1431 lines
225 KiB
XML
1431 lines
225 KiB
XML
<document id="7292DBAA30F9E60AB87A93B6BD31986A" ID-CLB-Dataset="59767" ID-DOI="10.11646/zootaxa.5319.4.2" ID-GBIF-Dataset="b0dd118c-22d4-43bd-8dd8-bac4d810d1fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="8203248" ID-ZooBank="20272586-E197-4DEA-811F-496572F121B7" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1690883685755" checkinUser="plazi" docAuthor="Kluge, Nikita, Sivaruban, T., Srinivasan, Pandiarajan, Barathy, S. & Isack, Rajasekaran" docDate="2023" docId="03E48787FFCEE640FF64F96B8FDCFD3B" docLanguage="en" docName="zootaxa.5319.4.2.pdf" docOrigin="Zootaxa 5319 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ecdyonurus (Rhithrogeniella) ornatus" docType="treatment" docVersion="6" lastPageNumber="522" masterDocId="FFDDFFFFFFC5E655FFF3FF928B39FFF6" masterDocTitle="Redescription of the subgenus Rhithrogeniella Ulmer 1939 (Ephemeroptera, Heptageniidae, genus Ecdyonurus) based on reared specimens from India and Thailand" masterLastPageNumber="523" masterPageNumber="501" pageNumber="512" updateTime="1699549378525" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="89776A5F64A36159CE7FDDE9486E51BB">Redescription of the subgenus Rhithrogeniella Ulmer 1939 (Ephemeroptera, Heptageniidae, genus Ecdyonurus) based on reared specimens from India and Thailand</mods:title>
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<mods:namePart id="897A8FE68E132A0B43E50650C413671E">Kluge, Nikita</mods:namePart>
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<mods:affiliation id="13F19ED7B296EA6448213052C27765EE">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia. Website: http: // insecta. bio. spbu. ru</mods:affiliation>
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<mods:namePart id="FC370DAFA37EDBD2A6A5CA74083C0AFE">Sivaruban, T.</mods:namePart>
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<mods:affiliation id="62097786E61C95CD49246297E3832263">PG and Research department of Zoology, the American College, Madurai- 625002, India.</mods:affiliation>
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<mods:namePart id="53E32AE2F2AA1BF1036DEA576533E9BF">Srinivasan, Pandiarajan</mods:namePart>
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<mods:affiliation id="391239CC3C0ADF6622C06AE5F5DF7B89">PG and Research department of Zoology, the American College, Madurai- 625002, India.</mods:affiliation>
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<mods:affiliation id="971807A22F42533F97D7806C98CD5AF6">Department of Zoology, Fatima College, Madurai- 625018, India.</mods:affiliation>
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<mods:namePart id="17CE4EC3F819A2DD8EE809EE240E79E3">Isack, Rajasekaran</mods:namePart>
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<mods:affiliation id="E908E53D260FA6C36958EB124A920021">PG and Research department of Zoology, the American College, Madurai- 625002, India.</mods:affiliation>
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<mods:date id="678CAC26AAECCA409858E34833000E77">2023</mods:date>
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<taxonomicName id="4C4D4D12FFCEE65EFF64F96B882DF8E5" authority="(Ulmer 1939)" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[151,788,1785,1812]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
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<emphasis id="B939EA83FFCEE65EFF64F96B8A13F8E2" bold="true" box="[151,298,1785,1812]" italics="true" pageId="11" pageNumber="512">Ecdyonurus</emphasis>
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(
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<emphasis id="B939EA83FFCEE65EFEC8F96B8AC3F8E2" bold="true" box="[315,506,1785,1812]" italics="true" pageId="11" pageNumber="512">Rhithrogeniella</emphasis>
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<emphasis id="B939EA83FFCEE65EFDF8F9688951F8E2" bold="true" box="[523,616,1786,1812]" italics="true" pageId="11" pageNumber="512">ornatus</emphasis>
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(
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<bibRefCitation id="EFDC4B60FFCEE65EFD84F96B8832F8E5" author="Ulmer, G." box="[631,779,1785,1811]" pageId="11" pageNumber="512" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939</bibRefCitation>
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(
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<figureCitation id="13762A14FFCEE65EFF6CF88F8A19F8C1" box="[159,288,1821,1847]" captionStart-0="FIGURES 1–10" captionStart-1="FIGURES 11–25" captionStart-10="FIGURES 91–96" captionStart-11="FIGURES 97–102" captionStart-2="FIGURES 26–33" captionStart-3="FIGURES 34–41" captionStart-4="FIGURES 42–52" captionStart-5="FIGURES 53–68" captionStart-6="FIGURES 69–72" captionStart-7="FIGURES 73–76" captionStart-8="FIGURES 77–84" captionStart-9="FIGURES 85–90" captionStartId-0="3.[151,264,1810,1834]" captionStartId-1="4.[151,264,1853,1877]" captionStartId-10="17.[151,264,1968,1992]" captionStartId-11="18.[151,264,1834,1858]" captionStartId-2="5.[151,264,1773,1797]" captionStartId-3="6.[151,264,1916,1940]" captionStartId-4="7.[151,264,1785,1809]" captionStartId-5="8.[151,264,1807,1831]" captionStartId-6="9.[151,264,1837,1861]" captionStartId-7="10.[151,264,1352,1376]" captionStartId-8="14.[151,264,1532,1556]" captionStartId-9="16.[151,264,1571,1595]" captionTargetBox-0="[151,1436,180,1772]" captionTargetBox-1="[151,1435,181,1829]" captionTargetBox-10="[177,1411,381,1945]" captionTargetBox-11="[151,1436,181,1810]" captionTargetBox-2="[151,1435,181,1749]" captionTargetBox-3="[151,1435,181,1893]" captionTargetBox-4="[151,1436,181,1762]" captionTargetBox-5="[164,1422,181,1784]" captionTargetBox-6="[151,1436,182,1813]" captionTargetBox-7="[151,1436,181,1328]" captionTargetBox-8="[151,1435,309,1509]" captionTargetBox-9="[164,1423,181,1546]" captionTargetId-0="figure-20@3.[151,1436,179,1784]" captionTargetId-1="figure-37@4.[151,1435,181,1829]" captionTargetId-10="figure-129@17.[177,1411,381,1945]" captionTargetId-11="figure-37@18.[151,1436,181,1810]" captionTargetId-2="figure-20@5.[151,1435,181,1749]" captionTargetId-3="figure-37@6.[151,1435,181,1893]" captionTargetId-4="figure-20@7.[151,1436,181,1762]" captionTargetId-5="figure-37@8.[164,1423,181,1784]" captionTargetId-6="figure-20@9.[151,1436,181,1813]" captionTargetId-7="figure-37@10.[151,1436,181,1328]" captionTargetId-8="figure-91@14.[151,1435,309,1509]" captionTargetId-9="figure-37@16.[164,1423,181,1546]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="17" captionTargetPageId-11="18" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="14" captionTargetPageId-9="16" captionText-0="FIGURES 1–10. Ecdyonurus (Rhithrogeniella) ornatus from Kodaikanal (India). 1–3, larvae; 4–10, tergalii I–VII." captionText-1="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." captionText-10="FIGURES 91–96. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from Pai (Thailand). 91–92, reared imago, specimen [XIII](4)2015; 93–94, reared imago, specimen [XIX](1)2015; 95–96, reared subimago, specimen [XVIII](4)2015." captionText-11="FIGURES 97–102. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs of reared imagines from Kwai-Yai (Thailand). 97–98, specimen [III](11)2015; 99–100, specimen [III](37)A2015; 101–102, specimen [IV](2)2015." captionText-2="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." captionText-3="FIGURES 34–41. Ecdyonurus (Rhithrogeniella) ornatus, exuviae of larval legs (reared specimen from Kodaikanal). 34, fore femur; 35, outer margin of fore femur; 36–38, fore, middle and hind legs; 39, hind tibia; 40, hind tarsus; 41, claw." captionText-4="FIGURES 42–52. Larvae. 42–50, Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae: 42–44, specimen from Agumbe (India), posterior margins of abdominal terga I, VIII and X; 45, specimen from Pai (Thailand), posterior margins of abdominal tergum VI; 46–48, cercus of specimen from Agumbe: 46, proximal portion; 47, middle portion in median-ventral view, to show bases of primary swimming setae on inner (median) side; 48, the same portion with focus on lateral-dorsal side, to show bases of secondary swimming setae on outer (lateral) side; 49–50, cercus of specimen from Kodaikanal (India). 51–52, Ecdyonurus (Rhithrogeniella) joernensis (Russia, Orenburg province, Tashla, 16.VIII.1986, coll. N. Kluge): two portions of cercus, dorsal view." captionText-5="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." captionText-6="FIGURES 69–72. Ecdyonurus (Rhithrogeniella) ornatus, genitalia of male imagines, ventral view. 69–70, specimen from Kodaikanal (India) in tense condition; 71, specimen from Agumbe (India); 72, specimen from Pai (Thailand)." captionText-7="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." captionText-8="FIGURES 77–84. Ecdyonurus (Rhithrogeniella) ornatus, female imagines. 77–78, specimen from Agumbe (India); 79–81, specimens from Kwai-Yai (Thailand); 82–84, specimens from Pai (Thailand)." captionText-9="FIGURES 85–90. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal." figureDoi-0="http://doi.org/10.5281/zenodo.8203250" figureDoi-1="http://doi.org/10.5281/zenodo.8203254" figureDoi-10="http://doi.org/10.5281/zenodo.8203272" figureDoi-11="http://doi.org/10.5281/zenodo.8203274" figureDoi-2="http://doi.org/10.5281/zenodo.8203256" figureDoi-3="http://doi.org/10.5281/zenodo.8203258" figureDoi-4="http://doi.org/10.5281/zenodo.8203260" figureDoi-5="http://doi.org/10.5281/zenodo.8203262" figureDoi-6="http://doi.org/10.5281/zenodo.8203264" figureDoi-7="http://doi.org/10.5281/zenodo.8203266" figureDoi-8="http://doi.org/10.5281/zenodo.8203268" figureDoi-9="http://doi.org/10.5281/zenodo.8203270" httpUri-0="https://zenodo.org/record/8203250/files/figure.png" httpUri-1="https://zenodo.org/record/8203254/files/figure.png" httpUri-10="https://zenodo.org/record/8203272/files/figure.png" httpUri-11="https://zenodo.org/record/8203274/files/figure.png" httpUri-2="https://zenodo.org/record/8203256/files/figure.png" httpUri-3="https://zenodo.org/record/8203258/files/figure.png" httpUri-4="https://zenodo.org/record/8203260/files/figure.png" httpUri-5="https://zenodo.org/record/8203262/files/figure.png" httpUri-6="https://zenodo.org/record/8203264/files/figure.png" httpUri-7="https://zenodo.org/record/8203266/files/figure.png" httpUri-8="https://zenodo.org/record/8203268/files/figure.png" httpUri-9="https://zenodo.org/record/8203270/files/figure.png" pageId="11" pageNumber="512">Figs 1–102</figureCitation>
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,
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<figureCitation id="13762A14FFCEE65EFEDFF88F8A6FF8C1" box="[300,342,1821,1847]" captionStart="FIGURES 103–110" captionStartId="19.[151,264,1692,1716]" captionTargetBox="[154,1436,345,1669]" captionTargetId="figure-106@19.[151,1436,345,1669]" captionTargetPageId="19" captionText="FIGURES 103–110. 103–109, Ecdyonurus (Rhithrogeniella) sp. 2: 103, male imago, 104, abdomen of male imago; 105, male subimago; 106, female subimago; 107, thorax of male subimago, ventral view; 108, male subimaginal exuviae; 109, enlarged part of its ventral arc of prealar bridge; 110, the same of E. (Rh.) ornatus." figureDoi="http://doi.org/10.5281/zenodo.8203276" httpUri="https://zenodo.org/record/8203276/files/figure.png" pageId="11" pageNumber="512">110</figureCitation>
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)
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<taxonomicName id="4C4D4D12FFCEE65EFF64F8F0897AF88C" authority="Ulmer 1939: 576" authorityName="Ulmer" authorityPageNumber="576" authorityYear="1939" box="[151,579,1890,1914]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFCEE65EFF64F8F08ABAF88C" box="[151,387,1890,1914]" italics="true" pageId="11" pageNumber="512">Rhithrogeniella ornata</emphasis>
|
||
<bibRefCitation id="EFDC4B60FFCEE65EFE7EF8F0897AF88C" author="Ulmer, G." box="[397,579,1890,1914]" pageId="11" pageNumber="512" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939: 576</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, figs 169–174 (♂ and
|
||
<specimenCount id="9D4BFD18FFCEE65EFCC4F8F08873F88F" box="[823,842,1890,1913]" pageId="11" pageNumber="512" type="female">♀</specimenCount>
|
||
imagines and subimagines);
|
||
<treatmentCitation id="0AEC1080FFCEE65EFB73F8F08E12F88C" author="Sartori, M." box="[1152,1323,1890,1914]" httpUri="http://treatment.plazi.org/id/EC87D9636F53EF803F3E86DAD2946C3C" page="49" pageId="11" pageNumber="512" year="2014">
|
||
<bibRefCitation id="EFDC4B60FFCEE65EFB73F8F08E12F88C" author="Sartori, M." box="[1152,1323,1890,1914]" pageId="11" pageNumber="512" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014: 49</bibRefCitation>
|
||
</treatmentCitation>
|
||
(♂ and
|
||
<specimenCount id="9D4BFD18FFCEE65EFA7AF8F08EA5F88F" box="[1417,1436,1890,1913]" pageId="11" pageNumber="512" type="female">♀</specimenCount>
|
||
</treatmentCitationGroup>
|
||
imagines and subimagines, eggs and larvae);
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFCEE65EFF64F80D899AF841" blockId="11.[151,1437,1890,2037]" box="[151,675,1951,1976]" pageId="11" pageNumber="512">
|
||
<taxonomicName id="4C4D4D12FFCEE65EFF64F80D89A6F841" ID-CoL="6X76L" authority=": Wang & McCafferty 2004: 21" authorityName=": Wang & McCafferty" authorityPageNumber="21" authorityYear="2004" box="[151,671,1951,1976]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFCEE65EFF64F80D8A66F841" box="[151,351,1951,1975]" italics="true" pageId="11" pageNumber="512">Rhithrogena ornata</emphasis>
|
||
:
|
||
<bibRefCitation id="EFDC4B60FFCEE65EFE9FF83289A6F841" author="Wang, T. - Q. & McCafferty, W. P." box="[364,671,1951,1976]" pageId="11" pageNumber="512" pagination="11 - 45" refId="ref10235" refString="Wang, T. - Q. & McCafferty, W. P. (2004) Heptageniidae (Ephemeroptera) of the world. Part I: Phylogenetic higher classification. Transactions of the American Entomological Society, 130 (1), 11 - 45." type="journal article" year="2004">Wang & McCafferty 2004: 21</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFCEE65EFF64F82C8A9AF803" blockId="11.[151,1437,1890,2037]" pageId="11" pageNumber="512">
|
||
<taxonomicName id="4C4D4D12FFCEE65EFF64F82C89DBF820" authority="Soldan & Braasch 1986: 206" authorityName="Soldan & Braasch" authorityPageNumber="206" authorityYear="1986" box="[151,738,1982,2006]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="B939EA83FFCEE65EFF64F82C8A96F820" box="[151,431,1982,2006]" italics="true" pageId="11" pageNumber="512">Rhithrogeniella tonkinensis</emphasis>
|
||
<bibRefCitation id="EFDC4B60FFCEE65EFE44F82C89DBF820" author="Soldan, T. & Braasch, D." box="[439,738,1982,2006]" pageId="11" pageNumber="512" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986: 206</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCount id="9D4BFD18FFCEE65EFD06F82C883FF820" box="[757,774,1982,2006]" pageId="11" pageNumber="512" type="female">♀</specimenCount>
|
||
imago, ♂ subimago, larva, egg)
|
||
<emphasis id="B939EA83FFCEE65EFBAEF82C8F9CF820" bold="true" box="[1117,1189,1982,2006]" pageId="11" pageNumber="512">syn. n.</emphasis>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFCEE65EFB47F82C8EA4F820" author="Boonsoong, B. & Braasch, D." box="[1204,1437,1982,2006]" pageId="11" pageNumber="512" pagination="61 - 93" refId="ref9226" refString="Boonsoong, B. & Braasch, D. (2013) Heptageniidae (Insecta, Ephemeroptera) of Thailand. ZooKeys, 272, 61 - 93. https: // doi. org / 10.3897 / zookeys. 272.3638" type="journal article" year="2013">Boonsoong & Braasch</bibRefCitation>
|
||
2013: 78 (larva, egg).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF64FF0589F5FF59" blockId="12.[151,886,151,205]" box="[151,716,151,175]" pageId="12" pageNumber="513">
|
||
<treatmentCitationGroup id="AB5D11BFFFC9E659FF64FF0589F5FF59" box="[151,716,151,175]" pageId="12" pageNumber="513">
|
||
<treatmentCitation id="0AEC1080FFC9E659FF64FF0589F1FF59" author="Wang, T. - Q. & McCafferty, W. P." box="[151,712,151,175]" page="21" pageId="12" pageNumber="513" year="2004">
|
||
<taxonomicName id="4C4D4D12FFC9E659FF64FF0589F1FF59" ID-CoL="6DSST" authority=": Wang & McCafferty 2004: 21" authorityName="Wang & McCafferty" authorityPageNumber="21" authorityYear="2004" box="[151,712,151,175]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="B939EA83FFC9E659FF64FF058AB0FF59" box="[151,393,151,175]" italics="true" pageId="12" pageNumber="513">Ecdyonurus tonkinensis</emphasis>
|
||
:
|
||
<bibRefCitation id="EFDC4B60FFC9E659FE66FF0589F1FF59" author="Wang, T. - Q. & McCafferty, W. P." box="[405,712,151,175]" pageId="12" pageNumber="513" pagination="11 - 45" refId="ref10235" refString="Wang, T. - Q. & McCafferty, W. P. (2004) Heptageniidae (Ephemeroptera) of the world. Part I: Phylogenetic higher classification. Transactions of the American Entomological Society, 130 (1), 11 - 45." type="journal article" year="2004">Wang & McCafferty 2004: 21</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF64FF27884FFF3B" blockId="12.[151,886,151,205]" box="[151,886,181,205]" pageId="12" pageNumber="513">
|
||
<treatmentCitationGroup id="AB5D11BFFFC9E659FF64FF27884FFF3B" box="[151,886,181,205]" pageId="12" pageNumber="513">
|
||
<treatmentCitation id="0AEC1080FFC9E659FF64FF2789CCFF3B" author="Kluge, N. J." box="[151,757,181,205]" page="168" pageId="12" pageNumber="513" year="2022">
|
||
<taxonomicName id="4C4D4D12FFC9E659FF64FF2789CCFF3B" ID-CoL="6DSST" authority=": Kluge 2022: 168" authorityName=": Kluge" authorityPageNumber="168" authorityYear="2022" box="[151,757,181,205]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="tonkinensis" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFC9E659FF64FF278A29FF3B" box="[151,272,181,205]" italics="true" pageId="12" pageNumber="513">Ecdyonurus</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFC9E659FEECFF278A84FF3B" box="[287,445,181,205]" italics="true" pageId="12" pageNumber="513">Rhithrogeniella</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFC9E659FE3FFF278907FF3B" box="[460,574,181,205]" italics="true" pageId="12" pageNumber="513">tonkinensis</emphasis>
|
||
:
|
||
<bibRefCitation id="EFDC4B60FFC9E659FDB8FF2789CCFF3B" author="Kluge, N. J." box="[587,757,181,205]" pageId="12" pageNumber="513" pagination="151 - 186" refId="ref9941" refString="Kluge, N. J. (2022) Taxonomic significance of microlepides on subimaginal tarsi of Ephemeroptera. Zootaxa, 5159 (2), 151 - 186. https: // doi. org / 10.11646 / zootaxa. 5159.2.1" type="journal article" year="2022">Kluge 2022: 168</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(subimago).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C357651AFFC9E659FF64FF6E888CFB6C" pageId="12" pageNumber="513" type="materials_examined">
|
||
<paragraph id="8BF23691FFC9E659FF64FF6E883DFEAB" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
<emphasis id="B939EA83FFC9E659FF64FF6E8A9CFEE3" bold="true" box="[151,421,251,278]" pageId="12" pageNumber="513">
|
||
<typeStatus id="54F68833FFC9E659FF64FF6E8A34FEE0" box="[151,269,252,278]" pageId="12" pageNumber="513" type="lectotype">Lectotype</typeStatus>
|
||
designation.
|
||
</emphasis>
|
||
<materialsCitation id="3B253CCCFFC9E659FE5EFF6E8839FEAB" ID-GBIF-Occurrence="4168147309" collectingDate="1932-07" country="Germany" county="This" location="Lieftinck in Buitenzorg" municipality="Biozentrum Grindel" pageId="12" pageNumber="513" specimenCount="1" stateProvince="Hamburg" typeStatus="lectotype">
|
||
<typeStatus id="54F68833FFC9E659FE5EFF6E8924FEE0" box="[429,541,252,278]" pageId="12" pageNumber="513" type="lectotype">Lectotype</typeStatus>
|
||
(designated here): male imago in alcohol, collected by
|
||
<location id="8E92604AFFC9E659FB67FF698EA5FEE3" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FB67FF698EA5FEE3" box="[1172,1436,251,277]" country="Germany" county="This" municipality="Biozentrum Grindel" name="Lieftinck in Buitenzorg" pageId="12" pageNumber="513" stateProvince="Hamburg">Lieftinck in Buitenzorg</location>
|
||
(currently Bogor) in
|
||
<date id="FFF31051FFC9E659FE8AFE8D8ADDFECC" box="[377,484,287,314]" pageId="12" pageNumber="513" value="1932-07">
|
||
<collectingDate id="EFB7E9B9FFC9E659FE8AFE8D8ADDFECC" box="[377,484,287,314]" pageId="12" pageNumber="513" value="1932-07">July 1932</collectingDate>
|
||
</date>
|
||
and deposited in
|
||
<location id="8E92604AFFC9E659FD53FE8D88A5FECC" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FD53FE8D88A5FECC" box="[672,924,287,314]" country="Germany" county="This" municipality="Biozentrum Grindel" name="Zoologisches Museum" pageId="12" pageNumber="513" stateProvince="Hamburg">Zoologisches Museum</location>
|
||
und
|
||
<collectingMunicipality id="6B96ACEBFFC9E659FC23FE8D8F88FECF" box="[976,1201,287,313]" pageId="12" pageNumber="513">Biozentrum Grindel</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="4989F873FFC9E659FB49FE8D8E1FFECF" box="[1210,1318,287,313]" country="Germany" name="Hamburg" pageId="12" pageNumber="513">Hamburg</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35A7601FFC9E659FADCFEB28EA0FECC" box="[1327,1433,288,314]" name="Germany" pageId="12" pageNumber="513">Germany</collectingCountry>
|
||
.
