId="31" pageNumber="32">kuthin</emphasis>
, “sinus” in Sanskrit, referring to the three-sinuous-pattern of the gonopods in lateral view.
</paragraph>
</subSubSection>
</treatment>
</document>
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