|
||
<collectingCounty id="62934E1DFFC9E659FF64FED18BF0FEAB" box="[151,201,323,349]" pageId="12" pageNumber="513">This</collectingCounty>
|
||
specimen is reported in literature as the following
|
||
</materialsCitation>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF34FEFA89AFFE50" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
1)
|
||
<materialsCitation id="3B253CCCFFC9E659FF15FEF589AFFE50" ID-GBIF-Occurrence="4168147302" collectingDate="1932-07" collectorName="Dr. LIEFTINCK" location="Spiritus" pageId="12" pageNumber="513" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="typen">
|
||
<bibRefCitation id="EFDC4B60FFC9E659FF15FEF58A49FE74" author="Ulmer, G." box="[230,368,359,386]" pageId="12" pageNumber="513" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939</bibRefCitation>
|
||
, p. 578: «
|
||
<specimenCount id="9D4BFD18FFC9E659FE12FEFA8932FE72" box="[481,523,360,388]" pageId="12" pageNumber="513" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4BFD18FFC9E659FDE4FEFA897AFE74" box="[535,579,360,386]" pageId="12" pageNumber="513" type="female">2 ♀</specimenCount>
|
||
. wohl frisch geschlüpft, in
|
||
<location id="8E92604AFFC9E659FC85FEF588E9FE77" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FC85FEF588E9FE77" box="[886,976,359,385]" name="Spiritus" pageId="12" pageNumber="513">Spiritus</location>
|
||
,
|
||
<location id="8E92604AFFC9E659FC2FFEF58F62FE77" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FC2FFEF58F62FE77" box="[988,1115,359,385]" name="Buitenzorg" pageId="12" pageNumber="513">Buitenzorg</location>
|
||
,
|
||
<date id="FFF31051FFC9E659FB95FEFA8FEFFE74" box="[1126,1238,360,386]" pageId="12" pageNumber="513" value="1932-07">
|
||
<collectingDate id="EFB7E9B9FFC9E659FB95FEFA8FEFFE74" box="[1126,1238,360,386]" pageId="12" pageNumber="513" value="1932-07">VII. 1932</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B85347FFC9E659FB12FEFA8EA5FE74" box="[1249,1436,360,386]" pageId="12" pageNumber="513">Dr. LIEFTINCK</collectorName>
|
||
leg. (
|
||
<specimenCount id="9D4BFD18FFC9E659FF21FE1E8BC4FE5E" box="[210,253,396,424]" pageId="12" pageNumber="513" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4BFD18FFC9E659FEFAFE1E8A0AFE50" box="[265,307,396,422]" pageId="12" pageNumber="513" type="female">1 ♀</specimenCount>
|
||
in meiner Sammlung,
|
||
<typeStatus id="54F68833FFC9E659FDC1FE1E8947FE50" box="[562,638,396,422]" pageId="12" pageNumber="513">Typen</typeStatus>
|
||
)»;
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF34FE228AA5FE18" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
<materialsCitation id="3B253CCCFFC9E659FF34FE228984FE3F" ID-GBIF-Occurrence="4168147311" box="[199,701,431,458]" collectingDate="1932-07" collectionCode="ZMH" collectorName="Dr. Lieftinck" country="Indonesia" location="Buitenzorg" pageId="12" pageNumber="513" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Timur" typeStatus="holotype">
|
||
2)
|
||
<bibRefCitation id="EFDC4B60FFC9E659FF14FE3D8A41FE3C" author="Sartori, M." box="[231,376,431,458]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
, p. 49: ́One male
|
||
<typeStatus id="54F68833FFC9E659FDA7FE3D8980FE3F" box="[596,697,431,457]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation id="3B253CCCFFC9E659FD35FE228ABFFE18" ID-GBIF-Occurrence="4168147301" collectingDate="1932-07" collectionCode="ZMH" collectorName="Dr. Lieftinck" country="Indonesia" location="Buitenzorg" pageId="12" pageNumber="513" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Timur" typeStatus="allotype">
|
||
<specimenCount id="9D4BFD18FFC9E659FD35FE22887DFE3F" box="[710,836,431,458]" pageId="12" pageNumber="513" type="female">one female</specimenCount>
|
||
<typeStatus id="54F68833FFC9E659FCBEFE3D8895FE3F" box="[845,940,431,457]" pageId="12" pageNumber="513" type="allotype">allotype</typeStatus>
|
||
:
|
||
<collectingCountry id="F35A7601FFC9E659FC4BFE3D8F11FE3F" box="[952,1064,431,457]" name="Indonesia" pageId="12" pageNumber="513">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="4989F873FFC9E659FBC7FE228F51FE3C" box="[1076,1128,432,458]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="513">Java</collectingRegion>
|
||
,
|
||
<location id="8E92604AFFC9E659FB85FE3D8FCCFE3F" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FB85FE3D8FCCFE3F" box="[1142,1269,431,457]" country="Indonesia" name="Buitenzorg" pageId="12" pageNumber="513" stateProvince="Jawa Timur">Buitenzorg</location>
|
||
,
|
||
<date id="FFF31051FFC9E659FAF2FE228E55FE3C" box="[1281,1388,432,458]" pageId="12" pageNumber="513" value="1932-07">
|
||
<collectingDate id="EFB7E9B9FFC9E659FAF2FE228E55FE3C" box="[1281,1388,432,458]" pageId="12" pageNumber="513" value="1932-07">VII 1932</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B85347FFC9E659FA8AFE228BC7FE1B" pageId="12" pageNumber="513">Dr. Lieftinck</collectorName>
|
||
leg. [
|
||
<collectionCode id="ED5CAE54FFC9E659FEC8FE468A45FE18" box="[315,380,468,494]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="12" pageNumber="513" type="Museum">ZMH</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
»;
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF34FE6A8FF0FDC3" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
<materialsCitation id="3B253CCCFFC9E659FF34FE6A8F81FDC3" ID-GBIF-Occurrence="4168147306" collectingDate="1932-07" collectorName="Dr. Lieftinck" country="Indonesia" location="Buitenzorg" pageId="12" pageNumber="513" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
|
||
3)
|
||
<bibRefCitation id="EFDC4B60FFC9E659FF14FE65895CFDE7" author="Sartori, M. & Kubiak, M. & Michalik, P." box="[231,613,503,530]" pageId="12" pageNumber="513" pagination="28 - 32" refId="ref10026" refString="Sartori, M., Kubiak, M. & Michalik, P. (2016) Deciphering genital anatomy of rare, delicate and precious specimens: first study of two type specimens of mayflies using micro-computed X-ray tomography (Ephemeroptera; Heptageniidae). Zoosymposia, 11, 28 - 32. https: // doi. org / 10.11646 / zoosymposia. 11.1.7" type="journal article" year="2016">Sartori, Kubiak & Michalik 2016</bibRefCitation>
|
||
, p. 30: ́The
|
||
<typeStatus id="54F68833FFC9E659FD0DFE658859FDE7" box="[766,864,503,529]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4D4D12FFC9E659FC7AFE658F3BFDE7" authorityName=": Wang & McCafferty" authorityYear="2004" box="[905,1026,503,529]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFC9E659FC7AFE658F3BFDE7" box="[905,1026,503,529]" italics="true" pageId="12" pageNumber="513">Rh. ornata</emphasis>
|
||
</taxonomicName>
|
||
is a specimen stored in ethanol and originating from
|
||
<collectingCountry id="F35A7601FFC9E659FEA4FD898AFEFDC3" box="[343,455,539,565]" name="Indonesia" pageId="12" pageNumber="513">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="4989F873FFC9E659FE22FD8E893CFDC0" box="[465,517,540,566]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="513">Java</collectingRegion>
|
||
,
|
||
<location id="8E92604AFFC9E659FDE2FD8989A9FDC3" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FDE2FD8989A9FDC3" box="[529,656,539,565]" country="Indonesia" name="Buitenzorg" pageId="12" pageNumber="513" stateProvince="Jawa Timur">Buitenzorg</location>
|
||
, [Bogor], collected in
|
||
<date id="FFF31051FFC9E659FC78FD8988C1FDC0" box="[907,1016,539,566]" pageId="12" pageNumber="513" value="1932-07">
|
||
<collectingDate id="EFB7E9B9FFC9E659FC78FD8988C1FDC0" box="[907,1016,539,566]" pageId="12" pageNumber="513" value="1932-07">July 1932</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26B85347FFC9E659FBD1FD8E8F81FDC3" box="[1058,1208,539,566]" pageId="12" pageNumber="513">Dr. Lieftinck</collectorName>
|
||
</materialsCitation>
|
||
».
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF34FDAD8EBBFC8F" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
Formerly (
|
||
<bibRefCitation id="EFDC4B60FFC9E659FEB3FDAD8AEBFDAC" author="Sartori, M." box="[320,466,575,602]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDC4B60FFC9E659FE2CFDAD8988FDAF" author="Sartori, M. & Kubiak, M. & Michalik, P." box="[479,689,575,602]" pageId="12" pageNumber="513" pagination="28 - 32" refId="ref10026" refString="Sartori, M., Kubiak, M. & Michalik, P. (2016) Deciphering genital anatomy of rare, delicate and precious specimens: first study of two type specimens of mayflies using micro-computed X-ray tomography (Ephemeroptera; Heptageniidae). Zoosymposia, 11, 28 - 32. https: // doi. org / 10.11646 / zoosymposia. 11.1.7" type="journal article" year="2016">
|
||
Sartori
|
||
<emphasis id="B939EA83FFC9E659FDC7FDD2895EFDAF" box="[564,615,575,601]" italics="true" pageId="12" pageNumber="513">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
) this specimen was regarded to be the
|
||
<typeStatus id="54F68833FFC9E659FB80FDAD8FE0FDAF" box="[1139,1241,575,601]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
. However
|
||
<bibRefCitation id="EFDC4B60FFC9E659FAA6FDAD8BDBFD88" author="Ulmer, G." pageId="12" pageNumber="513" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer (1939)</bibRefCitation>
|
||
did not designate a single specimen as the
|
||
<typeStatus id="54F68833FFC9E659FD34FDF18810FD8B" box="[711,809,611,637]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
and did not distinguish
|
||
<specimenCount id="9D4BFD18FFC9E659FBCAFDF68FE6FD8B" box="[1081,1247,611,638]" pageId="12" pageNumber="513" type="generic">two specimens</specimenCount>
|
||
(the male imago and the female imago) as
|
||
<typeStatus id="54F68833FFC9E659FE42FD15892AFD57" box="[433,531,647,673]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54F68833FFC9E659FDB4FD15899DFD57" box="[583,676,647,673]" pageId="12" pageNumber="513" type="allotype">allotype</typeStatus>
|
||
, but reported both them as types of equal status.According to Article 73 of the International Code of Zoological Nomenclature (4th edition), ́73.1.1. If an author when establishing a new nominal species-group taxon states in the original publication that
|
||
<emphasis id="B939EA83FFC9E659FC63FD428FE4FD1C" bold="true" box="[912,1245,719,746]" pageId="12" pageNumber="513">
|
||
<specimenCount id="9D4BFD18FFC9E659FC63FD428F0AFD1F" box="[912,1075,719,746]" pageId="12" pageNumber="513" type="generic">one specimen</specimenCount>
|
||
, and only one
|
||
</emphasis>
|
||
, is the
|
||
<typeStatus id="54F68833FFC9E659FAC0FD5D8EA1FD1F" box="[1331,1432,719,745]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
, or “the type”, or uses some equivalent expression, that specimen is the
|
||
<typeStatus id="54F68833FFC9E659FC40FD618F2CFCFB" box="[947,1045,755,781]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
fixed by original designation» and ́73.1.3. The
|
||
<typeStatus id="54F68833FFC9E659FEDDFC858AA9FCC7" box="[302,400,791,817]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
of a new nominal species-group taxon can only be fixed in the original publication and by the original author». Thus, according to the Article 73.2 of the Code, both specimens reported by
|
||
<bibRefCitation id="EFDC4B60FFC9E659FB17FCA98E44FCA0" author="Ulmer, G." box="[1252,1405,827,854]" pageId="12" pageNumber="513" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer (1939)</bibRefCitation>
|
||
as ́Typen» were
|
||
<typeStatus id="54F68833FFC9E659FECEFCF28A9BFC8C" box="[317,418,864,890]" pageId="12" pageNumber="513" type="syntype">syntypes</typeStatus>
|
||
, i.e. name-bearing types of equal status. Here one of them is designated as the
|
||
<typeStatus id="54F68833FFC9E659FAE0FCCD8E44FC8F" box="[1299,1405,863,889]" pageId="12" pageNumber="513" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF34FC1188CEFBFC" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
<emphasis id="B939EA83FFC9E659FF34FC118A88FC6B" bold="true" box="[199,433,899,925]" pageId="12" pageNumber="513">Material examined.</emphasis>
|
||
<materialsCitation id="3B253CCCFFC9E659FE4BFC168AB8FC10" ID-GBIF-Occurrence="4168147310" collectingDate="2013-01-11" collectingDateMax="2022-12-30" collectingDateMin="2013-01-11" collectionCode="ZIN, R" collectorName="N. Kluge & L. Sheyko" country="India" county="Shivamogga" location="Agumbe" municipality="Udupi" pageId="12" pageNumber="513" specimenCount="18" specimenCount-female="2" specimenCount-larva="16" stateProvince="Karnataka">
|
||
<collectingCountry id="F35A7601FFC9E659FE4BFC168935FC68" box="[440,524,900,926]" name="India" pageId="12" pageNumber="513">INDIA</collectingCountry>
|
||
: state
|
||
<collectingRegion id="4989F873FFC9E659FDA2FC1189FEFC6B" box="[593,711,899,925]" country="India" name="Karnataka" pageId="12" pageNumber="513">Karnataka</collectingRegion>
|
||
, border of Shivamogga and
|
||
<collectingMunicipality id="6B96ACEBFFC9E659FBF7FC118F73FC6B" box="[1028,1098,899,925]" pageId="12" pageNumber="513">Udupi</collectingMunicipality>
|
||
districts, near
|
||
<location id="8E92604AFFC9E659FB1DFC118E68FC6B" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FB1DFC118E68FC6B" box="[1262,1361,899,925]" country="India" county="Shivamogga" municipality="Udupi" name="Agumbe" pageId="12" pageNumber="513" stateProvince="Karnataka">Agumbe</location>
|
||
, rivers
|
||
<location id="8E92604AFFC9E659FF64FC358A34FC37" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FF64FC358A34FC37" box="[151,269,935,961]" country="India" county="Shivamogga" municipality="Udupi" name="Modi-hole" pageId="12" pageNumber="513" stateProvince="Karnataka">Modi-hole</location>
|
||
and
|
||
<location id="8E92604AFFC9E659FEB7FC358930FC37" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FEB7FC358930FC37" box="[324,521,935,961]" country="India" county="Shivamogga" municipality="Udupi" name="Seethanadhi-hole" pageId="12" pageNumber="513" stateProvince="Karnataka">Seethanadhi-hole</location>
|
||
,
|
||
<date id="FFF31051FFC9E659FDE7FC3A8992FC34" box="[532,683,936,962]" pageId="12" pageNumber="513" value="2013-01-11" valueMax="2013-01-31" valueMin="2013-01-11">
|
||
<collectingDate id="EFB7E9B9FFC9E659FDE7FC3A8992FC34" box="[532,683,936,962]" pageId="12" pageNumber="513" value="2013-01-11" valueMax="2013-01-31" valueMin="2013-01-11">11–31.I.2013</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26B85347FFC9E659FD1CFC3A886CFC37" box="[751,853,935,962]" pageId="12" pageNumber="513">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="26B85347FFC9E659FC8AFC3A88C9FC37" box="[889,1008,935,962]" pageId="12" pageNumber="513">L. Sheyko</collectorName>
|
||
: 1 L-S-I
|
||
<specimenCount id="9D4BFD18FFC9E659FBA2FC3B8F5DFC32" box="[1105,1124,937,964]" pageId="12" pageNumber="513">♂</specimenCount>
|
||
, 1 L-S
|
||
<specimenCount id="9D4BFD18FFC9E659FB40FC3B8FFEFC32" box="[1203,1223,937,964]" pageId="12" pageNumber="513">♂</specimenCount>
|
||
, 1 S
|
||
<specimenCount id="9D4BFD18FFC9E659FB04FC3B8E37FC32" box="[1271,1294,937,964]" pageId="12" pageNumber="513">♂</specimenCount>
|
||
, 2 L-S-I
|
||
<specimenCount id="9D4BFD18FFC9E659FA87FC3A8EBBFC34" box="[1396,1410,936,962]" pageId="12" pageNumber="513" type="female">♀</specimenCount>
|
||
, 1 S
|
||
<specimenCount id="9D4BFD18FFC9E659FF59FC5E8B85FC10" box="[170,188,972,998]" pageId="12" pageNumber="513" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D4BFD18FFC9E659FF39FC598A08FC13" box="[202,305,971,997]" pageId="12" pageNumber="513" type="larva">16 larvae</specimenCount>
|
||
(
|
||
<collectionCode id="ED5CAE54FFC9E659FEB2FC5E8A4CFC10" box="[321,373,972,998]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="513" type="Museum">ZIN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B253CCCFFC9E659FE7AFC5E88CAFBFC" ID-GBIF-Occurrence="4168147305" collectingDate="2013-01-11" collectingDateMax="2022-12-30" collectingDateMin="2013-01-11" collectionCode="ZIN, R" collectorName="P. Srinivasan" country="India" county="Shivamogga" location="Agumbe" municipality="Udupi" pageId="12" pageNumber="513" specimenCount="18" specimenCount-female="2" specimenCount-larva="16" specimenCount-male="2" stateProvince="Karnataka">
|
||
state
|
||
<collectingRegion id="4989F873FFC9E659FE31FC598970FC13" box="[450,585,971,997]" country="India" name="Tamil Nadu" pageId="12" pageNumber="513">Tamil Nadu</collectingRegion>
|
||
, Dindigul, Kodaikanal hills, Perumal malai stream,
|
||
<date id="FFF31051FFC9E659FB63FC5E8E7CFC10" box="[1168,1349,972,998]" pageId="12" pageNumber="513" value="2022-12-28" valueMax="2022-12-30" valueMin="2022-12-28">
|
||
<collectingDate id="EFB7E9B9FFC9E659FB63FC5E8E7CFC10" box="[1168,1349,972,998]" pageId="12" pageNumber="513" value="2022-12-28" valueMax="2022-12-30" valueMin="2022-12-28">28–30.XII.2022</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26B85347FFC9E659FA7AFC5E8A34FBFF" pageId="12" pageNumber="513">P. Srinivasan</collectorName>
|
||
&
|
||
<collectionCode id="ED5CAE54FFC9E659FEC2FC628A7EFBFC" box="[305,327,1008,1034]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="513">R</collectionCode>
|
||
. Isack: 1 L-S-I
|
||
<specimenCount id="9D4BFD18FFC9E659FE01FC63893CFBFA" box="[498,517,1009,1036]" pageId="12" pageNumber="513" type="male">♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4BFD18FFC9E659FDE3FC62890EFBFA" box="[528,567,1008,1036]" pageId="12" pageNumber="513" type="larva">2 L</specimenCount>
|
||
/S
|
||
<specimenCount id="9D4BFD18FFC9E659FDA2FC63895FFBFA" box="[593,614,1009,1036]" pageId="12" pageNumber="513" type="male">♂</specimenCount>
|
||
, 1 mature
|
||
<specimenCount id="9D4BFD18FFC9E659FD28FC7D882AFBFF" box="[731,787,1007,1033]" pageId="12" pageNumber="513" type="larva">larva</specimenCount>
|
||
<specimenCount id="9D4BFD18FFC9E659FCE9FC628816FBFC" box="[794,815,1008,1034]" pageId="12" pageNumber="513" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D4BFD18FFC9E659FCCEFC7D88AFFBFF" box="[829,918,1007,1034]" pageId="12" pageNumber="513" type="larva">5 larvae</specimenCount>
|
||
(
|
||
<collectionCode id="ED5CAE54FFC9E659FC56FC6288D2FBFC" box="[933,1003,1008,1034]" country="USA" httpUri="http://grbio.org/cool/2xyg-zxc2" name="Department of Biologics Research" pageId="12" pageNumber="513">AMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF34FB86888CFB6C" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
<materialsCitation id="3B253CCCFFC9E659FF34FB868BD9FB80" ID-GBIF-Occurrence="4168147308" collectingDate="2015-01-29" collectingDateMax="2015-02-14" collectingDateMin="2015-01-29" collectionCode="ZIN, IO" collectorName="N. Kluge & L. Sheyko" country="Thailand" county="river Kwai-Yai" location="Si Sawat" municipality="Khwae Yai" pageId="12" pageNumber="513" specimenCount="4" specimenCount-female="1" specimenCount-larva="3" stateProvince="Kanchanaburi">
|
||
<collectingCountry id="F35A7601FFC9E659FF34FB868A65FBD8" box="[199,348,1044,1070]" name="Thailand" pageId="12" pageNumber="513">THAILAND</collectingCountry>
|
||
:
|
||
<collectingRegion id="4989F873FFC9E659FE95FB818956FBDB" box="[358,623,1043,1069]" country="Thailand" name="Kanchanaburi" pageId="12" pageNumber="513">Kanchanaburi province</collectingRegion>
|
||
, river Kwai-Yai (=
|
||
<collectingMunicipality id="6B96ACEBFFC9E659FCB4FB8188FBFBDB" box="[839,962,1043,1069]" pageId="12" pageNumber="513">Khwae Yai</collectingMunicipality>
|
||
=
|
||
<location id="8E92604AFFC9E659FC2CFB818F7AFBD8" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FC2CFB818F7AFBD8" box="[991,1091,1043,1070]" country="Thailand" county="river Kwai-Yai" municipality="Khwae Yai" name="Si Sawat" pageId="12" pageNumber="513" stateProvince="Kanchanaburi">Si Sawat</location>
|
||
) and
|
||
<location id="8E92604AFFC9E659FB72FB818E26FBDB" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FB72FB818E26FBDB" box="[1153,1311,1043,1069]" country="Thailand" county="river Kwai-Yai" municipality="Khwae Yai" name="river Taphoen" pageId="12" pageNumber="513" stateProvince="Kanchanaburi">river Taphoen</location>
|
||
, Lad-Ya (=
|
||
<location id="8E92604AFFC9E659FF64FBAA8BDDFBA4" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FF64FBAA8BDDFBA4" box="[151,228,1080,1106]" country="Thailand" county="river Kwai-Yai" municipality="Khwae Yai" name="Lat Ya" pageId="12" pageNumber="513" stateProvince="Kanchanaburi">Lat Ya</location>
|
||
), resort «Island Resort River Kwai»,
|
||
<date id="FFF31051FFC9E659FD76FBA58827FBA4" box="[645,798,1079,1106]" pageId="12" pageNumber="513" value="2015-01-29" valueMax="2015-01-30" valueMin="2015-01-29">
|
||
<collectingDate id="EFB7E9B9FFC9E659FD76FBA58827FBA4" box="[645,798,1079,1106]" pageId="12" pageNumber="513" value="2015-01-29" valueMax="2015-01-30" valueMin="2015-01-29">29–30.I.2015</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26B85347FFC9E659FC92FBAA88FEFBA7" box="[865,967,1079,1106]" pageId="12" pageNumber="513">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="26B85347FFC9E659FC18FBAA8F5BFBA7" box="[1003,1122,1079,1106]" pageId="12" pageNumber="513">L. Sheyko</collectorName>
|
||
: 1 L-S
|
||
<specimenCount id="9D4BFD18FFC9E659FB5CFBAB8FFAFBA2" box="[1199,1219,1081,1108]" pageId="12" pageNumber="513">♂</specimenCount>
|
||
, 3 L-S-I
|
||
<specimenCount id="9D4BFD18FFC9E659FADAFBAA8E0EFBA4" box="[1321,1335,1080,1106]" pageId="12" pageNumber="513" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D4BFD18FFC9E659FAB0FBAA8EA5FBA7" box="[1347,1436,1079,1106]" pageId="12" pageNumber="513" type="larva">3 larvae</specimenCount>
|
||
(
|
||
<collectionCode id="ED5CAE54FFC9E659FF6CFBCE8BEAFB80" box="[159,211,1116,1142]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="513" type="Museum">ZIN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B253CCCFFC9E659FF14FBCE8888FB6C" ID-GBIF-Occurrence="4168147304" collectingDate="2015-01-29" collectingDateMax="2015-02-14" collectingDateMin="2015-01-29" collectionCode="ZIN, IO" collectorName="N. Kluge & L. Sheyko" country="Thailand" county="river Kwai-Yai" location="Si Sawat" municipality="Khwae Yai" pageId="12" pageNumber="513" specimenCount="4" specimenCount-female="1" specimenCount-larva="3" specimenCount-male="2" stateProvince="Kanchanaburi">
|
||
Mae-Hong-Son province,
|
||
<locationDeviation id="9A402FB3FFC9E659FDF8FBCE882FFB83" box="[523,790,1115,1142]" location="Chiang-Mai" pageId="12" pageNumber="513">
|
||
<quantity id="4CB59B74FFC9E659FDF8FBCE8968FB83" box="[523,593,1115,1142]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="513" unit="km" value="90.0">90 km</quantity>
|
||
NW Chiang-Mai
|
||
</locationDeviation>
|
||
,
|
||
<location id="8E92604AFFC9E659FCD2FBC988B8FB83" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFC9E659FCD2FBC988B8FB83" box="[801,897,1115,1141]" country="Thailand" county="river Kwai-Yai" municipality="Khwae Yai" name="river Pai" pageId="12" pageNumber="513" stateProvince="Kanchanaburi">river Pai</location>
|
||
upstream Pai,
|
||
<date id="FFF31051FFC9E659FBD6FBC98F81FB80" box="[1061,1208,1115,1142]" pageId="12" pageNumber="513" value="2015-02-05" valueMax="2015-02-14" valueMin="2015-02-05">
|
||
<collectingDate id="EFB7E9B9FFC9E659FBD6FBC98F81FB80" box="[1061,1208,1115,1142]" pageId="12" pageNumber="513" value="2015-02-05" valueMax="2015-02-14" valueMin="2015-02-05">5–14.II.2015</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26B85347FFC9E659FB08FBCE8E58FB83" box="[1275,1377,1115,1142]" pageId="12" pageNumber="513">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="26B85347FFC9E659FA77FBCE8BD6FB6F" pageId="12" pageNumber="513">L. Sheyko</collectorName>
|
||
: 2 L-S-I
|
||
<specimenCount id="9D4BFD18FFC9E659FEA3FB138A5AFB6A" box="[336,355,1153,1180]" pageId="12" pageNumber="513">♂</specimenCount>
|
||
, 1 L-S/I
|
||
<specimenCount id="9D4BFD18FFC9E659FE31FB138AEEFB6A" box="[450,471,1153,1180]" pageId="12" pageNumber="513">♂</specimenCount>
|
||
, 1 S-I
|
||
<specimenCount id="9D4BFD18FFC9E659FDEEFB138908FB6A" box="[541,561,1153,1180]" pageId="12" pageNumber="513" type="male">♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4BFD18FFC9E659FDCFFB12895AFB6A" box="[572,611,1152,1180]" pageId="12" pageNumber="513" type="larva">1 L</specimenCount>
|
||
/S
|
||
<specimenCount id="9D4BFD18FFC9E659FD8FFB1389A8FB6A" box="[636,657,1153,1180]" pageId="12" pageNumber="513" type="male">♂</specimenCount>
|
||
, 2 L-S-I
|
||
<specimenCount id="9D4BFD18FFC9E659FD04FB12883CFB6C" box="[759,773,1152,1178]" pageId="12" pageNumber="513" type="female">♀</specimenCount>
|
||
, 1 L-S
|
||
<specimenCount id="9D4BFD18FFC9E659FCA5FB12885CFB6C" box="[854,869,1152,1178]" pageId="12" pageNumber="513" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED5CAE54FFC9E659FC87FB128891FB6C" box="[884,936,1152,1178]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="513" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C357651AFFC9E658FF34FB318F00F9DB" lastPageId="13" lastPageNumber="514" pageId="12" pageNumber="513" type="description">
|
||
<paragraph id="8BF23691FFC9E659FF34FB318A65FB4B" blockId="12.[151,1437,251,2042]" box="[199,348,1187,1213]" pageId="12" pageNumber="513">
|
||
<emphasis id="B939EA83FFC9E659FF34FB318A65FB4B" bold="true" box="[199,348,1187,1213]" pageId="12" pageNumber="513">Descriptions</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF34FB5A8A9EF9BF" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
<emphasis id="B939EA83FFC9E659FF34FB5A8A2AFB14" bold="true" box="[199,275,1224,1250]" pageId="12" pageNumber="513">
|
||
<emphasis id="B939EA83FFC9E659FF34FB5A8A32FB14" bold="true" box="[199,267,1224,1250]" italics="true" pageId="12" pageNumber="513">Larva</emphasis>
|
||
.
|
||
</emphasis>
|
||
CUTICULAR COLORATION: Cuticle either pale ochre, or with more or less contrasting brown areas and ochre blanks (
|
||
<figureCitation id="13762A14FFC9E659FE40FB798901FAF0" box="[435,568,1259,1286]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="12" pageNumber="513">Figs 11–25</figureCitation>
|
||
). Head usually with pair of submedian blanks adjacent to anterior margin; often with more or less expressed pair of wider blanks laterad of them (
|
||
<figureCitation id="13762A14FFC9E659FC2FFA9D8F85FADC" box="[988,1212,1295,1322]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="12" pageNumber="513">Figs 11, 15, 19, 22</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFC9E659FB38FA9D8BEAFABB" author="Soldan, T. & Braasch, D." pageId="12" pageNumber="513" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 16;
|
||
<bibRefCitation id="EFDC4B60FFC9E659FEC4FAA18AFFFAB8" author="Sartori, M." box="[311,454,1331,1358]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: fig. 6); rarely submedian blanks absent (
|
||
<bibRefCitation id="EFDC4B60FFC9E659FC66FAA18F1EFAB8" author="Sartori, M." box="[917,1063,1331,1358]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: fig. 7). Each femur mostly light, with 3 dark maculae originating from two initial transverse bands: initial proximal transverse band is separated into two longitudinal maculae; distal transverse band integral, either V-shaped (
|
||
<figureCitation id="13762A14FFC9E659FBDCFAE98FBDFA60" box="[1071,1156,1403,1430]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="12" pageNumber="513">Fig. 17</figureCitation>
|
||
), or Z-shaped (
|
||
<figureCitation id="13762A14FFC9E659FACBFAE98EB6FA60" box="[1336,1423,1403,1430]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="12" pageNumber="513">Fig. 13</figureCitation>
|
||
). Tibia mostly light, with base darkened. Abdominal terga II–VII with following blanks: median blank and pair of submedian blanks (submedian sigilla) are either separated, or fused into integral median blank; pair of smaller or larger blanks are located laterad of it and adjacent to posterior margin; pair of smaller or larger lateral blanks are adjacent to anterior margin. Terga I and VIII–IX with more extensive blanks. Abdominal sterna with small blanks corresponding to sigilla.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF3BF9C68A90F964" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
HYPODERMAL COLORATION: Abdominal terga with or without reddish markings resembling that of winged stages (
|
||
<figureCitation id="13762A14FFC9E659FEB7F9E58AA5F964" box="[324,412,1655,1682]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="12" pageNumber="513">Figs 53</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC9E659FF3BF90E8F8DF80C" blockId="12.[151,1437,251,2042]" pageId="12" pageNumber="513">
|
||
SHAPE AND SETATION: Labrum moderately expanded laterally (
|
||
<figureCitation id="13762A14FFC9E659FC17F9098F02F943" box="[996,1083,1691,1717]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="12" pageNumber="513">Fig. 26</figureCitation>
|
||
); anterior portion sharply bent ventrally and invisible from above, with narrow median emargination; initial anterior margin (hidden on ventral side) with pair of regular setal rows separated by median emargination; setae of these rows stout, pointed, spine-like, inclined medially toward emargination (
|
||
<figureCitation id="13762A14FFC9E659FD62F89589DCF8D4" box="[657,741,1799,1826]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="12" pageNumber="513">Fig. 27</figureCitation>
|
||
); dorsal surface of labrum with irregular, long, hair-like setae. Mandibles as described by
|
||
<bibRefCitation id="EFDC4B60FFC9E659FE26F8B98808F8B0" author="Soldan, T. & Braasch, D." box="[469,817,1835,1862]" pageId="12" pageNumber="513" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch (1986: 208</bibRefCitation>
|
||
, figs 11–12) and
|
||
<bibRefCitation id="EFDC4B60FFC9E659FC0CF8B98FE8F8B0" author="Sartori, M." box="[1023,1233,1835,1862]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori (2014: 52)</bibRefCitation>
|
||
; prostheca of left mandible consists of 3–4 setae, prostheca of right mandible consists of 2–4 setae (
|
||
<figureCitation id="13762A14FFC9E659FBD9F8DD8F97F89C" box="[1066,1198,1871,1898]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="12" pageNumber="513">Figs 29–30</figureCitation>
|
||
). Maxilla with 10–15 comb-like setae, each with 4–11 denticles (
|
||
<figureCitation id="13762A14FFC9E659FD88F8E189C7F878" box="[635,766,1907,1934]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="12" pageNumber="513">Figs 28, 33</figureCitation>
|
||
). Distal dentiseta simple, proximal dentiseta bifurcate, with proximal branch pectinate (
|
||
<figureCitation id="13762A14FFC9E659FE3BF8058922F844" box="[456,539,1943,1970]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="12" pageNumber="513">Fig. 33</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFC9E659FDD5F805898DF844" author="Sartori, M." box="[550,692,1943,1970]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: fig. 12). Hypopharynx and superlinguae usual for
|
||
<taxonomicName id="4C4D4D12FFC9E659FB1AF8058E56F847" authorityName="Eaton" authorityYear="1868" box="[1257,1391,1943,1969]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B939EA83FFC9E659FB1AF8058E56F847" box="[1257,1391,1943,1969]" italics="true" pageId="12" pageNumber="513">Ecdyonurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A57F8FFC9E659FA86F80A8EA4F847" box="[1397,1437,1943,1970]" pageId="12" pageNumber="513" sensu="lato">s. l.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13762A14FFC9E659FF53F8298BCAF820" box="[160,243,1979,2006]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="12" pageNumber="513">Fig. 32</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFC9E659FF0DF8298935F823" author="Soldan, T. & Braasch, D." box="[254,524,1979,2006]" pageId="12" pageNumber="513" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 2;
|
||
<bibRefCitation id="EFDC4B60FFC9E659FDACF82989D7F820" author="Sartori, M." box="[607,750,1979,2006]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: fig. 11). Labium with paraglossae sharply expanded; glossae with oblique ridges on apex (
|
||
<figureCitation id="13762A14FFC9E659FE2DF84D8908F80C" box="[478,561,2015,2042]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="12" pageNumber="513">Fig. 31</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFC9E659FDCEF84D8872F80F" author="Soldan, T. & Braasch, D." box="[573,843,2015,2042]" pageId="12" pageNumber="513" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 9;
|
||
<bibRefCitation id="EFDC4B60FFC9E659FC53F84D8F16F80C" author="Sartori, M." box="[928,1071,2015,2042]" pageId="12" pageNumber="513" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: figs 9–10).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FF058F7FFEEB" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
Spatulate setae on dorsal surface of each femur flat, colorless, with narrow base, divergent margins and rounded apex (
|
||
<figureCitation id="13762A14FFC8E658FF29FF298A15FF23" box="[218,300,186,213]" captionStart="FIGURES 34–41" captionStartId="6.[151,264,1916,1940]" captionTargetBox="[151,1435,181,1893]" captionTargetId="figure-37@6.[151,1435,181,1893]" captionTargetPageId="6" captionText="FIGURES 34–41. Ecdyonurus (Rhithrogeniella) ornatus, exuviae of larval legs (reared specimen from Kodaikanal). 34, fore femur; 35, outer margin of fore femur; 36–38, fore, middle and hind legs; 39, hind tibia; 40, hind tarsus; 41, claw." figureDoi="http://doi.org/10.5281/zenodo.8203258" httpUri="https://zenodo.org/record/8203258/files/figure.png" pageId="13" pageNumber="514">Fig. 35</figureCitation>
|
||
). Tibia and tarsus of each leg with fewer similar setae (
|
||
<figureCitation id="13762A14FFC8E658FC79FF2988E4FF23" box="[906,989,187,213]" captionStart="FIGURES 34–41" captionStartId="6.[151,264,1916,1940]" captionTargetBox="[151,1435,181,1893]" captionTargetId="figure-37@6.[151,1435,181,1893]" captionTargetPageId="6" captionText="FIGURES 34–41. Ecdyonurus (Rhithrogeniella) ornatus, exuviae of larval legs (reared specimen from Kodaikanal). 34, fore femur; 35, outer margin of fore femur; 36–38, fore, middle and hind legs; 39, hind tibia; 40, hind tarsus; 41, claw." figureDoi="http://doi.org/10.5281/zenodo.8203258" httpUri="https://zenodo.org/record/8203258/files/figure.png" pageId="13" pageNumber="514">Fig. 40</figureCitation>
|
||
); middle and hind tibiae with sparse row of hair-like setae on outer margin (
|
||
<figureCitation id="13762A14FFC8E658FDE9FF4D8957FF0F" box="[538,622,223,249]" captionStart="FIGURES 34–41" captionStartId="6.[151,264,1916,1940]" captionTargetBox="[151,1435,181,1893]" captionTargetId="figure-37@6.[151,1435,181,1893]" captionTargetPageId="6" captionText="FIGURES 34–41. Ecdyonurus (Rhithrogeniella) ornatus, exuviae of larval legs (reared specimen from Kodaikanal). 34, fore femur; 35, outer margin of fore femur; 36–38, fore, middle and hind legs; 39, hind tibia; 40, hind tarsus; 41, claw." figureDoi="http://doi.org/10.5281/zenodo.8203258" httpUri="https://zenodo.org/record/8203258/files/figure.png" pageId="13" pageNumber="514">Fig. 39</figureCitation>
|
||
). Claw with rectangular projection at midlength and with one row of 3–6 denticles distad of it (
|
||
<figureCitation id="13762A14FFC8E658FE7AFE918AE5FEEB" box="[393,476,259,285]" captionStart="FIGURES 34–41" captionStartId="6.[151,264,1916,1940]" captionTargetBox="[151,1435,181,1893]" captionTargetId="figure-37@6.[151,1435,181,1893]" captionTargetPageId="6" captionText="FIGURES 34–41. Ecdyonurus (Rhithrogeniella) ornatus, exuviae of larval legs (reared specimen from Kodaikanal). 34, fore femur; 35, outer margin of fore femur; 36–38, fore, middle and hind legs; 39, hind tibia; 40, hind tarsus; 41, claw." figureDoi="http://doi.org/10.5281/zenodo.8203258" httpUri="https://zenodo.org/record/8203258/files/figure.png" pageId="13" pageNumber="514">Fig. 41</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFC8E658FE1BFE9189CEFEEB" author="Soldan, T. & Braasch, D." box="[488,759,259,285]" pageId="13" pageNumber="514" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 13;
|
||
<bibRefCitation id="EFDC4B60FFC8E658FCAAFE9188D0FEEB" author="Sartori, M." box="[857,1001,259,285]" pageId="13" pageNumber="514" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: fig. 18)
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FEB58E42FE27" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
Denticles on posterior margins of abdominal terga irregular, either narrow and sharply pointed, or dentate; denticles on terga I and X very small (
|
||
<figureCitation id="13762A14FFC8E658FDB6FED989FEFE93" box="[581,711,331,357]" captionStart="FIGURES 42–52" captionStartId="7.[151,264,1785,1809]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-20@7.[151,1436,181,1762]" captionTargetPageId="7" captionText="FIGURES 42–52. Larvae. 42–50, Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae: 42–44, specimen from Agumbe (India), posterior margins of abdominal terga I, VIII and X; 45, specimen from Pai (Thailand), posterior margins of abdominal tergum VI; 46–48, cercus of specimen from Agumbe: 46, proximal portion; 47, middle portion in median-ventral view, to show bases of primary swimming setae on inner (median) side; 48, the same portion with focus on lateral-dorsal side, to show bases of secondary swimming setae on outer (lateral) side; 49–50, cercus of specimen from Kodaikanal (India). 51–52, Ecdyonurus (Rhithrogeniella) joernensis (Russia, Orenburg province, Tashla, 16.VIII.1986, coll. N. Kluge): two portions of cercus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8203260" httpUri="https://zenodo.org/record/8203260/files/figure.png" pageId="13" pageNumber="514">Figs 42, 44</figureCitation>
|
||
), denticles on terga II–IX larger (
|
||
<figureCitation id="13762A14FFC8E658FBCCFED98FFBFE93" box="[1087,1218,330,357]" captionStart="FIGURES 42–52" captionStartId="7.[151,264,1785,1809]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-20@7.[151,1436,181,1762]" captionTargetPageId="7" captionText="FIGURES 42–52. Larvae. 42–50, Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae: 42–44, specimen from Agumbe (India), posterior margins of abdominal terga I, VIII and X; 45, specimen from Pai (Thailand), posterior margins of abdominal tergum VI; 46–48, cercus of specimen from Agumbe: 46, proximal portion; 47, middle portion in median-ventral view, to show bases of primary swimming setae on inner (median) side; 48, the same portion with focus on lateral-dorsal side, to show bases of secondary swimming setae on outer (lateral) side; 49–50, cercus of specimen from Kodaikanal (India). 51–52, Ecdyonurus (Rhithrogeniella) joernensis (Russia, Orenburg province, Tashla, 16.VIII.1986, coll. N. Kluge): two portions of cercus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8203260" httpUri="https://zenodo.org/record/8203260/files/figure.png" pageId="13" pageNumber="514">Figs 43, 45</figureCitation>
|
||
). Abdominal sterna without denticles. Tergalii increasing from II to V; tergalii I–VI with branched fibrillose portion, tergalii VII without fibrillose portion (
|
||
<figureCitation id="13762A14FFC8E658FE97FE018AE3FE5B" box="[356,474,403,429]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1810,1834]" captionTargetBox="[151,1436,180,1772]" captionTargetId="figure-20@3.[151,1436,179,1784]" captionTargetPageId="3" captionText="FIGURES 1–10. Ecdyonurus (Rhithrogeniella) ornatus from Kodaikanal (India). 1–3, larvae; 4–10, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8203250" httpUri="https://zenodo.org/record/8203250/files/figure.png" pageId="13" pageNumber="514">Figs 4–10</figureCitation>
|
||
). Number of branches of fibrillose portion either subequal on all tergalii I–VI, or less on tergalius VI (
|
||
<figureCitation id="13762A14FFC8E658FEBEFE258AABFE27" box="[333,402,439,465]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1810,1834]" captionTargetBox="[151,1436,180,1772]" captionTargetId="figure-20@3.[151,1436,179,1784]" captionTargetPageId="3" captionText="FIGURES 1–10. Ecdyonurus (Rhithrogeniella) ornatus from Kodaikanal (India). 1–3, larvae; 4–10, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8203250" httpUri="https://zenodo.org/record/8203250/files/figure.png" pageId="13" pageNumber="514">Fig. 9</figureCitation>
|
||
); among individuals examined, number of branches on tergalius VI varying from 2 to 12.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FE4888D5FD07" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34FE488A7AFE03" bold="true" box="[199,323,474,501]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34FE488A05FE03" bold="true" box="[199,316,474,501]" italics="true" pageId="13" pageNumber="514">Subimago</emphasis>
|
||
.
|
||
</emphasis>
|
||
CUTICULAR COLORATION: Head colorless, antennae light brown. Pronotum colorless. Mesonotum mostly colorless, with brown anterior chromozone (area anteriad of mesonotal suture), paired brown area including antelateroparapsidal suture and paired brown parascutellar chromozone (ovoid macula on parascutellum); paired lateroparapsidal chromozone (stretching along lateroparapsidal suture and widening posteriorly) either lighter brown (
|
||
<figureCitation id="13762A14FFC8E658FECFFDF98AB7FD73" box="[316,398,619,645]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 66</figureCitation>
|
||
), or colorless (
|
||
<figureCitation id="13762A14FFC8E658FDDCFDF989BBFD73" box="[559,642,619,645]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 67</figureCitation>
|
||
). Thoracic pleura and sterna colorless, with ventral ark of prealar bridge narrowly bordered with light brown (
|
||
<figureCitation id="13762A14FFC8E658FDB4FD1D8992FD5F" box="[583,683,655,681]" captionStart="FIGURES 103–110" captionStartId="19.[151,264,1692,1716]" captionTargetBox="[154,1436,345,1669]" captionTargetId="figure-106@19.[151,1436,345,1669]" captionTargetPageId="19" captionText="FIGURES 103–110. 103–109, Ecdyonurus (Rhithrogeniella) sp. 2: 103, male imago, 104, abdomen of male imago; 105, male subimago; 106, female subimago; 107, thorax of male subimago, ventral view; 108, male subimaginal exuviae; 109, enlarged part of its ventral arc of prealar bridge; 110, the same of E. (Rh.) ornatus." figureDoi="http://doi.org/10.5281/zenodo.8203276" httpUri="https://zenodo.org/record/8203276/files/figure.png" pageId="13" pageNumber="514">Fig. 110</figureCitation>
|
||
). Wing membrane colorless, microtrichia brown. Legs from light brown to nearly colorless; fore leg darker than middle and hind legs. Abdominal terga and sterna nearly colorless; gonostyli light brownish; cerci proximally colorless, distally light brownish.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF3BFD6989CDFCE3" blockId="13.[151,1437,151,2013]" box="[200,756,763,789]" pageId="13" pageNumber="514">HYPODERMAL COLORATION: As in imago.</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FC8D88ADFCAB" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
TEXTURE: In both sexes, on all leg pairs, all tarsomeres covered with blunt microlepides; microtrichia occupy only ventral-proximal part of first tarsomere (
|
||
<figureCitation id="13762A14FFC8E658FD65FCD189DEFCAB" box="[662,743,834,861]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 65</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFDC4B60FFC8E658FCF3FCD188B1FCAB" author="Kluge, N. J." box="[768,904,835,861]" pageId="13" pageNumber="514" pagination="151 - 186" refId="ref9941" refString="Kluge, N. J. (2022) Taxonomic significance of microlepides on subimaginal tarsi of Ephemeroptera. Zootaxa, 5159 (2), 151 - 186. https: // doi. org / 10.11646 / zootaxa. 5159.2.1" type="journal article" year="2022">Kluge 2022</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FCF588BEFC77" blockId="13.[151,1437,151,2013]" box="[199,903,870,897]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34FCF58A61FC77" bold="true" box="[199,344,870,897]" italics="true" pageId="13" pageNumber="514">Imago, male</emphasis>
|
||
(
|
||
<figureCitation id="13762A14FFC8E658FE94FCF58AD0FC77" box="[359,489,870,897]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Figs 57, 64</figureCitation>
|
||
). Head ochre with brown. Eyes gray.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FC198841FB8B" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
Pronotum ochre with brown macula medially (as in females —
|
||
<figureCitation id="13762A14FFC8E658FC65FC198F20FC53" box="[918,1049,907,933]" captionStart="FIGURES 77–84" captionStartId="14.[151,264,1532,1556]" captionTargetBox="[151,1435,309,1509]" captionTargetId="figure-91@14.[151,1435,309,1509]" captionTargetPageId="14" captionText="FIGURES 77–84. Ecdyonurus (Rhithrogeniella) ornatus, female imagines. 77–78, specimen from Agumbe (India); 79–81, specimens from Kwai-Yai (Thailand); 82–84, specimens from Pai (Thailand)." figureDoi="http://doi.org/10.5281/zenodo.8203268" httpUri="https://zenodo.org/record/8203268/files/figure.png" pageId="13" pageNumber="514">Figs 77–84</figureCitation>
|
||
). Mesonotum with ochre and light brown areas. Thoracic pleura and sterna light ochre (as in female —
|
||
<figureCitation id="13762A14FFC8E658FC73FC3D88E8FC3F" box="[896,977,943,969]" captionStart="FIGURES 77–84" captionStartId="14.[151,264,1532,1556]" captionTargetBox="[151,1435,309,1509]" captionTargetId="figure-91@14.[151,1435,309,1509]" captionTargetPageId="14" captionText="FIGURES 77–84. Ecdyonurus (Rhithrogeniella) ornatus, female imagines. 77–78, specimen from Agumbe (India); 79–81, specimens from Kwai-Yai (Thailand); 82–84, specimens from Pai (Thailand)." figureDoi="http://doi.org/10.5281/zenodo.8203268" httpUri="https://zenodo.org/record/8203268/files/figure.png" pageId="13" pageNumber="514">Fig. 77</figureCitation>
|
||
). Wing membrane colorless; veins mostly colorless, some longitudinal veins ochre or light brown (
|
||
<figureCitation id="13762A14FFC8E658FCFBFC4188B1FC1B" box="[776,904,978,1005]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Figs 57, 64</figureCitation>
|
||
; as in
|
||
<figureCitation id="13762A14FFC8E658FC3FFC418F74FC1B" box="[972,1101,978,1005]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Figs 58–61</figureCitation>
|
||
). Pterostigma with simple and complete crossveins perpendicular to Sc (
|
||
<figureCitation id="13762A14FFC8E658FD80FC6589F0FBE7" box="[627,713,1014,1041]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 58</figureCitation>
|
||
). Hind wing narrowing toward apex, with small, pointed costal projection (
|
||
<figureCitation id="13762A14FFC8E658FEE4FB898A53FBC3" box="[279,362,1051,1077]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 61</figureCitation>
|
||
). On fore leg, femur gradually changing color from light ochre base to dark reddish-brown apex, with reddish-brown darkening at midlength; tibia ochre with base and apex contrastingly dark brown; tarsus ochre with apex brown (
|
||
<figureCitation id="13762A14FFC8E658FE90FBF18A8EFB8B" box="[355,439,1123,1149]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 62</figureCitation>
|
||
). Middle and hind legs lighter (
|
||
<figureCitation id="13762A14FFC8E658FCE4FBF18853FB8B" box="[791,874,1123,1149]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 63</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FB158920FAC7" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
Abdomen mostly ochre, all terga I–X with reddish-brown paired submedian stripe and ochre submedian sigilla on its background (
|
||
<figureCitation id="13762A14FFC8E658FE88FB39893AFB33" box="[379,515,1194,1221]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Figs 54–57</figureCitation>
|
||
). Penis with proximal ⅔ unpaired, distal ¼ divided into pair of rounded lobes separated by wide space (
|
||
<figureCitation id="13762A14FFC8E658FE4FFB5D8978FB1F" box="[444,577,1231,1257]" captionStart-0="FIGURES 69–72" captionStart-1="FIGURES 73–76" captionStartId-0="9.[151,264,1837,1861]" captionStartId-1="10.[151,264,1352,1376]" captionTargetBox-0="[151,1436,182,1813]" captionTargetBox-1="[151,1436,181,1328]" captionTargetId-0="figure-20@9.[151,1436,181,1813]" captionTargetId-1="figure-37@10.[151,1436,181,1328]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 69–72. Ecdyonurus (Rhithrogeniella) ornatus, genitalia of male imagines, ventral view. 69–70, specimen from Kodaikanal (India) in tense condition; 71, specimen from Agumbe (India); 72, specimen from Pai (Thailand)." captionText-1="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi-0="http://doi.org/10.5281/zenodo.8203264" figureDoi-1="http://doi.org/10.5281/zenodo.8203266" httpUri-0="https://zenodo.org/record/8203264/files/figure.png" httpUri-1="https://zenodo.org/record/8203266/files/figure.png" pageId="13" pageNumber="514">Figs 69–73</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFC8E658FDBEFB5D8818FB1F" author="Sartori, M. & Kubiak, M. & Michalik, P." box="[589,801,1231,1257]" pageId="13" pageNumber="514" pagination="28 - 32" refId="ref10026" refString="Sartori, M., Kubiak, M. & Michalik, P. (2016) Deciphering genital anatomy of rare, delicate and precious specimens: first study of two type specimens of mayflies using micro-computed X-ray tomography (Ephemeroptera; Heptageniidae). Zoosymposia, 11, 28 - 32. https: // doi. org / 10.11646 / zoosymposia. 11.1.7" type="journal article" year="2016">
|
||
Sartori
|
||
<emphasis id="B939EA83FFC8E658FD51FB5D89ECFB1F" box="[674,725,1231,1257]" italics="true" pageId="13" pageNumber="514">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
: fig. 2); latero-dorsal and ventral (discal) spines absent. Pair of median titillators present or absent; if present, with single apical point (
|
||
<figureCitation id="13762A14FFC8E658FC0BFB618F72FAFB" box="[1016,1099,1267,1293]" captionStart="FIGURES 73–76" captionStartId="10.[151,264,1352,1376]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-37@10.[151,1436,181,1328]" captionTargetPageId="10" captionText="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi="http://doi.org/10.5281/zenodo.8203266" httpUri="https://zenodo.org/record/8203266/files/figure.png" pageId="13" pageNumber="514">Fig. 73</figureCitation>
|
||
). Cerci ochre, in proximal part with brown articulations (
|
||
<figureCitation id="13762A14FFC8E658FE4BFA858934FAC7" box="[440,525,1303,1329]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="13" pageNumber="514">Fig. 68</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FAA98878FAA3" blockId="13.[151,1437,151,2013]" box="[199,833,1338,1365]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34FAA98A57FAA3" bold="true" box="[199,366,1338,1365]" italics="true" pageId="13" pageNumber="514">Imago, female</emphasis>
|
||
. Coloration similar to male (
|
||
<figureCitation id="13762A14FFC8E658FD42FAA9880DFAA3" box="[689,820,1339,1365]" captionStart="FIGURES 77–84" captionStartId="14.[151,264,1532,1556]" captionTargetBox="[151,1435,309,1509]" captionTargetId="figure-91@14.[151,1435,309,1509]" captionTargetPageId="14" captionText="FIGURES 77–84. Ecdyonurus (Rhithrogeniella) ornatus, female imagines. 77–78, specimen from Agumbe (India); 79–81, specimens from Kwai-Yai (Thailand); 82–84, specimens from Pai (Thailand)." figureDoi="http://doi.org/10.5281/zenodo.8203268" httpUri="https://zenodo.org/record/8203268/files/figure.png" pageId="13" pageNumber="514">Figs 77–84</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34FACD8875F9FF" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34FACD8BC4FA8F" bold="true" box="[199,253,1375,1401]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34FACD8BCFFA8F" bold="true" box="[199,246,1375,1401]" italics="true" pageId="13" pageNumber="514">Egg</emphasis>
|
||
.
|
||
</emphasis>
|
||
Chorion with net-like relief bearing knob-terminated coiled thread (KTC) in each cell. Among individual females, ridges forming cells vary from integral (
|
||
<figureCitation id="13762A14FFC8E658FD4FFA118805FA6B" box="[700,828,1410,1437]" captionStart-0="FIGURES 85–90" captionStart-1="FIGURES 91–96" captionStartId-0="16.[151,264,1571,1595]" captionStartId-1="17.[151,264,1968,1992]" captionTargetBox-0="[164,1423,181,1546]" captionTargetBox-1="[177,1411,381,1945]" captionTargetId-0="figure-37@16.[164,1423,181,1546]" captionTargetId-1="figure-129@17.[177,1411,381,1945]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 85–90. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal." captionText-1="FIGURES 91–96. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from Pai (Thailand). 91–92, reared imago, specimen [XIII](4)2015; 93–94, reared imago, specimen [XIX](1)2015; 95–96, reared subimago, specimen [XVIII](4)2015." figureDoi-0="http://doi.org/10.5281/zenodo.8203270" figureDoi-1="http://doi.org/10.5281/zenodo.8203272" httpUri-0="https://zenodo.org/record/8203270/files/figure.png" httpUri-1="https://zenodo.org/record/8203272/files/figure.png" pageId="13" pageNumber="514">Figs 85–92</figureCitation>
|
||
) to interrupted (
|
||
<figureCitation id="13762A14FFC8E658FC02FA118FEAFA6B" box="[1009,1235,1411,1437]" captionStart-0="FIGURES 91–96" captionStart-1="FIGURES 97–102" captionStartId-0="17.[151,264,1968,1992]" captionStartId-1="18.[151,264,1834,1858]" captionTargetBox-0="[177,1411,381,1945]" captionTargetBox-1="[151,1436,181,1810]" captionTargetId-0="figure-129@17.[177,1411,381,1945]" captionTargetId-1="figure-37@18.[151,1436,181,1810]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 91–96. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from Pai (Thailand). 91–92, reared imago, specimen [XIII](4)2015; 93–94, reared imago, specimen [XIX](1)2015; 95–96, reared subimago, specimen [XVIII](4)2015." captionText-1="FIGURES 97–102. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs of reared imagines from Kwai-Yai (Thailand). 97–98, specimen [III](11)2015; 99–100, specimen [III](37)A2015; 101–102, specimen [IV](2)2015." figureDoi-0="http://doi.org/10.5281/zenodo.8203272" figureDoi-1="http://doi.org/10.5281/zenodo.8203274" httpUri-0="https://zenodo.org/record/8203272/files/figure.png" httpUri-1="https://zenodo.org/record/8203274/files/figure.png" pageId="13" pageNumber="514">Figs 96–97, 99–100</figureCitation>
|
||
) or irregular (
|
||
<figureCitation id="13762A14FFC8E658FA9CFA118A33FA37" captionStart-0="FIGURES 91–96" captionStart-1="FIGURES 97–102" captionStartId-0="17.[151,264,1968,1992]" captionStartId-1="18.[151,264,1834,1858]" captionTargetBox-0="[177,1411,381,1945]" captionTargetBox-1="[151,1436,181,1810]" captionTargetId-0="figure-129@17.[177,1411,381,1945]" captionTargetId-1="figure-37@18.[151,1436,181,1810]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 91–96. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from Pai (Thailand). 91–92, reared imago, specimen [XIII](4)2015; 93–94, reared imago, specimen [XIX](1)2015; 95–96, reared subimago, specimen [XVIII](4)2015." captionText-1="FIGURES 97–102. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs of reared imagines from Kwai-Yai (Thailand). 97–98, specimen [III](11)2015; 99–100, specimen [III](37)A2015; 101–102, specimen [IV](2)2015." figureDoi-0="http://doi.org/10.5281/zenodo.8203272" figureDoi-1="http://doi.org/10.5281/zenodo.8203274" httpUri-0="https://zenodo.org/record/8203272/files/figure.png" httpUri-1="https://zenodo.org/record/8203274/files/figure.png" pageId="13" pageNumber="514">Fig. 93, 95. 98</figureCitation>
|
||
); on eggs extracted from
|
||
<specimenCount id="9D4BFD18FFC8E658FDD7FA3589A6FA37" box="[548,671,1447,1473]" pageId="13" pageNumber="514" type="female">one female</specimenCount>
|
||
completely absent (
|
||
<figureCitation id="13762A14FFC8E658FC8DFA358F25FA37" box="[894,1052,1447,1473]" captionStart="FIGURES 97–102" captionStartId="18.[151,264,1834,1858]" captionTargetBox="[151,1436,181,1810]" captionTargetId="figure-37@18.[151,1436,181,1810]" captionTargetPageId="18" captionText="FIGURES 97–102. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs of reared imagines from Kwai-Yai (Thailand). 97–98, specimen [III](11)2015; 99–100, specimen [III](37)A2015; 101–102, specimen [IV](2)2015." figureDoi="http://doi.org/10.5281/zenodo.8203274" httpUri="https://zenodo.org/record/8203274/files/figure.png" pageId="13" pageNumber="514">Figs 101–102</figureCitation>
|
||
). Among individual females, KTC varying from small (
|
||
<figureCitation id="13762A14FFC8E658FE8FFA598AC5FA13" box="[380,508,1482,1509]" captionStart="FIGURES 85–90" captionStartId="16.[151,264,1571,1595]" captionTargetBox="[164,1423,181,1546]" captionTargetId="figure-37@16.[164,1423,181,1546]" captionTargetPageId="16" captionText="FIGURES 85–90. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal." figureDoi="http://doi.org/10.5281/zenodo.8203270" httpUri="https://zenodo.org/record/8203270/files/figure.png" pageId="13" pageNumber="514">Figs 85–87</figureCitation>
|
||
) to large (
|
||
<figureCitation id="13762A14FFC8E658FD9DFA5989D3FA13" box="[622,746,1483,1509]" captionStart="FIGURES 91–96" captionStartId="17.[151,264,1968,1992]" captionTargetBox="[177,1411,381,1945]" captionTargetId="figure-129@17.[177,1411,381,1945]" captionTargetPageId="17" captionText="FIGURES 91–96. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from Pai (Thailand). 91–92, reared imago, specimen [XIII](4)2015; 93–94, reared imago, specimen [XIX](1)2015; 95–96, reared subimago, specimen [XVIII](4)2015." figureDoi="http://doi.org/10.5281/zenodo.8203272" httpUri="https://zenodo.org/record/8203272/files/figure.png" pageId="13" pageNumber="514">Fig. 91–94</figureCitation>
|
||
) or large on one pole and small on another pole (
|
||
<figureCitation id="13762A14FFC8E658FAFEFA598EA8FA13" box="[1293,1425,1483,1509]" captionStart="FIGURES 85–90" captionStartId="16.[151,264,1571,1595]" captionTargetBox="[164,1423,181,1546]" captionTargetId="figure-37@16.[164,1423,181,1546]" captionTargetPageId="16" captionText="FIGURES 85–90. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal." figureDoi="http://doi.org/10.5281/zenodo.8203270" httpUri="https://zenodo.org/record/8203270/files/figure.png" pageId="13" pageNumber="514">Figs 89–90</figureCitation>
|
||
). Micropyle varying from small (
|
||
<figureCitation id="13762A14FFC8E658FE0BFA7D8941F9FF" box="[504,632,1518,1545]" captionStart="FIGURES 85–90" captionStartId="16.[151,264,1571,1595]" captionTargetBox="[164,1423,181,1546]" captionTargetId="figure-37@16.[164,1423,181,1546]" captionTargetPageId="16" captionText="FIGURES 85–90. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal." figureDoi="http://doi.org/10.5281/zenodo.8203270" httpUri="https://zenodo.org/record/8203270/files/figure.png" pageId="13" pageNumber="514">Figs 85–87</figureCitation>
|
||
) to large (
|
||
<figureCitation id="13762A14FFC8E658FD18FA7D8806F9FF" box="[747,831,1519,1545]" captionStart="FIGURES 85–90" captionStartId="16.[151,264,1571,1595]" captionTargetBox="[164,1423,181,1546]" captionTargetId="figure-37@16.[164,1423,181,1546]" captionTargetPageId="16" captionText="FIGURES 85–90. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal." figureDoi="http://doi.org/10.5281/zenodo.8203270" httpUri="https://zenodo.org/record/8203270/files/figure.png" pageId="13" pageNumber="514">Fig. 90</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34F9808F00F9DB" blockId="13.[151,1437,151,2013]" box="[199,1081,1554,1581]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34F9808A77F9DA" bold="true" box="[199,334,1554,1580]" pageId="13" pageNumber="514">Dimension.</emphasis>
|
||
Fore wing length
|
||
<quantity id="4CB59B74FFC8E658FDE8F9808941F9DB" box="[539,632,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="13" pageNumber="514" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
|
||
(up to
|
||
<quantity id="4CB59B74FFC8E658FD3BF9818830F9DB" box="[712,777,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="514" unit="mm" value="7.0">7 mm</quantity>
|
||
according to
|
||
<bibRefCitation id="EFDC4B60FFC8E658FC51F9818F15F9DB" author="Ulmer, G." box="[930,1068,1555,1581]" pageId="13" pageNumber="514" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C357651AFFC8E658FF34F9A48FC7F9A7" box="[199,1278,1590,1617]" pageId="13" pageNumber="514" type="distribution">
|
||
<paragraph id="8BF23691FFC8E658FF34F9A48FC7F9A7" blockId="13.[151,1437,151,2013]" box="[199,1278,1590,1617]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34F9A48A58F9A6" bold="true" box="[199,353,1590,1616]" pageId="13" pageNumber="514">Distribution.</emphasis>
|
||
Oriental Region; known from Southern
|
||
<collectingCountry id="F35A7601FFC8E658FCD4F9A5885AF9A7" box="[807,867,1591,1617]" name="India" pageId="13" pageNumber="514">India</collectingCountry>
|
||
, Indochina and Great Sunda Islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C357651AFFC8E641FF34F9C8899FFEB7" lastPageId="20" lastPageNumber="521" pageId="13" pageNumber="514" type="discussion">
|
||
<paragraph id="8BF23691FFC8E658FF34F9C88AEEF8BB" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34F9C88952F983" bold="true" box="[199,619,1626,1653]" pageId="13" pageNumber="514">Confusion about labrum structure.</emphasis>
|
||
<bibRefCitation id="EFDC4B60FFC8E658FD80F9C9889CF983" author="Soldan, T. & Braasch, D." box="[627,933,1627,1653]" pageId="13" pageNumber="514" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán and Braasch (1986)</bibRefCitation>
|
||
overlooked median emargination of labrum, and
|
||
<bibRefCitation id="EFDC4B60FFC8E658FF35F9ED8A5AF96F" author="Sartori, M." box="[198,355,1663,1689]" pageId="13" pageNumber="514" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori (2014)</bibRefCitation>
|
||
erroneously reported that labrum has ́no anteromedian emargination ... ventral face with shorter and stout setae along the anterior margin». On the photo (
|
||
<bibRefCitation id="EFDC4B60FFC8E658FCEBF9318890F94B" author="Sartori, M." box="[792,937,1699,1725]" pageId="13" pageNumber="514" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: fig. 8), focus is given on the dorsal side, but if enlarge the picture, it is possible to see the stout pointed setae bordering the initial anterior margin, which is bent ventrally; medially this paired setal row is interrupted by the median emargination of the initial anterior margin, but this emargination is invisible on the photo; shape of this labrum and its visible setal rows are exactly the same as in our specimens (
|
||
<figureCitation id="13762A14FFC8E658FEB4F8A18AF3F8BB" box="[327,458,1843,1869]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="13" pageNumber="514">Figs 26–27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E658FF34F8C489C5F84F" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="514">
|
||
<emphasis id="B939EA83FFC8E658FF34F8C48804F886" bold="true" box="[199,829,1878,1905]" pageId="13" pageNumber="514">Disagreements about presence / absence of titillators.</emphasis>
|
||
Originally, male imago of
|
||
<taxonomicName id="4C4D4D12FFC8E658FB99F8C58E53F886" authorityName="Ulmer" authorityYear="1939" box="[1130,1386,1879,1905]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFC8E658FB99F8C58E53F886" box="[1130,1386,1879,1905]" italics="true" pageId="13" pageNumber="514">Rhithrogeniella ornata</emphasis>
|
||
</taxonomicName>
|
||
was described as lacking titillators (
|
||
<bibRefCitation id="EFDC4B60FFC8E658FE01F8E9898BF863" author="Ulmer, G." box="[498,690,1914,1941]" pageId="13" pageNumber="514" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939: 575</bibRefCitation>
|
||
, fig. 169); male subimagines of this species were reported, but their genital structure was not described (
|
||
<bibRefCitation id="EFDC4B60FFC8E658FDDFF80D89C9F84F" author="Ulmer, G." box="[556,752,1950,1977]" pageId="13" pageNumber="514" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939: 577</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC8E65BFF34F8518BE4FEEB" blockId="13.[151,1437,151,2013]" lastBlockId="14.[151,1436,151,285]" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514">
|
||
<bibRefCitation id="EFDC4B60FFC8E658FF34F8518938F82B" author="Soldan, T. & Braasch, D." box="[199,513,1987,2013]" pageId="13" pageNumber="514" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán and Braasch (1986)</bibRefCitation>
|
||
described subimagines placed by then in a new species of
|
||
<taxonomicName id="4C4D4D12FFC8E658FB45F8518E5CF82B" authorityName="Ulmer" authorityYear="1939" box="[1206,1381,1987,2013]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B939EA83FFC8E658FB45F8518E5CF82B" box="[1206,1381,1987,2013]" italics="true" pageId="13" pageNumber="514">Rhithrogeniella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D12FFC8E65BFA85F8518A2CFF47" authorityName=": Kluge" authorityYear="2022" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" lastPageId="14" lastPageNumber="515" order="Ephemeroptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="B939EA83FFC8E65BFA85F8518A2CFF47" italics="true" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514">Rh. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
, which have well developed medial titillators with sclerotized apices. In the same paper, they reported that ́similar structures occur in subimagoes of
|
||
<taxonomicName id="4C4D4D12FFCBE65BFD33FF298814FF22" authorityName="Ulmer" authorityYear="1939" box="[704,813,187,212]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFCBE65BFD33FF298814FF22" box="[704,813,187,212]" italics="true" pageId="14" pageNumber="515">R. ornata</emphasis>
|
||
</taxonomicName>
|
||
(material in the Zoological Museum,
|
||
<collectingRegion id="4989F873FFCBE65BFB1AFF298E6FFF23" box="[1257,1366,187,213]" country="Germany" name="Hamburg" pageId="14" pageNumber="515">Hamburg</collectingRegion>
|
||
)» and wrote: ́Our opinion agrees with that of Jensen (pers. comm.) who supposes the existence of medial titillators as well».
|
||
</paragraph>
|
||
<caption id="DF326619FFCBE65BFF64FA6E8892F9CE" ID-DOI="http://doi.org/10.5281/zenodo.8203268" ID-Zenodo-Dep="8203268" httpUri="https://zenodo.org/record/8203268/files/figure.png" pageId="14" pageNumber="515" startId="14.[151,264,1532,1556]" targetBox="[151,1435,309,1509]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BF23691FFCBE65BFF64FA6E8892F9CE" blockId="14.[151,1436,1532,1593]" pageId="14" pageNumber="515">
|
||
<emphasis id="B939EA83FFCBE65BFF64FA6E8A6EF9E2" bold="true" box="[151,343,1532,1556]" pageId="14" pageNumber="515">FIGURES 77–84.</emphasis>
|
||
<taxonomicName id="4C4D4D12FFCBE65BFE93FA6E89DEF9E2" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[352,743,1532,1557]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFCBE65BFE93FA6E8AE0F9E2" box="[352,473,1532,1556]" italics="true" pageId="14" pageNumber="515">Ecdyonurus</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFCBE65BFE19FA6E89B1F9E2" box="[490,648,1532,1556]" italics="true" pageId="14" pageNumber="515">Rhithrogeniella</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFCBE65BFD69FA6F89DEF9E2" box="[666,743,1533,1556]" italics="true" pageId="14" pageNumber="515">ornatus</emphasis>
|
||
</taxonomicName>
|
||
, female imagines. 77–78, specimen from Agumbe (India); 79–81, specimens from Kwai-Yai (Thailand); 82–84, specimens from Pai (Thailand).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23691FFCBE65BFF34F9F58A33F8C3" blockId="14.[151,1437,1639,2025]" pageId="14" pageNumber="515">
|
||
<bibRefCitation id="EFDC4B60FFCBE65BFF34F9F58A51F977" author="Sartori, M." box="[199,360,1639,1665]" pageId="14" pageNumber="515" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori (2014)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFDC4B60FFCBE65BFE50F9F589B1F977" author="Sartori, M. & Kubiak, M. & Michalik, P." box="[419,648,1639,1665]" pageId="14" pageNumber="515" pagination="28 - 32" refId="ref10026" refString="Sartori, M., Kubiak, M. & Michalik, P. (2016) Deciphering genital anatomy of rare, delicate and precious specimens: first study of two type specimens of mayflies using micro-computed X-ray tomography (Ephemeroptera; Heptageniidae). Zoosymposia, 11, 28 - 32. https: // doi. org / 10.11646 / zoosymposia. 11.1.7" type="journal article" year="2016">
|
||
Sartori
|
||
<emphasis id="B939EA83FFCBE65BFE0AF9F58915F977" box="[505,556,1639,1665]" italics="true" pageId="14" pageNumber="515">et al</emphasis>
|
||
. (2016)
|
||
</bibRefCitation>
|
||
reexamined the single male imago and the male subimagines which belong to the
|
||
<typeStatus id="54F68833FFCBE65BFEC2F9198A58F953" box="[305,353,1675,1701]" pageId="14" pageNumber="515">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C4D4D12FFCBE65BFE3DF91989F6F952" authorityName="Ulmer" authorityYear="1939" box="[462,719,1675,1701]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFCBE65BFE3DF91989F6F952" box="[462,719,1675,1701]" italics="true" pageId="14" pageNumber="515">Rhithrogeniella ornata</emphasis>
|
||
</taxonomicName>
|
||
and found out that the male imago has no titillators, and a male subimago has titillators which exist on subimaginal cuticle only, but are absent on imaginal penis developing under the subimaginal cuticle (
|
||
<bibRefCitation id="EFDC4B60FFCBE65BFE41F941897FF91B" author="Sartori, M." box="[434,582,1747,1773]" pageId="14" pageNumber="515" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: fig. 4). Based on this fact, they concluded that
|
||
<taxonomicName id="4C4D4D12FFCBE65BFB8BF9418E1EF91B" authorityName="Ulmer" authorityYear="1939" box="[1144,1319,1747,1773]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B939EA83FFCBE65BFB8BF9418E1EF91B" box="[1144,1319,1747,1773]" italics="true" pageId="14" pageNumber="515">Rhithrogeniella</emphasis>
|
||
</taxonomicName>
|
||
(both
|
||
<taxonomicName id="4C4D4D12FFCBE65BFA85F9418BDBF8E6" authorityName="Ulmer" authorityYear="1939" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFCBE65BFA85F9418BDBF8E6" italics="true" pageId="14" pageNumber="515">Rh. ornata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D12FFCBE65BFEE9F9658AFFF8E7" authorityName=": Kluge" authorityYear="2022" box="[282,454,1783,1809]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="B939EA83FFCBE65BFEE9F9658AFFF8E7" box="[282,454,1783,1809]" italics="true" pageId="14" pageNumber="515">Rh. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
) have no titillators in imaginal stage, but have them in subimaginal stage only (
|
||
<bibRefCitation id="EFDC4B60FFCBE65BFABDF9658BC7F8C3" author="Sartori, M." pageId="14" pageNumber="515" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014: 59</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFCBE65AFF34F8AD8AF3FF23" blockId="14.[151,1437,1639,2025]" lastBlockId="15.[151,1436,151,213]" lastPageId="15" lastPageNumber="516" pageId="14" pageNumber="515">
|
||
<location id="8E92604AFFCBE65BFF34F8AD8BCAF8AF" LSID="urn:lsid:plazi:treatment:03E48787FFCEE640FF64F96B8FDCFD3B:8E92604AFFCBE65BFF34F8AD8BCAF8AF" box="[199,243,1855,1881]" country="India" municipality="All" name="Our" pageId="14" pageNumber="515">Our</location>
|
||
recent examination of the specimens from
|
||
<collectingCountry id="F35A7601FFCBE65BFD27F8AD8837F8AF" box="[724,782,1855,1881]" name="India" pageId="14" pageNumber="515">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F35A7601FFCBE65BFCB0F8AD889EF8AF" box="[835,935,1855,1881]" name="Thailand" pageId="14" pageNumber="515">Thailand</collectingCountry>
|
||
reveals that presence/absence of titillators in imaginal stage varies individually (
|
||
<tableCitation id="C6CF032AFFCBE65BFDD1F8F1894AF88B" box="[546,627,1891,1917]" captionStart="TABLE 1" captionStartId="15.[152,244,260,284]" captionTargetPageId="15" captionText="TABLE 1. Presence/absence of imaginal titillators in the specimens examined" httpUri="http://table.plazi.org/id/DF326619FFCAE65AFF6BFE9688E6FEEA" pageId="14" pageNumber="515" tableUuid="DF326619FFCAE65AFF6BFE9688E6FEEA">Table 1</tableCitation>
|
||
).
|
||
<collectingMunicipality id="6B96ACEBFFCBE65BFD79F8F18997F88B" box="[650,686,1891,1917]" pageId="14" pageNumber="515">All</collectingMunicipality>
|
||
6 examined males from
|
||
<collectingCountry id="F35A7601FFCBE65BFC30F8F188C4F88B" box="[963,1021,1891,1917]" name="India" pageId="14" pageNumber="515">India</collectingCountry>
|
||
(two imagines, two subimagines and
|
||
<specimenCount id="9D4BFD18FFCBE65BFF64F8158A34F857" box="[151,269,1927,1953]" pageId="14" pageNumber="515" type="larva">two larvae</specimenCount>
|
||
ready to molt to subimago) and one examined male subimago from Kwai-Yai in
|
||
<collectingCountry id="F35A7601FFCBE65BFB5EF8158E2AF857" box="[1197,1299,1927,1953]" name="Thailand" pageId="14" pageNumber="515">Thailand</collectingCountry>
|
||
, have either imaginal titillators (
|
||
<figureCitation id="13762A14FFCBE65BFE82F8398ACAF833" box="[369,499,1963,1989]" captionStart="FIGURES 69–72" captionStartId="9.[151,264,1837,1861]" captionTargetBox="[151,1436,182,1813]" captionTargetId="figure-20@9.[151,1436,181,1813]" captionTargetPageId="9" captionText="FIGURES 69–72. Ecdyonurus (Rhithrogeniella) ornatus, genitalia of male imagines, ventral view. 69–70, specimen from Kodaikanal (India) in tense condition; 71, specimen from Agumbe (India); 72, specimen from Pai (Thailand)." figureDoi="http://doi.org/10.5281/zenodo.8203264" httpUri="https://zenodo.org/record/8203264/files/figure.png" pageId="14" pageNumber="515">Figs 69–71</figureCitation>
|
||
), or tissues of future imaginal titillators located under cuticle of subimaginal titillator (
|
||
<figureCitation id="13762A14FFCBE65BFF6CF85D8BCAF81F" box="[159,243,1998,2025]" captionStart="FIGURES 73–76" captionStartId="10.[151,264,1352,1376]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-37@10.[151,1436,181,1328]" captionTargetPageId="10" captionText="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi="http://doi.org/10.5281/zenodo.8203266" httpUri="https://zenodo.org/record/8203266/files/figure.png" pageId="14" pageNumber="515">Fig. 75</figureCitation>
|
||
). Among specimens from Pai (
|
||
<collectingCountry id="F35A7601FFCBE65BFDB5F85D8997F81F" box="[582,686,1999,2025]" name="Thailand" pageId="14" pageNumber="515">Thailand</collectingCountry>
|
||
), 4 reared male imagines also have imaginal titillators (
|
||
<figureCitation id="13762A14FFCBE65BFAFDF85D8EA9F81F" box="[1294,1424,1999,2025]" captionStart-0="FIGURES 69–72" captionStart-1="FIGURES 73–76" captionStartId-0="9.[151,264,1837,1861]" captionStartId-1="10.[151,264,1352,1376]" captionTargetBox-0="[151,1436,182,1813]" captionTargetBox-1="[151,1436,181,1328]" captionTargetId-0="figure-20@9.[151,1436,181,1813]" captionTargetId-1="figure-37@10.[151,1436,181,1328]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 69–72. Ecdyonurus (Rhithrogeniella) ornatus, genitalia of male imagines, ventral view. 69–70, specimen from Kodaikanal (India) in tense condition; 71, specimen from Agumbe (India); 72, specimen from Pai (Thailand)." captionText-1="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi-0="http://doi.org/10.5281/zenodo.8203264" figureDoi-1="http://doi.org/10.5281/zenodo.8203266" httpUri-0="https://zenodo.org/record/8203264/files/figure.png" httpUri-1="https://zenodo.org/record/8203266/files/figure.png" pageId="14" pageNumber="515">Figs 72–73</figureCitation>
|
||
), but
|
||
<specimenCount id="9D4BFD18FFCAE65AFF37FF058A17FF47" box="[196,302,151,177]" pageId="15" pageNumber="516" type="larva">one larva</specimenCount>
|
||
ready to moult to subimago, has only subimaginal cuticle of titillators, without tissues of imaginal titillators under it (
|
||
<figureCitation id="13762A14FFCAE65AFE9AFF298A84FF23" box="[361,445,187,213]" captionStart="FIGURES 73–76" captionStartId="10.[151,264,1352,1376]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-37@10.[151,1436,181,1328]" captionTargetPageId="10" captionText="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi="http://doi.org/10.5281/zenodo.8203266" httpUri="https://zenodo.org/record/8203266/files/figure.png" pageId="15" pageNumber="516">Fig. 76</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF326619FFCAE65AFF6BFE9688E6FEEA" ID-Table-UUID="DF326619FFCAE65AFF6BFE9688E6FEEA" box="[152,991,260,284]" httpUri="http://table.plazi.org/id/DF326619FFCAE65AFF6BFE9688E6FEEA" pageId="15" pageNumber="516" startId="15.[152,244,260,284]" targetBox="[160,1383,298,783]" targetIsTable="true" targetPageId="15" targetType="table">
|
||
<paragraph id="8BF23691FFCAE65AFF6BFE9688E6FEEA" blockId="15.[152,991,260,284]" box="[152,991,260,284]" pageId="15" pageNumber="516">
|
||
<emphasis id="B939EA83FFCAE65AFF6BFE968A36FEEA" bold="true" box="[152,271,260,284]" pageId="15" pageNumber="516">TABLE 1.</emphasis>
|
||
Presence/absence of imaginal titillators in the specimens examined
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23691FFCAE65AFF53FEB88E6DFCFA" pageId="15" pageNumber="516">
|
||
<table id="F94DC431FFCA19AAFF53FEB88E5EFCF9" box="[160,1383,298,783]" gridcols="6" gridrows="13" pageId="15" pageNumber="516">
|
||
<tr id="357D34D3FFCA19AAFF53FEB88E5EFE96" box="[160,1383,298,352]" gridrow="0" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FEB88A50FE96" box="[160,361,298,352]" gridcol="0" gridrow="0" pageId="15" pageNumber="516">Locality</th>
|
||
<th id="76AC5DAFFFCA19AAFE6AFEB8896FFE96" box="[409,598,298,352]" gridcol="1" gridrow="0" pageId="15" pageNumber="516">Collection number</th>
|
||
<th id="76AC5DAFFFCA19AAFD8BFEB889C2FE96" box="[632,763,298,352]" gridcol="2" gridrow="0" pageId="15" pageNumber="516">Stage of development</th>
|
||
<th id="76AC5DAFFFCA19AAFCA0FEB888EAFE96" box="[851,979,298,352]" gridcol="3" gridrow="0" pageId="15" pageNumber="516">Subimaginal titillators</th>
|
||
<th id="76AC5DAFFFCA19AAFBC3FEB88FD5FE96" box="[1072,1260,298,352]" gridcol="4" gridrow="0" pageId="15" pageNumber="516">Imaginal titillators</th>
|
||
<th id="76AC5DAFFFCA19AAFADBFEB88E5EFE96" box="[1320,1383,298,352]" gridcol="5" gridrow="0" pageId="15" pageNumber="516">Fig.</th>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FEF98E5EFE70" box="[160,1383,363,390]" gridrow="1" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FEF98A50FE70" box="[160,361,363,390]" gridcol="0" gridrow="1" pageId="15" pageNumber="516">India: Agumbe</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFEF9896FFE70" box="[409,598,363,390]" gridcol="1" gridrow="1" pageId="15" pageNumber="516">[XII](1)2013</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFEF989C2FE70" box="[632,763,363,390]" gridcol="2" gridrow="1" pageId="15" pageNumber="516">L-S♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FEF988EAFE70" box="[851,979,363,390]" gridcol="3" gridrow="1" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FEF98FD5FE70" box="[1072,1260,363,390]" gridcol="4" gridrow="1" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFEF98E5EFE70" box="[1320,1383,363,390]" gridcol="5" gridrow="1" pageId="15" pageNumber="516">—</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FE1D8E5EFE5C" box="[160,1383,399,426]" gridrow="2" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FE1D8A50FE5C" box="[160,361,399,426]" gridcol="0" gridrow="2" pageId="15" pageNumber="516">India: Agumbe</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFE1D896FFE5C" box="[409,598,399,426]" gridcol="1" gridrow="2" pageId="15" pageNumber="516">[XV](6)2013</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFE1D89C2FE5C" box="[632,763,399,426]" gridcol="2" gridrow="2" pageId="15" pageNumber="516">L-S-I♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FE1D88EAFE5C" box="[851,979,399,426]" gridcol="3" gridrow="2" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FE1D8FD5FE5C" box="[1072,1260,399,426]" gridcol="4" gridrow="2" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFE1D8E5EFE5C" box="[1320,1383,399,426]" gridcol="5" gridrow="2" pageId="15" pageNumber="516">71</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FE218E5EFE3B" box="[160,1383,435,461]" gridrow="3" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FE218A50FE3B" box="[160,361,435,461]" gridcol="0" gridrow="3" pageId="15" pageNumber="516">India: Agumbe</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFE21896FFE3B" box="[409,598,435,461]" gridcol="1" gridrow="3" pageId="15" pageNumber="516">slide 21.II.2023-1</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFE2189C2FE3B" box="[632,763,435,461]" gridcol="2" gridrow="3" pageId="15" pageNumber="516">S♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FE2188EAFE3B" box="[851,979,435,461]" gridcol="3" gridrow="3" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FE218FD5FE3B" box="[1072,1260,435,461]" gridcol="4" gridrow="3" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFE218E5EFE3B" box="[1320,1383,435,461]" gridcol="5" gridrow="3" pageId="15" pageNumber="516">—</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FE448E5EFE07" box="[160,1383,470,497]" gridrow="4" pageId="15" pageNumber="516" rowspan-1="1">
|
||
<th id="76AC5DAFFFCA19AAFF53FE448A50FE07" box="[160,361,470,497]" gridcol="0" gridrow="4" pageId="15" pageNumber="516">India: Kodaikanal</th>
|
||
<td id="76AC5DAFFFCA19AAFD8BFE4489C2FE07" box="[632,763,470,497]" gridcol="2" gridrow="4" pageId="15" pageNumber="516">L-S-I♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FE4488EAFE07" box="[851,979,470,497]" gridcol="3" gridrow="4" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FE448FD5FE07" box="[1072,1260,470,497]" gridcol="4" gridrow="4" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFE448E5EFE07" box="[1320,1383,470,497]" gridcol="5" gridrow="4" pageId="15" pageNumber="516">69–70</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FE688E5EFDE3" box="[160,1383,506,533]" gridrow="5" pageId="15" pageNumber="516" rowspan-1="1">
|
||
<th id="76AC5DAFFFCA19AAFF53FE688A50FDE3" box="[160,361,506,533]" gridcol="0" gridrow="5" pageId="15" pageNumber="516">India: Kodaikanal</th>
|
||
<td id="76AC5DAFFFCA19AAFD8BFE6889C2FDE3" box="[632,763,506,533]" gridcol="2" gridrow="5" pageId="15" pageNumber="516">L/S♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FE6888EAFDE3" box="[851,979,506,533]" gridcol="3" gridrow="5" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FE688FD5FDE3" box="[1072,1260,506,533]" gridcol="4" gridrow="5" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFE688E5EFDE3" box="[1320,1383,506,533]" gridcol="5" gridrow="5" pageId="15" pageNumber="516">75</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FD8C8E5EFDCE" box="[160,1383,542,568]" gridrow="6" pageId="15" pageNumber="516" rowspan-1="1">
|
||
<th id="76AC5DAFFFCA19AAFF53FD8C8A50FDCE" box="[160,361,542,568]" gridcol="0" gridrow="6" pageId="15" pageNumber="516">India: Kodaikanal</th>
|
||
<td id="76AC5DAFFFCA19AAFD8BFD8C89C2FDCE" box="[632,763,542,568]" gridcol="2" gridrow="6" pageId="15" pageNumber="516">L/S♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FD8C88EAFDCE" box="[851,979,542,568]" gridcol="3" gridrow="6" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FD8C8FD5FDCE" box="[1072,1260,542,568]" gridcol="4" gridrow="6" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFD8C8E5EFDCE" box="[1320,1383,542,568]" gridcol="5" gridrow="6" pageId="15" pageNumber="516">—</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FDD38E5EFDAA" box="[160,1383,577,604]" gridrow="7" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FDD38A50FDAA" box="[160,361,577,604]" gridcol="0" gridrow="7" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFDD3896FFDAA" box="[409,598,577,604]" gridcol="1" gridrow="7" pageId="15" pageNumber="516">[XV](A)2025</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFDD389C2FDAA" box="[632,763,577,604]" gridcol="2" gridrow="7" pageId="15" pageNumber="516">S-I♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FDD388EAFDAA" box="[851,979,577,604]" gridcol="3" gridrow="7" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FDD38FD5FDAA" box="[1072,1260,577,604]" gridcol="4" gridrow="7" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFDD38E5EFDAA" box="[1320,1383,577,604]" gridcol="5" gridrow="7" pageId="15" pageNumber="516">—</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FDF78E5EFD76" box="[160,1383,613,640]" gridrow="8" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FDF78A50FD76" box="[160,361,613,640]" gridcol="0" gridrow="8" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFDF7896FFD76" box="[409,598,613,640]" gridcol="1" gridrow="8" pageId="15" pageNumber="516">[XVI](1)A2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFDF789C2FD76" box="[632,763,613,640]" gridcol="2" gridrow="8" pageId="15" pageNumber="516">L-S/I♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FDF788EAFD76" box="[851,979,613,640]" gridcol="3" gridrow="8" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FDF78FD5FD76" box="[1072,1260,613,640]" gridcol="4" gridrow="8" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFDF78E5EFD76" box="[1320,1383,613,640]" gridcol="5" gridrow="8" pageId="15" pageNumber="516">—</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FD1B8E5EFD55" box="[160,1383,649,675]" gridrow="9" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FD1B8A50FD55" box="[160,361,649,675]" gridcol="0" gridrow="9" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFD1B896FFD55" box="[409,598,649,675]" gridcol="1" gridrow="9" pageId="15" pageNumber="516">[XVI](2)A2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFD1B89C2FD55" box="[632,763,649,675]" gridcol="2" gridrow="9" pageId="15" pageNumber="516">L-S-I♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FD1B88EAFD55" box="[851,979,649,675]" gridcol="3" gridrow="9" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FD1B8FD5FD55" box="[1072,1260,649,675]" gridcol="4" gridrow="9" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFD1B8E5EFD55" box="[1320,1383,649,675]" gridcol="5" gridrow="9" pageId="15" pageNumber="516">—</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FD3F8E5EFD31" box="[160,1383,685,711]" gridrow="10" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FD3F8A50FD31" box="[160,361,685,711]" gridcol="0" gridrow="10" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFD3F896FFD31" box="[409,598,685,711]" gridcol="1" gridrow="10" pageId="15" pageNumber="516">[XIX](1)</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFD3F89C2FD31" box="[632,763,685,711]" gridcol="2" gridrow="10" pageId="15" pageNumber="516">L-S-I♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FD3F88EAFD31" box="[851,979,685,711]" gridcol="3" gridrow="10" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FD3F8FD5FD31" box="[1072,1260,685,711]" gridcol="4" gridrow="10" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFD3F8E5EFD31" box="[1320,1383,685,711]" gridcol="5" gridrow="10" pageId="15" pageNumber="516">72–74</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FD428E5EFD1D" box="[160,1383,720,747]" gridrow="11" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FD428A50FD1D" box="[160,361,720,747]" gridcol="0" gridrow="11" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFD42896FFD1D" box="[409,598,720,747]" gridcol="1" gridrow="11" pageId="15" pageNumber="516">slide 28.II.2023-1</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFD4289C2FD1D" box="[632,763,720,747]" gridcol="2" gridrow="11" pageId="15" pageNumber="516">L/S♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FD4288EAFD1D" box="[851,979,720,747]" gridcol="3" gridrow="11" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FD428FD5FD1D" box="[1072,1260,720,747]" gridcol="4" gridrow="11" pageId="15" pageNumber="516">–</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFD428E5EFD1D" box="[1320,1383,720,747]" gridcol="5" gridrow="11" pageId="15" pageNumber="516">76</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF53FD668E5EFCF9" box="[160,1383,756,783]" gridrow="12" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF53FD668A50FCF9" box="[160,361,756,783]" gridcol="0" gridrow="12" pageId="15" pageNumber="516">Thailand: Kwai-Yai</th>
|
||
<td id="76AC5DAFFFCA19AAFE6AFD66896FFCF9" box="[409,598,756,783]" gridcol="1" gridrow="12" pageId="15" pageNumber="516">[IV](2)B2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD8BFD6689C2FCF9" box="[632,763,756,783]" gridcol="2" gridrow="12" pageId="15" pageNumber="516">L-S♂</td>
|
||
<td id="76AC5DAFFFCA19AAFCA0FD6688EAFCF9" box="[851,979,756,783]" gridcol="3" gridrow="12" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFBC3FD668FD5FCF9" box="[1072,1260,756,783]" gridcol="4" gridrow="12" pageId="15" pageNumber="516">+</td>
|
||
<td id="76AC5DAFFFCA19AAFADBFD668E5EFCF9" box="[1320,1383,756,783]" gridcol="5" gridrow="12" pageId="15" pageNumber="516">—</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFCAE65AFF34FCAB8F54FC6D" blockId="15.[151,1436,825,924]" pageId="15" pageNumber="516">
|
||
Thus, in all cases subimaginal cuticle bears a pair of titillators in a form of very small, pointed denticles (
|
||
<figureCitation id="13762A14FFCAE65AFA99FCAB8BDBFC81" captionStart="FIGURES 73–76" captionStartId="10.[151,264,1352,1376]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-37@10.[151,1436,181,1328]" captionTargetPageId="10" captionText="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi="http://doi.org/10.5281/zenodo.8203266" httpUri="https://zenodo.org/record/8203266/files/figure.png" pageId="15" pageNumber="516">Figs 74–76</figureCitation>
|
||
); tissues of imaginal penis developing under these pointed denticles, either have a pair of larger imaginal titillators (
|
||
<figureCitation id="13762A14FFCAE65AFEFFFC138A66FC6A" box="[268,351,897,924]" captionStart="FIGURES 73–76" captionStartId="10.[151,264,1352,1376]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-37@10.[151,1436,181,1328]" captionTargetPageId="10" captionText="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi="http://doi.org/10.5281/zenodo.8203266" httpUri="https://zenodo.org/record/8203266/files/figure.png" pageId="15" pageNumber="516">Fig. 75</figureCitation>
|
||
), or lack them (
|
||
<figureCitation id="13762A14FFCAE65AFDFCFC13895AFC6D" box="[527,611,897,923]" captionStart="FIGURES 73–76" captionStartId="10.[151,264,1352,1376]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-37@10.[151,1436,181,1328]" captionTargetPageId="10" captionText="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." figureDoi="http://doi.org/10.5281/zenodo.8203266" httpUri="https://zenodo.org/record/8203266/files/figure.png" pageId="15" pageNumber="516">Fig. 76</figureCitation>
|
||
); in the last case imago will have no titillators.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFCAE65AFF64FC5B891EFBA6" blockId="15.[151,1436,969,1104]" pageId="15" pageNumber="516">
|
||
<emphasis id="B939EA83FFCAE65AFF64FC5B8AC9FC12" bold="true" box="[151,496,969,996]" pageId="15" pageNumber="516">Variability of egg structure.</emphasis>
|
||
Presence of mesh-like reticulate ridges was reported as diagnostic character of the subgenus
|
||
<taxonomicName id="4C4D4D12FFCAE65AFEC5FC7C891BFBFE" authority="(Flowers 1980)" baseAuthorityName="Flowers" baseAuthorityYear="1980" box="[310,546,1005,1032]" pageId="15" pageNumber="516" rank="subGenus" subGenus="Nixe">
|
||
<emphasis id="B939EA83FFCAE65AFEC5FC7C8A50FBF1" box="[310,361,1006,1031]" italics="true" pageId="15" pageNumber="516">Nixe</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDC4B60FFCAE65AFE88FC7F8920FBFE" author="Flowers, R. W." box="[379,537,1005,1032]" pageId="15" pageNumber="516" pagination="296 - 307" refId="ref9473" refString="Flowers, R. W. (1980) Two new genera of Nearctic Heptageniidae (Ephemeroptera). Florida Entomologist, 63 (3), 296 - 307. https: // doi. org / 10.2307 / 3494626" type="journal article" year="1980">Flowers 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and the genus
|
||
<taxonomicName id="4C4D4D12FFCAE65AFD25FC7F8F0AFBFE" authority="(Sartori 2014)" baseAuthorityName="Sartori" baseAuthorityYear="2014" box="[726,1075,1005,1032]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B939EA83FFCAE65AFD25FC7F88BCFBF1" box="[726,901,1005,1031]" italics="true" pageId="15" pageNumber="516">Rhithrogeniella</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDC4B60FFCAE65AFC65FC7F8F12FBFE" author="Sartori, M." box="[918,1067,1005,1032]" pageId="15" pageNumber="516" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Our recent examination of the specimens from
|
||
<collectingCountry id="F35A7601FFCAE65AFEA0FB838AB4FBDD" box="[339,397,1041,1067]" name="India" pageId="15" pageNumber="516">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F35A7601FFCAE65AFE36FB838910FBDD" box="[453,553,1041,1067]" name="Thailand" pageId="15" pageNumber="516">Thailand</collectingCountry>
|
||
reveals that presence/absence of this relief varies individually, at least among specimens from
|
||
<collectingCountry id="F35A7601FFCAE65AFEA3FBA78A8DFBB9" box="[336,436,1077,1103]" name="Thailand" pageId="15" pageNumber="516">Thailand</collectingCountry>
|
||
(
|
||
<tableCitation id="C6CF032AFFCAE65AFE37FBA78920FBA6" box="[452,537,1077,1104]" captionStart="TABLE 2" captionStartId="15.[152,244,1151,1175]" captionTargetPageId="15" captionText="TABLE 2. Relief of egg chorion in the specimens examined" httpUri="http://table.plazi.org/id/DF326619FFCAE65AFF6BFBED8819FB61" pageId="15" pageNumber="516" tableUuid="DF326619FFCAE65AFF6BFBED8819FB61">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF326619FFCAE65AFF6BFBED8819FB61" ID-Table-UUID="DF326619FFCAE65AFF6BFBED8819FB61" box="[152,800,1151,1176]" httpUri="http://table.plazi.org/id/DF326619FFCAE65AFF6BFBED8819FB61" pageId="15" pageNumber="516" startId="15.[152,244,1151,1175]" targetBox="[163,1392,1191,1578]" targetIsTable="true" targetPageId="15" targetType="table">
|
||
<paragraph id="8BF23691FFCAE65AFF6BFBED8819FB61" blockId="15.[152,800,1151,1176]" box="[152,800,1151,1176]" pageId="15" pageNumber="516">
|
||
<emphasis id="B939EA83FFCAE65AFF6BFBED8A36FB61" bold="true" box="[152,271,1151,1175]" pageId="15" pageNumber="516">TABLE 2.</emphasis>
|
||
Relief of egg chorion in the specimens examined
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23691FFCAE65AFF50FB358E49F9DC" pageId="15" pageNumber="516">
|
||
<table id="F94DC431FFCA19AAFF50FB358E49F9DC" box="[163,1392,1191,1578]" gridcols="5" gridrows="10" pageId="15" pageNumber="516">
|
||
<tr id="357D34D3FFCA19AAFF50FB358E49FB36" box="[163,1392,1191,1216]" gridrow="0" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FB358A52FB36" box="[163,363,1191,1216]" gridcol="0" gridrow="0" pageId="15" pageNumber="516">Locality</th>
|
||
<th id="76AC5DAFFFCA19AAFE5BFB35895CFB36" box="[424,613,1191,1216]" gridcol="1" gridrow="0" pageId="15" pageNumber="516">Collection number</th>
|
||
<th id="76AC5DAFFFCA19AAFD71FB358866FB36" box="[642,863,1191,1216]" gridcol="2" gridrow="0" pageId="15" pageNumber="516">Stage of development</th>
|
||
<th id="76AC5DAFFFCA19AAFC6BFB358FFCFB36" box="[920,1221,1191,1216]" gridcol="3" gridrow="0" pageId="15" pageNumber="516">Ridges forming net-like relief</th>
|
||
<th id="76AC5DAFFFCA19AAFAEBFB358E49FB36" box="[1304,1392,1191,1216]" gridcol="4" gridrow="0" pageId="15" pageNumber="516">Fig.</th>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FB428E49FB1E" box="[163,1392,1232,1256]" gridrow="1" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FB428A52FB1E" box="[163,363,1232,1256]" gridcol="0" gridrow="1" pageId="15" pageNumber="516">India: Agumbe</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BFB42895CFB1E" box="[424,613,1232,1256]" gridcol="1" gridrow="1" pageId="15" pageNumber="516">[XIII](2)2013</td>
|
||
<td id="76AC5DAFFFCA19AAFD71FB428866FB1E" box="[642,863,1232,1256]" gridcol="2" gridrow="1" pageId="15" pageNumber="516">L-S-I♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFB428FFCFB1E" box="[920,1221,1232,1256]" gridcol="3" gridrow="1" pageId="15" pageNumber="516">all regular</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFB428E49FB1E" box="[1304,1392,1232,1256]" gridcol="4" gridrow="1" pageId="15" pageNumber="516">85–86</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FB6A8E49FAE6" box="[163,1392,1272,1296]" gridrow="2" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FB6A8A52FAE6" box="[163,363,1272,1296]" gridcol="0" gridrow="2" pageId="15" pageNumber="516">India: Agumbe</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BFB6A895CFAE6" box="[424,613,1272,1296]" gridcol="1" gridrow="2" pageId="15" pageNumber="516">[IX](6)2013</td>
|
||
<td id="76AC5DAFFFCA19AAFD71FB6A8866FAE6" box="[642,863,1272,1296]" gridcol="2" gridrow="2" pageId="15" pageNumber="516">L-S-I♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFB6A8FFCFAE6" box="[920,1221,1272,1296]" gridcol="3" gridrow="2" pageId="15" pageNumber="516">all regular</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFB6A8E49FAE6" box="[1304,1392,1272,1296]" gridcol="4" gridrow="2" pageId="15" pageNumber="516">87–88</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FAB28E49FACE" box="[163,1392,1312,1336]" gridrow="3" pageId="15" pageNumber="516" rowspan-1="1">
|
||
<th id="76AC5DAFFFCA19AAFF50FAB28A52FACE" box="[163,363,1312,1336]" gridcol="0" gridrow="3" pageId="15" pageNumber="516">India: Kodaikanal</th>
|
||
<td id="76AC5DAFFFCA19AAFD71FAB28866FACE" box="[642,863,1312,1336]" gridcol="2" gridrow="3" pageId="15" pageNumber="516">L/S♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFAB28FFCFACE" box="[920,1221,1312,1336]" gridcol="3" gridrow="3" pageId="15" pageNumber="516">all regular</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFAB28E49FACE" box="[1304,1392,1312,1336]" gridcol="4" gridrow="3" pageId="15" pageNumber="516">89–90</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FADA8E49FA97" box="[163,1392,1352,1377]" gridrow="4" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FADA8A52FA97" box="[163,363,1352,1377]" gridcol="0" gridrow="4" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BFADA895CFA97" box="[424,613,1352,1377]" gridcol="1" gridrow="4" pageId="15" pageNumber="516">[XIII](4)2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD71FADA8866FA97" box="[642,863,1352,1377]" gridcol="2" gridrow="4" pageId="15" pageNumber="516">L-S-I♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFADA8FFCFA97" box="[920,1221,1352,1377]" gridcol="3" gridrow="4" pageId="15" pageNumber="516">regular or broken</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFADA8E49FA97" box="[1304,1392,1352,1377]" gridcol="4" gridrow="4" pageId="15" pageNumber="516">91–92</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FAE38E49FA7F" box="[163,1392,1393,1417]" gridrow="5" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FAE38A52FA7F" box="[163,363,1393,1417]" gridcol="0" gridrow="5" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BFAE3895CFA7F" box="[424,613,1393,1417]" gridcol="1" gridrow="5" pageId="15" pageNumber="516">[XIX](1)2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD71FAE38866FA7F" box="[642,863,1393,1417]" gridcol="2" gridrow="5" pageId="15" pageNumber="516">L-S-I♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFAE38FFCFA7F" box="[920,1221,1393,1417]" gridcol="3" gridrow="5" pageId="15" pageNumber="516">regular or smoothed</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFAE38E49FA7F" box="[1304,1392,1393,1417]" gridcol="4" gridrow="5" pageId="15" pageNumber="516">93–94</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FA0B8E49FA47" box="[163,1392,1433,1457]" gridrow="6" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FA0B8A52FA47" box="[163,363,1433,1457]" gridcol="0" gridrow="6" pageId="15" pageNumber="516">Thailand: Pai</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BFA0B895CFA47" box="[424,613,1433,1457]" gridcol="1" gridrow="6" pageId="15" pageNumber="516">[XVIII](4)2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD71FA0B8866FA47" box="[642,863,1433,1457]" gridcol="2" gridrow="6" pageId="15" pageNumber="516">L-S♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFA0B8FFCFA47" box="[920,1221,1433,1457]" gridcol="3" gridrow="6" pageId="15" pageNumber="516">broken or smoothed</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFA0B8E49FA47" box="[1304,1392,1433,1457]" gridcol="4" gridrow="6" pageId="15" pageNumber="516">95–96</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FA538E49FA2F" box="[163,1392,1473,1497]" gridrow="7" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FA538A52FA2F" box="[163,363,1473,1497]" gridcol="0" gridrow="7" pageId="15" pageNumber="516">Thailand: Kwai-Yai</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BFA53895CFA2F" box="[424,613,1473,1497]" gridcol="1" gridrow="7" pageId="15" pageNumber="516">[III](11)2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD71FA538866FA2F" box="[642,863,1473,1497]" gridcol="2" gridrow="7" pageId="15" pageNumber="516">L-S-I♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFA538FFCFA2F" box="[920,1221,1473,1497]" gridcol="3" gridrow="7" pageId="15" pageNumber="516">broken or smoothed</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFA538E49FA2F" box="[1304,1392,1473,1497]" gridcol="4" gridrow="7" pageId="15" pageNumber="516">97–98</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50FA7B8E49F9F4" box="[163,1392,1513,1538]" gridrow="8" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50FA7B8A52F9F4" box="[163,363,1513,1538]" gridcol="0" gridrow="8" pageId="15" pageNumber="516">Thailand: Kwai-Yai</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BFA7B895CF9F4" box="[424,613,1513,1538]" gridcol="1" gridrow="8" pageId="15" pageNumber="516">[III](37)A2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD71FA7B8866F9F4" box="[642,863,1513,1538]" gridcol="2" gridrow="8" pageId="15" pageNumber="516">L-S-I♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BFA7B8FFCF9F4" box="[920,1221,1513,1538]" gridcol="3" gridrow="8" pageId="15" pageNumber="516">broken</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBFA7B8E49F9F4" box="[1304,1392,1513,1538]" gridcol="4" gridrow="8" pageId="15" pageNumber="516">99–100</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50F9808E49F9DC" box="[163,1392,1554,1578]" gridrow="9" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50F9808A52F9DC" box="[163,363,1554,1578]" gridcol="0" gridrow="9" pageId="15" pageNumber="516">Thailand: Kwai-Yai</th>
|
||
<td id="76AC5DAFFFCA19AAFE5BF980895CF9DC" box="[424,613,1554,1578]" gridcol="1" gridrow="9" pageId="15" pageNumber="516">[IV](2)B2015</td>
|
||
<td id="76AC5DAFFFCA19AAFD71F9808866F9DC" box="[642,863,1554,1578]" gridcol="2" gridrow="9" pageId="15" pageNumber="516">L-S-I♀</td>
|
||
<td id="76AC5DAFFFCA19AAFC6BF9808FFCF9DC" box="[920,1221,1554,1578]" gridcol="3" gridrow="9" pageId="15" pageNumber="516">completely absent</td>
|
||
<td id="76AC5DAFFFCA19AAFAEBF9808E49F9DC" box="[1304,1392,1554,1578]" gridcol="4" gridrow="9" pageId="15" pageNumber="516">101–102</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFCAE65AFF64F9CB890FF961" blockId="15.[151,1437,1625,1687]" pageId="15" pageNumber="516">
|
||
<emphasis id="B939EA83FFCAE65AFF64F9CB8AF8F985" bold="true" box="[151,449,1625,1651]" pageId="15" pageNumber="516">Geographical variability.</emphasis>
|
||
Our small collection allows to make following assumptions about characters peculiar for certain geographical forms (
|
||
<tableCitation id="C6CF032AFFCAE65AFE27F9EF8911F961" box="[468,552,1661,1687]" captionStart="TABLE 3" captionStartId="15.[152,244,1735,1759]" captionTargetPageId="15" captionText="TABLE 3. Larval cuticular coloration" httpUri="http://table.plazi.org/id/DF326619FFCAE65AFF6BF955890BF929" pageId="15" pageNumber="516" tableUuid="DF326619FFCAE65AFF6BF955890BF929">Table 3</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF326619FFCAE65AFF6BF955890BF929" ID-Table-UUID="DF326619FFCAE65AFF6BF955890BF929" box="[152,562,1735,1759]" httpUri="http://table.plazi.org/id/DF326619FFCAE65AFF6BF955890BF929" pageId="15" pageNumber="516" startId="15.[152,244,1735,1759]" targetBox="[163,1374,1775,2001]" targetIsTable="true" targetPageId="15" targetType="table">
|
||
<paragraph id="8BF23691FFCAE65AFF6BF955890BF929" blockId="15.[152,562,1735,1759]" box="[152,562,1735,1759]" pageId="15" pageNumber="516">
|
||
<emphasis id="B939EA83FFCAE65AFF6BF9558A36F929" bold="true" box="[152,271,1735,1759]" pageId="15" pageNumber="516">TABLE 3.</emphasis>
|
||
Larval cuticular coloration
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23691FFCAE65AFF50F97D8FC1F826" pageId="15" pageNumber="516">
|
||
<table id="F94DC431FFCA19AAFF50F97D8E67F827" box="[163,1374,1775,2001]" gridcols="3" gridrows="6" pageId="15" pageNumber="516">
|
||
<tr id="357D34D3FFCA19AAFF50F97D8E67F8F1" box="[163,1374,1775,1799]" gridrow="0" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50F97D893FF8F1" box="[163,518,1775,1799]" gridcol="0" gridrow="0" pageId="15" pageNumber="516">Locality</th>
|
||
<th id="76AC5DAFFFCA19AAFD99F97D8F3DF8F1" box="[618,1028,1775,1799]" gridcol="1" gridrow="0" pageId="15" pageNumber="516">Medio-posterior blanks on frontal shield</th>
|
||
<th id="76AC5DAFFFCA19AAFBC0F97D8E67F8F1" box="[1075,1374,1775,1799]" gridcol="2" gridrow="0" pageId="15" pageNumber="516">Anterior margin of pronotum</th>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50F8858E67F8C6" box="[163,1374,1815,1840]" gridrow="1" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50F885893FF8C6" box="[163,518,1815,1840]" gridcol="0" gridrow="1" pageId="15" pageNumber="516">India (8 individuals)</th>
|
||
<td id="76AC5DAFFFCA19AAFD99F8858F3DF8C6" box="[618,1028,1815,1840]" gridcol="1" gridrow="1" pageId="15" pageNumber="516">absent</td>
|
||
<td id="76AC5DAFFFCA19AAFBC0F8858E67F8C6" box="[1075,1374,1815,1840]" gridcol="2" gridrow="1" pageId="15" pageNumber="516">light</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50F8D28E67F8AE" box="[163,1374,1856,1880]" gridrow="2" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50F8D2893FF8AE" box="[163,518,1856,1880]" gridcol="0" gridrow="2" pageId="15" pageNumber="516">Thailand: Pai (7 individuals)</th>
|
||
<td id="76AC5DAFFFCA19AAFD99F8D28F3DF8AE" box="[618,1028,1856,1880]" gridcol="1" gridrow="2" pageId="15" pageNumber="516">present</td>
|
||
<td id="76AC5DAFFFCA19AAFBC0F8D28E67F8AE" box="[1075,1374,1856,1880]" gridcol="2" gridrow="2" pageId="15" pageNumber="516">dark</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50F8FA8E67F876" box="[163,1374,1896,1920]" gridrow="3" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50F8FA893FF876" box="[163,518,1896,1920]" gridcol="0" gridrow="3" pageId="15" pageNumber="516">Thailand: Kwai-Yai (7 individuals)</th>
|
||
<td id="76AC5DAFFFCA19AAFD99F8FA8F3DF876" box="[618,1028,1896,1920]" gridcol="1" gridrow="3" pageId="15" pageNumber="516">coloration not expressed</td>
|
||
<td id="76AC5DAFFFCA19AAFBC0F8FA8E67F876" box="[1075,1374,1896,1920]" gridcol="2" gridrow="3" pageId="15" pageNumber="516">coloration not expressed</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50F8028E67F85E" box="[163,1374,1936,1960]" gridrow="4" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50F802893FF85E" box="[163,518,1936,1960]" gridcol="0" gridrow="4" pageId="15" pageNumber="516">
|
||
Vietnam (
|
||
<bibRefCitation id="EFDC4B60FFCAE65AFEF6F8028AC3F85E" author="Soldan, T. & Braasch, D." box="[261,506,1936,1960]" pageId="15" pageNumber="516" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
)
|
||
</th>
|
||
<td id="76AC5DAFFFCA19AAFD99F8028F3DF85E" box="[618,1028,1936,1960]" gridcol="1" gridrow="4" pageId="15" pageNumber="516">absent</td>
|
||
<td id="76AC5DAFFFCA19AAFBC0F8028E67F85E" box="[1075,1374,1936,1960]" gridcol="2" gridrow="4" pageId="15" pageNumber="516">dark</td>
|
||
</tr>
|
||
<tr id="357D34D3FFCA19AAFF50F82A8E67F827" box="[163,1374,1976,2001]" gridrow="5" pageId="15" pageNumber="516">
|
||
<th id="76AC5DAFFFCA19AAFF50F82A893FF827" box="[163,518,1976,2001]" gridcol="0" gridrow="5" pageId="15" pageNumber="516">
|
||
Sumatra (
|
||
<bibRefCitation id="EFDC4B60FFCAE65AFEF7F82A8ABCF827" author="Sartori, M." box="[260,389,1976,2001]" pageId="15" pageNumber="516" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
)
|
||
</th>
|
||
<td id="76AC5DAFFFCA19AAFD99F82A8F3DF827" box="[618,1028,1976,2001]" gridcol="1" gridrow="5" pageId="15" pageNumber="516">absent</td>
|
||
<td id="76AC5DAFFFCA19AAFBC0F82A8E67F827" box="[1075,1374,1976,2001]" gridcol="2" gridrow="5" pageId="15" pageNumber="516">dark</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF326619FFD5E645FF64F9B18E44F9A9" ID-DOI="http://doi.org/10.5281/zenodo.8203270" ID-Zenodo-Dep="8203270" httpUri="https://zenodo.org/record/8203270/files/figure.png" pageId="16" pageNumber="517" startId="16.[151,264,1571,1595]" targetBox="[164,1423,181,1546]" targetPageId="16" targetType="figure">
|
||
<paragraph id="8BF23691FFD5E645FF64F9B18E44F9A9" blockId="16.[151,1436,1571,1631]" pageId="16" pageNumber="517">
|
||
<emphasis id="B939EA83FFD5E645FF64F9B18A6DF9CD" bold="true" box="[151,340,1571,1595]" pageId="16" pageNumber="517">FIGURES 85–90.</emphasis>
|
||
<taxonomicName id="4C4D4D12FFD5E645FEAAF9B189E0F9CD" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[345,729,1571,1595]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD5E645FEAAF9B18AEBF9CD" box="[345,466,1571,1595]" italics="true" pageId="16" pageNumber="517">Ecdyonurus</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD5E645FE13F9B18947F9CD" box="[480,638,1571,1595]" italics="true" pageId="16" pageNumber="517">Rhithrogeniella</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD5E645FD7FF9B689E0F9CD" box="[652,729,1572,1595]" italics="true" pageId="16" pageNumber="517">ornatus</emphasis>
|
||
</taxonomicName>
|
||
, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23691FFD5E645FF34F9158E12F868" blockId="16.[151,1437,1671,2022]" pageId="16" pageNumber="517">
|
||
<emphasis id="B939EA83FFD5E645FF34F9158A01F954" bold="true" box="[199,312,1671,1698]" italics="true" pageId="16" pageNumber="517">
|
||
<collectingCountry id="F35A7601FFD5E645FF34F9158A0DF954" box="[199,308,1671,1698]" name="Thailand" pageId="16" pageNumber="517">Thailand</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
Denticles on posterior margins of abdominal terga II–IX are irregular, small and thin (
|
||
<figureCitation id="13762A14FFD5E645FAE5F91A8E53F955" box="[1302,1386,1672,1699]" captionStart="FIGURES 42–52" captionStartId="7.[151,264,1785,1809]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-20@7.[151,1436,181,1762]" captionTargetPageId="7" captionText="FIGURES 42–52. Larvae. 42–50, Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae: 42–44, specimen from Agumbe (India), posterior margins of abdominal terga I, VIII and X; 45, specimen from Pai (Thailand), posterior margins of abdominal tergum VI; 46–48, cercus of specimen from Agumbe: 46, proximal portion; 47, middle portion in median-ventral view, to show bases of primary swimming setae on inner (median) side; 48, the same portion with focus on lateral-dorsal side, to show bases of secondary swimming setae on outer (lateral) side; 49–50, cercus of specimen from Kodaikanal (India). 51–52, Ecdyonurus (Rhithrogeniella) joernensis (Russia, Orenburg province, Tashla, 16.VIII.1986, coll. N. Kluge): two portions of cercus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8203260" httpUri="https://zenodo.org/record/8203260/files/figure.png" pageId="16" pageNumber="517">Fig. 45</figureCitation>
|
||
), on some of these terga can be as small as on terga I and X. In all specimens from Kwai-Yai, larval cuticle is very pale and nearly lacks pigmentation, so that the features of cuticular coloration characteristic for individuals from Pai and India, are not expressed. All examined mature larvae from Pai have well expressed cuticular coloration. Frontal shield has a pair of blanks behind the pair of submedian blanks adjacent to anterior margin (
|
||
<figureCitation id="13762A14FFD5E645FB4DF88A8E49F8C4" box="[1214,1392,1816,1843]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="16" pageNumber="517">Figs 15, 19, 22</figureCitation>
|
||
), in contrast to the specimens from India (
|
||
<figureCitation id="13762A14FFD5E645FDCAF8AE89B3F8A0" box="[569,650,1852,1878]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="16" pageNumber="517">Fig. 11</figureCitation>
|
||
), Vietnam and Sumatra (
|
||
<bibRefCitation id="EFDC4B60FFD5E645FC6FF8AE8F93F8A0" author="Soldan, T. & Braasch, D." box="[924,1194,1852,1878]" pageId="16" pageNumber="517" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 16;
|
||
<bibRefCitation id="EFDC4B60FFD5E645FAF9F8AE8EA1F8A0" author="Sartori, M." box="[1290,1432,1852,1878]" pageId="16" pageNumber="517" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: figs 6–7). Cuticle of larval pronotum has dark area of this or that shape adjacent to anterior margin (
|
||
<figureCitation id="13762A14FFD5E645FB16F8F28EB7F88C" box="[1253,1422,1888,1914]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="16" pageNumber="517">Figs 16, 20, 23</figureCitation>
|
||
); the same in larvae from Vietnam (
|
||
<bibRefCitation id="EFDC4B60FFD5E645FDE0F816881DF868" author="Soldan, T. & Braasch, D." box="[531,804,1924,1950]" pageId="16" pageNumber="517" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 17) and Sumatra (
|
||
<bibRefCitation id="EFDC4B60FFD5E645FBD0F8168F8DF868" author="Sartori, M." box="[1059,1204,1924,1950]" pageId="16" pageNumber="517" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: figs 6–7).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFD5E644FF34F835899FFE93" blockId="16.[151,1437,1671,2022]" lastBlockId="17.[151,1436,151,357]" lastPageId="17" lastPageNumber="518" pageId="16" pageNumber="517">
|
||
<emphasis id="B939EA83FFD5E645FF34F8358A35F834" bold="true" box="[199,268,1959,1986]" italics="true" pageId="16" pageNumber="517">
|
||
<collectingCountry id="F35A7601FFD5E645FF34F8358A31F834" box="[199,264,1959,1986]" name="India" pageId="16" pageNumber="517">India</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
Denticles on posterior margins of abdominal terga II–IX are regularly alternating as large and small (
|
||
<figureCitation id="13762A14FFD5E645FA86F83A8B8FF810" captionStart="FIGURES 42–52" captionStartId="7.[151,264,1785,1809]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-20@7.[151,1436,181,1762]" captionTargetPageId="7" captionText="FIGURES 42–52. Larvae. 42–50, Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae: 42–44, specimen from Agumbe (India), posterior margins of abdominal terga I, VIII and X; 45, specimen from Pai (Thailand), posterior margins of abdominal tergum VI; 46–48, cercus of specimen from Agumbe: 46, proximal portion; 47, middle portion in median-ventral view, to show bases of primary swimming setae on inner (median) side; 48, the same portion with focus on lateral-dorsal side, to show bases of secondary swimming setae on outer (lateral) side; 49–50, cercus of specimen from Kodaikanal (India). 51–52, Ecdyonurus (Rhithrogeniella) joernensis (Russia, Orenburg province, Tashla, 16.VIII.1986, coll. N. Kluge): two portions of cercus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8203260" httpUri="https://zenodo.org/record/8203260/files/figure.png" pageId="16" pageNumber="517">Fig 43</figureCitation>
|
||
), well different from small denticles on terga I and X (
|
||
<figureCitation id="13762A14FFD5E645FCD5F85E88ECF810" box="[806,981,1996,2022]" captionStart="FIGURES 42–52" captionStartId="7.[151,264,1785,1809]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-20@7.[151,1436,181,1762]" captionTargetPageId="7" captionText="FIGURES 42–52. Larvae. 42–50, Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae: 42–44, specimen from Agumbe (India), posterior margins of abdominal terga I, VIII and X; 45, specimen from Pai (Thailand), posterior margins of abdominal tergum VI; 46–48, cercus of specimen from Agumbe: 46, proximal portion; 47, middle portion in median-ventral view, to show bases of primary swimming setae on inner (median) side; 48, the same portion with focus on lateral-dorsal side, to show bases of secondary swimming setae on outer (lateral) side; 49–50, cercus of specimen from Kodaikanal (India). 51–52, Ecdyonurus (Rhithrogeniella) joernensis (Russia, Orenburg province, Tashla, 16.VIII.1986, coll. N. Kluge): two portions of cercus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8203260" httpUri="https://zenodo.org/record/8203260/files/figure.png" pageId="16" pageNumber="517">Figs 42 and 44</figureCitation>
|
||
). All examined mature larvae from both localities in the states
|
||
<collectingRegion id="4989F873FFD4E644FE6AFF058935FF47" box="[409,524,151,177]" country="India" name="Karnataka" pageId="17" pageNumber="518">Karnataka</collectingRegion>
|
||
and
|
||
<collectingRegion id="4989F873FFD4E644FDB4FF0589F7FF47" box="[583,718,151,177]" country="India" name="Tamil Nadu" pageId="17" pageNumber="518">Tamil Nadu</collectingRegion>
|
||
have well expressed cuticular pigmentation (in contrast to the specimens from Kway-Yai in Thailand). In contrast to the individuals from Pai in Thailand, frontal shield has no additional blanks behind the pair of submedian blanks adjacent to anterior margin (
|
||
<figureCitation id="13762A14FFD4E644FBCAFF4D8FB3FF0F" box="[1081,1162,223,249]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="17" pageNumber="518">Fig. 11</figureCitation>
|
||
); the same is figured for the individuals from Vietnam (
|
||
<bibRefCitation id="EFDC4B60FFD4E644FE1CFE9189C6FEEB" author="Soldan, T. & Braasch, D." box="[495,767,259,285]" pageId="17" pageNumber="518" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 16) and from Sumatra (
|
||
<bibRefCitation id="EFDC4B60FFD4E644FBCEFE918FF7FEEB" author="Sartori, M." box="[1085,1230,259,285]" pageId="17" pageNumber="518" pagination="47 - 61" refId="ref9983" refString="Sartori, M. (2014) Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae). ZooKeys, 429, 47 - 61. https: // doi. org / 10.3897 / zookeys. 429.8116" type="journal article" year="2014">Sartori 2014</bibRefCitation>
|
||
: figs 6–7). Cuticle of larval pronotum has paired light blank adjacent to anterior margin (
|
||
<figureCitation id="13762A14FFD4E644FC5BFEB588C8FEB7" box="[936,1009,295,321]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1810,1834]" captionTargetBox="[151,1436,180,1772]" captionTargetId="figure-20@3.[151,1436,179,1784]" captionTargetPageId="3" captionText="FIGURES 1–10. Ecdyonurus (Rhithrogeniella) ornatus from Kodaikanal (India). 1–3, larvae; 4–10, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8203250" httpUri="https://zenodo.org/record/8203250/files/figure.png" pageId="17" pageNumber="518">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="13762A14FFD4E644FC0DFEB58F24FEB7" box="[1022,1053,295,321]" captionStart="FIGURES 11–25" captionStartId="4.[151,264,1853,1877]" captionTargetBox="[151,1435,181,1829]" captionTargetId="figure-37@4.[151,1435,181,1829]" captionTargetPageId="4" captionText="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." figureDoi="http://doi.org/10.5281/zenodo.8203254" httpUri="https://zenodo.org/record/8203254/files/figure.png" pageId="17" pageNumber="518">12</figureCitation>
|
||
); this differs from larvae from Pai in Thailand, Vietnam and Sumatra (see above).
|
||
</paragraph>
|
||
<caption id="DF326619FFD4E644FF64F8228EBAF81A" ID-DOI="http://doi.org/10.5281/zenodo.8203272" ID-Zenodo-Dep="8203272" httpUri="https://zenodo.org/record/8203272/files/figure.png" pageId="17" pageNumber="518" startId="17.[151,264,1968,1992]" targetBox="[177,1411,381,1945]" targetPageId="17" targetType="figure">
|
||
<paragraph id="8BF23691FFD4E644FF64F8228EBAF81A" blockId="17.[151,1436,1968,2028]" pageId="17" pageNumber="518">
|
||
<emphasis id="B939EA83FFD4E644FF64F8228A61F83E" bold="true" box="[151,344,1968,1992]" pageId="17" pageNumber="518">FIGURES 91–96.</emphasis>
|
||
<taxonomicName id="4C4D4D12FFD4E644FE92F82289D3F83E" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[353,746,1968,1992]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD4E644FE92F8228AE3F83E" box="[353,474,1968,1992]" italics="true" pageId="17" pageNumber="518">Ecdyonurus</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD4E644FE1FF82289B3F83E" box="[492,650,1968,1992]" italics="true" pageId="17" pageNumber="518">Rhithrogeniella</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD4E644FD6EF82389D3F83E" box="[669,746,1969,1992]" italics="true" pageId="17" pageNumber="518">ornatus</emphasis>
|
||
</taxonomicName>
|
||
, fully developed eggs from Pai (Thailand). 91–92, reared imago, specimen [XIII](4)2015; 93–94, reared imago, specimen [XIX](1)2015; 95–96, reared subimago, specimen [XVIII](4)2015.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF326619FFD7E647FF64F8B88FA9F890" ID-DOI="http://doi.org/10.5281/zenodo.8203274" ID-Zenodo-Dep="8203274" httpUri="https://zenodo.org/record/8203274/files/figure.png" pageId="18" pageNumber="519" startId="18.[151,264,1834,1858]" targetBox="[151,1436,181,1810]" targetPageId="18" targetType="figure">
|
||
<paragraph id="8BF23691FFD7E647FF64F8B88FA9F890" blockId="18.[151,1437,1834,1894]" pageId="18" pageNumber="519">
|
||
<emphasis id="B939EA83FFD7E647FF64F8B88A59F8B4" bold="true" box="[151,352,1834,1858]" pageId="18" pageNumber="519">FIGURES 97–102.</emphasis>
|
||
<taxonomicName id="4C4D4D12FFD7E647FE95F8B889DEF8B4" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[358,743,1834,1858]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="519" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD7E647FE95F8B88AE6F8B4" box="[358,479,1834,1858]" italics="true" pageId="18" pageNumber="519">Ecdyonurus</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD7E647FE1EF8B889B2F8B4" box="[493,651,1834,1858]" italics="true" pageId="18" pageNumber="519">Rhithrogeniella</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD7E647FD69F8B989DEF8B4" box="[666,743,1835,1858]" italics="true" pageId="18" pageNumber="519">ornatus</emphasis>
|
||
</taxonomicName>
|
||
, fully developed eggs of reared imagines from Kwai-Yai (Thailand). 97–98, specimen [III](11)2015; 99–100, specimen [III](37)A2015; 101–102, specimen [IV](2)2015.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23691FFD7E647FF34F8078A4FF824" blockId="18.[151,1436,1940,2003]" pageId="18" pageNumber="519">
|
||
In all three examined females, all eggs have regular mesh-like relief consisting of non-interrupted ridges (Figs 85–90 and
|
||
<tableCitation id="C6CF032AFFD7E647FEE0F82A8A50F824" box="[275,361,1976,2002]" captionStart="TABLE 2" captionStartId="15.[152,244,1151,1175]" captionTargetPageId="15" captionText="TABLE 2. Relief of egg chorion in the specimens examined" httpUri="http://table.plazi.org/id/DF326619FFCAE65AFF6BFBED8819FB61" pageId="18" pageNumber="519" tableUuid="DF326619FFCAE65AFF6BFBED8819FB61">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFD6E646FF64FF048941FEB7" blockId="19.[151,1437,150,321]" pageId="19" pageNumber="520">
|
||
<emphasis id="B939EA83FFD6E646FF64FF048811FF47" bold="true" box="[151,808,150,177]" pageId="19" pageNumber="520">
|
||
Synonymy of
|
||
<taxonomicName id="4C4D4D12FFD6E646FECFFF058AC2FF47" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[316,507,150,177]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="522" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD6E646FECFFF058A76FF47" bold="true" box="[316,335,151,177]" italics="true" pageId="19" pageNumber="520">E</emphasis>
|
||
. (
|
||
<emphasis id="B939EA83FFD6E646FE94FF048AB0FF47" bold="true" box="[359,393,150,177]" italics="true" pageId="19" pageNumber="520">Rh</emphasis>
|
||
.)
|
||
<emphasis id="B939EA83FFD6E646FE51FF058AC2FF47" bold="true" box="[418,507,151,177]" italics="true" pageId="19" pageNumber="520">ornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B939EA83FFD6E646FDC4FF058973FF47" bold="true" box="[567,586,151,177]" italics="true" pageId="19" pageNumber="520">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C4D4D12FFD6E646FDAAFF048818FF47" authorityName=": Kluge" authorityYear="2022" box="[601,801,150,177]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="tonkinensis" subGenus="Rhithrogeniella">
|
||
(
|
||
<emphasis id="B939EA83FFD6E646FD91FF0489BDFF47" bold="true" box="[610,644,150,177]" italics="true" pageId="19" pageNumber="520">Rh</emphasis>
|
||
.)
|
||
<emphasis id="B939EA83FFD6E646FD6EFF048818FF47" bold="true" box="[669,801,150,177]" italics="true" pageId="19" pageNumber="520">tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Originally
|
||
<taxonomicName id="4C4D4D12FFD6E646FC58FF058F1DFF46" authorityName="Ulmer" authorityYear="1939" box="[939,1060,151,177]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFD6E646FC58FF058F1DFF46" box="[939,1060,151,177]" italics="true" pageId="19" pageNumber="520">Rh. ornata</emphasis>
|
||
</taxonomicName>
|
||
was described from Great Sunda Islands (Java and Sumatra), and
|
||
<taxonomicName id="4C4D4D12FFD6E646FDF2FF298995FF23" authorityName=": Kluge" authorityYear="2022" box="[513,684,187,213]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="B939EA83FFD6E646FDF2FF298995FF23" box="[513,684,187,213]" italics="true" pageId="19" pageNumber="520">Rh. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
was described from Indochinese Peninsula (
|
||
<collectingCountry id="F35A7601FFD6E646FB6DFF298E3BFF23" box="[1182,1282,187,213]" name="Vietnam" pageId="19" pageNumber="520">Vietnam</collectingCountry>
|
||
). The original differential diagnosis of
|
||
<taxonomicName id="4C4D4D12FFD6E646FE55FF4D8969FF0F" authorityName=": Kluge" authorityYear="2022" box="[422,592,223,249]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="B939EA83FFD6E646FE55FF4D8969FF0F" box="[422,592,223,249]" italics="true" pageId="19" pageNumber="520">Rh. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
contained the following statement: ́Since the adult male of
|
||
<taxonomicName id="4C4D4D12FFD6E646FB1DFF4D8A2CFEEB" authority="Kluge & Sivaruban & Srinivasan & Barathy & Isack, 2023" authorityName="Kluge & Sivaruban & Srinivasan & Barathy & Isack" authorityYear="2023" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="tonkinensis" status="sp. nov.">
|
||
<emphasis id="B939EA83FFD6E646FB1DFF4D8A2CFEEB" italics="true" pageId="19" pageNumber="520">Rhithrogeniella tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A57F8FFD6E646FED3FE918A59FEEB" box="[288,352,259,285]" pageId="19" pageNumber="520" rank="species">sp. n.</taxonomicNameLabel>
|
||
and nymphs of
|
||
<taxonomicName id="4C4D4D12FFD6E646FDD0FE9189DDFEEB" authority="Ulmer" authorityName="Ulmer" authorityYear="1939" box="[547,740,259,285]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFD6E646FDD0FE9189A8FEEA" box="[547,657,259,284]" italics="true" pageId="19" pageNumber="520">R. ornata</emphasis>
|
||
Ulmer
|
||
</taxonomicName>
|
||
are unknown only subimagoes of these two species can be compared» (
|
||
<bibRefCitation id="EFDC4B60FFD6E646FED1FEB58952FEB7" author="Soldan, T. & Braasch, D." box="[290,619,295,321]" pageId="19" pageNumber="520" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986: 209</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF326619FFD6E646FF64F90E8844F90A" ID-DOI="http://doi.org/10.5281/zenodo.8203276" ID-Zenodo-Dep="8203276" httpUri="https://zenodo.org/record/8203276/files/figure.png" pageId="19" pageNumber="520" startId="19.[151,264,1692,1716]" targetBox="[154,1436,345,1669]" targetPageId="19" targetType="figure">
|
||
<paragraph id="8BF23691FFD6E646FF64F90E8844F90A" blockId="19.[151,1436,1692,1789]" pageId="19" pageNumber="520">
|
||
<emphasis id="B939EA83FFD6E646FF64F90E8A55F942" bold="true" box="[151,364,1692,1716]" pageId="19" pageNumber="520">FIGURES 103–110.</emphasis>
|
||
103–109,
|
||
<taxonomicName id="4C4D4D12FFD6E646FE2BF90E883FF943" box="[472,774,1692,1717]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="subGenus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD6E646FE2BF90E8968F942" box="[472,593,1692,1716]" italics="true" pageId="19" pageNumber="520">Ecdyonurus</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD6E646FD93F90E89C7F942" box="[608,766,1692,1716]" italics="true" pageId="19" pageNumber="520">Rhithrogeniella</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
sp. 2: 103, male imago, 104, abdomen of male imago; 105, male subimago; 106, female subimago; 107, thorax of male subimago, ventral view; 108, male subimaginal exuviae; 109, enlarged part of its ventral arc of prealar bridge; 110, the same of
|
||
<taxonomicName id="4C4D4D12FFD6E646FD27F977884FF90A" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[724,886,1764,1789]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD6E646FD27F97789D3F90A" box="[724,746,1765,1788]" italics="true" pageId="19" pageNumber="520">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD6E646FD0BF9768822F90A" box="[760,795,1764,1788]" italics="true" pageId="19" pageNumber="520">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD6E646FCDAF977884FF90A" box="[809,886,1765,1788]" italics="true" pageId="19" pageNumber="520">ornatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23691FFD6E646FF34F8B98BC6F89F" blockId="19.[151,1437,1835,2041]" pageId="19" pageNumber="520">Subimagines of these species were said to differ by abdominal coloration, leg proportions and shape of male genitalia.</paragraph>
|
||
<paragraph id="8BF23691FFD6E641FF34F8E1899FFEB7" blockId="19.[151,1437,1835,2041]" lastBlockId="20.[151,1436,150,321]" lastPageId="20" lastPageNumber="521" pageId="19" pageNumber="520">
|
||
The difference in abdominal coloration was caused by the fact that the Soldán’s drawing of subimaginal abdomen (
|
||
<bibRefCitation id="EFDC4B60FFD6E646FEF8F8058921F846" author="Soldan, T. & Braasch, D." box="[267,536,1942,1969]" pageId="19" pageNumber="520" pagination="202 - 212" refId="ref10092" refString="Soldan, T. & Braasch, D. (1986) Rhithrogeniella tonkinensis sp. n. (Ephemeroptera, Heptageniidae) from Vietnam, with descriptions of the nymphal stages and biology of the genus. Acta Entomologica Bohemoslovaca, 83, 202 - 212." type="journal article" year="1986">Soldán & Braasch 1986</bibRefCitation>
|
||
: fig. 19) was probably made from subimago just emerged and not fully colored; it resembles the subimaginal abdomen extracted from larva ready to molt (
|
||
<figureCitation id="13762A14FFD6E646FC44F8298F33F823" box="[951,1034,1978,2005]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="19" pageNumber="520">Fig. 53</figureCitation>
|
||
), while the Ulmer’s drawing (
|
||
<bibRefCitation id="EFDC4B60FFD6E646FAA6F8298BEAF80F" author="Ulmer, G." pageId="19" pageNumber="520" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939</bibRefCitation>
|
||
: fig. 170) was probably made from an older subimago, whose coloration is approximated to the imaginal one (
|
||
<figureCitation id="13762A14FFD1E641FF6CFF058A1CFF47" box="[159,293,150,177]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="20" pageNumber="521">Figs 54–57</figureCitation>
|
||
). Female abdomen of
|
||
<taxonomicName id="4C4D4D12FFD1E641FDDBFF0589EFFF47" authorityName="Kluge & Sivaruban & Srinivasan & Barathy & Isack" authorityYear="2023" box="[552,726,151,177]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="tonkinensis">
|
||
<emphasis id="B939EA83FFD1E641FDDBFF0589EFFF47" box="[552,726,151,177]" italics="true" pageId="20" pageNumber="521">Rh. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
was said to be ́without tergal markings», while abdomen of female
|
||
<taxonomicName id="4C4D4D12FFD1E641FF1FFF298A5CFF22" authorityName="Ulmer" authorityYear="1939" box="[236,357,187,213]" class="Insecta" family="Heptageniidae" genus="Rhithrogeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B939EA83FFD1E641FF1FFF298A5CFF22" box="[236,357,187,213]" italics="true" pageId="20" pageNumber="521">Rh. ornata</emphasis>
|
||
</taxonomicName>
|
||
has tergal markings (
|
||
<bibRefCitation id="EFDC4B60FFD1E641FDAFFF2989D1FF23" author="Ulmer, G." box="[604,744,187,213]" pageId="20" pageNumber="521" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939</bibRefCitation>
|
||
: fig. 171). Actually abdominal markings of female are worse expressed than in male (
|
||
<figureCitation id="13762A14FFD1E641FE54FF4D8912FF0F" box="[423,555,223,249]" captionStart="FIGURES 77–84" captionStartId="14.[151,264,1532,1556]" captionTargetBox="[151,1435,309,1509]" captionTargetId="figure-91@14.[151,1435,309,1509]" captionTargetPageId="14" captionText="FIGURES 77–84. Ecdyonurus (Rhithrogeniella) ornatus, female imagines. 77–78, specimen from Agumbe (India); 79–81, specimens from Kwai-Yai (Thailand); 82–84, specimens from Pai (Thailand)." figureDoi="http://doi.org/10.5281/zenodo.8203268" httpUri="https://zenodo.org/record/8203268/files/figure.png" pageId="20" pageNumber="521">Figs 78–84</figureCitation>
|
||
). Hypodermal coloration of male abdomen, female pronotum, mesonotum and abdomen of our specimens from
|
||
<collectingCountry id="F35A7601FFD1E641FDF8FE91897CFEEB" box="[523,581,259,285]" name="India" pageId="20" pageNumber="521">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F35A7601FFD1E641FD8FFE9189D9FEEB" box="[636,736,259,285]" name="Thailand" pageId="20" pageNumber="521">Thailand</collectingCountry>
|
||
are the same as in the
|
||
<typeStatus id="54F68833FFD1E641FC12FE918FBAFEEB" box="[993,1155,259,285]" pageId="20" pageNumber="521" type="paralectotype">paralectotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4D4D12FFD1E641FB5AFE918E66FEEA" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[1193,1375,259,285]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD1E641FB5AFE918FF8FEEA" box="[1193,1217,259,284]" italics="true" pageId="20" pageNumber="521">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD1E641FB21FE918FC1FEEB" box="[1234,1272,259,285]" italics="true" pageId="20" pageNumber="521">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD1E641FAFAFE918E66FEEA" box="[1289,1375,259,284]" italics="true" pageId="20" pageNumber="521">ornatus</emphasis>
|
||
</taxonomicName>
|
||
from Java and Sumatra (
|
||
<bibRefCitation id="EFDC4B60FFD1E641FE9EFEB58AC1FEB7" author="Ulmer, G." box="[365,504,295,321]" pageId="20" pageNumber="521" pagination="443 - 692" refId="ref10209" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="journal article" year="1939">Ulmer 1939</bibRefCitation>
|
||
: figs 170–172).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF326619FFD1E641FF64F8DF8915F87C" ID-DOI="http://doi.org/10.5281/zenodo.8203280" ID-Zenodo-Dep="8203280" httpUri="https://zenodo.org/record/8203280/files/figure.png" pageId="20" pageNumber="521" startId="20.[151,264,1869,1893]" targetBox="[151,1436,366,1846]" targetPageId="20" targetType="figure">
|
||
<paragraph id="8BF23691FFD1E641FF64F8DF8915F87C" blockId="20.[151,1436,1869,1930]" pageId="20" pageNumber="521">
|
||
<emphasis id="B939EA83FFD1E641FF64F8DF8A51F890" bold="true" box="[151,360,1869,1894]" pageId="20" pageNumber="521">FIGURES 111–116.</emphasis>
|
||
<taxonomicName id="4C4D4D12FFD1E641FE9CF8DC89A4F890" box="[367,669,1870,1894]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="subGenus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD1E641FE9CF8DC8AD1F890" box="[367,488,1870,1894]" italics="true" pageId="20" pageNumber="521">Ecdyonurus</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD1E641FE04F8DC89ACF890" box="[503,661,1870,1894]" italics="true" pageId="20" pageNumber="521">Rhithrogeniella</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
sp. 2: 111, genitalia of male imago; 112, penis of male subimago; 113–114, penes of male imagines; 115–116, eggs.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C357651AFFD1E640FF34F82A8FDCFD3B" lastPageId="21" lastPageNumber="522" pageId="20" pageNumber="521" type="diagnosis">
|
||
<paragraph id="8BF23691FFD1E640FF34F82A89ABFE93" blockId="20.[151,1436,1976,2038]" lastBlockId="21.[151,1437,151,717]" lastPageId="21" lastPageNumber="522" pageId="20" pageNumber="521">
|
||
<emphasis id="B939EA83FFD1E641FF34F82A8A5BF824" bold="true" box="[199,354,1976,2002]" pageId="20" pageNumber="521">Comparison.</emphasis>
|
||
Beside
|
||
<emphasis id="B939EA83FFD1E641FE45F82B8AF8F824" box="[438,449,1977,2002]" italics="true" pageId="20" pageNumber="521">s</emphasis>
|
||
<taxonomicName id="4C4D4D12FFD1E641FE3AF82B89B9F824" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[457,640,1976,2002]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD1E641FE3AF82B8ADBF824" box="[457,482,1977,2002]" italics="true" pageId="20" pageNumber="521">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD1E641FE00F82A8923F824" box="[499,538,1976,2002]" italics="true" pageId="20" pageNumber="521">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD1E641FDD9F82B89B9F824" box="[554,640,1977,2002]" italics="true" pageId="20" pageNumber="521">ornatus</emphasis>
|
||
</taxonomicName>
|
||
, three other Oriental species were described based on larvae and eggs from
|
||
<collectingCountry id="F35A7601FFD1E641FF2BF84E8A14F800" box="[216,301,2012,2038]" name="Taiwan" pageId="20" pageNumber="521">Taiwan</collectingCountry>
|
||
, under the names
|
||
<taxonomicName id="4C4D4D12FFD1E641FDF7F84F89C2F800" authorityName="Kang & Yang" authorityYear="1994" box="[516,763,2012,2038]" class="Insecta" family="Heptageniidae" genus="Nixe" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="littorosus" subGenus="Nixe">
|
||
<emphasis id="B939EA83FFD1E641FDF7F84F890EF800" box="[516,567,2013,2038]" italics="true" pageId="20" pageNumber="521">Nixe</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD1E641FDB8F84F8947F800" box="[587,638,2013,2038]" italics="true" pageId="20" pageNumber="521">Nixe</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD1E641FD61F84E89C2F800" box="[658,763,2012,2038]" italics="true" pageId="20" pageNumber="521">littorosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D12FFD1E641FCFFF84F88C3F800" authorityName="Kang & Yang" authorityYear="1994" box="[780,1018,2012,2038]" class="Insecta" family="Heptageniidae" genus="Nixe" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="mitificus" subGenus="Nixe">
|
||
<emphasis id="B939EA83FFD1E641FCFFF84F8806F800" box="[780,831,2013,2038]" italics="true" pageId="20" pageNumber="521">Nixe</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD1E641FCA0F84F88BFF800" box="[851,902,2013,2038]" italics="true" pageId="20" pageNumber="521">Nixe</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD1E641FC69F84E88C3F800" box="[922,1018,2012,2038]" italics="true" pageId="20" pageNumber="521">mitificus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D12FFD1E640FBCBF84F8A27FF47" authority="(Kang & Yang 1994)" baseAuthorityName="Kang & Yang" baseAuthorityYear="1994" class="Insecta" family="Heptageniidae" genus="Nixe" kingdom="Animalia" lastPageId="21" lastPageNumber="522" order="Ephemeroptera" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="obscurus" subGenus="Nixe">
|
||
<emphasis id="B939EA83FFD1E641FBCBF84F8F52F800" box="[1080,1131,2013,2038]" italics="true" pageId="20" pageNumber="521">Nixe</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD1E641FB8CF84F8F8BF800" box="[1151,1202,2013,2038]" italics="true" pageId="20" pageNumber="521">Nixe</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD1E641FB35F84E8E12F800" box="[1222,1323,2012,2038]" italics="true" pageId="20" pageNumber="521">obscurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDC4B60FFD1E640FACCF84E8A2CFF47" author="Kang, S. C. & Yang, C. T." lastPageId="21" lastPageNumber="522" pageId="20" pageNumber="521" pagination="5 - 36" refId="ref9662" refString="Kang, S. C. & Yang, C. T. (1994) Heptageniidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 5 - 36." type="journal article" year="1994">Kang & Yang 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Originally, the subgenus
|
||
<taxonomicName id="4C4D4D12FFD0E640FDB7FF05894EFF46" baseAuthorityName="Flowers" baseAuthorityYear="1980" box="[580,631,151,176]" pageId="21" pageNumber="522" rank="subGenus" subGenus="Nixe">
|
||
<emphasis id="B939EA83FFD0E640FDB7FF05894EFF46" box="[580,631,151,176]" italics="true" pageId="21" pageNumber="522">Nixe</emphasis>
|
||
</taxonomicName>
|
||
was characterized by absence of fibrilliform portion on tergalius VI of larva and ́mesh-like reticulate ridges» on egg (
|
||
<bibRefCitation id="EFDC4B60FFD0E640FD43FF298876FF23" author="Flowers, R. W." box="[688,847,187,213]" pageId="21" pageNumber="522" pagination="296 - 307" refId="ref9473" refString="Flowers, R. W. (1980) Two new genera of Nearctic Heptageniidae (Ephemeroptera). Florida Entomologist, 63 (3), 296 - 307. https: // doi. org / 10.2307 / 3494626" type="journal article" year="1980">Flowers 1980</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFDC4B60FFD0E640FC90FF298F69FF23" author="Kang, S. C. & Yang, C. T." box="[867,1104,187,213]" pageId="21" pageNumber="522" pagination="5 - 36" refId="ref9662" refString="Kang, S. C. & Yang, C. T. (1994) Heptageniidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 5 - 36." type="journal article" year="1994">Kang & Yang (1994)</bibRefCitation>
|
||
reported only the ́mesh-like reticular ridges» and ignored the structure of tergalii. It is unclear from the descriptions of these species, if they have the fibrilliform portion on tergalius VI or not. Characters reported as diagnostic for these species, vary individually in
|
||
<taxonomicName id="4C4D4D12FFD0E640FF40FEB58A5EFEB6" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[179,359,295,321]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FF40FEB58BF5FEB6" box="[179,204,295,320]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FF28FEB58A3BFEB7" box="[219,258,295,321]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FEE2FEB58A5EFEB6" box="[273,359,295,320]" italics="true" pageId="21" pageNumber="522">ornatus</emphasis>
|
||
</taxonomicName>
|
||
. Since imagines of these species are unknown, it is unclear if they really represent three different species distinct from
|
||
<taxonomicName id="4C4D4D12FFD0E640FE75FED98902FE92" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[390,571,331,357]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FE75FED98AA7FE92" box="[390,414,331,356]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FE5DFED98AECFE93" box="[430,469,331,357]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FE16FED98902FE92" box="[485,571,331,356]" italics="true" pageId="21" pageNumber="522">ornatus</emphasis>
|
||
</taxonomicName>
|
||
, or not.
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFD0E640FF34FEFD8FDCFD3B" blockId="21.[151,1437,151,717]" pageId="21" pageNumber="522">
|
||
Among species examined,
|
||
<taxonomicName id="4C4D4D12FFD0E640FE03FEFD8987FE7E" authorityName="Bengtsson" authorityYear="1909" box="[496,702,367,393]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="joernensis" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FE03FEFD8931FE7E" box="[496,520,367,392]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FDE5FEFD8904FE7F" box="[534,573,367,393]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FDB8FEFD8987FE7E" box="[587,702,367,392]" italics="true" pageId="21" pageNumber="522">joernensis</emphasis>
|
||
</taxonomicName>
|
||
(widely distributed from Scandinavia and northern part of Russian Plain to Siberia,
|
||
<collectingCountry id="F35A7601FFD0E640FEA3FE018AEFFE5B" box="[336,470,403,429]" name="Kazakhstan" pageId="21" pageNumber="522">Kazakhstan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35A7601FFD0E640FE13FE018974FE5B" box="[480,589,403,429]" name="Mongolia" pageId="21" pageNumber="522">Mongolia</collectingCountry>
|
||
and Russian Far East) is most closely related to
|
||
<taxonomicName id="4C4D4D12FFD0E640FB94FE018E22FE5A" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[1127,1307,403,429]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FB94FE018FB9FE5A" box="[1127,1152,403,428]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FB7CFE018F8FFE5B" box="[1167,1206,403,429]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FB36FE018E22FE5A" box="[1221,1307,403,428]" italics="true" pageId="21" pageNumber="522">ornatus</emphasis>
|
||
</taxonomicName>
|
||
. Eggs of
|
||
<taxonomicName id="4C4D4D12FFD0E640FA77FE018A73FE26" authorityName="Bengtsson" authorityYear="1909" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="joernensis" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FA77FE018EA4FE5A" box="[1412,1437,403,428]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FF52FE258BFEFE27" box="[161,199,439,465]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FF24FE258A73FE26" box="[215,330,439,464]" italics="true" pageId="21" pageNumber="522">joernensis</emphasis>
|
||
</taxonomicName>
|
||
have the same mesh-like ridges (
|
||
<bibRefCitation id="EFDC4B60FFD0E640FD33FE258866FE27" author="Flowers, R. W." box="[704,863,439,465]" pageId="21" pageNumber="522" pagination="193 - 197" refId="ref9514" refString="Flowers, R. W. (1986) Holarctic distributions of three taxa of Heptageniidae (Ephemeroptera). Entomological News, 97 (5), 193 - 197." type="journal article" year="1986">Flowers 1986</bibRefCitation>
|
||
: fig. 3), larva has the same cuticular coloration and leg setation; in both species paraglossae are greatly projected laterally (
|
||
<figureCitation id="13762A14FFD0E640FC4CFE498F2DFE03" box="[959,1044,475,501]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="21" pageNumber="522">Fig. 31</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B60FFD0E640FBD3FE498F90FE03" author="Kluge, N. J." box="[1056,1193,475,501]" pageId="21" pageNumber="522" pagination="176 - 220" refId="ref9805" refString="Kluge, N. J. (1997) Ephemeroptera. In: Tsalolikhin, S. J. (Ed.), Key to freshwater invertebrates of Russia and adjacent lands, S- Petersburg, 3, pp. 176 - 220. [in Russian]" type="journal article" year="1997">Kluge 1997</bibRefCitation>
|
||
: tab. 15: fig. 7). Both species have similar individual variability of subimaginal mesonotum (compare
|
||
<figureCitation id="13762A14FFD0E640FBD3FE6D8FA7FDEF" box="[1056,1182,511,537]" captionStart="FIGURES 53–68" captionStartId="8.[151,264,1807,1831]" captionTargetBox="[164,1422,181,1784]" captionTargetId="figure-37@8.[164,1423,181,1784]" captionTargetPageId="8" captionText="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." figureDoi="http://doi.org/10.5281/zenodo.8203262" httpUri="https://zenodo.org/record/8203262/files/figure.png" pageId="21" pageNumber="522">Figs 66–67</figureCitation>
|
||
with
|
||
<bibRefCitation id="EFDC4B60FFD0E640FB2CFE6D8E5EFDEF" author="Kluge, N. J." box="[1247,1383,511,537]" pageId="21" pageNumber="522" pagination="561 - 579" refId="ref9699" refString="Kluge, N. J. (1980) To the knowledge of mayflies (Ephemeroptera) of Taimyr National District. Entomologicheskoe Obozrenie, 59 (3), 561 - 579. [in Russian]" type="journal article" year="1980">Kluge 1980</bibRefCitation>
|
||
: figs 93–94). Imagines of these species have similar general appearance, including coloration of the body and legs, shape of hind wing, proportion of tarsal segments. Male imago of
|
||
<taxonomicName id="4C4D4D12FFD0E640FCDDFDD588DBFD96" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[814,994,583,609]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FCDDFDD5887EFD96" box="[814,839,583,608]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FCA5FDD58845FD97" box="[854,892,583,609]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FC7FFDD588DBFD96" box="[908,994,583,608]" italics="true" pageId="21" pageNumber="522">ornatus</emphasis>
|
||
</taxonomicName>
|
||
well differs from
|
||
<taxonomicName id="4C4D4D12FFD0E640FB5AFDD58E43FD96" authorityName="Bengtsson" authorityYear="1909" box="[1193,1402,583,609]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="joernensis" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FB5AFDD58FFBFD96" box="[1193,1218,583,608]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FB22FDD58FC1FD97" box="[1233,1272,583,609]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FAF4FDD58E43FD96" box="[1287,1402,583,608]" italics="true" pageId="21" pageNumber="522">joernensis</emphasis>
|
||
</taxonomicName>
|
||
by more simple shape of penis and by absence of discal spines on ventral side. Larva of
|
||
<taxonomicName id="4C4D4D12FFD0E640FBAAFDF98E36FD72" baseAuthorityName="Ulmer" baseAuthorityYear="1939" box="[1113,1295,619,645]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="ornatus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FBAAFDF98F48FD72" box="[1113,1137,619,644]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FB71FDF98F91FD73" box="[1154,1192,619,645]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FB4AFDF98E36FD72" box="[1209,1295,619,644]" italics="true" pageId="21" pageNumber="522">ornatus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4D4D12FFD0E640FF64FD1D8A53FD5E" authorityName="Bengtsson" authorityYear="1909" box="[151,362,655,681]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="joernensis" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FF64FD1D8B89FD5E" box="[151,176,655,680]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FF33FD1D8BDEFD5F" box="[192,231,655,681]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FF04FD1D8A53FD5E" box="[247,362,655,680]" italics="true" pageId="21" pageNumber="522">joernensis</emphasis>
|
||
</taxonomicName>
|
||
by size of tergalii, which increase from tergalius II to tergalius V (
|
||
<figureCitation id="13762A14FFD0E640FBABFD1D8FF4FD5F" box="[1112,1229,655,681]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1810,1834]" captionTargetBox="[151,1436,180,1772]" captionTargetId="figure-20@3.[151,1436,179,1784]" captionTargetPageId="3" captionText="FIGURES 1–10. Ecdyonurus (Rhithrogeniella) ornatus from Kodaikanal (India). 1–3, larvae; 4–10, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8203250" httpUri="https://zenodo.org/record/8203250/files/figure.png" pageId="21" pageNumber="522">Figs 4–10</figureCitation>
|
||
), while in
|
||
<taxonomicName id="4C4D4D12FFD0E640FAB7FD1D8A33FD3A" authorityName="Bengtsson" authorityYear="1909" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="joernensis" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFD0E640FAB7FD1D8E64FD5E" box="[1348,1373,655,680]" italics="true" pageId="21" pageNumber="522">E.</emphasis>
|
||
(
|
||
<emphasis id="B939EA83FFD0E640FA9EFD1D8EAAFD5F" box="[1389,1427,655,681]" italics="true" pageId="21" pageNumber="522">Rh.</emphasis>
|
||
)
|
||
<emphasis id="B939EA83FFD0E640FF64FD218A33FD3A" box="[151,266,691,716]" italics="true" pageId="21" pageNumber="522">joernensis</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C4D4D12FFD0E640FEC2FD218AEBFD3B" box="[305,466,691,717]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="mongolicus">
|
||
<emphasis id="B939EA83FFD0E640FEC2FD218AEBFD3B" box="[305,466,691,717]" italics="true" pageId="21" pageNumber="522">E. mongolicus</emphasis>
|
||
</taxonomicName>
|
||
) they decrease from tergalius II to tergalius VII (
|
||
<bibRefCitation id="EFDC4B60FFD0E640FC0AFD218FB9FD3B" author="Kluge, N. J." box="[1017,1152,691,717]" pageId="21" pageNumber="522" pagination="561 - 579" refId="ref9699" refString="Kluge, N. J. (1980) To the knowledge of mayflies (Ephemeroptera) of Taimyr National District. Entomologicheskoe Obozrenie, 59 (3), 561 - 579. [in Russian]" type="journal article" year="1980">Kluge 1980</bibRefCitation>
|
||
: fig. 84).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |